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<document id="A11FA3AA084563803C294B304637C670" ID-DOI="10.24199/j.mmv.2018.77.04" ID-ISSN="1447-2554" ID-Zenodo-Dep="12208545" ID-ZooBank="urn:lsid:zoobank.org:pub:22334107-0784-466E-8288-D6E29F87F6E2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718989371731" checkinUser="felipe" docAuthor="Melville, Jane, Ritchie, Euan G., Chapple, Stephanie N. J., Glor, Richard E. &amp; Schulte, James A." docDate="2018" docId="942187EC4B6AFF8592A8F8CCC663FEC4" docLanguage="en" docName="MemMusVictoria.77.41-61.pdf" docOrigin="Memoirs of Museum Victoria (Mem. Mus. Vic.) 77" docSource="http://dx.doi.org/10.24199/j.mmv.2018.77.04" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Tropicagama Melville, Ritchie, Chapple, Glor &amp; Schulte, 2018, gen. nov." docType="treatment" docUuid="F534B4D5-CBD7-41E0-950A-B95B14F1D858" docUuidSource="ZooBank" docVersion="2" lastPageNumber="57" masterDocId="6818FF944B65FF95923BFFD9C358FFAA" masterDocTitle="Diversity in Australias tropical savannas: An integrative taxonomic revision of agamid lizards from the genera Amphibolurus and Lophognathus (Lacertilia: Agamidae)" masterLastPageNumber="61" masterPageNumber="41" pageNumber="56" updateTime="1718992566665" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="363794FFC7432AA77F02C162116057C2">Diversity in Australias tropical savannas: An integrative taxonomic revision of agamid lizards from the genera Amphibolurus and Lophognathus (Lacertilia: Agamidae)</mods:title>
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<mods:namePart id="892EBF07D3DA3F8876EB8BEA8435DA82">Melville, Jane</mods:namePart>
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<mods:namePart id="480C066CD4AF5B1C9ACB0EBFE0F3101C">Ritchie, Euan G.</mods:namePart>
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<mods:namePart id="0CE93DB381021DB22941B86B6FF69D0B">Chapple, Stephanie N. J.</mods:namePart>
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<mods:namePart id="4290B93EB4AB631DD225096C754B8CFE">Glor, Richard E.</mods:namePart>
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<mods:namePart id="87F212D0ADDDDA4F9F11C65E48978A0D">Schulte, James A.</mods:namePart>
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<paragraph id="1C3736FA4B6AFF9A92A8F8CCC295F880" blockId="15.[147,461,1813,1835]" box="[147,461,1813,1835]" pageId="15" pageNumber="56">
<heading id="477F81964B6AFF9A92A8F8CCC295F880" bold="true" box="[147,461,1813,1835]" fontSize="9" level="2" pageId="15" pageNumber="56" reason="2">
<emphasis id="2EFCEAE84B6AFF9A92A8F8CCC295F880" bold="true" box="[147,461,1813,1835]" pageId="15" pageNumber="56">
Genus
<taxonomicName id="DB884D794B6AFF9A92E5F8CCC231F880" ID-CoL="8GRSN" ID-ENA="2849038" authorityName="Melville &amp; Ritchie &amp; Chapple &amp; Glor &amp; Schulte" authorityYear="2018" box="[222,361,1813,1834]" class="Reptilia" family="Agamidae" genus="Tropicagama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="56" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="2EFCEAE84B6AFF9A92E5F8CCC231F880" bold="true" box="[222,361,1813,1834]" italics="true" pageId="15" pageNumber="56">Tropicagama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35CF57934B6AFF9A934BF8CCC295F880" box="[368,461,1813,1834]" pageId="15" pageNumber="56" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1C3736FA4B6AFF9A9282F899C052F8F9" blockId="15.[147,778,1856,1902]" box="[185,778,1856,1876]" pageId="15" pageNumber="56">ZooBank LSID: urn:lsid:zoobank.org:act: F534B4D5-CBD7-</paragraph>
</subSubSection>
<subSubSection id="549265714B6AFF8592A8F882C663FEC4" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56" type="description">
<paragraph id="1C3736FA4B6AFF9A92A8F882C2C1F8C4" blockId="15.[147,778,1856,1902]" box="[147,409,1883,1902]" pageId="15" pageNumber="56">41E0-950A-B95B14F1D858.</paragraph>
<paragraph id="1C3736FA4B6AFF9A916CFF3FC716FEC8" blockId="15.[817,1449,229,354]" pageId="15" pageNumber="56">
<emphasis id="2EFCEAE84B6AFF9A916CFF3FC67AFF52" box="[855,1314,229,248]" italics="true" pageId="15" pageNumber="56">
Type-species.
