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<document id="332CEE83D74C36857ECBEF9DB0C65025" ID-CLB-Dataset="28107" ID-DOI="10.11646/zootaxa.4514.1.9" ID-GBIF-Dataset="80715276-706f-4d06-a8c2-49e32c03f155" ID-ISSN="1175-5326" ID-Zenodo-Dep="2605858" ID-ZooBank="C4BCDC9D-BCB2-4DF0-8EFB-7DBC61CB3E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553575677981" checkinUser="plazi" docAuthor="Diez, Yander L., Hernández, Claudia Sanjuan, Reygel, Patrick, Roosen, Paulien &amp; Artois, Tom" docDate="2018" docId="B21887C6E358B153E69392025148FD81" docLanguage="en" docName="zootaxa.4514.1.9.pdf" docOrigin="Zootaxa 4514 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polycydora intermedia Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois 2018, sp. n." docType="treatment" docUuid="27E54D7D-337A-4B8B-BACD-21D56EE0E0EB" docUuidSource="ZooBank" docVersion="10" lastPageNumber="120" masterDocId="4E21FFBEE352B15EE60493065517FF99" masterDocTitle="First record of Polycystididae (Platyhelminthes, Kalyptorhynchia) from Cuba, with the description of a new genus and five new species, and remarks and the description of one new species from Panama" masterLastPageNumber="125" masterPageNumber="107" pageNumber="117" updateTime="1698712660214" updateUser="ExternalLinkService">
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<mods:title id="619D172A30FFAAC665237262F092A77B">First record of Polycystididae (Platyhelminthes, Kalyptorhynchia) from Cuba, with the description of a new genus and five new species, and remarks and the description of one new species from Panama</mods:title>
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<mods:namePart id="2CE3EE08EAFB30241442509AC7ED7C72">Diez, Yander L.</mods:namePart>
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<mods:namePart id="8A34AE760E4187F45272081F45744C05">Hernández, Claudia Sanjuan</mods:namePart>
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<mods:namePart id="486060BB5C71FDE5B821D1C200D3EB31">Reygel, Patrick</mods:namePart>
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<mods:namePart id="D4E5833AFFDCD8FB2C25D0F8350D33DA">Roosen, Paulien</mods:namePart>
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<mods:namePart id="DD5B008EE3FDA381F1C20A15E69999DB">Artois, Tom</mods:namePart>
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<mods:title id="EDDBDEB7D2F053FB83B6A1941E7BB118">Zootaxa</mods:title>
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<mods:date id="95DE5CDA19851EBA164E56C26C759A14">2018</mods:date>
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<mods:number id="8C6D2730D7F3D71ABC45E457BA5F2492">2018-11-06</mods:number>
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<mods:number id="4577818379677D52E222BD35E5DF2042">4514</mods:number>
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<treatment id="B21887C6E358B153E69392025148FD81" ID-DOI="http://doi.org/10.5281/zenodo.5992805" ID-GBIF-Taxon="154494515" ID-Zenodo-Dep="5992805" LSID="urn:lsid:zoobank.org:act:27E54D7D-337A-4B8B-BACD-21D56EE0E0EB" httpUri="http://treatment.plazi.org/id/B21887C6E358B153E69392025148FD81" lastPageId="13" lastPageNumber="120" pageId="10" pageNumber="117">
<subSubSection id="72AB655BE358B154E69392025668FE87" box="[151,895,260,286]" pageId="10" pageNumber="117" type="nomenclature">
<paragraph id="3A0E36D0E358B154E69392025668FE87" blockId="10.[151,895,260,319]" box="[151,895,260,286]" pageId="10" pageNumber="117">
<heading id="614681BCE358B154E69392025668FE87" bold="true" box="[151,895,260,286]" fontSize="11" level="1" pageId="10" pageNumber="117" reason="1">
<emphasis id="08C5EAC2E358B154E69392025668FE87" bold="true" box="[151,895,260,286]" pageId="10" pageNumber="117">
<taxonomicName id="FDB14D53E358B154E69392025622FE87" ID-CoL="4L4Y4" authority="Diez, Sanjuan, Reygel &amp; Artois" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[151,821,260,286]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="10" pageNumber="117" phylum="Platyhelminthes" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="08C5EAC2E358B154E693920254B0FE87" bold="true" box="[151,423,260,286]" italics="true" pageId="10" pageNumber="117">Polycydora intermedia</emphasis>
Diez, Sanjuan, Reygel &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="13F657B9E358B154E53992025668FE87" box="[829,895,260,286]" pageId="10" pageNumber="117" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="72AB655BE358B154E693922055FCFEA6" box="[151,235,294,319]" pageId="10" pageNumber="117" type="description">
<paragraph id="3A0E36D0E358B154E693922055FCFEA6" blockId="10.[151,895,260,319]" box="[151,235,294,319]" pageId="10" pageNumber="117">
(
<figureCitation id="A28A2A55E358B154E69B922055F4FEA6" box="[159,227,294,319]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72AB655BE358B154E693926857B9FDA2" pageId="10" pageNumber="117" type="materials_examined">
<paragraph id="3A0E36D0E358B154E693926857B9FDA2" blockId="10.[151,1437,366,1975]" pageId="10" pageNumber="117">
<emphasis id="08C5EAC2E358B154E69392685723FE1E" bold="true" box="[151,564,366,391]" pageId="10" pageNumber="117">
Material and distribution in
<materialsCitation id="8AD93C8DE358B154E7E892685723FE1E" ID-GBIF-Occurrence="2028137444" box="[492,564,366,391]" collectingDate="2016-05-15" collectingDateMax="2018-02-06" collectingDateMin="2016-05-15" collectionCode="FMNH, KV, VIII" collectorName="de Gibara" country="Cuba" county="Bahia de Gibara" elevation="6" location="Observations" municipality="Five" pageId="10" pageNumber="117" specimenCount="1" typeStatus="holotype">
<collectingCountry id="42A67640E358B154E7E892685738FE1E" box="[492,559,366,391]" name="Cuba" pageId="10" pageNumber="117">Cuba</collectingCountry>
.
</materialsCitation>
</emphasis>
<location id="3F6E600BE358B154E43F926857D8FE1E" LSID="urn:lsid:plazi:treatment:B21887C6E358B153E69392025148FD81:3F6E600BE358B154E43F926857D8FE1E" box="[571,719,366,391]" country="Cuba" county="Bahia de Gibara" municipality="Five" name="Observations" pageId="10" pageNumber="117">Observations</location>
on live animals.
<collectingMunicipality id="DA6AACAAE358B154E58A926856D7FE1E" box="[910,960,366,391]" pageId="10" pageNumber="117">Five</collectingMunicipality>
whole mounts, one of which is designated
<typeStatus id="E50A8872E358B154E693929455EEFE32" box="[151,249,402,427]" pageId="10" pageNumber="117" type="holotype">holotype</typeStatus>
(FMNH http://id.luomus.fi/KV.604), and ten serially-sectioned specimens collected in
<collectingCounty id="D36F4E5CE358B154E2E59294508BFE32" box="[1249,1436,402,427]" pageId="10" pageNumber="117">
Bahía
<collectorName id="97445306E358B154E3289294508BFE32" box="[1324,1436,402,427]" pageId="10" pageNumber="117">de Gibara</collectorName>
</collectingCounty>
(type locality) (
<collectingDate id="5E4BE9F8E358B154E74092B054EBFE56" box="[324,508,438,463]" pageId="10" pageNumber="117" value="2017-03-26">
<date id="4E0F1010E358B154E74092B054EBFE56" box="[324,508,438,463]" pageId="10" pageNumber="117" value="2017-03-26">March 26, 2017</date>
</collectingDate>
), sublittoral, in coarse sand,
<elevation id="B19CD1E3E358B154E53992B0566AFE57" box="[829,893,438,463]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="117" unit="m" value="0.6">
<quantity id="FD499B35E358B154E53992B0566AFE57" box="[829,893,438,463]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="117" unit="m" value="0.6">0.6 m</quantity>
</elevation>
depth, salinity 35 ‰. Three whole mounts from Siboney (
<collectingDate id="5E4BE9F8E358B154E70792DD54B4FE6A" box="[259,419,474,499]" pageId="10" pageNumber="117" value="2016-05-15">
<date id="4E0F1010E358B154E70792DD54B4FE6A" box="[259,419,474,499]" pageId="10" pageNumber="117" value="2016-05-15">May 15, 2016</date>
</collectingDate>
&amp;
<collectingDate id="5E4BE9F8E358B154E7CE92DD5748FE6A" box="[458,607,474,499]" pageId="10" pageNumber="117" value="2017-06-05">
<date id="4E0F1010E358B154E7CE92DD5748FE6A" box="[458,607,474,499]" pageId="10" pageNumber="117" value="2017-06-05">June 5, 2017</date>
</collectingDate>
), intertidal, upper
<quantity id="FD499B35E358B154E53492DD5661FE6B" box="[816,886,475,499]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="117" unit="cm" value="10.0">10 cm</quantity>
of fine sand with organic matter, salinity 35 ‰. One whole mount from Bueycabón (
<collectingDate id="5E4BE9F8E358B154E43A92F8561EFD8E" box="[574,777,510,535]" pageId="10" pageNumber="117" value="2018-02-06">
<date id="4E0F1010E358B154E43A92F8561EFD8E" box="[574,777,510,535]" pageId="10" pageNumber="117" value="2018-02-06">February 6, 2018</date>
</collectingDate>
), intertidal, upper
<quantity id="FD499B35E358B154E5DA92F95131FD8F" box="[990,1062,511,535]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="117" unit="cm" value="10.0">10 cm</quantity>
of fine sand, salinity 33 ‰. All reference material in HU (VIII.3.15VIII.3.32).
