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<document ID-DOI="10.11646/zootaxa.5207.1.1" ID-GBIF-Dataset="a1bc8125-ad00-4d07-b9a1-4e6420c6bd98" ID-ISSN="1175­5334" ID-Zenodo-Dep="7328080" ID-ZooBank="A57EB27B-8524-42D8-8B85-564F879A5938" checkinTime="1668423676913" checkinUser="plazi" docAuthor="TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG &amp; WANG, XINGMIN" docDate="2022" docId="314287B0F639FFA9FF7BFDAE0EDCF9D3" docLanguage="en" docName="zootaxa.5207.1.1.pdf" docOrigin="Zootaxa 5207 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5207.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.4:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="4" docTitle="Sasajiscymnus sylvaticus" docType="key" docVersion="2" lastPageNumber="9" masterDocId="CD7BFFC8F631FFA1FFECFFE20B40FF81" masterDocTitle="A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae)" masterLastPageNumber="104" masterPageNumber="1" pageNumber="9" updateTime="1668629162570" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>TONG, JUNBO</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CHEN, XIAOSHENG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ZHANG, XIAONING</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>HUO, LIZHI</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WU, GANG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>PENG, ZHENGQIANG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WANG, XINGMIN</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5207</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5207.1.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5207.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">a1bc8125-ad00-4d07-b9a1-4e6420c6bd98</mods:identifier>
<mods:identifier type="ISSN">1175­5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">7328080</mods:identifier>
<mods:identifier type="ZooBank">A57EB27B-8524-42D8-8B85-564F879A5938</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7328100" ID-Zenodo-Dep="7328100" LSID="urn:lsid:plazi:treatment:314287B0F639FFA9FF7BFDAE0EDCF9D3" httpUri="http://treatment.plazi.org/id/314287B0F639FFA9FF7BFDAE0EDCF9D3" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection box="[151,629,588,615]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph blockId="8.[151,629,588,615]" box="[151,629,588,615]" pageId="8" pageNumber="9">
<heading bold="true" box="[151,629,588,615]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis bold="true" box="[151,629,588,615]" pageId="8" pageNumber="9">
<taxonomicName baseAuthorityName="Lewis" baseAuthorityYear="1896" box="[151,453,588,615]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sylvaticus">
<emphasis bold="true" box="[151,453,588,615]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus sylvaticus</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="discussion">
<paragraph blockId="8.[151,1437,660,831]" pageId="8" pageNumber="9">
The
<taxonomicName baseAuthorityName="Lewis" baseAuthorityYear="1896" box="[204,486,660,687]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sylvaticus">
<emphasis box="[204,486,660,687]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus sylvaticus</emphasis>
</taxonomicName>
species group is characterized by the long and slender penis with short outer arm, apex of penis simple without appendage, penis guide sometimes bifurcated apically, interocular distance greater than eye width in frontal view, abdominal postcoxal lines recurved toward the base of ventrite, and triangular coxites usually elongate.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="multiple">
<paragraph blockId="8.[151,1437,660,831]" box="[199,1106,804,831]" pageId="8" pageNumber="9">
12 species of
<taxonomicName baseAuthorityName="Lewis" baseAuthorityYear="1896" box="[352,462,804,830]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sylvaticus">
<emphasis box="[352,462,804,830]" italics="true" pageId="8" pageNumber="9">sylvaticus</emphasis>
</taxonomicName>
group are recorded from
<collectingCountry box="[750,820,804,830]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
, including 5 new species.
</paragraph>
<paragraph blockId="8.[151,1242,912,939]" box="[151,1242,912,939]" pageId="8" pageNumber="9">
<heading bold="true" box="[151,1242,912,939]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis bold="true" box="[151,1242,912,939]" pageId="8" pageNumber="9">
Key to the species of
<taxonomicName baseAuthorityName="Lewis" baseAuthorityYear="1896" box="[413,530,912,939]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sylvaticus">
<emphasis bold="true" box="[413,530,912,939]" italics="true" pageId="8" pageNumber="9">sylvaticus</emphasis>
</taxonomicName>
group within the genus
<taxonomicName authority="Vandenberg, 2004" authorityName="Vandenberg" authorityYear="2004" box="[832,1242,912,939]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[832,1010,912,938]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus</emphasis>
<bibRefCitation author="Vandenberg, N. J." box="[1016,1242,912,939]" pageId="8" pageNumber="9" pagination="483 - 484" refId="ref65879" refString="Vandenberg, N. J. (2004) Homonymy in the Coccinellidae (Coleoptera), or something fishy about Pseudoscymnus Chapin. Proceedings of the Entomological Society of Washington, 106 (2), 483 - 484." type="journal article" year="2004">Vandenberg, 2004</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="key">
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,981,1003]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,981,1003]" pageId="8" pageNumber="9">1. Elytra entirely black................................................................................... 2</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1011,1033]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1011,1033]" pageId="8" pageNumber="9">- Elytra black with yellowish brown apices.................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1040,1062]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1040,1062]" pageId="8" pageNumber="9">2. Penis guide widest at base in ventral view.................................................................. 3</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1069,1091]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1069,1091]" pageId="8" pageNumber="9">
- Penis guide approximately widest at basal 1/
<quantity box="[653,691,1069,1091]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="9" unit="in" value="3.0">3 in</quantity>
ventral view (
<figureCitation box="[829,899,1069,1091]" captionStart="FIGURE 4" captionStartId="9.[151,250,1833,1857]" captionTargetBox="[163,1425,750,1806]" captionTargetId="figure-225@9.[151,1436,741,1810]" captionTargetPageId="9" captionText="FIGURE 4. S. fusinus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–g male genitalia: e penis, f tegmen, lateral view, g tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–g: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328090" httpUri="https://zenodo.org/record/7328090/files/figure.png" pageId="8" pageNumber="9">
Fig.