<taxonomicName id="DB884D794B6AFF9A91E1FF3CC784FF52" authorityName="Gunther" authorityYear="1867" box="[986,1244,229,248]" class="Reptilia" family="Agamidae" genus="Grammatophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="temporalis">Grammatophora temporalis</taxonomicName>
(part.)
</emphasis>
<materialsCitation id="ACE03CA74B6AFF9A9714FF3CC711FEC8" collectionCode="BMNH" collectorName="Gunther, A." location="Magazine of Natural History" pageId="15" pageNumber="56" specimenCode="BMNH 1946.8" specimenCount="1" typeStatus="syntype">
<collectorName id="B17D532C4B6AFF9A9714FF3CC6FCFF52" box="[1327,1444,229,248]" pageId="15" pageNumber="56">Günther, A.</collectorName>
, 1867. Additions to the knowledge of Australian reptiles and fishes.
<emphasis id="2EFCEAE84B6AFF9A910AFEC3C7C5FE87" box="[817,1181,282,301]" italics="true" pageId="15" pageNumber="56">
Annals and
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</emphasis>
20(3): 4568 [52] [one of the original
<typeStatus id="C33388584B6AFF9A9144FEECC09EFEE2" box="[895,966,309,328]" pageId="15" pageNumber="56" type="syntype">syntype</typeStatus>
series,
<specimenCode id="4C2E9E814B6AFF9A9630FEECC7CBFEED" box="[1035,1171,308,328]" collectionCode="BMNH" pageId="15" pageNumber="56">BMNH 1946.8</specimenCode>
.12.73 (Nickol Bay), represents
<taxonomicName id="DB884D794B6AFF9A910AFE96C0A0FEC8" authorityName="Melville &amp; Ritchie &amp; Chapple &amp; Glor &amp; Schulte" authorityYear="2018" box="[817,1016,335,354]" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="horneri" status="sp. nov.">
<emphasis id="2EFCEAE84B6AFF9A910AFE96C0A0FEC8" box="[817,1016,335,354]" italics="true" pageId="15" pageNumber="56">Lophognathus horneri</emphasis>
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<taxonomicNameLabel id="35CF57934B6AFF9A91C6FE96C71DFEC8" box="[1021,1093,335,354]" pageId="15" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
]
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.
</paragraph>
<paragraph id="1C3736FA4B6AFF9A910AFEA3C712FCEE" blockId="15.[815,1449,377,837]" pageId="15" pageNumber="56">
<emphasis id="2EFCEAE84B6AFF9A910AFEA3C0C5FE25" box="[817,925,378,399]" italics="true" pageId="15" pageNumber="56">Diagnosis.</emphasis>
A monotypic genus consisting of a large agamid lizard in the subfamily
<taxonomicName id="DB884D794B6AFF9A966FFE4EC7A5FE07" box="[1108,1277,407,429]" pageId="15" pageNumber="56" rank="subFamily" subFamily="Amphibolurinae">Amphibolurinae</taxonomicName>
, with exposed tympanum, gular scales smooth to weakly keeled, ventral scales smooth to weakly keeled. Very long-limbed, prominent erectable nuchal crest. Long tail and head relatively narrow for length. Dorsal scales uniform, with keels converging posteriorly toward midline. Prominent pale dorsolateral stripes that are broadly continuous with wide pale stripe along upper and lower jaw. Lacks well-defined pale stripe between eye and ear. Upper portion of head usually dark grey or black and uniformly coloured. Under the head, on the chin, gular and neck areas, there is dark grey or black uniform pigmentation in adult males, with two narrow white stripes extending from the back of the jaw anteriorly under the chin, parallel to the jaw, ending approximately half way along the jaw. Femoral pores 16; preanal pores 2 (range 13).
</paragraph>
<paragraph id="1C3736FA4B6AFF9A910AFC82C0B1FC24" blockId="15.[817,1448,857,910]" pageId="15" pageNumber="56">
<emphasis id="2EFCEAE84B6AFF9A910AFC82C6F0FCDB" box="[817,1448,857,881]" italics="true" pageId="15" pageNumber="56">
Included species.
<taxonomicName id="DB884D794B6AFF9A963DFC80C611FCDB" authorityName="Gunther" authorityYear="1867" box="[1030,1353,857,881]" class="Reptilia" family="Agamidae" genus="Grammatophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="temporalis">Grammatophora temporalis</taxonomicName>
(part.)
</emphasis>
Günther, A., 1867.
