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<document id="7827F3269FFA68C5840ACCA732D67EC9" ID-CLB-Dataset="30418" ID-DOI="10.1649/0010-065X(2001)055[0411:DAPOSA]2.0.CO;2" ID-GBIF-Dataset="e4f989d2-2dcb-4d3a-9886-95a337ff4c31" ID-Zenodo-Dep="1208234" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522135981945" checkinUser="plazi" docAuthor="Nico M. Franz" docDate="2001" docId="405B4E03FFA4292755DA0124FCFAFEDB" docLanguage="en" docName="theColeopteristsBulletin.55.4.411.pdf" docOrigin="The Coleopterists Bulletin 55 (4)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Staminodeus denticulatus Nico M. Franz 2001, new species" docType="treatment" docVersion="7" lastPageNumber="424" masterDocId="BC62367BFFA929295409005EFF99FF82" masterDocTitle="Description And Phylogeny Of Staminodeus, A New Genus Of Derelomini (Coleoptera: Curculionidae) Associated With Cyclanthaceae" masterLastPageNumber="432" masterPageNumber="411" pageNumber="424" updateTime="1698533443887" updateUser="ExternalLinkService">
<mods:mods id="8A939941C684BA0628495DD82BEB978E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DB2E463BC1F424B6F0A547DF2AABCD70">
<mods:title id="564E59D5711DD640BA5DDA31A42FECC2">Description And Phylogeny Of Staminodeus, A New Genus Of Derelomini (Coleoptera: Curculionidae) Associated With Cyclanthaceae</mods:title>
</mods:titleInfo>
<mods:name id="E4300C70E25D56018857DEC18F9D7617" type="personal">
<mods:role id="07A9683C36A5B5286E0CCE3BC304A2D5">
<mods:roleTerm id="BF2D8CF3FF07EF1ABC2CFE3FD061A65C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10CC8A8CA676A29178583A0C163D650D">Nico M. Franz</mods:namePart>
</mods:name>
<mods:typeOfResource id="551B897A36C15FD54866046E0873F2D1">text</mods:typeOfResource>
<mods:relatedItem id="28326967ECF524DAC10B17EFEEF30ADB" type="host">
<mods:titleInfo id="E32379829EB1441A76E9987C79AA5AD3">
<mods:title id="A3926FC9E716DF784858D01A91DDEBA9">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="32664EF06F4F548C155D3E56C2942AA2">
<mods:date id="8DAD7CBE6BFE740A82BA387F6AF9157C">2001</mods:date>
<mods:detail id="7526E51695E8224473FE56ECAEAED4B8" type="volume">
<mods:number id="1C1AA7F679B60AECA00071C4C18F9F4F">55</mods:number>
</mods:detail>
<mods:detail id="0E6B7B1EF7B96686022BFE1CBD681B74" type="issue">
<mods:number id="6A059A4031EA6C51F290325FA97E9033">4</mods:number>
</mods:detail>
<mods:extent id="C91CDAD3F3A4EF7A5D9B7742BD0723F3" unit="page">
<mods:start id="9CDD3AEDD3CCD2E4EBDD50D72EE8143B">411</mods:start>
<mods:end id="43860CD4EF6E72EEE954196616554B43">432</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D7FDB30E1FF9E0E563139E29F92BF825">journal article</mods:classification>
<mods:identifier id="9462BB2CFD9DD01B4149EC16A622D6C7" type="CLB-Dataset">30418</mods:identifier>
<mods:identifier id="A441079FD38360D11AF1614DE22DA4C3" type="DOI">10.1649/0010-065X(2001)055[0411:DAPOSA]2.0.CO;2</mods:identifier>
<mods:identifier id="6E42A346E8DD89681FB865DC41A2D8ED" type="GBIF-Dataset">e4f989d2-2dcb-4d3a-9886-95a337ff4c31</mods:identifier>
<mods:identifier id="B098CB8E26FFBA8BBBE004E43F041338" type="Zenodo-Dep">1208234</mods:identifier>
</mods:mods>
<treatment id="405B4E03FFA4292755DA0124FCFAFEDB" ID-DOI="http://doi.org/10.5281/zenodo.6279370" ID-GBIF-Taxon="142656681" ID-Zenodo-Dep="6279370" LSID="urn:lsid:plazi:treatment:405B4E03FFA4292755DA0124FCFAFEDB" httpUri="http://treatment.plazi.org/id/405B4E03FFA4292755DA0124FCFAFEDB" lastPageId="14" lastPageNumber="424" pageId="13" pageNumber="424">
<subSubSection id="80E8AC9EFFA4292455DA0124FCEAFE0C" box="[467,883,377,399]" pageId="13" pageNumber="424" type="nomenclature">
<paragraph id="C84DFF15FFA4292455DA0124FCEAFE0C" blockId="13.[467,883,377,399]" box="[467,883,377,399]" pageId="13" pageNumber="424">
<heading id="93054879FFA4292455DA0124FCEAFE0C" box="[467,883,377,399]" fontSize="9" level="3" pageId="13" pageNumber="424" reason="1">
<emphasis id="FA862307FFA4292455DA0124FD71FE0D" box="[467,744,378,399]" italics="true" pageId="13" pageNumber="424">
<taxonomicName id="0FF28496FFA4292455DA0124FD7AFE0D" authority="Nico M. Franz, 2001" authorityName="Nico M. Franz" authorityYear="2001" box="[467,739,378,399]" class="Insecta" family="Curculionidae" genus="Staminodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="424" phylum="Arthropoda" rank="species" species="denticulatus" status="sp. nov.">Staminodeus denticulatus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="E1B59E7CFFA4292456F90127FCEAFE0C" box="[752,883,377,398]" pageId="13" pageNumber="424" rank="species">
<emphasis id="FA862307FFA4292456F90127FCEAFE0C" bold="true" box="[752,883,377,398]" pageId="13" pageNumber="424">new species</emphasis>
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="80E8AC9EFFA42924551901FAFEF5FDA6" pageId="13" pageNumber="424" type="diagnosis">
<paragraph id="C84DFF15FFA42924551901FAFEF5FDA6" blockId="13.[246,1078,420,548]" pageId="13" pageNumber="424">
