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<document id="69745A37C7F73EA0B44421B2304BF4C2" ID-DOI="10.11646/zootaxa.5399.5.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10517546" ID-ZooBank="09003490-4E1B-49B0-A73E-93C87EBA34E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1705393972181" checkinUser="plazi" docAuthor="Pelingen, Arthien Lovell" docDate="2024" docId="664887A7FFA8FF9CFF0FF99BFBE4FAD9" docLanguage="en" docName="zootaxa.5399.5.6.pdf" docOrigin="Zootaxa 5399 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoperla philippinensis Pelingen 2024, sp. nov." docType="treatment" docUuid="A1F061BA-25B9-43C8-AFA9-9EDD37F8455D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="575" masterDocId="9A71FFDFFFA9FF99FF98FF96FFC2FFA2" masterDocTitle="Description of Neoperla philippinensis sp. nov. confirmed using integrative taxonomy and some notes on Neoperla oculata Banks, 1924 (Plecoptera: Perlidae)" masterLastPageNumber="578" masterPageNumber="570" pageNumber="571" updateTime="1705602983961" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="06B6AD9BB09EF744B91DED27AFFF172F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E35E227B8A981A93F4FCF5DDC0A4FBF9">Description of Neoperla philippinensis sp. nov. confirmed using integrative taxonomy and some notes on Neoperla oculata Banks, 1924 (Plecoptera: Perlidae)</mods:title>
</mods:titleInfo>
<mods:name id="F673F032A5C6397F1B58DC6C81D21D9C" type="personal">
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<mods:roleTerm id="E76191FC1A69AAE4E9C7B1CEF2BCD462">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E95C18B7B2C5AFCF6DC59898DCED2A69">Pelingen, Arthien Lovell</mods:namePart>
<mods:affiliation id="45BF4529E4331B7D85BD6AC569C07D6B">MS Graduate Fellow, Department of Science and Technology- Science Education Institute, Taguig City 1631, Philippines &amp; Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City 1108, Philippines</mods:affiliation>
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<mods:title id="DCECC740D2474470D80E480B0BD96C0E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="06AB89C1D5C58EBB9D7558918254C1E8">
<mods:date id="D40E764F093F30EFAFB39B6479CCB7A5">2024</mods:date>
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<mods:number id="E2D2A6E1ECDD06337CB20309F1B9AB99">2024-01-16</mods:number>
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<mods:detail id="B8BA92F53A9FAFA180E727B033ACB87F" type="volume">
<mods:number id="05189432A7C1DAE555457A05B41D9E9A">5399</mods:number>
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<mods:detail id="C4415B0D6083D07500E1A615C993991D" type="issue">
<mods:number id="885A242610C6892347D0B812FD16DA67">5</mods:number>
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<mods:url id="B696FEFEF303DFCD624B77ABBE5D4092">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
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<mods:classification id="D6BC878FA09685F9022960145A30090F">journal article</mods:classification>
<mods:identifier id="C97BAD46769B86523FB10D83AED5F500" type="DOI">10.11646/zootaxa.5399.5.6</mods:identifier>
<mods:identifier id="544481FC8360E267B17E3C76F0358C57" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1F756DA17546A7A243354F6054A4AA15" type="Zenodo-Dep">10517546</mods:identifier>
<mods:identifier id="23583B99DE710794E82DC8C7D0A8A158" type="ZooBank">09003490-4E1B-49B0-A73E-93C87EBA34E3</mods:identifier>
</mods:mods>
<treatment id="664887A7FFA8FF9CFF0FF99BFBE4FAD9" ID-DOI="http://doi.org/10.5281/zenodo.10517480" ID-Zenodo-Dep="10517480" LSID="urn:lsid:zoobank.org:act:A1F061BA-25B9-43C8-AFA9-9EDD37F8455D" httpUri="http://treatment.plazi.org/id/664887A7FFA8FF9CFF0FF99BFBE4FAD9" lastPageId="5" lastPageNumber="575" pageId="1" pageNumber="571">
<subSubSection id="A6FB653AFFA8FF98FF0FF99BFC5EF9EF" pageId="1" pageNumber="571" type="nomenclature">
<paragraph id="EE5E36B1FFA8FF98FF0FF99BFD4DF98A" blockId="1.[151,924,1549,1613]" box="[151,655,1549,1576]" pageId="1" pageNumber="571">
<heading id="B51681DDFFA8FF98FF0FF99BFD4DF98A" bold="true" box="[151,655,1549,1576]" fontSize="11" level="1" pageId="1" pageNumber="571" reason="1">
<emphasis id="DC95EAA3FFA8FF98FF0FF99BFD4DF98A" bold="true" box="[151,655,1549,1576]" pageId="1" pageNumber="571">
<taxonomicName id="29E14D32FFA8FF98FF0FF99BFDE8F98A" authority="Pelingen" authorityName="Pelingen" authorityYear="2024" box="[151,554,1549,1576]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="571" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFA8FF98FF0FF99BFE75F98A" bold="true" box="[151,439,1549,1576]" italics="true" pageId="1" pageNumber="571">Neoperla philippinensis</emphasis>
Pelingen
</taxonomicName>
<taxonomicNameLabel id="C7A657D8FFA8FF98FDA9F998FD4DF98A" box="[561,655,1550,1576]" pageId="1" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EE5E36B1FFA8FF98FF0FF9A4FC5EF9EF" blockId="1.[151,924,1549,1613]" box="[151,924,1586,1613]" pageId="1" pageNumber="571">
https://zoobank.org/
<uuid id="9A470C64FFA8FF98FEE5F9A4FC5EF9EF" box="[381,924,1586,1613]" pageId="1" pageNumber="571">A1F061BA-25B9-43C8-AFA9-9EDD37F8455D</uuid>
</paragraph>
</subSubSection>
<subSubSection id="A6FB653AFFA8FF98FF0FF9E5FE0CF95B" pageId="1" pageNumber="571" type="materials_examined">
<paragraph id="EE5E36B1FFA8FF98FF0FF9E5FE0CF95B" blockId="1.[151,1437,1651,2037]" pageId="1" pageNumber="571">
<emphasis id="DC95EAA3FFA8FF98FF0FF9E5FE42F92F" bold="true" box="[151,384,1651,1677]" pageId="1" pageNumber="571">Material examined.</emphasis>