</paragraph>
<paragraph id="1C3736FA4B6AFF9A910AFC7CC662FBB8" blockId="15.[816,1448,933,1042]" pageId="15" pageNumber="56">
<emphasis id="2EFCEAE84B6AFF9A910AFC7CC0E9FC10" box="[817,945,933,954]" italics="true" pageId="15" pageNumber="56">Distribution.</emphasis>
Far northern Australian coastal regions in the Northern Territory and western Cape York. Also occurs on northern offshore islands, including Indonesian islands close to Australian waters and southern
<collectingCountry id="649F766A4B6AFF9A964AFC24C66EFBB8" box="[1137,1334,1020,1042]" name="Papua New Guinea" pageId="15" pageNumber="56">Papua New Guinea</collectingCountry>
.
</paragraph>
<paragraph id="1C3736FA4B6AFF9A910AFBE1C772FBE7" blockId="15.[817,1066,1080,1101]" box="[817,1066,1080,1101]" pageId="15" pageNumber="56">
<taxonomicName id="DB884D794B6AFF9A910AFBE1C772FBE7" authorityName="Melville &amp; Ritchie &amp; Chapple &amp; Glor &amp; Schulte" authorityYear="2018" box="[817,1066,1080,1101]" class="Reptilia" family="Agamidae" genus="Tropicagama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="temporalis">
<emphasis id="2EFCEAE84B6AFF9A910AFBE1C772FBE7" box="[817,1066,1080,1101]" italics="true" pageId="15" pageNumber="56">Tropicagama temporalis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="1C3736FA4B6AFF9A910AFBBBC0E2FBD0" blockId="15.[817,954,1122,1146]" box="[817,954,1122,1146]" pageId="15" pageNumber="56">
<emphasis id="2EFCEAE84B6AFF9A910AFBBBC0E2FBD0" box="[817,954,1122,1146]" italics="true" pageId="15" pageNumber="56">
(
<figureCitation id="84B32A7F4B6AFF9A9102FBBBC0DBFBD3" box="[825,899,1122,1145]" captionStart="Figure 5" captionStartId="9.[149,205,1574,1593]" captionTargetBox="[152,772,231,1554]" captionTargetId="figure-313@9.[147,779,227,1555]" captionTargetPageId="9" captionText="Figure 5. Primary type specimens:a,Amphibolurus centralis (MCZ 35207); b, Gowidon longirostris (BMNH 1946.8.28.73); c, Lophognathus gilbert (BMNH1946.8.28.69); d,Tropicagama temporalis (BMNH 1946.8.28.72)." figureDoi="http://doi.org/10.5281/zenodo.12208559" httpUri="https://zenodo.org/record/12208559/files/figure.png" pageId="15" pageNumber="56">Figs. 5</figureCitation>
&amp;
<figureCitation id="84B32A7F4B6AFF9A919FFBBBC0EAFBD0" box="[932,946,1122,1146]" captionStart="Figure 8" captionStartId="11.[149,209,1777,1796]" captionTargetBox="[176,1398,244,1677]" captionTargetId="figure-15@11.[147,1447,227,1718]" captionTargetPageId="11" captionText="Figure 8. Photos in life of species under revision: a, Lophognathus horneri sp. nov., adult male with breeding colouration, 80 mile beach, Western Australia (photo: R. Glor); b, Lophognathus gilberti, Katherine, Northern Territory (photo: R. Glor); c, Amphibolurus centralis, adult male with breeding colouration, West MacDonnell Ranges, Northern Territory (photo: J. Melville); d, Amphibolurus burnsi, adult male with breeding colouration, Westmar, Queensland (photo: S. Wilson); e, Tropicagama temporalis, Jabiru, Northern Territory (photo: S. Wilson); f, Gowidon longirostris, adult male with breeding colouration, Ormiston Gorge, Northern Territory (photo: R. Glor)." figureDoi="http://doi.org/10.5281/zenodo.12208563" httpUri="https://zenodo.org/record/12208563/files/figure.png" pageId="15" pageNumber="56">8</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="1C3736FA4B6AFF9A916CFB56C70DFAA7" blockId="15.[816,1450,1167,1902]" pageId="15" pageNumber="56">
<taxonomicName id="DB884D794B6AFF9A916CFB56C702FB08" authorityName="Gunther" authorityYear="1867" box="[855,1114,1167,1186]" class="Reptilia" family="Agamidae" genus="Grammatophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="temporalis">
<emphasis id="2EFCEAE84B6AFF9A916CFB56C702FB08" box="[855,1114,1167,1186]" italics="true" pageId="15" pageNumber="56">Grammatophora temporalis</emphasis>
</taxonomicName>
(part.)
<materialsCitation id="ACE03CA74B6AFF9A969AFB56C709FAA7" collectionCode="BMNH" collectorName="Gunther, A." location="Magazine of Natural History" pageId="15" pageNumber="56" specimenCode="BMNH 1946.8" specimenCount="1" typeStatus="syntype">
<collectorName id="B17D532C4B6AFF9A969AFB56C657FB08" box="[1185,1295,1167,1186]" pageId="15" pageNumber="56">Günther, A.</collectorName>
, 1867. Additions to the knowledge of Australian reptiles and fishes.