<emphasis id="FA862307FFA42924551901FAFE1BFE3B" bold="true" box="[272,386,420,441]" pageId="13" pageNumber="424">Diagnosis.</emphasis>
<taxonomicName id="0FF28496FFA42924558301FBFD0EFE38" authorityName="Nico M. Franz" authorityYear="2001" box="[394,663,421,442]" class="Insecta" family="Curculionidae" genus="Staminodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="424" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="FA862307FFA42924558301FBFD0EFE38" box="[394,663,421,442]" italics="true" pageId="13" pageNumber="424">Staminodeus denticulatus</emphasis>
</taxonomicName>
is distinguished from the other species of
<taxonomicName id="0FF28496FFA42924551A019EFE00FE57" authorityName="Nico M. Franz" authorityYear="2001" box="[275,409,448,469]" class="Insecta" family="Curculionidae" genus="Staminodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="FA862307FFA42924551A019EFE00FE57" box="[275,409,448,469]" italics="true" pageId="13" pageNumber="424">Staminodeus</emphasis>
</taxonomicName>
in general, and from
<taxonomicName id="0FF28496FFA429245674019EFC8FFE57" authorityName="Nico M. Franz" authorityYear="2001" box="[637,790,448,469]" class="Insecta" family="Curculionidae" genus="Staminodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="424" phylum="Arthropoda" rank="species" species="curvitibialis">
<emphasis id="FA862307FFA429245674019EFC8FFE57" box="[637,790,448,469]" italics="true" pageId="13" pageNumber="424">S. curvitibialis</emphasis>
</taxonomicName>
in particular, by the frontal spinule and the high number of small denticuli along the apically inermous protibia in males; as well as the fairly long and abruptly deflexed frontal spine in females.
</paragraph>
</subSubSection>
<subSubSection id="80E8AC9EFFA4292455040267FC70FB97" pageId="13" pageNumber="424" type="description">
<paragraph id="C84DFF15FFA4292455040267FC22FC07" blockId="13.[246,1079,569,1045]" pageId="13" pageNumber="424">
<emphasis id="FA862307FFA4292455040267FEA6FDCE" bold="true" box="[269,319,569,588]" pageId="13" pageNumber="424">Male</emphasis>
. Length
<quantity id="0F0A52F0FFA42924559E0264FD93FDCF" box="[407,522,570,589]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="13" pageNumber="424" unit="mm" value="2.1" valueMax="2.4" valueMin="1.8">1.8–2.4 mm</quantity>
, width
<quantity id="0F0A52F0FFA42924565A0264FD5EFDCF" box="[595,711,570,589]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="13" pageNumber="424" unit="mm" value="1.05" valueMax="1.2" valueMin="0.9">0.9–1.2 mm</quantity>
, l/
<geoCoordinate id="ADC699D2FFA4292456E90264FCB6FDCF" box="[736,815,570,589]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="5555" value="-1.9">w 1.9</geoCoordinate>
–2.2. Rostrum
<quantity id="0F0A52F0FFA4292457B70264FBABFDCF" box="[958,1074,570,589]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="13" pageNumber="424" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 mm</quantity>
, r/p 0.7–0.8, laterobasal 1/3 carinulate. Eyes separated by distance slightly greater than breadth of antennal club. Frons with 1 apically directed, arcuate spinule, nearly 1/4 as long as diameter of eye, anteriorly with long, fine, erect setae. Pronotum l/
<geoCoordinate id="ADC699D2FFA4292457B202DCFB91FD17" box="[955,1032,642,661]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="5555" value="-0.8">w 0.8</geoCoordinate>
–0.9, anteriorly distinctly narrowed, distinctly convex. Prosternum distinctly longer than mesosternum, equilateral, distinctly convex. Profemur (
<figureCitation id="50C9E390FFA4292456E102ECFCB5FD47" box="[744,812,690,709]" captionStart="Fig. 3" captionStartId="7.[269,303,1467,1486]" captionTargetBox="[257,1072,272,1441]" captionTargetId="figure@7.[246,1078,265,1446]" captionTargetPageId="7" captionText="Fig. 3. Staminodeus, male, right profemur and protibia, anterior view, a) S. inermis; b) S. curvitibialis; c) S. denticulatus; d) S. bispinosus; e) S. forcipis; f) S. dilatatus; g) S. vectoris. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1208240/files/figure.png" pageId="13" pageNumber="424">Fig. 3c</figureCitation>
) slightly arcuate, f/p 1.0/ 1.3, width similar throughout, ventrally denticulate; protibia (
<figureCitation id="50C9E390FFA4292457350294FCE6FD5F" box="[828,895,714,733]" captionStart="Fig. 3" captionStartId="7.[269,303,1467,1486]" captionTargetBox="[257,1072,272,1441]" captionTargetId="figure@7.[246,1078,265,1446]" captionTargetPageId="7" captionText="Fig. 3. Staminodeus, male, right profemur and protibia, anterior view, a) S. inermis; b) S. curvitibialis; c) S. denticulatus; d) S. bispinosus; e) S. forcipis; f) S. dilatatus; g) S. vectoris. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1208240/files/figure.png" pageId="13" pageNumber="424">Fig. 3c</figureCitation>
) slightly arcuate, t/ f 1.0, anteroventral margin with row of 12–18 apically directed, arcuate, acute denticuli, apically inermous; mesotibia apically inermous. Elytron l/