<materialsCitation id="5E893CECFFA8FF98FE1FF9E5FBF9F913" collectingDate="2019-01-13" collectionCode="NMP" collectorName="Siglos" country="Philippines" elevation="50" latitude="6.57" location="Pasonanca" longitude="122.066666" municipality="Mindanao" pageId="1" pageNumber="571" specimenCode="PL150" specimenCount="1" specimenCount-female="1" stateProvince="Zamboanga" typeStatus="holotype">
<emphasis id="DC95EAA3FFA8FF98FE1FF9E5FE3FF92F" bold="true" box="[391,509,1651,1677]" pageId="1" pageNumber="571">
<typeStatus id="315A8813FFA8FF98FE1FF9E5FE3AF92F" box="[391,504,1651,1677]" pageId="1" pageNumber="571" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="F8E7FD38FFA8FF98FD9CF9E5FDAFF92F" box="[516,621,1651,1680]" pageId="1" pageNumber="571">1 ♁ adult</specimenCount>
(
<specimenCode id="BE479ECAFFA8FF98FDE3F9E5FD09F92C" box="[635,715,1651,1678]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" pageNumber="571" type="Museum">PL150</specimenCode>
);
<emphasis id="DC95EAA3FFA8FF98FD47F9E5FC5EF92F" bold="true" box="[735,924,1651,1677]" pageId="1" pageNumber="571">
<collectingCountry id="96F67621FFA8FF98FD47F9E5FC54F92F" box="[735,918,1651,1677]" name="Philippines" pageId="1" pageNumber="571">PHILIPPINES</collectingCountry>
:
</emphasis>
<collectingMunicipality id="0E3AACCBFFA8FF98FC3AF9E5FBD4F92F" box="[930,1046,1651,1677]" pageId="1" pageNumber="571">Mindanao</collectingMunicipality>
,
<collectingRegion id="2C25F853FFA8FF98FBBAF9E5FB68F92F" box="[1058,1194,1651,1677]" country="Philippines" name="Zamboanga" pageId="1" pageNumber="571">Zamboanga</collectingRegion>
;
<location id="EB3E606AFFA8FF98FB2CF9E2FAE8F92C" LSID="urn:lsid:plazi:treatment:664887A7FFA8FF9CFF0FF99BFBE4FAD9:EB3E606AFFA8FF98FB2CF9E2FAE8F92C" box="[1204,1322,1652,1678]" country="Philippines" longitude="122.066666" municipality="Mindanao" name="Pasonanca" pageId="1" pageNumber="571" stateProvince="Zamboanga">Pasonanca</location>
[899]; ca.
<geoCoordinate id="8BD55076FFA8FF98FF0FF901FF2DF910" box="[151,239,1687,1714]" degrees="6" direction="north" minutes="57" pageId="1" pageNumber="571">6°57”N</geoCoordinate>
,
<geoCoordinate id="8BD55076FFA8FF98FF65F901FEA9F913" box="[253,363,1687,1713]" degrees="122" direction="east" minutes="04" orientation="longitude" pageId="1" pageNumber="571" precision="925" value="122.066666">122°04’E</geoCoordinate>
; ca.
<elevation id="65CCD182FFA8FF98FE39F901FDD9F913" box="[417,539,1687,1714]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="571" unit="m" value="50.0">
<quantity id="29199B54FFA8FF98FE39F901FE1EF910" box="[417,476,1687,1714]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="571" unit="m" value="50.0">50 m</quantity>
a.s.l.
</elevation>
;
<date id="9A5F1071FFA8FF98FDB1F901FD2BF913" box="[553,745,1687,1714]" pageId="1" pageNumber="571" value="2019-01-13">
<collectingDate id="8A1BE999FFA8FF98FDB1F901FD2BF913" box="[553,745,1687,1714]" pageId="1" pageNumber="571" value="2019-01-13">13 January 2019</collectingDate>
</date>
; leg.
<collectorName id="43145367FFA8FF98FCB0F901FCB3F913" box="[808,881,1687,1713]" pageId="1" pageNumber="571">Siglos</collectorName>
;
<collectionCode id="88F0AE74FFA8FF98FCE6F90EFC78F910" box="[894,954,1688,1714]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="1" pageNumber="571">NMP</collectionCode>
in ethanol.
</materialsCitation>
<materialsCitation id="5E893CECFFA8FF98FBDBF901FE0CF95B" collectingDate="2019-01-13" collectionCode="NMP" collectorName="Siglos" country="Philippines" elevation="50" location="Pasonanca" longitude="122.066666" municipality="Mindanao" pageId="1" pageNumber="571" specimenCode="PL151" specimenCount="1" specimenCount-female="1" stateProvince="Zamboanga" typeStatus="paratype">
<emphasis id="DC95EAA3FFA8FF98FBDBF901FB7BF913" bold="true" box="[1091,1209,1687,1713]" pageId="1" pageNumber="571">
<typeStatus id="315A8813FFA8FF98FBDBF901FB76F913" box="[1091,1204,1687,1713]" pageId="1" pageNumber="571" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="F8E7FD38FFA8FF98FB5AF901FAECF913" box="[1218,1326,1687,1714]" pageId="1" pageNumber="571">1 ♀ adult</specimenCount>
(
<specimenCode id="BE479ECAFFA8FF98FAA7F901FA4DF910" box="[1343,1423,1687,1714]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" pageNumber="571" type="Museum">PL151</specimenCode>
);
<emphasis id="DC95EAA3FFA8FF98FF0FF92DFE96F977" bold="true" box="[151,340,1723,1749]" pageId="1" pageNumber="571">
<collectingCountry id="96F67621FFA8FF98FF0FF92DFE8CF977" box="[151,334,1723,1749]" name="Philippines" pageId="1" pageNumber="571">PHILIPPINES</collectingCountry>
:
</emphasis>
<collectingMunicipality id="0E3AACCBFFA8FF98FEC2F92DFE0CF977" box="[346,462,1723,1749]" pageId="1" pageNumber="571">Mindanao</collectingMunicipality>
,
<collectingRegion id="2C25F853FFA8FF98FE41F92DFDA3F977" box="[473,609,1723,1749]" country="Philippines" name="Zamboanga" pageId="1" pageNumber="571">Zamboanga</collectingRegion>
;
<location id="EB3E606AFFA8FF98FDF3F92AFD23F974" LSID="urn:lsid:plazi:treatment:664887A7FFA8FF9CFF0FF99BFBE4FAD9:EB3E606AFFA8FF98FDF3F92AFD23F974" box="[619,737,1724,1750]" country="Philippines" longitude="122.066666" municipality="Mindanao" name="Pasonanca" pageId="1" pageNumber="571" stateProvince="Zamboanga">Pasonanca</location>
[899]; ca.
<geoCoordinate id="8BD55076FFA8FF98FCC1F92DFC73F974" box="[857,945,1723,1750]" pageId="1" pageNumber="571">6°57”N</geoCoordinate>
,
<geoCoordinate id="8BD55076FFA8FF98FC24F92DFBE8F977" box="[956,1066,1723,1749]" degrees="122" direction="east" minutes="04" orientation="longitude" pageId="1" pageNumber="571" precision="925" value="122.066666">122°04’E</geoCoordinate>
; ca.