<emphasis id="2EFCEAE84B6AFF9A970CFB73C719FB7D" italics="true" pageId="15" pageNumber="56">
Annals and
<location id="195760214B6AFF9A910AFB1CC719FB7D" LSID="urn:lsid:plazi:treatment:942187EC4B6AFF8592A8F8CCC663FEC4:195760214B6AFF9A910AFB1CC719FB7D" box="[817,1089,1220,1239]" name="Magazine of Natural History" pageId="15" pageNumber="56">Magazine of Natural History</location>
</emphasis>
20(3): 4568 [52] [one of the original
<typeStatus id="C33388584B6AFF9A910AFB06C020FB58" box="[817,888,1247,1266]" pageId="15" pageNumber="56" type="syntype">syntype</typeStatus>
series,
<specimenCode id="4C2E9E814B6AFF9A91ECFB06C734FB58" box="[983,1132,1247,1266]" collectionCode="BMNH" pageId="15" pageNumber="56">BMNH 1946.8</specimenCode>
.12.73 (Nickol Bay), represents
<taxonomicName id="DB884D794B6AFF9A910AFB20C0A7FAA6" authorityName="Melville &amp; Ritchie &amp; Chapple &amp; Glor &amp; Schulte" authorityYear="2018" box="[817,1023,1273,1292]" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="horneri" status="sp. nov.">
<emphasis id="2EFCEAE84B6AFF9A910AFB20C0A7FAA6" box="[817,1023,1273,1292]" italics="true" pageId="15" pageNumber="56">Lophognathus horneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35CF57934B6AFF9A963EFB23C715FAA7" box="[1029,1101,1274,1293]" pageId="15" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
]
</materialsCitation>
.
</paragraph>
<paragraph id="1C3736FA4B6AFF9A916CFACDC0F2FA3B" blockId="15.[816,1450,1167,1902]" pageId="15" pageNumber="56">
<taxonomicName id="DB884D794B6AFF9A916CFACDC7B0FA8D" authority="Macleay, W., 1877" authorityName="Macleay, W." authorityYear="1877" box="[855,1256,1300,1319]" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="2EFCEAE84B6AFF9A916CFACDC773FA8D" box="[855,1067,1300,1319]" italics="true" pageId="15" pageNumber="56">Lophognathus lateralis</emphasis>
<materialsCitation id="ACE03CA74B6AFF9A960DFACDC7B0FA8D" box="[1078,1256,1300,1319]" collectionCode="AM" country="Papua New Guinea" county="Binaturi River" location="Linnean Society of New South Wales" municipality="Mawatta" pageId="15" pageNumber="56" specimenCode="R31882" specimenCount="1" typeStatus="holotype">Macleay, W., 1877</materialsCitation>
</taxonomicName>
. The lizards of the Chevert Expedition. Second paper.
<emphasis id="2EFCEAE84B6AFF9A964BFAF7C0BEFAF6" italics="true" pageId="15" pageNumber="56">
Proceedings of the
<location id="195760214B6AFF9A9722FAF6C0BEFAF6" LSID="urn:lsid:plazi:treatment:942187EC4B6AFF8592A8F8CCC663FEC4:195760214B6AFF9A9722FAF6C0BEFAF6" country="Papua New Guinea" county="Binaturi River" municipality="Mawatta" name="Linnean Society of New South Wales" pageId="15" pageNumber="56">Linnean Society of New South Wales</location>
</emphasis>
2: 97104 [1878 on title page] [103]. Type data.
<typeStatus id="C33388584B6AFF9A910AFABAC0DDFADC" box="[817,901,1379,1398]" pageId="15" pageNumber="56" type="holotype">Holotype</typeStatus>
AM
<specimenCode id="4C2E9E814B6AFF9A9187FABAC75CFADC" box="[956,1028,1379,1398]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="56">R31882</specimenCode>
,
<collectingMunicipality id="FC53AC804B6AFF9A962BFABDC739FADD" box="[1040,1121,1380,1399]" pageId="15" pageNumber="56">Mawatta</collectingMunicipality>
,
<collectingCounty id="F5564E764B6AFF9A9655FABAC7ADFADC" box="[1134,1269,1379,1398]" pageId="15" pageNumber="56">Binaturi River</collectingCounty>
(as Katow),
<collectingCountry id="649F766A4B6AFF9A974BFABDC0FDFA3B" name="Papua New Guinea" pageId="15" pageNumber="56">Papua New Guinea</collectingCountry>
.