</paragraph>
</subSubSection>
<subSubSection id="72AB655BE358B154E6C3914054AAFD1B" pageId="10" pageNumber="117" type="etymology">
<paragraph id="3A0E36D0E358B154E6C3914054AAFD1B" blockId="10.[151,1437,366,1975]" pageId="10" pageNumber="117">
<emphasis id="08C5EAC2E358B154E6C391405459FDC6" bold="true" box="[199,334,582,607]" pageId="10" pageNumber="117">Etymology.</emphasis>
The epithet refers to the fact that the new species is morphologically intermediate between
<taxonomicName id="FDB14D53E358B154E693916C541EFD1A" authorityName="Kolliker" authorityYear="1845" box="[151,265,618,643]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="10" pageNumber="117" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E358B154E693916C541EFD1A" box="[151,265,618,643]" italics="true" pageId="10" pageNumber="117">Polycystis</emphasis>
</taxonomicName>
and
<taxonomicName id="FDB14D53E358B154E73B916C54A0FD1A" authorityName="Marcus" authorityYear="1948" box="[319,439,618,643]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="10" pageNumber="117" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E358B154E73B916C54A0FD1A" box="[319,439,618,643]" italics="true" pageId="10" pageNumber="117">Paulodora</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="72AB655BE358B154E6C391885612FD76" pageId="10" pageNumber="117" type="diagnosis">
<paragraph id="3A0E36D0E358B154E6C391885612FD76" blockId="10.[151,1437,366,1975]" pageId="10" pageNumber="117">
<emphasis id="08C5EAC2E358B154E6C391885456FD3E" bold="true" box="[199,321,654,679]" pageId="10" pageNumber="117">Diagnosis.</emphasis>
Species of
<taxonomicName id="FDB14D53E358B154E7D091885742FD3E" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[468,597,654,679]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="10" pageNumber="117" phylum="Platyhelminthes" rank="genus" status="gen. nov.">
<emphasis id="08C5EAC2E358B154E7D091885742FD3E" box="[468,597,654,679]" italics="true" pageId="10" pageNumber="117">Polycydora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13F657B9E358B154E466918857A0FD3E" box="[610,695,654,679]" pageId="10" pageNumber="117" rank="genus">
<emphasis id="08C5EAC2E358B154E466918857A0FD3E" bold="true" box="[610,695,654,679]" pageId="10" pageNumber="117">gen. n.</emphasis>
</taxonomicNameLabel>
with a 75-µm-long prostate stylet
<typeStatus id="E50A8872E358B154E2599189519AFD3E" box="[1117,1165,655,679]" pageId="10" pageNumber="117">type</typeStatus>
II. The outer stylet is proximally funnel shaped, ±18 µm wide, then is constricted to a tube 4 µm wide. At one-third of its length it shows a 180° turn. Stylet with an asymmetrical distal opening.
</paragraph>
</subSubSection>
<subSubSection id="72AB655BE358B152E6C391FC51E7FD31" lastPageId="12" lastPageNumber="119" pageId="10" pageNumber="117" type="description">
<paragraph id="3A0E36D0E358B154E6C391FC5195FCE6" blockId="10.[151,1437,366,1975]" pageId="10" pageNumber="117">
<emphasis id="08C5EAC2E358B154E6C391FC544EFC8A" bold="true" box="[199,345,762,787]" pageId="10" pageNumber="117">Description.</emphasis>
Living animals
<quantity id="FD499B35E358B154E41691FC577DFC8B" box="[530,618,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="117" unit="mm" value="1.5">1.5 mm</quantity>
long. Length measured on whole mounts
<quantity id="FD499B35E358B154E24D91FC51C6FC8B" box="[1097,1233,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="10" pageNumber="117" unit="mm" value="1.2" valueMax="1.5" valueMin="0.9">0.91.5 mm</quantity>
(
<emphasis id="08C5EAC2E358B154E2E691FA51E0FC8A" box="[1250,1271,764,787]" italics="true" pageId="10" pageNumber="117"></emphasis>
=
<quantity id="FD499B35E358B154E30B91FD507EFC8B" box="[1295,1385,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="117" unit="mm" value="1.2">1.2 mm</quantity>
; n = 4). Translucent, with a pair of eyes. Syncytial epidermis 7 µm thick and ciliated, cilia 67 µm long. Two
<typeStatus id="E50A8872E358B154E3469019506AFCAE" box="[1346,1405,799,823]" pageId="10" pageNumber="117">types</typeStatus>
of vacuoles present in the epidermis: empty ones (support structures) and others filled with small granules. Rhabdites as large as the height of the epidermis, only present in the posterior two-third of the body.
</paragraph>
<paragraph id="3A0E36D0E358B154E6C3908C5499FB5A" blockId="10.[151,1437,366,1975]" pageId="10" pageNumber="117">
Proboscis of the typical polycystidid
<typeStatus id="E50A8872E358B154E47D908D57BEFC3A" box="[633,681,907,931]" pageId="10" pageNumber="117">type</typeStatus>
(
<figureCitation id="A28A2A55E358B154E4B9908C5614FC3A" box="[701,771,906,931]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4</figureCitation>
AB: pr,
<figureCitation id="A28A2A55E358B154E576908C56DBFC3A" box="[882,972,906,931]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
; for a detailed description see
<bibRefCitation id="5E204B21E358B154E333908C55C5FC5E" author="Meixner, J." pageId="10" pageNumber="117" pagination="255 - 343" refId="ref12046" refString="Meixner, J. (1925) Beitrag zur Morphologie und zum System der Turbellaria-Rhabdocoela: 1. Die Kalyptorhynchia. Zeitschrift fur Morphologie und Okologie der Tiere, 3, 255 - 343. https: // doi. org / 10.1007 / BF 00408146" type="journal article" year="1925">Meixner, 1925</bibRefCitation>
), 1/5 of the body length, with six pairs of fixator muscles (
<figureCitation id="A28A2A55E358B154E59D90A856E0FC5E" box="[921,1015,942,967]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: pfix). With four pairs of proboscis retractors (
<figureCitation id="A28A2A55E358B154E71590D4547CFC72" box="[273,363,978,1003]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: pret). Cone retractors well-developed (
<figureCitation id="A28A2A55E358B154E52890D45691FC72" box="[812,902,978,1003]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: cret). As in most
<taxonomicName id="FDB14D53E358B154E25290D451ECFC72" authorityName="Graff" authorityYear="1905" box="[1110,1275,978,1003]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="10" pageNumber="117" phylum="Platyhelminthes" rank="family">Polycystididae</taxonomicName>
with a pair of ventral integument retractors (
<figureCitation id="A28A2A55E358B154E7FF90F0574DFB96" box="[507,602,1014,1039]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: ir). Proboscis sheath surrounded by an internal circular and an outer longitudinal muscle layer. Without nuclei at the junction between sheath and cone epithelium. Thin dilator muscles are present in the distal half of the proboscis sheath (
<figureCitation id="A28A2A55E358B154E4FF97385641FBCE" box="[763,854,1086,1111]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: dil). Proboscis opening surrounded by a sphincter (
<figureCitation id="A28A2A55E358B154E69B976455E8FBE2" box="[159,255,1122,1147]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: sph). Proboscis glands can be observed just caudally from the proboscis: one central large gland producing an eosinophilic fine-grained secretion (
<figureCitation id="A28A2A55E358B154E4D99780562CFB06" box="[733,827,1158,1183]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: peg), and surrounding it, coarse-grained basophilic glands (
<figureCitation id="A28A2A55E358B154E6EB97AC545FFB5A" box="[239,328,1194,1219]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4C</figureCitation>
: pbg).