<elevation id="65CCD182FFA8FF98FBC4F92DFB13F977" box="[1116,1233,1723,1750]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="571" unit="m" value="50.0">
<quantity id="29199B54FFA8FF98FBC4F92DFB56F974" box="[1116,1172,1723,1750]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="571" unit="m" value="50.0">50 m</quantity>
a.s.l.
</elevation>
;
<date id="9A5F1071FFA8FF98FB45F92DFA55F977" box="[1245,1431,1723,1750]" pageId="1" pageNumber="571" value="2019-01-13">
<collectingDate id="8A1BE999FFA8FF98FB45F92DFA55F977" box="[1245,1431,1723,1750]" pageId="1" pageNumber="571" value="2019-01-13">13 January 2019</collectingDate>
</date>
; leg.
<collectorName id="43145367FFA8FF98FF5FF949FED2F95B" box="[199,272,1759,1785]" pageId="1" pageNumber="571">Siglos</collectorName>
;
<collectionCode id="88F0AE74FFA8FF98FE84F976FE9AF958" box="[284,344,1760,1786]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="1" pageNumber="571">NMP</collectionCode>
in ethanol
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A6FB653AFFA8FF9BFF5FF895FEECF9DB" lastPageId="2" lastPageNumber="572" pageId="1" pageNumber="571" type="description">
<paragraph id="EE5E36B1FFA8FF98FF5FF895FF24F857" blockId="1.[151,1437,1651,2037]" pageId="1" pageNumber="571">
<emphasis id="DC95EAA3FFA8FF98FF5FF895FE6EF8BF" bold="true" box="[199,428,1795,1821]" pageId="1" pageNumber="571">
Description.
<emphasis id="DC95EAA3FFA8FF98FEF9F895FE6EF8BF" bold="true" box="[353,428,1795,1821]" italics="true" pageId="1" pageNumber="571">Imago</emphasis>
</emphasis>
(
<figureCitation id="76DA2A34FFA8FF98FE24F895FDD4F8BF" box="[444,534,1795,1821]" captionStart="FIGURE 1" captionStartId="2.[152,255,1165,1189]" captionTargetBox="[305,1283,181,1141]" captionTargetId="figure-15@2.[305,1283,181,1141]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus. a. Neoperla philippinensis sp. nov. b. Neoperla oculata Banks, 1924." figureDoi="http://doi.org/10.5281/zenodo.10517548" httpUri="https://zenodo.org/record/10517548/files/figure.png" pageId="1" pageNumber="571">Fig. 1A</figureCitation>
,
<figureCitation id="76DA2A34FFA8FF98FDBBF895FDF3F8BF" box="[547,561,1795,1821]" captionStart="FIGURE 3" captionStartId="4.[152,255,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-15@4.[214,1373,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Female habitus. Neoperla philippinensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10517554" httpUri="https://zenodo.org/record/10517554/files/figure.png" pageId="1" pageNumber="571">3</figureCitation>
): Small sized species. Forewing length of male: 10.0 mm, female: 12.0 mm. General color pale brownish yellow. Ocelli large, separated by a small gap. Head wider than pronotum, pale yellow, mottlings pale brown. M-line clearly visible. Antenna and palpi color same as head. Pronotum anterior corners angular, posterior corners curved, ground color same as head, has a median light brown stripe, area adjacent to the median line are paler; sublaterally, is a broad area of darker rugose pigmentation. Meso- and metanotum slightly darker than pronotum. Legs light brown, proportion 1.0: 1.2: 1.4, tibia darker than femur. Wings hyaline, veins brown.
</paragraph>
<caption id="BA9E6639FFABFF9BFF00FB1BFBB5FB07" ID-DOI="http://doi.org/10.5281/zenodo.10517548" ID-Zenodo-Dep="10517548" box="[152,1143,1165,1189]" httpUri="https://zenodo.org/record/10517548/files/figure.png" pageId="2" pageNumber="572" startId="2.[152,255,1165,1189]" targetBox="[305,1283,181,1141]" targetPageId="2" targetType="figure">
<paragraph id="EE5E36B1FFABFF9BFF00FB1BFBB5FB07" blockId="2.[152,1143,1165,1189]" box="[152,1143,1165,1189]" pageId="2" pageNumber="572">
<emphasis id="DC95EAA3FFABFF9BFF00FB1BFE70FB07" bold="true" box="[152,434,1165,1189]" pageId="2" pageNumber="572">FIGURE 1. Male habitus.</emphasis>
a.
<taxonomicName id="29E14D32FFABFF9BFE49FB1BFD01FB07" authority="Pelingen, 2024" authorityName="Pelingen" authorityYear="2024" box="[465,707,1165,1189]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFABFF9BFE49FB1BFD01FB07" box="[465,707,1165,1189]" italics="true" pageId="2" pageNumber="572">Neoperla philippinensis</emphasis>
</taxonomicName>
<emphasis id="DC95EAA3FFABFF9BFD52FB1BFCD8FB07" bold="true" box="[714,794,1165,1189]" pageId="2" pageNumber="572">
<taxonomicNameLabel id="C7A657D8FFABFF9BFD52FB1BFCD8FB07" box="[714,794,1165,1189]" pageId="2" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
b.
<taxonomicName id="29E14D32FFABFF9BFCA2FB1BFBB1FB07" authority="Banks, 1924" authorityName="Banks" authorityYear="1924" box="[826,1139,1165,1189]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFABFF9BFCA2FB1BFC29FB07" box="[826,1003,1165,1189]" italics="true" pageId="2" pageNumber="572">Neoperla oculata</emphasis>
<bibRefCitation id="8A704B40FFABFF9BFC69FB1BFBB1FB07" author="Banks, N." box="[1009,1139,1165,1189]" pageId="2" pageNumber="572" pagination="421 - 455" refId="ref3433" refString="Banks, N. (1924) Descriptions of new Neuropteroid insects. Bulletin of the Museum Comparative Zoology, 65, 421 - 455. [https: // www. biodiversitylibrary. org / page / 4774296]" type="journal article" year="1924">Banks, 1924</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="EE5E36B1FFABFF9BFF5FFB61FF11FA03" blockId="2.[151,1437,1271,2018]" pageId="2" pageNumber="572">
<emphasis id="DC95EAA3FFABFF9BFF5FFB61FE41FAB0" bold="true" box="[199,387,1271,1298]" italics="true" pageId="2" pageNumber="572">Male terminalia</emphasis>
(
<figureCitation id="76DA2A34FFABFF9BFE0BFB61FE2FFAB3" box="[403,493,1271,1297]" captionStart="FIGURE 2" captionStartId="3.[152,255,1963,1987]" captionTargetBox="[314,1274,595,1936]" captionTargetId="figure-135@3.[314,1274,595,1939]" captionTargetPageId="3" captionText="FIGURE 2. Male terminalia dorsal and aedeagus lateral. a. &amp; c. Neoperla philippinensis sp. nov. b. &amp; d. Neoperla oculata Banks, 1924 (typical form)." figureDoi="http://doi.org/10.5281/zenodo.10517552" httpUri="https://zenodo.org/record/10517552/files/figure.png" pageId="2" pageNumber="572">Fig. 2A</figureCitation>
): Sterna and terga 2–6 simple. Posterior of tergum 7 with small median hump less prominent than that of
<taxonomicName id="29E14D32FFABFF9BFE38FA8DFDA4FA97" authorityName="Banks" authorityYear="1924" box="[416,614,1307,1333]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFABFF9BFE38FA8DFDA4FA97" box="[416,614,1307,1333]" italics="true" pageId="2" pageNumber="572">Neoperla oculata</emphasis>
</taxonomicName>
with fewer sensilla basiconica. Tergum 8 without process, median with sensilla basiconica and long setae. Tergum 9 median with sparsely arranged long setae. Hemitergal lobe with simple dark edges and sparse quite long setae. Hemitergal lobe process curved sinuously and slightly raised in lateral view.