</paragraph>
<paragraph id="1C3736FA4B6AFF9A916CFA41C675FA51" blockId="15.[816,1450,1167,1902]" pageId="15" pageNumber="56">
<taxonomicName id="DB884D794B6AFF9A916CFA41C65FFA01" authority="Boulenger, G. A., 1883" authorityName="Boulenger, G. A." authorityYear="1883" box="[855,1287,1432,1451]" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="labialis">
<emphasis id="2EFCEAE84B6AFF9A916CFA41C779FA01" box="[855,1057,1432,1451]" italics="true" pageId="15" pageNumber="56">Lophognathus labialis</emphasis>
<materialsCitation id="ACE03CA74B6AFF9A9611FA41C65FFA01" box="[1066,1287,1432,1451]" collectionCode="BMNH" county="Magazine of Natural History" location="Remarks" municipality="Annals" pageId="15" pageNumber="56" specimenCode="BMNH 1946.8" specimenCount="1" typeStatus="syntype">Boulenger, G. A., 1883</materialsCitation>
</taxonomicName>
.
<location id="195760214B6AFF9A9729FA41C63BFA01" LSID="urn:lsid:plazi:treatment:942187EC4B6AFF8592A8F8CCC663FEC4:195760214B6AFF9A9729FA41C63BFA01" box="[1298,1379,1432,1451]" county="Magazine of Natural History" municipality="Annals" name="Remarks" pageId="15" pageNumber="56">Remarks</location>
on the lizards of the genus
<taxonomicName id="DB884D794B6AFF9A91CBFA6AC737FA6C" box="[1008,1135,1459,1478]" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="2EFCEAE84B6AFF9A91CBFA6AC737FA6C" box="[1008,1135,1459,1478]" italics="true" pageId="15" pageNumber="56">Lophognathus</emphasis>
</taxonomicName>
.
<emphasis id="2EFCEAE84B6AFF9A9646FA6AC02CFA4A" italics="true" pageId="15" pageNumber="56">
<collectingMunicipality id="FC53AC804B6AFF9A9646FA6AC7E3FA6C" box="[1149,1211,1459,1478]" pageId="15" pageNumber="56">Annals</collectingMunicipality>
and
<collectingCounty id="F5564E764B6AFF9A96D0FA6AC02CFA4A" pageId="15" pageNumber="56">Magazine of Natural History</collectingCounty>
</emphasis>
5 12: 225-226 [225]. Type data.
<typeStatus id="C33388584B6AFF9A96CAFA14C665FA4A" box="[1265,1341,1485,1504]" pageId="15" pageNumber="56" type="syntype">Syntype</typeStatus>
(s)
<specimenCode id="4C2E9E814B6AFF9A9758FA17C028FA51" collectionCode="BMNH" pageId="15" pageNumber="56">BMNH 1946.8</specimenCode>
.
<materialsCitation id="ACE03CA74B6AFF9A9148FA31C7AAFA51" box="[883,1266,1512,1531]" collectionCode="BMNH" collectorName="Port Essington, NT" location="Port Essington" pageId="15" pageNumber="56" specimenCode="BMNH 1946.8" specimenCount="1" typeStatus="syntype">
12.72,
<collectorName id="B17D532C4B6AFF9A918BFA31C738FA51" box="[944,1120,1512,1531]" pageId="15" pageNumber="56">
<location id="195760214B6AFF9A918BFA31C76FFA51" LSID="urn:lsid:plazi:treatment:942187EC4B6AFF8592A8F8CCC663FEC4:195760214B6AFF9A918BFA31C76FFA51" box="[944,1079,1512,1531]" name="Port Essington" pageId="15" pageNumber="56">Port Essington</location>
, NT
</collectorName>
;
<specimenCode id="4C2E9E814B6AFF9A9652FA31C7AAFA51" box="[1129,1266,1512,1531]" collectionCode="BMNH" pageId="15" pageNumber="56">BMNH 1946.8</specimenCode>
</materialsCitation>
.12.63.
</paragraph>
<paragraph id="1C3736FA4B6AFF9A916CF9DBC0C8F9D5" blockId="15.[816,1450,1167,1902]" pageId="15" pageNumber="56">
<taxonomicName id="DB884D794B6AFF9A916CF9DBC66AF9BF" authority="Boulenger, G. A., 1883" authorityName="Boulenger, G. A." authorityYear="1883" box="[855,1330,1538,1557]" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="species" species="maculilabris">
<emphasis id="2EFCEAE84B6AFF9A916CF9DBC70EF9BF" box="[855,1110,1538,1557]" italics="true" pageId="15" pageNumber="56">Lophognathus maculilabris</emphasis>
<materialsCitation id="ACE03CA74B6AFF9A9666F9DBC66AF9BF" box="[1117,1330,1538,1557]" collectionCode="BMNH" country="Indonesia" location="Magazine of Natural History" municipality="Timor Laut Islands" pageId="15" pageNumber="56" specimenCount="1" typeStatus="syntype">Boulenger, G. A., 1883</materialsCitation>
</taxonomicName>
. Remarks on the lizards of the genus
<taxonomicName id="DB884D794B6AFF9A961FF9C4C7FFF99A" box="[1060,1191,1565,1584]" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="genus">
<emphasis id="2EFCEAE84B6AFF9A961FF9C4C7FFF99A" box="[1060,1191,1565,1584]" italics="true" pageId="15" pageNumber="56">Lophognathus</emphasis>
</taxonomicName>
.