</paragraph>
<paragraph id="3A0E36D0E358B154E6C397C85716F90E" blockId="10.[151,1437,366,1975]" pageId="10" pageNumber="117">
Pharynx located at about 35% (
<figureCitation id="A28A2A55E358B154E42397C8577DFB7E" box="[551,618,1230,1255]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4</figureCitation>
AB: ph,
<figureCitation id="A28A2A55E358B154E4D397C85627FB7E" box="[727,816,1230,1255]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
), measuring 153174 µm in diameter (
<emphasis id="08C5EAC2E358B154E2E297D651EDFB7E" box="[1254,1274,1232,1255]" italics="true" pageId="10" pageNumber="117"></emphasis>
= 162 µm; n = 4). Prepharyngeal cavity (
<figureCitation id="A28A2A55E358B154E7C397F45730FA92" box="[455,551,1266,1291]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: ppc) lined with a nucleated epithelium surrounded by an internal layer of circular muscles and an external layer of longitudinal muscles, opening to the outside through the mouth (
<figureCitation id="A28A2A55E358B154E3399610508EFAB6" box="[1341,1433,1302,1327]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: m). Pharynx bulb with the normal polycystidid construction, showing the four teeth surrounding the proximal pharynx opening, typical of the family (
<figureCitation id="A28A2A55E358B154E451965857B8FAEE" box="[597,687,1374,1399]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: th). Three
<typeStatus id="E50A8872E358B154E52B9659567DFAEE" box="[815,874,1375,1399]" pageId="10" pageNumber="117">types</typeStatus>
of glands open in the distal part of the pharyngeal lumen, which are, from distal to proximal: eosinophilic glands (stained light pink) (
<figureCitation id="A28A2A55E358B154E26D968451DFFA02" box="[1129,1224,1410,1435]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: phg1), basophilic glands with a coarse-grained secretion (stained dark blue-black) (
<figureCitation id="A28A2A55E358B154E58096A056F6FA26" box="[900,993,1446,1471]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: phg2), and eosinophilic glands with a fine-grained secretion (stained brownish) (
<figureCitation id="A28A2A55E358B154E47D96CC57C1FA7A" box="[633,726,1482,1507]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: phg3). At its distal side the pharynx has a muscular sphincter (
<figureCitation id="A28A2A55E358B154E69B96E855ECF99E" box="[159,251,1518,1543]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: sph), just distal from the opening of the glands. The musculature of the pharynx consists of an external layer of circular muscles (
<figureCitation id="A28A2A55E358B154E7BE95145702F9B2" box="[442,533,1554,1579]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: cm), just inside of the septum and a longitudinal muscle (
<figureCitation id="A28A2A55E358B154E2A6951451EAF9B2" box="[1186,1277,1554,1579]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: lm) layer just outside of it, which is continuous with the longitudinal muscle layer of the prepharyngeal cavity. The pharynx lumen is surrounded by a circular muscle layer and a longitudinal one. Radial muscles run between the internal and the external walls (
<figureCitation id="A28A2A55E358B154E768957854D0F90E" box="[364,455,1662,1687]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4D</figureCitation>
: rm).
</paragraph>
<paragraph id="3A0E36D0E358B154E6C395A4579DF82E" blockId="10.[151,1437,366,1975]" pageId="10" pageNumber="117">
Genital organs situated in the middle section of the body. Testes (
<figureCitation id="A28A2A55E358B154E5A895A45111F922" box="[940,1030,1698,1723]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4B</figureCitation>
: t) situated caudally to the pharynx. Seminal vesicles (
<figureCitation id="A28A2A55E358B154E76595C054E9F946" box="[353,510,1734,1759]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4B &amp; 4E</figureCitation>
: sv) located between the testes and delineated by a thin nucleated epithelium. Both seminal vesicles fuse to form a thin ejaculatory duct (
<figureCitation id="A28A2A55E358B154E50595EC564CF89A" box="[769,859,1770,1795]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4E</figureCitation>
: ed), covered by circular muscles. The ejaculatory duct runs aside the prostate vesicle, entering the male genital atrium next to the proximal funnel of the prostate stylet
<typeStatus id="E50A8872E358B154E6DF9435541CF8D2" box="[219,267,1843,1867]" pageId="10" pageNumber="117">type</typeStatus>
II. Prostate vesicle
<typeStatus id="E50A8872E358B154E7ED9435570EF8D2" box="[489,537,1843,1867]" pageId="10" pageNumber="117">type</typeStatus>
II elongated (
<figureCitation id="A28A2A55E358B154E4B294345645F8D2" box="[694,850,1842,1867]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4B &amp; 4E</figureCitation>
: pv2), 6377 µm long (
<emphasis id="08C5EAC2E358B154E25E94325179F8D2" box="[1114,1134,1844,1867]" italics="true" pageId="10" pageNumber="117"></emphasis>
= 69 µm; n = 4) and 1830 µm wide (
<emphasis id="08C5EAC2E358B154E70B945E5432F8F6" box="[271,293,1880,1903]" italics="true" pageId="10" pageNumber="117"></emphasis>
= 23 µm; n = 4), filled with an eosinophilic secretion. Prostate vesicle surrounded by two muscular layers: external oblique fibres, and internal longitudinal fibres. Prostate vesicle connected proximally to brownish accessory glands
<typeStatus id="E50A8872E358B154E75F9499549CF82E" box="[347,395,1951,1975]" pageId="10" pageNumber="117">type</typeStatus>
I (
<figureCitation id="A28A2A55E358B154E7AF94985750F82E" box="[427,583,1950,1975]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="10" pageNumber="117">Fig. 4B &amp; 4E</figureCitation>
: ag1).
</paragraph>
<caption id="6ECE6658E359B155E6939599564BF888" httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="11" pageNumber="118" startId="11.[151,250,1695,1717]" targetBox="[177,1405,224,1662]" targetPageId="11">
<paragraph id="3A0E36D0E359B155E6939599564BF888" blockId="11.[151,1436,1695,1809]" pageId="11" pageNumber="118">
<emphasis id="08C5EAC2E359B155E69395995405F92D" bold="true" box="[151,274,1695,1717]" pageId="11" pageNumber="118">FIGURE 4.</emphasis>
<taxonomicName id="FDB14D53E359B155E71C95995716F92C" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[280,513,1695,1717]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="11" pageNumber="118" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E359B155E71C95995716F92C" box="[280,513,1695,1717]" italics="true" pageId="11" pageNumber="118">Polycydora intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E359B155E40295A6579CF92C" bold="true" box="[518,651,1696,1717]" pageId="11" pageNumber="118">
<taxonomicNameLabel id="13F657B9E359B155E40295A6575AF92C" box="[518,589,1696,1717]" pageId="11" pageNumber="118" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand.