<geoCoordinate id="ADC699D2FFA42924576802A4FC29FC8F" box="[865,944,762,781]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="5555" value="-1.3">w 1.3</geoCoordinate>
–1.5 (N 4). Median lobe (
<figureCitation id="50C9E390FFA429245577034CFE5AFCA7" box="[382,451,786,805]" captionStart="Fig. 4" captionStartId="10.[272,306,1472,1491]" captionTargetBox="[257,1079,271,1448]" captionTargetId="figure@10.[246,1084,265,1452]" captionTargetPageId="10" captionText="Fig. 4. Staminodeus, male, aedeagus, ventral and lateral view, a) S. inermis; b) S. curvitibialis; c) S. denticulatus; d) S. bispinosus; e) S. forcipis; f) S. dilatatus; g) S. vectoris. Illustrations are interpretative: denticulation and setation are omitted. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/1208242/files/figure.png" pageId="13" pageNumber="424">Fig. 4c</figureCitation>
) l/
<geoCoordinate id="ADC699D2FFA4292455E8034CFDABFCA7" box="[481,562,786,805]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="5555" value="-2.3">w 2.3</geoCoordinate>
–2.3 (N 3), apical margin medially projected, laterobasal sclerites weakly sclerotized, short, spinulose, median sclerite heavily sclerotized, short, basally rotundate, impressed, apically spinulose, with 2 elongate, slightly diverging projections, apical rami broad, apically deflexed, with small, elongate, spinulose region between apical rami, aedeagal apodemes slightly longer than median lobe.
</paragraph>
<paragraph id="C84DFF15FFA42924550403D7FB82FC7F" blockId="13.[246,1079,569,1045]" pageId="13" pageNumber="424">
<emphasis id="FA862307FFA42924550403D7FECCFC1E" bold="true" box="[269,341,905,924]" pageId="13" pageNumber="424">Female</emphasis>
. Length
<quantity id="0F0A52F0FFA4292455A003D4FD85FC1F" box="[425,540,906,925]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="13" pageNumber="424" unit="mm" value="2.15" valueMax="2.4" valueMin="1.9">1.9–2.4 mm</quantity>
, width 1.0–
<quantity id="0F0A52F0FFA42924568203D4FD4DFC1F" box="[651,724,906,925]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="424" unit="mm" value="1.2">1.2 mm</quantity>
, l/
<geoCoordinate id="ADC699D2FFA4292456E503D4FCAFFC1F" box="[748,822,906,925]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="5555" value="-1.8">w 1.8</geoCoordinate>
–2.0. Rostrum
<quantity id="0F0A52F0FFA4292457C803D4FBABFC1F" box="[961,1074,906,925]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="13" pageNumber="424" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
, r/p 0.9–1.0, laterobasal 1/3 carinulate. Frontal spine nearly 2/3–3/4 as long as diameter of eye, basally subrectate, perpendicular to frons, apically abruptly deflexed (not gradually arcuate). Pronotum l/
<geoCoordinate id="ADC699D2FFA4292455F5038CFDC9FC67" box="[508,592,978,997]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="5555" value="-0.8">w 0.8</geoCoordinate>
–0.9, greatest width near anterior 1/3. Prosternum longer than mesosternum. F/p 0.9–1.2; t/f 0.9–1.1. Elytron l/
<geoCoordinate id="ADC699D2FFA42924577703B4FC49FC7F" box="[894,976,1002,1021]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="5555" value="-1.2">w 1.2</geoCoordinate>
–1.5 (N
</paragraph>
<paragraph id="C84DFF15FFA4292454FF045CFC70FB97" blockId="13.[246,1079,569,1045]" box="[246,1001,1026,1045]" pageId="13" pageNumber="424">
12). Spermatheca (
<figureCitation id="50C9E390FFA4292455A0045CFE74FB97" box="[425,493,1026,1045]" captionStart="Fig. 5" captionStartId="11.[269,303,1502,1521]" captionTargetBox="[285,1052,274,1473]" captionTargetId="figure@11.[269,1056,265,1481]" captionTargetPageId="11" captionText="Fig. 5. Staminodeus, female, spermatheca and gland reservoir, a) S. inermis; b) S. curvitibialis; c) S. denticulatus; d) S. bispinosus; e) S. forcipis; f) S. dilatatus; g) S. vectoris. Scale bar 0.05 mm." httpUri="https://zenodo.org/record/1208244/files/figure.png" pageId="13" pageNumber="424">Fig. 5c</figureCitation>
) deflexed nearly 90, apex slightly separated, obtuse.
</paragraph>
</subSubSection>
<subSubSection id="80E8AC9EFFA4292455190470FE2DFA1C" pageId="13" pageNumber="424" type="materials_examined">
<paragraph id="C84DFF15FFA4292455190470FE2DFA1C" blockId="13.[246,1079,1070,1652]" pageId="13" pageNumber="424">
<materialsCitation id="789AF548FFA4292455190470FBABFBDC" ID-GBIF-Occurrence="1836754051" collectingDate="1994-12" collectionCode="CMNC" collectorName="L. Gomez" country="Costa Rica" elevation="1100" location="Las Cruces" municipality="Type Information. Male" pageId="13" pageNumber="424" specimenCount="2" stateProvince="Puntarenas" typeStatus="holotype">
<collectingMunicipality id="2829656FFFA4292455190470FDBCFBC6" box="[272,549,1070,1092]" pageId="13" pageNumber="424">
<emphasis id="FA862307FFA4292455190470FE7AFBC1" bold="true" box="[272,483,1070,1091]" pageId="13" pageNumber="424">
<typeStatus id="174941B7FFA4292455190470FED0FBC1" box="[272,329,1070,1091]" pageId="13" pageNumber="424">Type</typeStatus>
Information.