<emphasis id="2EFCEAE84B6AFF9A9683F9C4C09CF9E0" italics="true" pageId="15" pageNumber="56">
Annals and
<location id="195760214B6AFF9A970AF9C4C09CF9E0" LSID="urn:lsid:plazi:treatment:942187EC4B6AFF8592A8F8CCC663FEC4:195760214B6AFF9A970AF9C4C09CF9E0" country="Indonesia" municipality="Timor Laut Islands" name="Magazine of Natural History" pageId="15" pageNumber="56">Magazine of Natural History</location>
</emphasis>
12(5): 225226 [226]. Type data.
<typeStatus id="C33388584B6AFF9A9738F9EEC602F9E0" box="[1283,1370,1591,1610]" pageId="15" pageNumber="56" type="syntype">Syntypes</typeStatus>
: BMNH 1946.8.28.70-71 (BMNH 1919.8.26.13-14),
<collectingMunicipality id="FC53AC804B6AFF9A96E5F98BC6FDF9CF" box="[1246,1445,1618,1637]" pageId="15" pageNumber="56">Timor Laut Islands</collectingMunicipality>
,
<collectingCountry id="649F766A4B6AFF9A910AF9B5C0D4F9D5" box="[817,908,1644,1663]" name="Indonesia" pageId="15" pageNumber="56">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="1C3736FA4B6AFF9A916CF95EC03EF8AE" blockId="15.[816,1450,1167,1902]" pageId="15" pageNumber="56">
Synonymy that of: Melville, J., this work; Cogger, H.G. 1983, in Cogger, H.G., Cameron, E.E., and Cogger, H.M.
<taxonomicName id="DB884D794B6AFF9A971AF978C624F91E" box="[1313,1404,1697,1716]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="class">Amphibia</taxonomicName>
and
<taxonomicName id="DB884D794B6AFF9A910AF965C026F965" authorityName="Linnaeus" authorityYear="1758" box="[817,894,1724,1743]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptiles" pageId="15" pageNumber="56" phylum="Chordata" rank="order">Reptiles</taxonomicName>
. Pp.
<quantity id="DB709B1F4B6AFF9A9197F965C74BF965" box="[940,1043,1724,1743]" metricMagnitude="0" metricUnit="m" metricValue="3.0861" metricValueMax="3.0987999999999998" metricValueMin="3.0734" pageId="15" pageNumber="56" unit="in" value="121.5" valueMax="122.0" valueMin="121.0">121122 in</quantity>
: Walton, D.W. (ed.)
<emphasis id="2EFCEAE84B6AFF9A96F7F965C0DFF943" italics="true" pageId="15" pageNumber="56">
Zoological catalogue of
<collectingCountry id="649F766A4B6AFF9A9109F90FC0DFF943" box="[818,903,1750,1769]" name="Australia" pageId="15" pageNumber="56">Australia</collectingCountry>
</emphasis>
. Vol. 1. Netley,
<collectingRegion id="DE4CF8184B6AFF9A962EF90FC7FEF943" box="[1045,1190,1750,1769]" country="Australia" name="South Australia" pageId="15" pageNumber="56">South Australia</collectingRegion>
: Griffin Press Ltd. vi 313 pp. [122].
</paragraph>
<paragraph id="1C3736FA4B6AFF85916CF8D5C1C3FEB9" blockId="15.[816,1450,1167,1902]" lastBlockId="16.[140,676,229,275]" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56">
<emphasis id="2EFCEAE84B6AFF9A916CF8D5C0ECF8B4" box="[855,948,1804,1822]" italics="true" pageId="15" pageNumber="56">
<typeStatus id="C33388584B6AFF9A916CF8D5C0E8F8B4" box="[855,944,1804,1822]" pageId="15" pageNumber="56" type="lectotype">Lectotype</typeStatus>
.
</emphasis>
BMNH 1946.8.28.72, Port Essington,
<collectingRegion id="DE4CF8184B6AFF9A976FF8D2C0DCF893" country="Australia" name="Northern Territory" pageId="15" pageNumber="56">Northern Territory</collectingRegion>
. Designation by Cogger, H.G., 1983, in Cogger, H.G., Cameron, E.E., and Cogger, H.M.