</paragraph>
</caption>
<paragraph id="3A0E36D0E359B152E6C3943956D4FF4D" blockId="11.[151,1436,1854,2023]" lastBlockId="12.[151,1437,151,2013]" lastPageId="12" lastPageNumber="119" pageId="11" pageNumber="118">
Prostate stylet
<typeStatus id="E50A8872E359B155E7689439548BF8CE" box="[364,412,1855,1879]" pageId="11" pageNumber="118">type</typeStatus>
II double walled and sickle shaped (
<figureCitation id="A28A2A55E359B155E53194385660F8CE" box="[821,887,1854,1879]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="11" pageNumber="118">Fig. 4</figureCitation>
AB &amp; 4E: ps2,
<figureCitation id="A28A2A55E359B155E23694385190F8CE" box="[1074,1159,1854,1879]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="11" pageNumber="118">Fig. 4F</figureCitation>
), 6087 µm long (
<emphasis id="08C5EAC2E359B155E3529439507DF8CF" box="[1366,1386,1855,1878]" italics="true" pageId="11" pageNumber="118"></emphasis>
= 75 µm; n = 8), with a proximal funnel, 1621 µm wide (
<emphasis id="08C5EAC2E359B155E4F794625610F8E2" box="[755,775,1892,1915]" italics="true" pageId="11" pageNumber="118"></emphasis>
= 18 µm; n = 8). Distally from the funnel, the stylet is 45 µm wide (
<emphasis id="08C5EAC2E359B155E70F94815409F807" box="[267,286,1927,1950]" italics="true" pageId="11" pageNumber="118"></emphasis>
= 4 µm; n = 8), showing a 180° twist. Male bursa (
<figureCitation id="A28A2A55E359B155E550948056E6F806" box="[852,1009,1926,1951]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="11" pageNumber="118">Fig. 4B &amp; 4E</figureCitation>
: mb) well developed, surrounded by a thick muscular coat, filled with sperm and separated from its muscular stalk by a sphincter (
<figureCitation id="A28A2A55E359B155E2F394AC5043F85A" box="[1271,1364,1962,1987]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="11" pageNumber="118">Fig. 4E</figureCitation>
: sph). Epithelium with nuclei surrounds the bursal stalk and the male atrium. Bursal stalk entering the common genital atrium (
<figureCitation id="A28A2A55E35EB152E6F59391545DFF29" box="[241,330,151,176]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4E</figureCitation>
: cga) dorsally, at the same place as does the male genital atrium. The internal vesicle of the male bursa is surrounded by a tick sheath of syncytial epithelium (
<figureCitation id="A28A2A55E35EB152E54593BA568FFF4D" box="[833,920,188,213]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4E</figureCitation>
: se)
</paragraph>
<paragraph id="3A0E36D0E35EB152E6C393D951E7FD31" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="119">
Ovaries kidney shaped (
<figureCitation id="A28A2A55E35EB152E7D693D9577AFF61" box="[466,621,223,248]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4B &amp; 4E</figureCitation>
: ov) and located in the posterior third of the body. Oocytes organised in a row, diminishing in size from the most distal one to the most proximal one. Paired vitellaria extend at both sides from the middle to the end of the body (
<figureCitation id="A28A2A55E35EB152E42892215767FED9" box="[556,624,295,320]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4</figureCitation>
AB &amp; 4E: vi). Oviducts very short, each receiving a vitelloduct. Both oviducts unite to form a short common female duct
<typeStatus id="E50A8872E35EB152E4DE924B561EFEFC" box="[730,777,333,357]" pageId="12" pageNumber="119">type</typeStatus>
I that enters the common genital atrium caudally. Oviducts and female duct are lined by a nucleated epithelium. Muscles are not well defined around the oviducts and the female duct, however, there seems to be an internal layer of circular muscles and an external layer of longitudinal ones. Large female bursa 50 µm long (measured in one sectioned specimen), contains a central space with sperm and nuclei. It connects to the female duct by a bursal stalk at the junction of the two oviducts. Bursal stalk lined by a nucleated epithelium. Muscle layers are not observable around the female bursa and the stalk. A uterus (
<figureCitation id="A28A2A55E35EB152E2DE92F95024FD81" box="[1242,1331,511,536]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4E</figureCitation>
: ut) with nucleated epithelium connects at the anterior side of the common genital atrium. Two rings of glands can be observed at the distal end of the uterus: one coarse-granulated basophilic gland (
<figureCitation id="A28A2A55E35EB152E5DF91415121FDF9" box="[987,1078,583,608]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4E</figureCitation>
: ubg), and one fine-granulated eosinophilic gland (
<figureCitation id="A28A2A55E35EB152E76A916A54D4FD1D" box="[366,451,620,645]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4E</figureCitation>
: ueg) separated from the nucleated epithelium by a dense structure, which could be a thick sphincter (
<figureCitation id="A28A2A55E35EB152E70E91895448FD31" box="[266,351,655,680]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4E</figureCitation>
: x). Common genital atrium opens externally by a common gonopore (
<figureCitation id="A28A2A55E35EB152E263918951ABFD31" box="[1127,1212,655,680]" captionStart="FIGURE 4" captionStartId="11.[151,250,1695,1717]" captionTargetBox="[177,1405,224,1662]" captionTargetId="figure@11.[151,1436,193,1674]" captionTargetPageId="11" captionText="FIGURE 4. Polycydora intermedia gen. n. sp. n. AB, Habitus (from a live animal). C, Schematic reconstruction from sagittal sections of the proboscis. D, Schematic reconstruction from sagittal sections of the pharynx from the right side. E, Schematic reconstruction from sagittal sections of the genital system from the right side. F, Drawing of the stylet in the holotype. The scale bar only corresponds to the stylet (F), all other drawings are freehand." httpUri="https://zenodo.org/record/2605866/files/figure.png" pageId="12" pageNumber="119">Fig. 4E</figureCitation>
: cg).
</paragraph>
</subSubSection>
<subSubSection id="72AB655BE35EB152E6C391B25057FA05" pageId="12" pageNumber="119" type="discussion">
<paragraph id="3A0E36D0E35EB152E6C391B25700FB89" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="119">
<emphasis id="08C5EAC2E35EB152E6C391B2545BFD54" bold="true" box="[199,332,692,717]" pageId="12" pageNumber="119">Discussion.</emphasis>
The new species shows all diagnostic features of
<taxonomicName id="FDB14D53E35EB152E57E91B25135FD54" authorityName="Graff" authorityYear="1905" box="[890,1058,692,717]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="family">Polycystididae</taxonomicName>
, the most typical of which are the presence of four teeth around the proximal pharynx opening and of three pairs of proboscis fixators (
<bibRefCitation id="5E204B21E35EB152E2F291D15087FD69" author="Karling, T. G." box="[1270,1424,727,752]" pageId="12" pageNumber="119" pagination="159 - 183" refId="ref11790" refString="Karling, T. G. (1964) Uber einige neue und ungenugend bekannte Turbellaria Eukalyptorhynchia. Zoologischer Anzeiger, 172, 159 - 183." type="journal article" year="1964">Karling 1964</bibRefCitation>
). It lacks the typical features of the subtaxa
<taxonomicName id="FDB14D53E35EB152E49491FA560BFC8C" authorityName="Graff" authorityYear="1905" box="[656,796,764,789]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="subFamily" subFamily="Gyratricinae">Gyratricinae</taxonomicName>
[unpaired ovaries; terminal (male) gonopore; unpaired seminal vesicle],
<taxonomicName id="FDB14D53E35EB152E7839019575FFCA1" authorityName="Tessens, Janssen &amp; Artois" authorityYear="2014" box="[391,584,799,824]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="subFamily" subFamily="Scanorhynchinae">Scanorhynchinae</taxonomicName>
(position of external muscles of proboscis sheath inverted),
<taxonomicName id="FDB14D53E35EB152E69390425468FCC4" authorityName="Tessens, Janssen &amp; Artois" authorityYear="2014" box="[151,383,836,861]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="subFamily" subFamily="Phonorhynchoidinae">Phonorhynchoidinae</taxonomicName>
(proboscis sheath without circular muscles, subterminal gonopore combined with a very long genital atrium),
<taxonomicName id="FDB14D53E35EB152E75B90615774FC19" authorityName="Evdonin" authorityYear="1977" box="[351,611,871,896]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="subFamily" subFamily="Psammopolycystidinae">Psammopolycystidinae</taxonomicName>
(syncytial epidermis) and
<taxonomicName id="FDB14D53E35EB152E5B3906151BEFC19" authorityName="Evdonin" authorityYear="1977" box="[951,1193,871,896]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="subFamily" subFamily="Typhlopolycystidinae">Typhlopolycystidinae</taxonomicName>
(unpaired ovarium, enlarged proximal dilators of the proboscis sheath, typical seminal receptacle on the bursal stalk) (for details see