</emphasis>
Male
</collectingMunicipality>
<typeStatus id="174941B7FFA4292456380471FD15FBC6" box="[561,652,1071,1092]" pageId="13" pageNumber="424" type="holotype">holotype</typeStatus>
‘‘
<collectingCountry id="B0E5BF85FFA4292456AB0471FCBBFBC6" box="[674,802,1071,1092]" name="Costa Rica" pageId="13" pageNumber="424">Costa Rica</collectingCountry>
,
<collectingRegion id="0A3631F7FFA4292457380471FC3FFBC6" box="[817,934,1071,1092]" country="Costa Rica" name="Puntarenas" pageId="13" pageNumber="424">Puntarenas</collectingRegion>
,
<location id="CD2DA9CEFFA4292457BF0471FBABFBC6" LSID="urn:lsid:plazi:treatment:405B4E03FFA4292755DA0124FCFAFEDB:CD2DA9CEFFA4292457BF0471FBABFBC6" box="[950,1074,1071,1092]" country="Costa Rica" municipality="Type Information. Male" name="Las Cruces" pageId="13" pageNumber="424" stateProvince="Puntarenas">Las Cruces</location>
,
<elevation id="43DF1826FFA4292454FF0417FECAFBDC" box="[246,339,1097,1118]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="424" unit="m" value="1100.0">
<quantity id="0F0A52F0FFA4292454FF0417FECAFBDC" box="[246,339,1097,1118]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="424" unit="m" value="1100.0">1,100 m</quantity>
</elevation>
,
<collectedFrom id="7BEEBCE8FFA42924556A0417FD92FBDD" box="[355,523,1097,1119]" pageId="13" pageNumber="424">
on
<taxonomicName id="0FF28496FFA4292455810414FD92FBDD" box="[392,523,1098,1119]" class="Liliopsida" family="Cyclanthaceae" genus="Carludovica" higherTaxonomySource="CoL" kingdom="Plantae" order="Pandanales" pageId="13" pageNumber="424" phylum="Liliopsida" rank="genus">
<emphasis id="FA862307FFA4292455810414FD92FBDD" box="[392,523,1098,1119]" italics="true" pageId="13" pageNumber="424">Carludovica</emphasis>
</taxonomicName>
</collectedFrom>
subpalmata, leg.
<collectorName id="65079AC3FFA4292456DC0417FCDCFBDC" box="[725,837,1097,1118]" pageId="13" pageNumber="424">L. Gómez</collectorName>
,
<collectingDate id="AC08203DFFA42924575D0417FC25FBDC" box="[852,956,1097,1118]" pageId="13" pageNumber="424" value="1994-12">
<date id="BC4CD9D5FFA42924575D0417FC25FBDC" box="[852,956,1097,1118]" pageId="13" pageNumber="424" value="1994-12">XII­1994</date>
</collectingDate>
’’ (
<collectionCode id="AEE367D0FFA4292457D10417FBB2FBDC" box="[984,1067,1097,1118]" pageId="13" pageNumber="424">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="789AF548FFA4292454FF043AFCA3FBFB" ID-GBIF-Occurrence="1836754065" box="[246,826,1124,1145]" collectionCode="CMNC" pageId="13" pageNumber="424" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<specimenCount id="DEF4349CFFA4292454FF043AFEA4FBFB" box="[246,317,1124,1145]" pageId="13" pageNumber="424" type="female">female</specimenCount>
<typeStatus id="174941B7FFA42924554C043AFE05FBFB" box="[325,412,1124,1145]" pageId="13" pageNumber="424" type="allotype">allotype</typeStatus>
, same label as male holotype (
<collectionCode id="AEE367D0FFA4292456E9043AFCAAFBFB" box="[736,819,1124,1145]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="13" pageNumber="424">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="789AF548FFA42924574F043AFEE5FB66" ID-GBIF-Occurrence="1836754057" collectingDate="1977-08-09" collectingDateMax="1978-04-22" collectingDateMin="1977-08-09" collectionCode="CWOB, MIUP, MUCR" collectorName="H. Wolda" country="Costa Rica" latitude="8.733334" location="Collector'" longLatPrecision="1300" longitude="-82.25" municipality="Fortuna" pageId="13" pageNumber="424" specimenCount="1" stateProvince="Chiriqui" typeStatus="paratype">
male
<typeStatus id="174941B7FFA429245789043AFC7DFBFB" box="[896,996,1124,1145]" pageId="13" pageNumber="424" type="paratype">paratypes</typeStatus>
‘‘PAN­ AMA,
<collectingRegion id="0A3631F7FFA42924554F0421FE39FB16" box="[326,416,1151,1172]" country="Panama" name="Chiriqui" pageId="13" pageNumber="424">Chiriqui</collectingRegion>
,
<collectingMunicipality id="2829656FFFA4292455B90421FD9DFB16" box="[432,516,1151,1172]" pageId="13" pageNumber="424">Fortuna</collectingMunicipality>