</paragraph>
<paragraph id="EE5E36B1FFABFF9BFF5FFA3DFDEEF9AF" blockId="2.[151,1437,1271,2018]" pageId="2" pageNumber="572">
<emphasis id="DC95EAA3FFABFF9BFF5FFA3DFEF5FA64" bold="true" box="[199,311,1451,1478]" italics="true" pageId="2" pageNumber="572">Aedeagus</emphasis>
(
<figureCitation id="76DA2A34FFABFF9BFEDEFA3DFE5DFA67" box="[326,415,1451,1477]" captionStart="FIGURE 2" captionStartId="3.[152,255,1963,1987]" captionTargetBox="[314,1274,595,1936]" captionTargetId="figure-135@3.[314,1274,595,1939]" captionTargetPageId="3" captionText="FIGURE 2. Male terminalia dorsal and aedeagus lateral. a. &amp; c. Neoperla philippinensis sp. nov. b. &amp; d. Neoperla oculata Banks, 1924 (typical form)." figureDoi="http://doi.org/10.5281/zenodo.10517552" httpUri="https://zenodo.org/record/10517552/files/figure.png" pageId="2" pageNumber="572">Fig. 2C</figureCitation>
): Aedeagal tube plump, larger second medioventral lobe heavily covered with spines, small but more distinct basolateral lobes covered with fine spines, distinctly smaller but densely covered midlateral lobes, and aedeagal sac bent less ventrally.
</paragraph>
<paragraph id="EE5E36B1FFABFF9BFF5FF981FEECF9DB" blockId="2.[151,1437,1271,2018]" pageId="2" pageNumber="572">
<emphasis id="DC95EAA3FFABFF9BFF5FF981FF34F993" bold="true" box="[199,246,1559,1585]" pageId="2" pageNumber="572">Egg</emphasis>
(
<figureCitation id="76DA2A34FFABFF9BFE9EF981FE48F993" box="[262,394,1559,1585]" captionStart="FIGURE 4" captionStartId="5.[152,255,1162,1186]" captionTargetBox="[151,1435,181,1139]" captionTargetId="figure-28@5.[151,1435,181,1139]" captionTargetPageId="5" captionText="FIGURE 4. Eggs. a. &amp; b. Neoperla oculata Banks, 1924. c. &amp; d. Neoperla philippinensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10517556" httpUri="https://zenodo.org/record/10517556/files/figure.png" pageId="2" pageNumber="572">Figs 4C–D</figureCitation>
): Color dark brown, shape spherical ovoid, hatching line visible. Length ca. 315 μm, width ca. 210 μm. Punctate chorion with more proportionate hexagons in the collar end. Micropyles small, nearly unnoticeable.
</paragraph>
</subSubSection>
<subSubSection id="A6FB653AFFABFF9CFF5FF915FE66FAAD" lastPageId="5" lastPageNumber="575" pageId="2" pageNumber="572" type="diagnosis">
<paragraph id="EE5E36B1FFABFF9BFF5FF915FBFBF843" blockId="2.[151,1437,1271,2018]" pageId="2" pageNumber="572">
<emphasis id="DC95EAA3FFABFF9BFF5FF915FE1BF93F" bold="true" box="[199,473,1667,1693]" pageId="2" pageNumber="572">Differential Diagnosis.</emphasis>
At first glance, the aedeagus resembles that of
<taxonomicName id="29E14D32FFABFF9BFBBAF912FB5FF93F" authorityName="Banks" authorityYear="1924" box="[1058,1181,1667,1693]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFABFF9BFBBAF912FB5FF93F" box="[1058,1181,1667,1693]" italics="true" pageId="2" pageNumber="572">N. oculata</emphasis>
</taxonomicName>
because the defining characteristic in the key provided in
<bibRefCitation id="8A704B40FFABFF9BFDA4F931FD0CF963" author="Sivec, I." box="[572,718,1703,1729]" pageId="2" pageNumber="572" pagination="1 - 44" refId="ref4528" refString="Sivec, I. (1984) Study of genus Neoperla (Plecoptera: Perlidae) from the Philippines. Scopolia, 7, 1 - 44. [https: // www. zobodat. at / pdf / Scopolia _ 7 _ 0001 - 0044. pdf]" type="journal article" year="1984">Sivec (1984)</bibRefCitation>
is “two ventral outgrowths, a pair of subterminal lateroventral outgrowths, and a pair of dorsolateral outgrowths.” The illustration in
<bibRefCitation id="8A704B40FFABFF9BFC51F95DFB48F947" author="Sivec, I." box="[969,1162,1739,1765]" pageId="2" pageNumber="572" pagination="1 - 44" refId="ref4528" refString="Sivec, I. (1984) Study of genus Neoperla (Plecoptera: Perlidae) from the Philippines. Scopolia, 7, 1 - 44. [https: // www. zobodat. at / pdf / Scopolia _ 7 _ 0001 - 0044. pdf]" type="journal article" year="1984">Sivec (1984: 31)</bibRefCitation>
with caption “variation from Rizal’’ more closely resembles the aedeagus of the specimen at hand. However, the specimens examined in this study were found in Mindanao. This is also different from the “typical form” of
<taxonomicName id="29E14D32FFABFF9BFBC2F882FB12F88F" authorityName="Banks" authorityYear="1924" box="[1114,1232,1811,1837]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFABFF9BFBC2F882FB12F88F" box="[1114,1232,1811,1837]" italics="true" pageId="2" pageNumber="572">N. oculata</emphasis>
</taxonomicName>
aedeagus. Hence, it is distinct from both forms of
<taxonomicName id="29E14D32FFABFF9BFD9AF8AEFDBAF8F3" authorityName="Banks" authorityYear="1924" box="[514,632,1847,1873]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFABFF9BFD9AF8AEFDBAF8F3" box="[514,632,1847,1873]" italics="true" pageId="2" pageNumber="572">N. oculata</emphasis>
</taxonomicName>
. Morphologically, the obvious differences of
<taxonomicName id="29E14D32FFABFF9BFBE4F8AEFAF8F8F3" authority="Pelingen, 2024" authorityName="Pelingen" authorityYear="2024" box="[1148,1338,1847,1873]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFABFF9BFBE4F8AEFAF8F8F3" box="[1148,1338,1847,1873]" italics="true" pageId="2" pageNumber="572">N. philippinensis</emphasis>