<bibRefCitation id="5E204B21E35EB152E69390B65461FC51" author="Tessens, B. &amp; Janssen, T. &amp; Artois, T." box="[151,374,943,968]" pageId="12" pageNumber="119" pagination="519 - 530" refId="ref12637" refString="Tessens, B., Janssen, T. &amp; Artois, T. (2014) Molecular phylogeny of Kalyptorhynchia (Rhabdocoela, Platyhelminthes) inferred from ribosomal sequence data. Zoologica Scripta, 43 (5), 519 - 530. https: // doi. org / 10.1111 / zsc. 12066" type="journal article" year="2014">
Tessens
<emphasis id="08C5EAC2E35EB152E6FC90B7543AFC51" box="[248,301,943,968]" italics="true" pageId="12" pageNumber="119">et al</emphasis>
. 2014
</bibRefCitation>
and references therein). Therefore, it can be included in
<taxonomicName id="FDB14D53E35EB152E21290A951A9FC51" authorityName="Schockaert &amp; Karling" authorityYear="1970" box="[1046,1214,943,968]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="subFamily" subFamily="Polycystidinae">Polycystidinae</taxonomicName>
, a morphologically heterogeneous group, comprising most of the polycystidid taxa for which at this moment no clear apomorphies can be given (see
<bibRefCitation id="5E204B21E35EB152E73690FE571CFB89" author="Tessens, B. &amp; Janssen, T. &amp; Artois, T." box="[306,523,1015,1040]" pageId="12" pageNumber="119" pagination="519 - 530" refId="ref12637" refString="Tessens, B., Janssen, T. &amp; Artois, T. (2014) Molecular phylogeny of Kalyptorhynchia (Rhabdocoela, Platyhelminthes) inferred from ribosomal sequence data. Zoologica Scripta, 43 (5), 519 - 530. https: // doi. org / 10.1111 / zsc. 12066" type="journal article" year="2014">
Tessens
<emphasis id="08C5EAC2E35EB152E78B90FF54D7FB89" box="[399,448,1015,1040]" italics="true" pageId="12" pageNumber="119">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
<paragraph id="3A0E36D0E35EB152E6C3971A5057FA05" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="119">
The male atrial organs of
<taxonomicName id="FDB14D53E35EB152E7EF971B5796FBAC" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[491,641,1052,1077]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E7EF971B5796FBAC" box="[491,641,1052,1077]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E48D971A5637FBAC" bold="true" box="[649,800,1052,1077]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E48D971A57CFFBAC" box="[649,728,1052,1077]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
include a double-walled prostate stylet
<typeStatus id="E50A8872E35EB152E2E7971B5004FBAC" box="[1251,1299,1053,1077]" pageId="12" pageNumber="119">type</typeStatus>
II, found in several representatives of
<taxonomicName id="FDB14D53E35EB152E7D09739576EFBC1" authorityName="Schockaert &amp; Karling" authorityYear="1970" box="[468,633,1087,1112]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="subFamily" subFamily="Polycystidinae">Polycystidinae</taxonomicName>
(
<taxonomicName id="FDB14D53E35EB152E49697395109FBC1" authority="Karling, 1952" authorityName="Karling" authorityYear="1952" box="[658,1054,1087,1112]" class="Rhabditophora" family="Polycystididae" genus="Antiboreorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E49697395670FBC1" box="[658,871,1087,1112]" italics="true" pageId="12" pageNumber="119">Antiboreorhynchus</emphasis>
<bibRefCitation id="5E204B21E35EB152E57397395109FBC1" author="Karling, T. G." box="[887,1054,1087,1112]" pageId="12" pageNumber="119" pagination="1 - 50" refId="ref11688" refString="Karling, T. G. (1952) Kalyptorhynchia (Turbellaria). Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, 4 (9), 1 - 50." type="journal article" year="1952">Karling, 1952</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FDB14D53E35EB152E2359739508FFBC1" authority="Karling, 1952" authorityName="Karling" authorityYear="1952" box="[1073,1432,1087,1112]" class="Rhabditophora" family="Polycystididae" genus="Austrorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E235973951F6FBC1" box="[1073,1249,1087,1112]" italics="true" pageId="12" pageNumber="119">Austrorhynchus</emphasis>
<bibRefCitation id="5E204B21E35EB152E2F59739508FFBC1" author="Karling, T. G." box="[1265,1432,1087,1112]" pageId="12" pageNumber="119" pagination="1 - 50" refId="ref11688" refString="Karling, T. G. (1952) Kalyptorhynchia (Turbellaria). Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, 4 (9), 1 - 50." type="journal article" year="1952">Karling, 1952</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FDB14D53E35EB152E69397625703FBE5" authority="Evdonin, 1970" authorityName="Evdonin" authorityYear="1970" box="[151,532,1124,1149]" class="Rhabditophora" family="Polycystididae" genus="Cincturorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E69397625477FBE4" box="[151,352,1124,1149]" italics="true" pageId="12" pageNumber="119">Cincturorhynchus</emphasis>
<bibRefCitation id="5E204B21E35EB152E76D97625703FBE5" author="Evdonin, L. A." box="[361,532,1124,1149]" pageId="12" pageNumber="119" pagination="781 - 784" refId="ref11537" refString="Evdonin, L. A. (1970) A new genus of Turbellarians from the family Polycystididae. Zoological Journal, 49 (5), 781 - 784." type="journal article" year="1970">Evdonin, 1970</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FDB14D53E35EB152E426976257EFFBE4" authorityName="Karling &amp; Schockaert" authorityYear="1977" box="[546,760,1124,1149]" class="Rhabditophora" family="Polycystididae" genus="Paraustrorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E426976257EFFBE4" box="[546,760,1124,1149]" italics="true" pageId="12" pageNumber="119">Paraustrorhynchus</emphasis>
</taxonomicName>
,
<taxonomicName id="FDB14D53E35EB152E50D97625150FBE4" authority="Graff, 1905" authorityName="Graff" authorityYear="1905" box="[777,1095,1124,1149]" class="Rhabditophora" family="Polycystididae" genus="Phonorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E50D976256AFFBE4" box="[777,952,1124,1149]" italics="true" pageId="12" pageNumber="119">Phonorhynchus</emphasis>
<bibRefCitation id="5E204B21E35EB152E5C697625150FBE4" author="Graff, L. von" box="[962,1095,1124,1149]" pageId="12" pageNumber="119" pagination="68 - 148" refId="ref11603" refString="Graff, L. von (1905) Marine Turbellarien Orotavas und der Kusten Europas. Zeitschrift fur wissenschaftliche Zoologie, 83, 68 - 148." type="journal article" year="1905">Graff, 1905</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FDB14D53E35EB152E25C9763508FFBE4" authority="Reisinger, 1926" authorityName="Reisinger" authorityYear="1926" box="[1112,1432,1124,1149]" class="Rhabditophora" family="Polycystididae" genus="Porrocystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E25C976351CFFBE5" box="[1112,1240,1125,1148]" italics="true" pageId="12" pageNumber="119">Porrocystis</emphasis>
<bibRefCitation id="5E204B21E35EB152E2E69762508FFBE4" author="Reisinger, E." box="[1250,1432,1124,1149]" pageId="12" pageNumber="119" pagination="417 - 463" refId="ref12254" refString="Reisinger, E. (1926) Zur Turbellarienfauna der Antarktis. Deutsche Sudpolar-Expedition Zoologie, 18 (10), 417 - 463." type="journal article" year="1926">Reisinger, 1926</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FDB14D53E35EB152E693978F54B9FB39" authority="Sekera, 1901" authorityName="Sekera" authorityYear="1901" box="[151,430,1159,1184]" class="Rhabditophora" family="Polycystididae" genus="Progyrator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E693978F5403FB39" box="[151,276,1161,1184]" italics="true" pageId="12" pageNumber="119">Progyrator</emphasis>
<bibRefCitation id="5E204B21E35EB152E71E978154B9FB39" author="Sekera, E." box="[282,430,1159,1184]" pageId="12" pageNumber="119" pagination="79 - 81" refId="ref12612" refString="Sekera, E. (1901) Uber eine marine Art der Gattung Gyrator Ehrb. Zoologischer Anzeiger, 24, 79 - 81." type="journal article" year="1901">Sekera, 1901</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="FDB14D53E35EB152E7ED978156D8FB39" authority="Artois &amp; Schockaert, 1999" authorityName="Artois &amp; Schockaert" authorityYear="1999" box="[489,975,1159,1184]" class="Rhabditophora" family="Polycystididae" genus="Pygmorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E7ED9781578AFB39" box="[489,669,1159,1184]" italics="true" pageId="12" pageNumber="119">Pygmorhynchus</emphasis>
<bibRefCitation id="5E204B21E35EB152E4A0978156D8FB39" author="Artois, T. J. &amp; Schockaert, E. R." box="[676,975,1159,1184]" pageId="12" pageNumber="119" pagination="309 - 324" refId="ref10758" refString="Artois, T. J. &amp; Schockaert, E. R. (1999) Interstitial fauna of the Galapagos: Porrocystidinae (Plathelminthes, Polycystididae). Tropical Zoology, 12, 309 - 324. https: // doi. org / 10.1080 / 03946975.1999.10539397" type="journal article" year="1999">Artois &amp; Schockaert, 1999</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="5E204B21E35EB152E20D97815024FB39" author="Artois, T. J. &amp; Schockaert, E. R." box="[1033,1331,1159,1184]" pageId="12" pageNumber="119" pagination="179 - 190" refId="ref10907" refString="Artois, T. J. &amp; Schockaert, E. R. (2003) Primary homology assessment in the male atrial system of the Polycystididae (Platyhelminthes: Eukalyptorhynchia). Zoologischer Anzeiger, 242 (2), 179 - 190. https: // doi. org / 10.1078 / 0044 - 5231 - 00095" type="journal article" year="2003">Artois &amp; Schockaert 2003</bibRefCitation>