, (
<geoCoordinate id="ADC699D2FFA4292456140421FDF3FB16" box="[541,618,1151,1172]" direction="north" orientation="latitude" pageId="13" pageNumber="424" precision="925" value="8.733334">8 44 N</geoCoordinate>
,
<geoCoordinate id="ADC699D2FFA4292456730421FD40FB16" box="[634,729,1151,1172]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="925" value="-82.25">82 15 W</geoCoordinate>
),
<collectingMethod id="11B38702FFA4292457090421FCC4FB16" box="[768,861,1151,1172]" pageId="13" pageNumber="424">UV trap</collectingMethod>
,
<collectingDate id="AC08203DFFA4292457670421FB91FB16" box="[878,1032,1151,1172]" pageId="13" pageNumber="424" value="1978-04-22">
<date id="BC4CD9D5FFA4292457670421FB91FB16" box="[878,1032,1151,1172]" pageId="13" pageNumber="424" value="1978-04-22">Apr. 22, 1978</date>
</collectingDate>
/
<collectorName id="65079AC3FFA4292450140421FEA5FB2C" pageId="13" pageNumber="424">H. Wolda</collectorName>
,
<location id="CD2DA9CEFFA42924554004C7FE2FFB2C" LSID="urn:lsid:plazi:treatment:405B4E03FFA4292755DA0124FCFAFEDB:CD2DA9CEFFA42924554004C7FE2FFB2C" box="[329,438,1177,1198]" country="Costa Rica" latitude="8.733334" longLatPrecision="1300" longitude="-82.25" municipality="Fortuna" name="Collector'" pageId="13" pageNumber="424" stateProvince="Chiriqui">Collector’</location>
’ (
<collectionCode id="AEE367D0FFA4292455C304C7FD85FB2C" box="[458,540,1177,1198]" pageId="13" pageNumber="424">CWOB</collectionCode>
, 2), same label as previous specimens with different date ‘‘
<collectingDate id="AC08203DFFA42924555704EAFE6CFB4B" box="[350,501,1204,1225]" pageId="13" pageNumber="424" value="1977-08-09">
<date id="BC4CD9D5FFA42924555704EAFE6CFB4B" box="[350,501,1204,1225]" pageId="13" pageNumber="424" value="1977-08-09">Aug. 9, 1977</date>
</collectingDate>
’’ (
<collectionCode id="AEE367D0FFA42924561904EAFDCBFB4B" box="[528,594,1204,1225]" pageId="13" pageNumber="424">MIUP</collectionCode>
, 1); same label as male holoype (
<collectionCode id="AEE367D0FFA4292457B504EAFB94FB4B" box="[956,1037,1204,1225]" country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="13" pageNumber="424">MUCR</collectionCode>
, 1); ‘‘
<collectingCountry id="B0E5BF85FFA4292455090491FEE5FB66" box="[256,380,1231,1252]" name="Costa Rica" pageId="13" pageNumber="424">Costa Rica</collectingCountry>
</materialsCitation>
,
<materialsCitation id="789AF548FFA42924558E0491FDC0FA9B" ID-GBIF-Occurrence="1836754062" collectingDate="1994-12-29" collectionCode="MUCR, CMNC" collectorName="L. Gomez" country="Costa Rica" elevation="1000" location="Las Cruces" pageId="13" pageNumber="424" specimenCount="2" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="0A3631F7FFA42924558E0491FE65FB66" box="[391,508,1231,1252]" country="Costa Rica" name="Puntarenas" pageId="13" pageNumber="424">Puntarenas</collectingRegion>
,
<location id="CD2DA9CEFFA4292456010491FD18FB66" LSID="urn:lsid:plazi:treatment:405B4E03FFA4292755DA0124FCFAFEDB:CD2DA9CEFFA4292456010491FD18FB66" box="[520,641,1231,1252]" country="Costa Rica" name="Las Cruces" pageId="13" pageNumber="424" stateProvince="Puntarenas">Las Cruces</location>
,
<elevation id="43DF1826FFA4292456840491FD7FFB66" box="[653,742,1231,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="424" unit="m" value="1000.0">
<quantity id="0F0A52F0FFA4292456840491FD7FFB66" box="[653,742,1231,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="424" unit="m" value="1000.0">1,000 m</quantity>
</elevation>
,
<collectedFrom id="7BEEBCE8FFA4292456FB0491FC0EFB66" box="[754,919,1231,1252]" pageId="13" pageNumber="424">
on
<taxonomicName id="0FF28496FFA42924571D0491FC0EFB66" box="[788,919,1231,1252]" class="Liliopsida" family="Cyclanthaceae" genus="Evodianthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="13" pageNumber="424" phylum="Tracheophyta" rank="genus">
<emphasis id="FA862307FFA42924571D0491FC0EFB66" box="[788,919,1231,1252]" italics="true" pageId="13" pageNumber="424">Evodianthus</emphasis>
</taxonomicName>
</collectedFrom>
funifer, leg.