</taxonomicName>
<emphasis id="DC95EAA3FFABFF9BFADAF8A1FA5EF8F3" bold="true" box="[1346,1436,1847,1873]" pageId="2" pageNumber="572">
<taxonomicNameLabel id="C7A657D8FFABFF9BFADAF8A1FA5EF8F3" box="[1346,1436,1847,1873]" pageId="2" pageNumber="572" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="29E14D32FFABFF9BFF4CF8CAFE8BF8D7" authorityName="Banks" authorityYear="1924" box="[212,329,1883,1909]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFABFF9BFF4CF8CAFE8BF8D7" box="[212,329,1883,1909]" italics="true" pageId="2" pageNumber="572">N. oculata</emphasis>
</taxonomicName>
in the aedeagus are as follows: larger second medioventral lobe heavily covered with spines, small but more distinct basolateral lobes covered with fine spines, distinctly smaller but densely covered mediolateral lobes, and a less ventrad-bend to the aedeagal sac. The genetic divergence distances (
<tableCitation id="A363030AFFABFF9BFBD6F835FB66F81F" box="[1102,1188,1955,1981]" captionStart="TABLE 2" captionStartId="3.[152,245,151,175]" captionTargetPageId="3" captionText="TABLE 2. Pairwise distances of Neoperla partial COI sequences (620 bp) based on the Kimura-two-parameter (K2P) model." httpUri="http://table.plazi.org/id/BA9E6639FFAAFF9AFF00FF01FF23FF71" pageId="2" pageNumber="572" tableUuid="BA9E6639FFAAFF9AFF00FF01FF23FF71">Table 2</tableCitation>
) also show that this is a different species from the
<taxonomicName id="29E14D32FFABFF9BFE48F85EFD87F843" authorityName="Banks" authorityYear="1924" box="[464,581,1991,2017]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="572" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFABFF9BFE48F85EFD87F843" box="[464,581,1991,2017]" italics="true" pageId="2" pageNumber="572">N. oculata</emphasis>
</taxonomicName>
(6.49–7.58% interspecific genetic distance).
</paragraph>
<caption id="BA9E6639FFAAFF9AFF00FF01FF23FF71" ID-Table-UUID="BA9E6639FFAAFF9AFF00FF01FF23FF71" httpUri="http://table.plazi.org/id/BA9E6639FFAAFF9AFF00FF01FF23FF71" pageId="3" pageNumber="573" startId="3.[152,245,151,175]" targetBox="[171,1393,227,534]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="EE5E36B1FFAAFF9AFF00FF01FF23FF71" blockId="3.[151,1436,151,211]" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF00FF01FED2FF0D" bold="true" box="[152,272,151,175]" pageId="3" pageNumber="573">TABLE 2.</emphasis>
Pairwise distances of
<taxonomicName id="29E14D32FFAAFF9AFD9DFF01FDAAFF0D" authorityName="Needham" authorityYear="1905" box="[517,616,151,175]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="DC95EAA3FFAAFF9AFD9DFF01FDAAFF0D" box="[517,616,151,175]" italics="true" pageId="3" pageNumber="573">Neoperla</emphasis>
</taxonomicName>
partial COI sequences (620 bp) based on the Kimura-two-parameter (K2P) model.
</paragraph>
</caption>
<paragraph id="EE5E36B1FFAAFF9AFEFCFF72FADFFDB4" pageId="3" pageNumber="573">
<table id="9CE1C411FFAA0066FF33FF75FAB3FDB4" box="[171,1393,227,534]" gridcols="9" gridrows="8" pageId="3" pageNumber="573">
<tr id="50D134F3FFAA0066FF33FF75FAB3FF5E" box="[171,1393,227,252]" gridrow="0" pageId="3" pageNumber="573" rowspan-0="1">
<th id="13005D8FFFAA0066FF7CFF75FDF0FF5E" box="[228,562,227,252]" gridcol="1" gridrow="0" pageId="3" pageNumber="573">Species</th>
<th id="13005D8FFFAA0066FDFEFF75FD6EFF5E" box="[614,684,227,252]" gridcol="2" gridrow="0" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFD1BFF75FD52FF59" bold="true" box="[643,656,227,251]" pageId="3" pageNumber="573">1</emphasis>
</th>
<th id="13005D8FFFAA0066FD7BFF75FCEBFF5E" box="[739,809,227,252]" gridcol="3" gridrow="0" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFC98FF75FCCFFF59" bold="true" box="[768,781,227,251]" pageId="3" pageNumber="573">2</emphasis>
</th>
<th id="13005D8FFFAA0066FCF8FF75FC64FF5E" box="[864,934,227,252]" gridcol="4" gridrow="0" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFCE5FF75FC48FF59" bold="true" box="[893,906,227,251]" pageId="3" pageNumber="573">3</emphasis>
</th>
<th id="13005D8FFFAA0066FC45FF75FBE1FF5E" box="[989,1059,227,252]" gridcol="5" gridrow="0" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFC62FF75FBC5FF59" bold="true" box="[1018,1031,227,251]" pageId="3" pageNumber="573">4</emphasis>
</th>
<th id="13005D8FFFAA0066FBC2FF75FB62FF5E" box="[1114,1184,227,252]" gridcol="6" gridrow="0" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFBEFFF75FB46FF59" bold="true" box="[1143,1156,227,251]" pageId="3" pageNumber="573">5</emphasis>
</th>
<th id="13005D8FFFAA0066FB4FFF75FADFFF5E" box="[1239,1309,227,252]" gridcol="7" gridrow="0" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFB6BFF75FAC2FF59" bold="true" box="[1267,1280,227,251]" pageId="3" pageNumber="573">6</emphasis>
</th>
<th id="13005D8FFFAA0066FAFCFF75FAB3FF5E" box="[1380,1393,227,252]" gridcol="8" gridrow="0" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFAFCFF75FAB3FF59" bold="true" box="[1380,1393,227,251]" pageId="3" pageNumber="573">7</emphasis>
</th>
</tr>