). In these genera the prostate stylet is connected to a prostate vesicle
<typeStatus id="E50A8872E35EB152E53F97AB567CFB5C" box="[827,875,1197,1221]" pageId="12" pageNumber="119">type</typeStatus>
II. However, all these genera (except
<taxonomicName id="FDB14D53E35EB152E31B97AB508BFB5D" authorityName="Sekera" authorityYear="1901" box="[1311,1436,1197,1220]" class="Rhabditophora" family="Polycystididae" genus="Progyrator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E31B97AB508BFB5D" box="[1311,1436,1197,1220]" italics="true" pageId="12" pageNumber="119">Progyrator</emphasis>
</taxonomicName>
and
<taxonomicName id="FDB14D53E35EB152E6C297C9546DFB71" authorityName="Artois &amp; Schockaert" authorityYear="1999" box="[198,378,1231,1256]" class="Rhabditophora" family="Polycystididae" genus="Pygmorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E6C297C9546DFB71" box="[198,378,1231,1256]" italics="true" pageId="12" pageNumber="119">Pygmorhynchus</emphasis>
</taxonomicName>
) include an accessory stylet in the male atrium, a structure lacking in
<taxonomicName id="FDB14D53E35EB152E28D97D75009FB71" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[1161,1310,1231,1256]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E28D97D75009FB71" box="[1161,1310,1231,1256]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E32197C955B1FA94" bold="true" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E32197C95063FB71" box="[1317,1396,1231,1256]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n
</emphasis>
. The prostate vesicle
<typeStatus id="E50A8872E35EB152E79B97F354D8FA94" box="[415,463,1269,1293]" pageId="12" pageNumber="119">type</typeStatus>
II of
<taxonomicName id="FDB14D53E35EB152E41697F357B0FA94" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[530,679,1268,1293]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E41697F357B0FA94" box="[530,679,1268,1293]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E4AB97F2565FFA94" bold="true" box="[687,840,1268,1293]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E4AB97F257E8FA94" box="[687,767,1268,1293]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
contains only an eosinophilic secretion, while in the other genera of the subfamily with this kind of prostate vesicle the secretion is basophilic (
<bibRefCitation id="5E204B21E35EB152E2AB961155C3FACD" author="Artois, T. J. &amp; Schockaert, E. R." pageId="12" pageNumber="119" pagination="179 - 190" refId="ref10907" refString="Artois, T. J. &amp; Schockaert, E. R. (2003) Primary homology assessment in the male atrial system of the Polycystididae (Platyhelminthes: Eukalyptorhynchia). Zoologischer Anzeiger, 242 (2), 179 - 190. https: // doi. org / 10.1078 / 0044 - 5231 - 00095" type="journal article" year="2003">Artois &amp; Schockaert 2003</bibRefCitation>
). Only in species of
<taxonomicName id="FDB14D53E35EB152E7B8963B570BFACD" authorityName="Ehrenberg" authorityYear="1831" box="[444,540,1341,1364]" class="Rhabditophora" family="Polycystididae" genus="Gyratrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E7B8963B570BFACD" box="[444,540,1341,1364]" italics="true" pageId="12" pageNumber="119">Gyratrix</emphasis>
</taxonomicName>
,
<taxonomicName id="FDB14D53E35EB152E428963A564AFACC" authority="Karling, 1955" authorityName="Karling" authorityYear="1955" box="[556,861,1340,1365]" class="Rhabditophora" family="Polycystididae" genus="Gyratricella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E428963A57A0FACC" box="[556,695,1340,1365]" italics="true" pageId="12" pageNumber="119">Gyratricella</emphasis>
<bibRefCitation id="5E204B21E35EB152E4BA963A564AFACC" author="Karling, T. G." box="[702,861,1340,1365]" pageId="12" pageNumber="119" pagination="1 - 39" refId="ref11723" refString="Karling, T. G. (1955) Studien uber Kalyptorhynchien (Turbellaria) 5. Der Verwandtschaftskreis von Gyratrix Ehrenberg. Acta Zoologica Fennica, 88, 1 - 39." type="journal article" year="1955">Karling, 1955</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="FDB14D53E35EB152E59D963B5196FACC" authority="Karling, 1956" authorityName="Karling" authorityYear="1956" box="[921,1153,1340,1365]" class="Rhabditophora" family="Polycystididae" genus="Papia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E59D963B56CBFACD" box="[921,988,1341,1364]" italics="true" pageId="12" pageNumber="119">Papia</emphasis>
<bibRefCitation id="5E204B21E35EB152E5E0963A5196FACC" author="Karling, T. G." box="[996,1153,1340,1365]" pageId="12" pageNumber="119" pagination="187 - 279" refId="ref11756" refString="Karling, T. G. (1956) Morphologisch-histologische Untersuchungen an den mannlichen Atrialorganen der Kalyptorhynchia (Turbellaria). Arkiv for Zoologi, 9 (7), 187 - 279." type="journal article" year="1956">Karling, 1956</bibRefCitation>
</taxonomicName>
does the prostate vesicle
<typeStatus id="E50A8872E35EB152E693966755D0FAE0" box="[151,199,1377,1401]" pageId="12" pageNumber="119">type</typeStatus>
II contain an eosinophilic secretion (
<bibRefCitation id="5E204B21E35EB152E46D96595682FAE1" author="Artois, T. J. &amp; Schockaert, E. R." box="[617,917,1375,1400]" pageId="12" pageNumber="119" pagination="179 - 190" refId="ref10907" refString="Artois, T. J. &amp; Schockaert, E. R. (2003) Primary homology assessment in the male atrial system of the Polycystididae (Platyhelminthes: Eukalyptorhynchia). Zoologischer Anzeiger, 242 (2), 179 - 190. https: // doi. org / 10.1078 / 0044 - 5231 - 00095" type="journal article" year="2003">Artois &amp; Schockaert 2003</bibRefCitation>
), but these taxa differ from
<taxonomicName id="FDB14D53E35EB152E2CA96675074FAE1" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[1230,1379,1375,1400]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E2CA96675074FAE1" box="[1230,1379,1375,1400]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E36F965955E3FA04" bold="true" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E36F965955B9FA04" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
by many other diagnostic characters, as they belong to different subfamilies (
<bibRefCitation id="5E204B21E35EB152E25D96825023FA05" author="Tessens, B. &amp; Janssen, T. &amp; Artois, T." box="[1113,1332,1412,1437]" pageId="12" pageNumber="119" pagination="519 - 530" refId="ref12637" refString="Tessens, B., Janssen, T. &amp; Artois, T. (2014) Molecular phylogeny of Kalyptorhynchia (Rhabdocoela, Platyhelminthes) inferred from ribosomal sequence data. Zoologica Scripta, 43 (5), 519 - 530. https: // doi. org / 10.1111 / zsc. 12066" type="journal article" year="2014">
Tessens
<emphasis id="08C5EAC2E35EB152E2BC968351FEFA04" box="[1208,1257,1412,1437]" italics="true" pageId="12" pageNumber="119">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72AB655BE35EB153E6C396A151AAFEFC" lastPageId="13" lastPageNumber="120" pageId="12" pageNumber="119" type="reference_group">
<paragraph id="3A0E36D0E35EB152E6C396A15077F979" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="119">
<taxonomicName id="FDB14D53E35EB152E6C396A154D9FA59" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[199,462,1447,1472]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E6C396A154D9FA59" box="[199,462,1447,1472]" italics="true" pageId="12" pageNumber="119">Polycydora intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E7DD96A1576BFA59" bold="true" box="[473,636,1447,1472]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E7DD96A1573BFA59" box="[473,556,1447,1472]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
resembles species of
<taxonomicName id="FDB14D53E35EB152E58796A156E2FA59" ID-CoL="7PTJF" authorityName="Kolliker" authorityYear="1845" box="[899,1013,1447,1472]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E58796A156E2FA59" box="[899,1013,1447,1472]" italics="true" pageId="12" pageNumber="119">Polycystis</emphasis>
</taxonomicName>
as well as of
<taxonomicName id="FDB14D53E35EB152E2A196A1500AFA59" ID-CoL="7PQJK" authorityName="Marcus" authorityYear="1948" box="[1189,1309,1447,1472]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E2A196A1500AFA59" box="[1189,1309,1447,1472]" italics="true" pageId="12" pageNumber="119">Paulodora</emphasis>
</taxonomicName>
. However, representatives of these two genera have a prostate stylet
<typeStatus id="E50A8872E35EB152E52896CB564BFA7C" box="[812,860,1485,1509]" pageId="12" pageNumber="119">type</typeStatus>
I and a prostate vesicle