<taxonomicName id="DB884D794B6AFF9A964EF899C788F8F9" box="[1141,1232,1856,1875]" class="Amphibia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="56" phylum="Chordata" rank="class">Amphibia</taxonomicName>
and
<taxonomicName id="DB884D794B6AFF9A973BF899C615F8F9" authorityName="Linnaeus" authorityYear="1758" box="[1280,1357,1856,1875]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptiles" pageId="15" pageNumber="56" phylum="Chordata" rank="order">Reptiles</taxonomicName>
. Pp.
<quantity id="DB709B1F4B6AFF9A9746F899C036F8C4" metricMagnitude="0" metricUnit="m" metricValue="3.0861" metricValueMax="3.0987999999999998" metricValueMin="3.0734" pageId="15" pageNumber="56" unit="in" value="121.5" valueMax="122.0" valueMin="121.0">121 122 in</quantity>
: Walton, D.W. (ed.)
<emphasis id="2EFCEAE84B6AFF9A9613F882C607F8C4" box="[1064,1375,1883,1902]" italics="true" pageId="15" pageNumber="56">
Zoological catalogue of
<collectingCountry id="649F766A4B6AFF9A9731F882C607F8C4" box="[1290,1375,1883,1902]" name="Australia" pageId="15" pageNumber="56">Australia</collectingCountry>
</emphasis>
. Vol. 1. Netley,
<collectingRegion id="DE4CF8184B75FF8592EAFF3CC23BFF52" box="[209,355,229,248]" country="Australia" name="South Australia" pageId="16" pageNumber="57">South Australia</collectingRegion>
: Griffin Press Ltd. vi 313 pp. [122].
<emphasis id="2EFCEAE84B75FF859289FED9C26DFEB9" box="[178,309,256,275]" italics="true" pageId="16" pageNumber="57">
<typeStatus id="C33388584B75FF859289FED9C269FEB9" box="[178,305,256,275]" pageId="16" pageNumber="57" type="paralectotype">Paralectotype</typeStatus>
.
</emphasis>
BMNH 1946.8.12.63, Nickol Bay, WA.
</paragraph>
<paragraph id="1C3736FA4B75FF8592B7FEF2C2DBFEEA" blockId="16.[140,387,298,320]" box="[140,387,298,320]" pageId="16" pageNumber="57">
<emphasis id="2EFCEAE84B75FF8592B7FEF2C3A0FEEA" box="[140,248,299,320]" italics="true" pageId="16" pageNumber="57">Diagnosis.</emphasis>
As for genus.
</paragraph>
<paragraph id="1C3736FA4B75FF8592B7FE8EC39AFBE2" blockId="16.[138,773,343,1096]" pageId="16" pageNumber="57">
<emphasis id="2EFCEAE84B75FF8592B7FE8EC2C1FEC6" box="[140,409,343,364]" italics="true" pageId="16" pageNumber="57">
Description of
<typeStatus id="C33388584B75FF859309FE8EC2CEFEC6" box="[306,406,343,364]" pageId="16" pageNumber="57" type="lectotype">Lectotype</typeStatus>
.
</emphasis>
Adult. Moderately sized slender agamid lizard with relatively short rounded snout; head narrow and moderately elongated. Prominent canthal ridge consisting of row of enlarged heavily keeled scales. Small nasal scale and nares below ridge. Visible tympanum. A distinct neck, long limbs, long and slender tail, which is damaged and missing its end. Infralabials 11; supralabials 12. Labials elongate unkeeled. Scales on dorsal surface of head moderately heterogeneous, strongly keeled. Prominent nuchal crest of 10 enlarged scales, from ear to shoulder and extending along the back as a row of enlarge scales to base of tail. Head scales heterogeneous and strongly keeled; 45 enlarged white spinose scales protruding from rear of head, posterior to the jaw. Dorsal scales on body and tail mostly homogeneous and weakly to moderately keeled. Scales on thighs homogeneous and strongly keeled. Scales on the ventral surface of head strongly keeled and are weakly keeled on the body. Upper portion of head dark grey-brown and uniformly coloured. Under the head, on the chin, gular and neck dark grey pigmentation, with two pale stripes extending from back of the jaw anteriorly under chin, parallel to jaw, ending approximately half way along jaw. Broad white lip stripe, equally wide on lower and narrow upper lip, extends below ear and continues as two broad white dorsoventral stripes, extending to shoulder. Broad dorsoventral stripes intersected by three dark bands at neck, shoulders and upper back.