<collectorName id="65079AC3FFA4292450160491FEDDFB7C" pageId="13" pageNumber="424">L. Gómez</collectorName>
,
<collectingDate id="AC08203DFFA42924555804B7FE79FB7C" box="[337,480,1257,1278]" pageId="13" pageNumber="424" value="1994-12-29">
<date id="BC4CD9D5FFA42924555804B7FE79FB7C" box="[337,480,1257,1278]" pageId="13" pageNumber="424" value="1994-12-29">XII­29–1994</date>
</collectingDate>
’’ (
<collectionCode id="AEE367D0FFA4292455F304B7FDD2FB7C" box="[506,587,1257,1278]" country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="13" pageNumber="424">MUCR</collectionCode>
, 1); female
<typeStatus id="174941B7FFA4292456C404B7FCADFB7C" box="[717,820,1257,1278]" pageId="13" pageNumber="424" type="paratype">paratypes</typeStatus>
, same label as male
<typeStatus id="174941B7FFA42924501A04B7FEAEFA9B" pageId="13" pageNumber="424" type="holotype">holotype</typeStatus>
(
<collectionCode id="AEE367D0FFA429245540055AFE03FA9B" box="[329,410,1284,1305]" pageId="13" pageNumber="424">CMNC</collectionCode>
, 1); ‘‘
<collectingCountry id="B0E5BF85FFA4292455D3055AFDC0FA9B" box="[474,601,1284,1305]" name="Costa Rica" pageId="13" pageNumber="424">Costa Rica</collectingCountry>
</materialsCitation>
,
<materialsCitation id="789AF548FFA42924566E055AFEAFFACC" ID-GBIF-Occurrence="1836754046" collectingDate="1994-12-29" collectionCode="CMNC, MUCR" collectorName="L. Gomez" country="Costa Rica" elevation="1000" location="Las Cruces" pageId="13" pageNumber="424" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="0A3631F7FFA42924566E055AFD45FA9B" box="[615,732,1284,1305]" country="Costa Rica" name="Puntarenas" pageId="13" pageNumber="424">Puntarenas</collectingRegion>
,
<location id="CD2DA9CEFFA4292456E2055AFCFFFA9B" LSID="urn:lsid:plazi:treatment:405B4E03FFA4292755DA0124FCFAFEDB:CD2DA9CEFFA4292456E2055AFCFFFA9B" box="[747,870,1284,1305]" country="Costa Rica" name="Las Cruces" pageId="13" pageNumber="424" stateProvince="Puntarenas">Las Cruces</location>
,
<elevation id="43DF1826FFA42924577C055AFC48FA9B" box="[885,977,1284,1305]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="424" unit="m" value="1000.0">
<quantity id="0F0A52F0FFA42924577C055AFC48FA9B" box="[885,977,1284,1305]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="424" unit="m" value="1000.0">1,000 m</quantity>
</elevation>
,
<collectedFrom id="7BEEBCE8FFA4292457E9055AFEC9FAB6" pageId="13" pageNumber="424">
on
<taxonomicName id="0FF28496FFA42924500C055AFEC9FAB6" class="Liliopsida" family="Cyclanthaceae" genus="Evodianthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="13" pageNumber="424" phylum="Tracheophyta" rank="genus">
<emphasis id="FA862307FFA42924500C055AFEC9FAB6" italics="true" pageId="13" pageNumber="424">Evodianthus</emphasis>
</taxonomicName>
</collectedFrom>
funifer, leg.
<collectorName id="65079AC3FFA4292455D10541FDDAFAB6" box="[472,579,1311,1332]" pageId="13" pageNumber="424">L. Gómez</collectorName>
,
<collectingDate id="AC08203DFFA4292456470541FD47FAB6" box="[590,734,1311,1332]" pageId="13" pageNumber="424" value="1994-12-29">
<date id="BC4CD9D5FFA4292456470541FD47FAB6" box="[590,734,1311,1332]" pageId="13" pageNumber="424" value="1994-12-29">XII­29–1994</date>
</collectingDate>
’’ (
<collectionCode id="AEE367D0FFA4292456FF0541FCDEFAB6" box="[758,839,1311,1332]" pageId="13" pageNumber="424">CMNC</collectionCode>
, 1;
<collectionCode id="AEE367D0FFA4292457640541FC24FAB6" box="[877,957,1311,1332]" country="Costa Rica" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="13" pageNumber="424">MUCR</collectionCode>
, 3); ‘‘PAN­ AMA
</materialsCitation>
,
<materialsCitation id="789AF548FFA4292455480567FE2DFA1C" ID-GBIF-Occurrence="1836754064" collectingDate="1978-04-22" collectingDateMax="1978-08-02" collectingDateMin="1978-04-22" collectionCode="CWOB, MIUP" country="Costa Rica" county="Fortuna" latitude="8.733334" location="Collector'" longLatPrecision="1300" longitude="-82.25" municipality="Wolda" pageId="13" pageNumber="424" specimenCount="1" stateProvince="Chiriqui" typeStatus="paratype">
<collectingRegion id="0A3631F7FFA4292455480567FE02FACC" box="[321,411,1337,1358]" country="Panama" name="Chiriqui" pageId="13" pageNumber="424">Chiriqui</collectingRegion>
,
<collectingCounty id="212C8799FFA4292455AE0567FE62FACC" box="[423,507,1337,1358]" pageId="13" pageNumber="424">Fortuna</collectingCounty>
, (
<geoCoordinate id="ADC699D2FFA4292456190567FDC5FACC" box="[528,604,1337,1358]" direction="north" orientation="latitude" pageId="13" pageNumber="424" precision="925" value="8.733334">8 44 N</geoCoordinate>
,
<geoCoordinate id="ADC699D2FFA4292456610567FD50FACC" box="[616,713,1337,1358]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="925" value="-82.25">82 15 W</geoCoordinate>
),
<collectingMethod id="11B38702FFA4292456D50567FCACFACC" box="[732,821,1337,1358]" pageId="13" pageNumber="424">UV trap</collectingMethod>
,
<collectingDate id="AC08203DFFA4292457480567FC48FACC" box="[833,977,1337,1358]" pageId="13" pageNumber="424" value="1978-04-22">
<date id="BC4CD9D5FFA4292457480567FC48FACC" box="[833,977,1337,1358]" pageId="13" pageNumber="424" value="1978-04-22">Apr. 22, 1978</date>
</collectingDate>
/ H.