<typeStatus id="E50A8872E35EB152E27396CB51B0FA7C" box="[1143,1191,1485,1509]" pageId="12" pageNumber="119">type</typeStatus>
I. The prostate stylet
<typeStatus id="E50A8872E35EB152E69396F755D0F990" box="[151,199,1521,1545]" pageId="12" pageNumber="119">type</typeStatus>
I in
<taxonomicName id="FDB14D53E35EB152E6FD96E9547CF991" ID-CoL="7PTJF" authorityName="Kolliker" authorityYear="1845" box="[249,363,1519,1544]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E6FD96E9547CF991" box="[249,363,1519,1544]" italics="true" pageId="12" pageNumber="119">Polycystis</emphasis>
</taxonomicName>
is short and wide, with the distal rim of the outer stylet toothed [except in
<taxonomicName id="FDB14D53E35EB152E2A796F7549CF9B4" ID-CoL="4L57F" authority="(Marcus, 1948) Karling, 1952" authorityName="Karling" authorityYear="1952" baseAuthorityName="Marcus" baseAuthorityYear="1948" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="gabriellae">
<emphasis id="08C5EAC2E35EB152E2A796F75026F991" box="[1187,1329,1519,1544]" italics="true" pageId="12" pageNumber="119">P. gabriellae</emphasis>
(Marcus, 1948)
<bibRefCitation id="5E204B21E35EB152E6E09512549CF9B4" author="Karling, T. G." box="[228,395,1556,1581]" pageId="12" pageNumber="119" pagination="1 - 50" refId="ref11688" refString="Karling, T. G. (1952) Kalyptorhynchia (Turbellaria). Further Zoological Results of the Swedish Antarctic Expedition 1901 - 1903, 4 (9), 1 - 50." type="journal article" year="1952">Karling, 1952</bibRefCitation>
</taxonomicName>
]. In
<taxonomicName id="FDB14D53E35EB152E7C09512572BF9B4" ID-CoL="7PQJK" authorityName="Marcus" authorityYear="1948" box="[452,572,1556,1581]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E7C09512572BF9B4" box="[452,572,1556,1581]" italics="true" pageId="12" pageNumber="119">Paulodora</emphasis>
</taxonomicName>
the outer stylet is rather elongated and complex, with folds and gutters, probably to drain the sperm. The prostate stylet of
<taxonomicName id="FDB14D53E35EB152E4ED953F5696F9C9" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[745,897,1591,1616]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E4ED953F5696F9C9" box="[745,897,1591,1616]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E58F95315136F9C9" bold="true" box="[907,1057,1591,1616]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E58F953156CAF9C9" box="[907,989,1591,1616]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n
</emphasis>
. is (probably) of
<typeStatus id="E50A8872E35EB152E2F4953E5037F9C9" box="[1264,1312,1592,1616]" pageId="12" pageNumber="119">type</typeStatus>
II, as it is connected to a typical prostate vesicle of
<typeStatus id="E50A8872E35EB152E487955B57A4F9EC" box="[643,691,1629,1653]" pageId="12" pageNumber="119">type</typeStatus>
II. It is rather long, but does resemble that of some species of
<taxonomicName id="FDB14D53E35EB152E69395795418F901" ID-CoL="7PQJK" authorityName="Marcus" authorityYear="1948" box="[151,271,1663,1688]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E69395795418F901" box="[151,271,1663,1688]" italics="true" pageId="12" pageNumber="119">Paulodora</emphasis>
</taxonomicName>
(e.g.
<taxonomicName id="FDB14D53E35EB152E752958757D8F901" ID-CoL="6TVBD" authority="Schockaert, 1982" authorityName="Schockaert" authorityYear="1982" box="[342,719,1663,1688]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="subcontorta">
<emphasis id="08C5EAC2E35EB152E75295875473F901" box="[342,356,1665,1688]" italics="true" pageId="12" pageNumber="119">P</emphasis>
.
<emphasis id="08C5EAC2E35EB152E772957954EBF901" box="[374,508,1663,1688]" italics="true" pageId="12" pageNumber="119">subcontorta</emphasis>
<bibRefCitation id="5E204B21E35EB152E403957957D8F901" author="Schockaert, E. R." box="[519,719,1663,1688]" pageId="12" pageNumber="119" pagination="81 - 96" refId="ref12404" refString="Schockaert, E. R. (1982) Turbellaria from Somalia 2. Kalyptorhynchia. Part 2. Monitore Zoologico Italiano, 18 (2), 81 - 96. https: // doi. org / 10.1080 / 03749444.1982.10736660" type="journal article" year="1982">Schockaert, 1982</bibRefCitation>
</taxonomicName>
) as to its general construction, i.e. with gutter-like folds, but without further ornamentations. Apart from the above-mentioned differences/resemblances in the construction of the stylet,
<taxonomicName id="FDB14D53E35EB152E70F95CF54B7F979" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[267,416,1735,1760]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E70F95CF54B7F979" box="[267,416,1735,1760]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E7A295C1572CF979" bold="true" box="[422,571,1735,1760]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E7A295C154E1F979" box="[422,502,1735,1760]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
lacks several of the typical, diagnostic features of the other two genera.
</paragraph>
<paragraph id="3A0E36D0E35EB153E6C395EA51AAFEFC" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1437,151,536]" lastPageId="13" lastPageNumber="120" pageId="12" pageNumber="119">
Diagnostic features of species of
<taxonomicName id="FDB14D53E35EB152E43C95EA57BEF89C" ID-CoL="7PTJF" authorityName="Kolliker" authorityYear="1845" box="[568,681,1772,1797]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E43C95EA57BEF89C" box="[568,681,1772,1797]" italics="true" pageId="12" pageNumber="119">Polycystis</emphasis>
</taxonomicName>
lacking in
<taxonomicName id="FDB14D53E35EB152E52095EB56AEF89C" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[804,953,1772,1797]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E52095EB56AEF89C" box="[804,953,1772,1797]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E5BB95EA5145F89C" bold="true" box="[959,1106,1772,1797]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E5BB95EA5119F89C" box="[959,1038,1772,1797]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
are the presence of accessory glands
<typeStatus id="E50A8872E35EB152E6ED9416540EF8B1" box="[233,281,1808,1832]" pageId="12" pageNumber="119">type</typeStatus>
I (also present in species of
<taxonomicName id="FDB14D53E35EB152E46C94095629F8B1" ID-CoL="7PQ7Q" authorityName="Karling &amp; Schockaert" authorityYear="1977" box="[616,830,1807,1832]" class="Rhabditophora" family="Polycystididae" genus="Paraustrorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E46C94095629F8B1" box="[616,830,1807,1832]" italics="true" pageId="12" pageNumber="119">Paraustrorhynchus</emphasis>
</taxonomicName>
) and a large, asymmetrical, mostly excentric muscle bulb on the stalk of the male bursa (Artois &amp; Schokaert 1998, 2003;
<bibRefCitation id="5E204B21E35EB152E5AA94325183F8D4" author="Willems, W. R. &amp; Schockaert, E. R. &amp; Artois, T. J." box="[942,1172,1844,1869]" pageId="12" pageNumber="119" pagination="329 - 355" refId="ref12693" refString="Willems, W. R., Schockaert, E. R. &amp; Artois, T. J. (2006) Report on the Polycystididae (Rhabdocoela, Kalyptorhynchia) from Australia, with the description of twelve new species and six new genera. Hydrobiologia, 563, 329 - 355. https: // doi. org / 10.1007 / s 10750 - 006 - 0041 - 3" type="journal article" year="2006">
Willems
<emphasis id="08C5EAC2E35EB152E2119433515FF8D4" box="[1045,1096,1844,1869]" italics="true" pageId="12" pageNumber="119">et al</emphasis>
. 2006
</bibRefCitation>
). The stalk of the male bursa is heavily muscular in
<taxonomicName id="FDB14D53E35EB152E7F2945F5786F8E9" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[502,657,1879,1904]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E7F2945F5786F8E9" box="[502,657,1879,1904]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E49A94515652F8E9" bold="true" box="[670,837,1879,1904]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E49A945157E4F8E9" box="[670,755,1879,1904]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
, but it is never asymmetrical, let alone excentric. Furthermore, the tick syncytial epithelium surrounding the internal vesicle of the male bursa in
<taxonomicName id="FDB14D53E35EB152E2CA947B5074F80C" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[1230,1379,1916,1941]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E2CA947B5074F80C" box="[1230,1379,1916,1941]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E36F947A55F8F820" bold="true" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E36F947A55B9F820" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n
</emphasis>
. is not found in any species of
<taxonomicName id="FDB14D53E35EB152E45894A657C3F820" ID-CoL="7PQJK" authorityName="Marcus" authorityYear="1948" box="[604,724,1952,1977]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E45894A657C3F820" box="[604,724,1952,1977]" italics="true" pageId="12" pageNumber="119">Paulodora</emphasis>
</taxonomicName>
or