<tr id="50D134F3FFAA0066FF33FE9DFAB3FE86" box="[171,1393,267,292]" gridrow="1" pageId="3" pageNumber="573" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="13005D8FFFAA0066FF33FE9DFF7AFE86" box="[171,184,267,292]" gridcol="0" gridrow="1" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF33FE9DFF7AFE81" bold="true" box="[171,184,267,291]" pageId="3" pageNumber="573">1</emphasis>
</th>
<td id="13005D8FFFAA0066FF7CFE9DFDF0FE86" box="[228,562,267,292]" gridcol="1" gridrow="1" pageId="3" pageNumber="573">
<taxonomicName id="29E14D32FFAAFF9AFF7CFE9BFE52FE86" authority="Pelingen, 2024" authorityName="Pelingen" authorityYear="2024" box="[228,400,268,292]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFAAFF9AFF7CFE9BFE52FE86" box="[228,400,268,292]" italics="true" pageId="3" pageNumber="573">N. philippinensis</emphasis>
</taxonomicName>
<emphasis id="DC95EAA3FFAAFF9AFE0EFE9AFE25FE86" bold="true" box="[406,487,268,292]" pageId="3" pageNumber="573">
<taxonomicNameLabel id="C7A657D8FFAAFF9AFE0EFE9AFE25FE86" box="[406,487,268,292]" pageId="3" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
PL150
</td>
</tr>
<tr id="50D134F3FFAA0066FF33FEA2FAB3FEEE" box="[171,1393,308,332]" gridrow="2" pageId="3" pageNumber="573" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="13005D8FFFAA0066FF33FEA2FF7AFEEE" box="[171,184,308,332]" gridcol="0" gridrow="2" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF33FEA2FF7AFEEE" bold="true" box="[171,184,308,332]" pageId="3" pageNumber="573">2</emphasis>
</th>
<td id="13005D8FFFAA0066FF7CFEA2FDF0FEEE" box="[228,562,308,332]" gridcol="1" gridrow="2" pageId="3" pageNumber="573">
<taxonomicName id="29E14D32FFAAFF9AFF7CFEA3FE52FEEE" authority="Pelingen, 2024" authorityName="Pelingen" authorityYear="2024" box="[228,400,308,332]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFAAFF9AFF7CFEA3FE52FEEE" box="[228,400,308,332]" italics="true" pageId="3" pageNumber="573">N. philippinensis</emphasis>
</taxonomicName>
<emphasis id="DC95EAA3FFAAFF9AFE0EFEA2FE25FEEE" bold="true" box="[406,487,308,332]" pageId="3" pageNumber="573">
<taxonomicNameLabel id="C7A657D8FFAAFF9AFE0EFEA2FE25FEEE" box="[406,487,308,332]" pageId="3" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
PL151
</td>
<td id="13005D8FFFAA0066FDFEFEA2FD6EFEEE" box="[614,684,308,332]" gridcol="2" gridrow="2" pageId="3" pageNumber="573">0.0097</td>
</tr>
<tr id="50D134F3FFAA0066FF33FECAFAB3FED7" box="[171,1393,348,373]" gridrow="3" pageId="3" pageNumber="573" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="13005D8FFFAA0066FF33FECAFF7AFED7" box="[171,184,348,373]" gridcol="0" gridrow="3" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF33FECAFF7AFED6" bold="true" box="[171,184,348,372]" pageId="3" pageNumber="573">3</emphasis>
</th>
<td id="13005D8FFFAA0066FF7CFECAFDF0FED7" box="[228,562,348,373]" gridcol="1" gridrow="3" pageId="3" pageNumber="573">
<taxonomicName id="29E14D32FFAAFF9AFEAFFECBFE04FED7" authority="PL" authorityName="PL" box="[311,454,349,373]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFAAFF9AFEAFFECBFE63FED7" box="[311,417,349,373]" italics="true" pageId="3" pageNumber="573">N. oculata</emphasis>
PL
</taxonomicName>
82
</td>
<td id="13005D8FFFAA0066FDFEFECAFD6EFED7" box="[614,684,348,373]" gridcol="2" gridrow="3" pageId="3" pageNumber="573">0.0684</td>
<td id="13005D8FFFAA0066FD7BFECAFCEBFED7" box="[739,809,348,373]" gridcol="3" gridrow="3" pageId="3" pageNumber="573">0.0649</td>
</tr>
<tr id="50D134F3FFAA0066FF33FE12FAB3FE3F" box="[171,1393,388,413]" gridrow="4" pageId="3" pageNumber="573" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="13005D8FFFAA0066FF33FE12FF7AFE3F" box="[171,184,388,413]" gridcol="0" gridrow="4" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF33FE12FF7AFE3E" bold="true" box="[171,184,388,412]" pageId="3" pageNumber="573">4</emphasis>
</th>
<td id="13005D8FFFAA0066FF7CFE12FDF0FE3F" box="[228,562,388,413]" gridcol="1" gridrow="4" pageId="3" pageNumber="573">
<taxonomicName id="29E14D32FFAAFF9AFEAFFE10FE04FE3F" authority="PL" authorityName="PL" box="[311,454,389,413]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFAAFF9AFEAFFE10FE63FE3F" box="[311,417,389,413]" italics="true" pageId="3" pageNumber="573">N. oculata</emphasis>
PL
</taxonomicName>
91
</td>
<td id="13005D8FFFAA0066FDFEFE12FD6EFE3F" box="[614,684,388,413]" gridcol="2" gridrow="4" pageId="3" pageNumber="573">0.0758</td>
<td id="13005D8FFFAA0066FD7BFE12FCEBFE3F" box="[739,809,388,413]" gridcol="3" gridrow="4" pageId="3" pageNumber="573">0.0722</td>
<td id="13005D8FFFAA0066FCF8FE12FC64FE3F" box="[864,934,388,413]" gridcol="4" gridrow="4" pageId="3" pageNumber="573">0.0098</td>
</tr>
<tr id="50D134F3FFAA0066FF33FE3BFAB3FE67" box="[171,1393,429,453]" gridrow="5" pageId="3" pageNumber="573" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="13005D8FFFAA0066FF33FE3BFF7AFE67" box="[171,184,429,453]" gridcol="0" gridrow="5" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF33FE3BFF7AFE67" bold="true" box="[171,184,429,453]" pageId="3" pageNumber="573">5</emphasis>
</th>
<td id="13005D8FFFAA0066FF7CFE3BFDF0FE67" box="[228,562,429,453]" gridcol="1" gridrow="5" pageId="3" pageNumber="573">