<collectingMunicipality id="2829656FFFA42924500A0567FE88FAEB" pageId="13" pageNumber="424">Wolda</collectingMunicipality>
,
<location id="CD2DA9CEFFA429245529050AFE14FAEB" LSID="urn:lsid:plazi:treatment:405B4E03FFA4292755DA0124FCFAFEDB:CD2DA9CEFFA429245529050AFE14FAEB" box="[288,397,1364,1385]" country="Costa Rica" county="Fortuna" latitude="8.733334" longLatPrecision="1300" longitude="-82.25" municipality="Wolda" name="Collector'" pageId="13" pageNumber="424" stateProvince="Chiriqui">Collector’</location>
’ (
<collectionCode id="AEE367D0FFA4292455AB050AFE6DFAEB" box="[418,500,1364,1385]" pageId="13" pageNumber="424">CWOB</collectionCode>
, 1), same labels as previous specimen with different dates ‘‘
<collectingDate id="AC08203DFFA4292455480531FE43FA06" box="[321,474,1391,1412]" pageId="13" pageNumber="424" value="1978-05-02">
<date id="BC4CD9D5FFA4292455480531FE43FA06" box="[321,474,1391,1412]" pageId="13" pageNumber="424" value="1978-05-02">May 2, 1978</date>
</collectingDate>
’’ (
<collectionCode id="AEE367D0FFA4292455F10531FDD3FA06" box="[504,586,1391,1412]" pageId="13" pageNumber="424">CWOB</collectionCode>
, 1), ‘‘
<collectingDate id="AC08203DFFA4292456850531FCB4FA06" box="[652,813,1391,1412]" pageId="13" pageNumber="424" value="1978-07-11">
<date id="BC4CD9D5FFA4292456850531FCB4FA06" box="[652,813,1391,1412]" pageId="13" pageNumber="424" value="1978-07-11">July 11, 1978</date>
</collectingDate>
’’ (
<collectionCode id="AEE367D0FFA4292457420531FC04FA06" box="[843,925,1391,1412]" pageId="13" pageNumber="424">CWOB</collectionCode>
, 1), ‘‘
<collectingDate id="AC08203DFFA4292457D60531FEB6FA1C" pageId="13" pageNumber="424" value="1978-08-02">
<date id="BC4CD9D5FFA4292457D60531FEB6FA1C" pageId="13" pageNumber="424" value="1978-08-02">Aug. 2, 1978</date>
</collectingDate>
’’ (
<collectionCode id="AEE367D0FFA42924554305D7FE15FA1C" box="[330,396,1417,1438]" pageId="13" pageNumber="424">MIUP</collectionCode>
, 2).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="80E8AC9EFFA42924551905FAFD22FA56" pageId="13" pageNumber="424" type="etymology">
<paragraph id="C84DFF15FFA42924551905FAFD22FA56" blockId="13.[246,1079,1070,1652]" pageId="13" pageNumber="424">
<emphasis id="FA862307FFA42924551905FAFE16FA3B" bold="true" box="[272,399,1444,1465]" pageId="13" pageNumber="424">Etymology.</emphasis>
Named for the denticulate protibia in males—
<taxonomicName id="0FF28496FFA42924578E05FAFB9EFA3B" authorityName="Nico M. Franz" authorityYear="2001" box="[903,1031,1444,1465]" class="Insecta" family="Curculionidae" genus="Staminodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="424" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="FA862307FFA42924578E05FAFB9EFA3B" box="[903,1031,1444,1465]" italics="true" pageId="13" pageNumber="424">denticulatus</emphasis>
</taxonomicName>
signifying ‘‘with small teeth’’ (
<bibRefCitation id="AC6382E4FFA42924562C05E1FD36FA56" author="Brown, R. W." bookContentInfo="882 pp." box="[549,687,1471,1492]" journalOrPublisher="Smithsonian Institution Press, Washington, D. C." pageId="13" pageNumber="424" refId="ref11994" refString="Brown, R. W. 1956. Composition of scientific words. Revised edition. Smithsonian Institution Press, Washington, D. C. 882 pp." title="Composition of scientific words. Revised edition" type="book" year="1956">Brown 1956</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="80E8AC9EFFA4292455190587FED4F9DB" pageId="13" pageNumber="424" type="distribution">
<paragraph id="C84DFF15FFA4292455190587FED4F9DB" blockId="13.[246,1079,1070,1652]" pageId="13" pageNumber="424">
<emphasis id="FA862307FFA4292455190587FE06FA6C" bold="true" box="[272,415,1497,1518]" pageId="13" pageNumber="424">Distribution.</emphasis>
<taxonomicName id="0FF28496FFA4292455AE0584FD2CFA6D" authorityName="Nico M. Franz" authorityYear="2001" box="[423,693,1498,1519]" class="Insecta" family="Curculionidae" genus="Staminodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="424" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="FA862307FFA4292455AE0584FD2CFA6D" box="[423,693,1498,1519]" italics="true" pageId="13" pageNumber="424">Staminodeus denticulatus</emphasis>
</taxonomicName>
has been collected in the Cordillera de Talamanca of
<collectingCountry id="B0E5BF85FFA4292455B205AAFDAFF98B" box="[443,566,1524,1545]" name="Costa Rica" pageId="13" pageNumber="424">Costa Rica</collectingCountry>
,
<collectingRegion id="0A3631F7FFA42924564E05AAFCCBF98B" box="[583,850,1524,1545]" country="Costa Rica" name="Puntarenas" pageId="13" pageNumber="424">Provincia de Puntarenas</collectingRegion>
, San Vito:
<geoCoordinate id="ADC699D2FFA4292457D005AAFBABF98B" box="[985,1074,1524,1545]" direction="north" orientation="latitude" pageId="13" pageNumber="424" precision="15" value="0.14638889">0 8 47 N</geoCoordinate>