<taxonomicName id="FDB14D53E35EB152E4F894A65679F820" ID-CoL="7PTJF" authorityName="Kolliker" authorityYear="1845" box="[764,878,1952,1977]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E4F894A65679F820" box="[764,878,1952,1977]" italics="true" pageId="12" pageNumber="119">Polycystis</emphasis>
</taxonomicName>
. Moreover, most of the species of
<taxonomicName id="FDB14D53E35EB152E2FB94A65066F820" ID-CoL="7PTJF" authorityName="Kolliker" authorityYear="1845" box="[1279,1393,1952,1977]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35EB152E2FB94A65066F820" box="[1279,1393,1952,1977]" italics="true" pageId="12" pageNumber="119">Polycystis</emphasis>
</taxonomicName>
are darkly pigmented (
<bibRefCitation id="5E204B21E35EB152E76D94C25784F845" author="Artois, T. J. &amp; Schockaert, E. R." box="[361,659,1988,2013]" pageId="12" pageNumber="119" pagination="309 - 324" refId="ref10758" refString="Artois, T. J. &amp; Schockaert, E. R. (1999) Interstitial fauna of the Galapagos: Porrocystidinae (Plathelminthes, Polycystididae). Tropical Zoology, 12, 309 - 324. https: // doi. org / 10.1080 / 03946975.1999.10539397" type="journal article" year="1999">Artois &amp; Schockaert 1999</bibRefCitation>
), which is not the case in
<taxonomicName id="FDB14D53E35EB152E5B594C35151F844" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[945,1094,1988,2013]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="12" pageNumber="119" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35EB152E5B594C35151F844" box="[945,1094,1988,2013]" italics="true" pageId="12" pageNumber="119">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35EB152E24894C251F7F844" bold="true" box="[1100,1248,1988,2013]" pageId="12" pageNumber="119">
<taxonomicNameLabel id="13F657B9E35EB152E24894C2518DF844" box="[1100,1178,1988,2013]" pageId="12" pageNumber="119" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
. Most species of
<taxonomicName id="FDB14D53E35FB153E69393915418FF29" ID-CoL="7PQJK" authorityName="Marcus" authorityYear="1948" box="[151,271,151,176]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35FB153E69393915418FF29" box="[151,271,151,176]" italics="true" pageId="13" pageNumber="120">Paulodora</emphasis>
</taxonomicName>
typically have elongated, kidney-shaped ovaries (see
<bibRefCitation id="5E204B21E35FB153E58B939151B6FF29" author="Artois, T. J. &amp; Tessens, B. S." box="[911,1185,151,176]" pageId="13" pageNumber="120" pagination="1 - 27" refId="ref11031" refString="Artois, T. J. &amp; Tessens, B. S. (2008) Polycystididae (Rhabditophora: Rhabdocoela: Kalyptorhynchia) from the Indian Ocean, with the description of twelve species. Zootaxa, 1849, 1 - 27." type="journal article" year="2008">Artois &amp; Tessens 2008</bibRefCitation>
), as is the case in
<taxonomicName id="FDB14D53E35FB153E383939F5407FF4C" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35FB153E383939F5407FF4C" italics="true" pageId="13" pageNumber="120">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35FB153E71993BA54D5FF4C" bold="true" box="[285,450,188,213]" pageId="13" pageNumber="120">
<taxonomicNameLabel id="13F657B9E35FB153E71993BA5465FF4C" box="[285,370,188,213]" pageId="13" pageNumber="120" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
, but
<taxonomicName id="FDB14D53E35FB153E40293BB57B6FF4C" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[518,673,188,213]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35FB153E40293BB57B6FF4C" box="[518,673,188,213]" italics="true" pageId="13" pageNumber="120">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35FB153E4A993BA5644FF4C" bold="true" box="[685,851,188,213]" pageId="13" pageNumber="120">
<taxonomicNameLabel id="13F657B9E35FB153E4A993BA5615FF4C" box="[685,770,188,213]" pageId="13" pageNumber="120" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
lacks the hard, umbrella-shaped &quot;nozzles&quot; at the connection between male bursa and each ovary that are so typical of species of
<taxonomicName id="FDB14D53E35FB153E22C93D955CCFE85" ID-CoL="7PQJK" authority="(Artois &amp; Schockaert 1998)" authorityName="Artois &amp; Schockaert" authorityYear="1998" baseAuthorityName="Artois &amp; Schockaert" baseAuthorityYear="1998" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35FB153E22C93D951B7FF61" box="[1064,1184,223,248]" italics="true" pageId="13" pageNumber="120">Paulodora</emphasis>
(Artois &amp; Schockaert 1998)
</taxonomicName>
. Moreover, in species of
<taxonomicName id="FDB14D53E35FB153E4159202579EFE84" ID-CoL="7PQJK" baseAuthorityName="Artois &amp; Schockaert" baseAuthorityYear="1998" box="[529,649,260,285]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35FB153E4159202579EFE84" box="[529,649,260,285]" italics="true" pageId="13" pageNumber="120">Paulodora</emphasis>
</taxonomicName>
the oviducts (mostly) merge with the male bursal tissue, and the ejaculatory duct makes a 270° turn around the prostate vesicle before it ends in the male atrium next to the stylet. In
<taxonomicName id="FDB14D53E35FB153E693924B543CFEFC" ID-CoL="4L4Y4" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[151,299,332,357]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35FB153E693924B543CFEFC" box="[151,299,332,357]" italics="true" pageId="13" pageNumber="120">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35FB153E737924A54DFFEFC" bold="true" box="[307,456,332,357]" pageId="13" pageNumber="120">
<taxonomicNameLabel id="13F657B9E35FB153E737924A5495FEFC" box="[307,386,332,357]" pageId="13" pageNumber="120" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
, the male bursal stalk is free, and the ejaculatory duct runs straight.
</paragraph>
</subSubSection>
<subSubSection id="72AB655BE35FB153E6C392765148FD81" pageId="13" pageNumber="120" type="description">
<paragraph id="3A0E36D0E35FB153E6C3927651A0FE49" blockId="13.[151,1437,151,536]" pageId="13" pageNumber="120">
A typical feature of
<taxonomicName id="FDB14D53E35FB153E7A29277572DFE11" authority="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois, 2018" authorityName="Diez &amp; Hernández &amp; Reygel &amp; Roosen &amp; Artois" authorityYear="2018" box="[422,570,367,392]" class="Rhabditophora" family="Polycystididae" genus="Polycydora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="species" species="intermedia" status="gen. nov.">
<emphasis id="08C5EAC2E35FB153E7A29277572DFE11" box="[422,570,367,392]" italics="true" pageId="13" pageNumber="120">P. intermedia</emphasis>
</taxonomicName>
<emphasis id="08C5EAC2E35FB153E444926957C4FE11" bold="true" box="[576,723,367,392]" pageId="13" pageNumber="120">
<taxonomicNameLabel id="13F657B9E35FB153E44492695798FE11" box="[576,655,367,392]" pageId="13" pageNumber="120" rank="genus">gen. n.</taxonomicNameLabel>
sp. n.
</emphasis>
is the presence of two bursae, a male and a female one, with the female bursa being large and with a long stalk. In species of
<taxonomicName id="FDB14D53E35FB153E563929256C8FE34" baseAuthorityName="Artois &amp; Schockaert" baseAuthorityYear="1998" box="[871,991,404,429]" class="Rhabditophora" family="Polycystididae" genus="Paulodora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35FB153E563929256C8FE34" box="[871,991,404,429]" italics="true" pageId="13" pageNumber="120">Paulodora</emphasis>
</taxonomicName>
a female bursa is lacking, whereas in species of
<taxonomicName id="FDB14D53E35FB153E70992B15468FE49" authorityName="Kolliker" authorityYear="1845" box="[269,383,439,464]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="genus">
<emphasis id="08C5EAC2E35FB153E70992B15468FE49" box="[269,383,439,464]" italics="true" pageId="13" pageNumber="120">Polycystis</emphasis>
</taxonomicName>
it is very small (or absent, as in
<taxonomicName id="FDB14D53E35FB153E4EE92BF566FFE49" authorityName="Karling" authorityYear="1952" baseAuthorityName="Marcus" baseAuthorityYear="1948" box="[746,888,439,464]" class="Rhabditophora" family="Polycystididae" genus="Polycystis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="species" species="gabriellae">
<emphasis id="08C5EAC2E35FB153E4EE92BF566FFE49" box="[746,888,439,464]" italics="true" pageId="13" pageNumber="120">P. gabriellae</emphasis>
</taxonomicName>
) and almost without a stalk.
</paragraph>
<paragraph id="3A0E36D0E35FB153E6C392DB5148FD81" blockId="13.[151,1437,151,536]" pageId="13" pageNumber="120">
Because of the issues discussed above, the new species cannot unambiguously be placed in any of the existing genera within
<taxonomicName id="FDB14D53E35FB153E73392F954C8FD81" authorityName="Graff" authorityYear="1905" box="[311,479,511,536]" class="Rhabditophora" family="Polycystididae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="13" pageNumber="120" phylum="Platyhelminthes" rank="family">Polycystididae</taxonomicName>
, and, therefore, the erection of a new genus is warranted.
</paragraph>
</subSubSection>
</treatment>
</document>