<taxonomicName id="29E14D32FFAAFF9AFEA9FE38FE03FE67" authority="PL" authorityName="PL" box="[305,449,429,453]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFAAFF9AFEA9FE38FE59FE67" box="[305,411,429,453]" italics="true" pageId="3" pageNumber="573">N. oculata</emphasis>
PL
</taxonomicName>
101
</td>
<td id="13005D8FFFAA0066FDFEFE3BFD6EFE67" box="[614,684,429,453]" gridcol="2" gridrow="5" pageId="3" pageNumber="573">0.0684</td>
<td id="13005D8FFFAA0066FD7BFE3BFCEBFE67" box="[739,809,429,453]" gridcol="3" gridrow="5" pageId="3" pageNumber="573">0.0649</td>
<td id="13005D8FFFAA0066FCF8FE3BFC64FE67" box="[864,934,429,453]" gridcol="4" gridrow="5" pageId="3" pageNumber="573">0.0000</td>
<td id="13005D8FFFAA0066FC45FE3BFBE1FE67" box="[989,1059,429,453]" gridcol="5" gridrow="5" pageId="3" pageNumber="573">0.0098</td>
</tr>
<tr id="50D134F3FFAA0066FF33FE43FAB3FE4F" box="[171,1393,469,493]" gridrow="6" pageId="3" pageNumber="573" rowspan-7="1" rowspan-8="1">
<th id="13005D8FFFAA0066FF33FE43FF7AFE4F" box="[171,184,469,493]" gridcol="0" gridrow="6" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF33FE43FF7AFE4F" bold="true" box="[171,184,469,493]" pageId="3" pageNumber="573">6</emphasis>
</th>
<td id="13005D8FFFAA0066FF7CFE43FDF0FE4F" box="[228,562,469,493]" gridcol="1" gridrow="6" pageId="3" pageNumber="573">
<taxonomicName id="29E14D32FFAAFF9AFEA9FE40FE02FE4F" authority="PL" authorityName="PL" box="[305,448,469,493]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFAAFF9AFEA9FE40FE59FE4F" box="[305,411,469,493]" italics="true" pageId="3" pageNumber="573">N. oculata</emphasis>
PL
</taxonomicName>
108
</td>
<td id="13005D8FFFAA0066FDFEFE43FD6EFE4F" box="[614,684,469,493]" gridcol="2" gridrow="6" pageId="3" pageNumber="573">0.0684</td>
<td id="13005D8FFFAA0066FD7BFE43FCEBFE4F" box="[739,809,469,493]" gridcol="3" gridrow="6" pageId="3" pageNumber="573">0.0649</td>
<td id="13005D8FFFAA0066FCF8FE43FC64FE4F" box="[864,934,469,493]" gridcol="4" gridrow="6" pageId="3" pageNumber="573">0.0000</td>
<td id="13005D8FFFAA0066FC45FE43FBE1FE4F" box="[989,1059,469,493]" gridcol="5" gridrow="6" pageId="3" pageNumber="573">0.0098</td>
<td id="13005D8FFFAA0066FBC2FE43FB62FE4F" box="[1114,1184,469,493]" gridcol="6" gridrow="6" pageId="3" pageNumber="573">0.0000</td>
</tr>
<tr id="50D134F3FFAA0066FF33FE6BFAB3FDB4" box="[171,1393,509,534]" gridrow="7" pageId="3" pageNumber="573" rowspan-8="1">
<th id="13005D8FFFAA0066FF33FE6BFF7AFDB4" box="[171,184,509,534]" gridcol="0" gridrow="7" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF33FE6BFF7AFDB7" bold="true" box="[171,184,509,533]" pageId="3" pageNumber="573">7</emphasis>
</th>
<td id="13005D8FFFAA0066FF7CFE6BFDF0FDB4" box="[228,562,509,534]" gridcol="1" gridrow="7" pageId="3" pageNumber="573">
<taxonomicName id="29E14D32FFAAFF9AFEA9FE68FE02FDB4" authority="PL" authorityName="PL" box="[305,448,510,534]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFAAFF9AFEA9FE68FE59FDB4" box="[305,411,510,534]" italics="true" pageId="3" pageNumber="573">N. oculata</emphasis>
PL
</taxonomicName>
162
</td>
<td id="13005D8FFFAA0066FDFEFE6BFD6EFDB4" box="[614,684,509,534]" gridcol="2" gridrow="7" pageId="3" pageNumber="573">0.0740</td>
<td id="13005D8FFFAA0066FD7BFE6BFCEBFDB4" box="[739,809,509,534]" gridcol="3" gridrow="7" pageId="3" pageNumber="573">0.0704</td>
<td id="13005D8FFFAA0066FCF8FE6BFC64FDB4" box="[864,934,509,534]" gridcol="4" gridrow="7" pageId="3" pageNumber="573">0.0081</td>
<td id="13005D8FFFAA0066FC45FE6BFBE1FDB4" box="[989,1059,509,534]" gridcol="5" gridrow="7" pageId="3" pageNumber="573">0.0049</td>
<td id="13005D8FFFAA0066FBC2FE6BFB62FDB4" box="[1114,1184,509,534]" gridcol="6" gridrow="7" pageId="3" pageNumber="573">0.0081</td>
<td id="13005D8FFFAA0066FB4FFE6BFADFFDB4" box="[1239,1309,509,534]" gridcol="7" gridrow="7" pageId="3" pageNumber="573">0.0081</td>
</tr>
</table>
</paragraph>
<caption id="BA9E6639FFAAFF9AFF00F83DFE73F845" ID-DOI="http://doi.org/10.5281/zenodo.10517552" ID-Zenodo-Dep="10517552" httpUri="https://zenodo.org/record/10517552/files/figure.png" pageId="3" pageNumber="573" startId="3.[152,255,1963,1987]" targetBox="[314,1274,595,1936]" targetPageId="3" targetType="figure">
<paragraph id="EE5E36B1FFAAFF9AFF00F83DFE73F845" blockId="3.[151,1436,1963,2023]" pageId="3" pageNumber="573">
<emphasis id="DC95EAA3FFAAFF9AFF00F83DFCC2F861" bold="true" box="[152,768,1963,1987]" pageId="3" pageNumber="573">FIGURE 2. Male terminalia dorsal and aedeagus lateral.</emphasis>
a. &amp; c.
<taxonomicName id="29E14D32FFAAFF9AFCC8F83DFB80F861" authority="Pelingen, 2024" authorityName="Pelingen" authorityYear="2024" box="[848,1090,1963,1987]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFAAFF9AFCC8F83DFB80F861" box="[848,1090,1963,1987]" italics="true" pageId="3" pageNumber="573">Neoperla philippinensis</emphasis>
</taxonomicName>
<emphasis id="DC95EAA3FFAAFF9AFBD1F83DFB5BF861" bold="true" box="[1097,1177,1963,1987]" pageId="3" pageNumber="573">
<taxonomicNameLabel id="C7A657D8FFAAFF9AFBD1F83DFB5BF861" box="[1097,1177,1963,1987]" pageId="3" pageNumber="573" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
b. &amp; d.