,
<geoCoordinate id="ADC699D2FFA4292454FF0651FECDF9A6" box="[246,340,1551,1572]" direction="west" orientation="longitude" pageId="13" pageNumber="424" precision="925" value="-82.95">82 57 W</geoCoordinate>
; on the Carribean slope of
<collectingRegion id="0A3631F7FFA42924568B0651FD4EF9A6" box="[642,727,1551,1572]" country="Panama" name="Panama" pageId="13" pageNumber="424">
<collectingCountry id="B0E5BF85FFA42924568B0651FD4EF9A6" box="[642,727,1551,1572]" name="Panama" pageId="13" pageNumber="424">Panama</collectingCountry>
</collectingRegion>
,
<collectingRegion id="0A3631F7FFA4292456EC0651FC60F9A6" box="[741,1017,1551,1572]" country="Panama" name="Bocas del Toro" pageId="13" pageNumber="424">Provincia Bocas del Toro</collectingRegion>
,
<collectingRegion id="0A3631F7FFA4292450010651FEB1F9BC" country="Panama" name="Chiriqui" pageId="13" pageNumber="424">Chiriquí</collectingRegion>
Grande; and on the Pacific slope of
<collectingRegion id="0A3631F7FFA4292456BB0677FC9EF9BC" box="[690,775,1577,1598]" country="Panama" name="Panama" pageId="13" pageNumber="424">
<collectingCountry id="B0E5BF85FFA4292456BB0677FC9EF9BC" box="[690,775,1577,1598]" name="Panama" pageId="13" pageNumber="424">Panama</collectingCountry>
</collectingRegion>
,
<collectingRegion id="0A3631F7FFA42924571A0677FC56F9BC" box="[787,975,1577,1598]" country="Panama" name="Chiriqui" pageId="13" pageNumber="424">Provincia Chiriquı</collectingRegion>
´, Fortuna (
<figureCitation id="50C9E390FFA4292454F7061AFED9F9DB" box="[254,320,1604,1625]" captionStart="Fig. 6" captionStartId="12.[272,306,544,563]" captionTargetBox="[254,1079,273,520]" captionTargetId="figure@12.[248,1083,265,524]" captionTargetPageId="12" captionText="Fig. 6. Distribution of Staminodeus in Costa Rica, Panama, and Venezuela, S. bispinosus; S. curvitibialis (distributional record for Colombia, Provincia Nariño, La Planada is omitted); S. denticulatus; S. dilatatus; S. forcipis; S. inermis; S. vectoris." httpUri="https://zenodo.org/record/1208246/files/figure.png" pageId="13" pageNumber="424">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="80E8AC9EFFA4292755190600FCFAFEDB" lastPageId="14" lastPageNumber="425" pageId="13" pageNumber="424" type="biology_ecology">
<paragraph id="C84DFF15FFA4292755190600FCFAFEDB" blockId="13.[246,1079,1070,1652]" lastBlockId="14.[249,1081,271,345]" lastPageId="14" lastPageNumber="425" pageId="13" pageNumber="424">
<emphasis id="FA862307FFA4292455190600FE55F9F1" bold="true" box="[272,460,1630,1651]" pageId="13" pageNumber="424">Natural History.</emphasis>
<taxonomicName id="0FF28496FFA4292455DC0601FD7AF9F6" authorityName="Nico M. Franz" authorityYear="2001" box="[469,739,1631,1652]" class="Insecta" family="Curculionidae" genus="Staminodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="424" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="FA862307FFA4292455DC0601FD7AF9F6" box="[469,739,1631,1652]" italics="true" pageId="13" pageNumber="424">Staminodeus denticulatus</emphasis>
</taxonomicName>
is associated with
<emphasis id="FA862307FFA4292757BA0601FED6FEA6" italics="true" lastPageId="14" lastPageNumber="425" pageId="13" pageNumber="424">
<taxonomicName id="0FF28496FFA4292457BA0601FBAFF9F6" box="[947,1078,1631,1652]" class="Liliopsida" family="Cyclanthaceae" genus="Carludovica" higherTaxonomySource="CoL" kingdom="Plantae" order="Pandanales" pageId="13" pageNumber="424" phylum="Liliopsida" rank="genus">Carludovica</taxonomicName>
palmata
</emphasis>
Ruíz &amp; Pavón and
<emphasis id="FA862307FFA7292756290151FD69FEA6" box="[544,752,271,292]" italics="true" pageId="14" pageNumber="425">
<taxonomicName id="0FF28496FFA7292756290151FD3AFEA6" box="[544,675,271,292]" class="Liliopsida" family="Cyclanthaceae" genus="Evodianthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="425" phylum="Tracheophyta" rank="genus">Evodianthus</taxonomicName>
funifer
</emphasis>
(Poit.) Lindman in Costa Rica, and with
<taxonomicName id="0FF28496FFA7292755540177FE5EFEBC" authorityName="Harling" authorityYear="1954" box="[349,455,297,318]" class="Liliopsida" family="Cyclanthaceae" genus="Asplundia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="425" phylum="Tracheophyta" rank="genus">
<emphasis id="FA862307FFA7292755540177FE5EFEBC" box="[349,455,297,318]" italics="true" pageId="14" pageNumber="425">Asplundia</emphasis>
</taxonomicName>
sp. in Panama, Provincia Bocas del Toro, Chiriquí Grande (label information by Windsor &amp; Stockwell,
<date id="BC4CD9D5FFA7292756D5011AFCCEFEDB" box="[732,855,324,345]" pageId="14" pageNumber="425" value="1996-10-13">X­13–1996</date>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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