</paragraph>
<paragraph id="1C3736FA4B75FF8592B7FBB9C2D9FA7E" blockId="16.[139,772,1119,1492]" pageId="16" pageNumber="57">
<emphasis id="2EFCEAE84B75FF8592B7FBB9C3B7FBDF" box="[140,239,1120,1141]" italics="true" pageId="16" pageNumber="57">Variation.</emphasis>
Adult males have dark grey to black uniform colour on top of head and below chin, onto neck and ventral surface of shoulders. However, females and juveniles often lack this uniform colour and instead have brown and black patterning on top of the head and only have flecks of grey, brown or black on their ventral surface. However, they still have two continuous dorsolateral stripes from the jaw onto the back and lack a well-defined pale stripe between the eye and the ear. Also, it is common for
<taxonomicName id="DB884D794B75FF859322FA93C140FAF4" authorityName="Melville &amp; Ritchie &amp; Chapple &amp; Glor &amp; Schulte" authorityYear="2018" box="[281,536,1353,1375]" class="Reptilia" family="Agamidae" genus="Tropicagama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="57" phylum="Chordata" rank="species" species="temporalis">
<emphasis id="2EFCEAE84B75FF859322FA93C140FAF4" box="[281,536,1353,1375]" italics="true" pageId="16" pageNumber="57">Tropicagama temporalis</emphasis>
</taxonomicName>
to have one or more broad dark lateral bands across the back at the shoulders. Commonly, the white lower lip stripe does not extend the full length of the jaw; instead, it is only present on the posterior section of the lower jaw.
</paragraph>
<paragraph id="1C3736FA4B75FF8592B7FA32C147F8A2" blockId="16.[139,771,1515,1800]" pageId="16" pageNumber="57">
<emphasis id="2EFCEAE84B75FF8592B7FA32C2C4F9AA" box="[140,412,1515,1536]" italics="true" pageId="16" pageNumber="57">Distribution and ecology.</emphasis>
Far northern Australian coastal regions in the Northern Territory and western Cape York. Also occurs on northern offshore islands, including Indonesian islands close to Australian waters and southern
<collectingCountry id="649F766A4B75FF8590B7F99AC381F9DC" name="Papua New Guinea" pageId="16" pageNumber="57">Papua New Guinea</collectingCountry>
. Semi-arboreal, occurring in dry tropical woodland habitats, particularly associated with coastal pandanus and paperbark watercourses. Genetic data has not yet found evidence of this species occurring on the Western Australian mainland. However, future work may find this species occurring in coastal Kimberley regions.
</paragraph>
<paragraph id="1C3736FA4B75FF8592B7F8C7C663FEC4" blockId="16.[140,770,1822,1903]" lastBlockId="16.[808,1440,229,366]" pageId="16" pageNumber="57">
<emphasis id="2EFCEAE84B75FF8592B7F8C7C2B7F899" box="[140,495,1822,1844]" italics="true" pageId="16" pageNumber="57">
Remarks.
<taxonomicName id="DB884D794B75FF8592C9F8C6C2B7F899" authorityName="Melville &amp; Ritchie &amp; Chapple &amp; Glor &amp; Schulte" authorityYear="2018" box="[242,495,1822,1844]" class="Reptilia" family="Agamidae" genus="Tropicagama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="57" phylum="Chordata" rank="species" species="temporalis">Tropicagama temporalis</taxonomicName>
</emphasis>
is superficially similar to, and has extensive distributional overlap with,
<taxonomicName id="DB884D794B75FF85904BF8E5C397F8C4" authorityName="Gray" authorityYear="1842" class="Squamata" family="Agamidae" genus="Lophognathus" kingdom="Animalia" pageId="16" pageNumber="57" phylum="Chordata" rank="species" species="gilbert">
<emphasis id="2EFCEAE84B75FF85904BF8E5C397F8C4" italics="true" pageId="16" pageNumber="57">Lophognathus gilbert</emphasis>
</taxonomicName>
(fig. 4).
<taxonomicName id="DB884D794B75FF859309F883C2E4F8C4" authorityName="Melville &amp; Ritchie &amp; Chapple &amp; Glor &amp; Schulte" authorityYear="2018" box="[306,444,1881,1903]" class="Reptilia" family="Agamidae" genus="Tropicagama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="57" phylum="Chordata" rank="species" species="temporalis">
<emphasis id="2EFCEAE84B75FF859309F883C2E4F8C4" box="[306,444,1881,1903]" italics="true" pageId="16" pageNumber="57">T. temporalis</emphasis>
</taxonomicName>
is readily separated from this species on body proportions (fig. 2) and by having two or fewer preanal pores, uniform dorsal scales with keels converging posteriorly toward midline, a prominent pale dorsolateral stripe that is broadly continuous with stripe along jaw, and lacking a well-defined pale stripe between the eye and the ear.
</paragraph>
</subSubSection>
</treatment>
</document>