<taxonomicName id="29E14D32FFAAFF9AFB74F83DFED4F845" authority="Banks, 1924" authorityName="Banks" authorityYear="1924" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="573" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFAAFF9AFB74F83DFA5EF861" box="[1260,1436,1963,1987]" italics="true" pageId="3" pageNumber="573">Neoperla oculata</emphasis>
<bibRefCitation id="8A704B40FFAAFF9AFF0FF859FED4F845" author="Banks, N." box="[151,278,1999,2023]" pageId="3" pageNumber="573" pagination="421 - 455" refId="ref3433" refString="Banks, N. (1924) Descriptions of new Neuropteroid insects. Bulletin of the Museum Comparative Zoology, 65, 421 - 455. [https: // www. biodiversitylibrary. org / page / 4774296]" type="journal article" year="1924">Banks, 1924</bibRefCitation>
</taxonomicName>
(typical form).
</paragraph>
</caption>
<caption id="BA9E6639FFADFF9DFF00F801FCCDF812" ID-DOI="http://doi.org/10.5281/zenodo.10517554" ID-Zenodo-Dep="10517554" box="[152,783,1943,1968]" httpUri="https://zenodo.org/record/10517554/files/figure.png" pageId="4" pageNumber="574" startId="4.[152,255,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="4" targetType="figure">
<paragraph id="EE5E36B1FFADFF9DFF00F801FCCDF812" blockId="4.[152,783,1943,1968]" box="[152,783,1943,1968]" pageId="4" pageNumber="574">
<emphasis id="DC95EAA3FFADFF9DFF00F801FEDAF80D" bold="true" box="[152,280,1943,1967]" pageId="4" pageNumber="574">FIGURE 3.</emphasis>
Female habitus.
<taxonomicName id="29E14D32FFADFF9DFE5DF80EFD7AF812" authority="Pelingen, 2024" authorityName="Pelingen" authorityYear="2024" box="[453,696,1944,1968]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="574" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFADFF9DFE5DF80EFD7AF812" box="[453,696,1944,1968]" italics="true" pageId="4" pageNumber="574">Neoperla philippinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D8FFADFF9DFD27F80EFCCDF812" box="[703,783,1944,1968]" pageId="4" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="BA9E6639FFACFF9CFF00FB1CFB40FB01" ID-DOI="http://doi.org/10.5281/zenodo.10517556" ID-Zenodo-Dep="10517556" box="[152,1154,1162,1187]" httpUri="https://zenodo.org/record/10517556/files/figure.png" pageId="5" pageNumber="575" startId="5.[152,255,1162,1186]" targetBox="[151,1435,181,1139]" targetPageId="5" targetType="figure">
<paragraph id="EE5E36B1FFACFF9CFF00FB1CFB40FB01" blockId="5.[152,1154,1162,1187]" box="[152,1154,1162,1187]" pageId="5" pageNumber="575">
<emphasis id="DC95EAA3FFACFF9CFF00FB1CFE98FB01" bold="true" box="[152,346,1162,1187]" pageId="5" pageNumber="575">FIGURE 4. Eggs.</emphasis>
a. &amp; b.
<taxonomicName id="29E14D32FFACFF9CFE33FB1DFD26FB01" authority="Banks, 1924" authorityName="Banks" authorityYear="1924" box="[427,740,1163,1187]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="oculata">
<emphasis id="DC95EAA3FFACFF9CFE33FB1DFD99FB01" box="[427,603,1163,1187]" italics="true" pageId="5" pageNumber="575">Neoperla oculata</emphasis>
<bibRefCitation id="8A704B40FFACFF9CFDFAFB1DFD26FB01" author="Banks, N." box="[610,740,1163,1187]" pageId="5" pageNumber="575" pagination="421 - 455" refId="ref3433" refString="Banks, N. (1924) Descriptions of new Neuropteroid insects. Bulletin of the Museum Comparative Zoology, 65, 421 - 455. [https: // www. biodiversitylibrary. org / page / 4774296]" type="journal article" year="1924">Banks, 1924</bibRefCitation>
</taxonomicName>
. c. &amp; d.
<taxonomicName id="29E14D32FFACFF9CFCA0FB1DFBE9FB01" authority="Pelingen, 2024" authorityName="Pelingen" authorityYear="2024" box="[824,1067,1163,1187]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="575" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="DC95EAA3FFACFF9CFCA0FB1DFBE9FB01" box="[824,1067,1163,1187]" italics="true" pageId="5" pageNumber="575">Neoperla philippinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D8FFACFF9CFBA9FB1DFB40FB01" box="[1073,1154,1163,1187]" pageId="5" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="EE5E36B1FFACFF9CFF5FFB63FE66FAAD" blockId="5.[151,1436,1269,1403]" box="[199,420,1269,1295]" pageId="5" pageNumber="575">
<emphasis id="DC95EAA3FFACFF9CFF5FFB63FEE2FAAD" bold="true" box="[199,288,1269,1295]" pageId="5" pageNumber="575">Nymph</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A6FB653AFFACFF9CFF5FFA8FFD11FAF5" pageId="5" pageNumber="575" type="etymology">
<paragraph id="EE5E36B1FFACFF9CFF5FFA8FFD11FAF5" blockId="5.[151,1436,1269,1403]" pageId="5" pageNumber="575">
<emphasis id="DC95EAA3FFACFF9CFF5FFA8FFE8CFA91" bold="true" box="[199,334,1305,1331]" pageId="5" pageNumber="575">Etymology.</emphasis>
The specific epithet refers to the name of the country of origin,
<collectingCountry id="96F67621FFACFF9CFB8DFA8FFB54FA91" box="[1045,1174,1305,1331]" name="Philippines" pageId="5" pageNumber="575">Philippines</collectingCountry>
, where stonefly species diversity has been overlooked for the past decades.
</paragraph>
</subSubSection>
<subSubSection id="A6FB653AFFACFF9CFF5FFAF6FBE4FAD9" box="[199,1062,1376,1403]" pageId="5" pageNumber="575" type="distribution">
<paragraph id="EE5E36B1FFACFF9CFF5FFAF6FBE4FAD9" blockId="5.[151,1436,1269,1403]" box="[199,1062,1376,1403]" pageId="5" pageNumber="575">
<emphasis id="DC95EAA3FFACFF9CFF5FFAF6FEA3FAD8" bold="true" box="[199,353,1376,1402]" pageId="5" pageNumber="575">Distribution.</emphasis>
This species is known so far only from Mindanao,
<collectingCountry id="96F67621FFACFF9CFC39FAF7FBE1FAD9" box="[929,1059,1377,1403]" name="Philippines" pageId="5" pageNumber="575">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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