<quantity box="[874,899,1070,1091]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="8" pageNumber="9" unit="g" value="4.0">4g</quantity>
</figureCitation>
)......
<emphasis bold="true" box="[980,1436,1069,1091]" pageId="8" pageNumber="9">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[980,1353,1069,1091]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fusinus" status="sp. nov.">
<emphasis bold="true" box="[980,1205,1069,1091]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus fusinus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1070,1091]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1099,1121]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1099,1121]" pageId="8" pageNumber="9">3. Parameres about 1/2 as long as penis guide................................................................. 4</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1437,1128,1150]" pageId="8" pageNumber="9">
<keyLead box="[151,1437,1128,1150]" pageId="8" pageNumber="9">
- Parameres about 3/4 as long as penis guide.................................
<taxonomicName authority="(Ohta, 1929)" baseAuthorityName="Ohta" baseAuthorityYear="1929" box="[1037,1437,1128,1150]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tainanensis">
<emphasis bold="true" box="[1037,1437,1128,1150]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[1037,1302,1128,1149]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus tainanensis</emphasis>
(
<bibRefCitation author="Ohta, Y." box="[1315,1429,1128,1150]" pageId="8" pageNumber="9" pagination="1 - 16" refId="ref65137" refString="Ohta, Y. (1929) Scymninen Japans. Insecta Matsumurana, 4 (1 - 2), 1 - 16." type="journal article" year="1929">Ohta, 1929</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1157,1179]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1157,1179]" pageId="8" pageNumber="9">
4. Penis guide apically pointed in ventral view (
<figureCitation box="[660,730,1157,1179]" captionStart="FIGURE 5" captionStartId="11.[151,250,1273,1297]" captionTargetBox="[160,1430,184,1248]" captionTargetId="figure-18@11.[151,1436,181,1249]" captionTargetPageId="11" captionText="FIGURE 5. S. acutus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328092" httpUri="https://zenodo.org/record/7328092/files/figure.png" pageId="8" pageNumber="9">Fig. 5h</figureCitation>
).....................
<emphasis bold="true" box="[988,1436,1157,1179]" pageId="8" pageNumber="9">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[988,1353,1157,1179]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="acutus" status="sp. nov.">
<emphasis bold="true" box="[988,1205,1157,1179]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus acutus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1158,1179]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1186,1209]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1186,1209]" pageId="8" pageNumber="9">
- Penis guide apically rounded in ventral view (
<figureCitation box="[667,737,1187,1209]" captionStart="FIGURE 6" captionStartId="12.[151,250,1472,1496]" captionTargetBox="[163,1426,383,1448]" captionTargetId="figure-93@12.[151,1436,381,1448]" captionTargetPageId="12" captionText="FIGURE 6. S. anmashanus (Yang, 1971): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328096" httpUri="https://zenodo.org/record/7328096/files/figure.png" pageId="8" pageNumber="9">Fig. 6h</figureCitation>
).......................
<taxonomicName authority="(Yang, 1971)" baseAuthorityName="Yang" baseAuthorityYear="1971" box="[1024,1436,1186,1208]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="anmashanus">
<emphasis bold="true" box="[1024,1436,1186,1208]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[1024,1302,1186,1208]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus anmashanus</emphasis>
(
<bibRefCitation author="Yang, C. T." box="[1315,1429,1186,1208]" pageId="8" pageNumber="9" pagination="85 - 96" refId="ref66136" refString="Yang, C. T. (1971) Notes on the species of genus Pseudoscymnus from Taiwan. Journal of Agriculture and Forestry, 20: 85 - 96." type="journal article" year="1971">Yang, 1971</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1216,1238]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1216,1238]" pageId="8" pageNumber="9">5. Pronotum entirely yellow to yellowish brown............................................................... 6</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1245,1267]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1245,1267]" pageId="8" pageNumber="9">- Pronotum partly or entirely black......................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1274,1297]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1274,1297]" pageId="8" pageNumber="9">
6. Pronotum yellow; scutellar shield black (
<figureCitation box="[620,688,1275,1297]" captionStart="FIGURE 7" captionStartId="13.[151,250,1796,1820]" captionTargetBox="[158,1430,710,1772]" captionTargetId="figure-196@13.[151,1436,705,1772]" captionTargetPageId="13" captionText="FIGURE 7. S. cultratus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f penis capsule, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328098" httpUri="https://zenodo.org/record/7328098/files/figure.png" pageId="8" pageNumber="9">Fig. 7a</figureCitation>
)......................
<emphasis bold="true" box="[966,1436,1274,1296]" pageId="8" pageNumber="9">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[966,1353,1274,1296]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cultratus" status="sp. nov.">
<emphasis bold="true" box="[966,1205,1274,1296]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus cultratus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1275,1296]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1437,1304,1326]" pageId="8" pageNumber="9">
<keyLead box="[151,1437,1304,1326]" pageId="8" pageNumber="9">
- Pronotum yellowish brown; scutellar shield dark brown (
<figureCitation box="[774,842,1304,1326]" captionStart="FIGURE 8" captionStartId="15.[151,250,1251,1275]" captionTargetBox="[161,1431,168,1233]" captionTargetId="figure-18@15.[151,1436,166,1235]" captionTargetPageId="15" captionText="FIGURE 8. S. sylvaticus (Lewis, 1896): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328102" httpUri="https://zenodo.org/record/7328102/files/figure.png" pageId="8" pageNumber="9">Fig. 8a</figureCitation>
)................
<taxonomicName authority="(Lewis, 1896)" baseAuthorityName="Lewis" baseAuthorityYear="1896" box="[1047,1437,1304,1326]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sylvaticus">
<emphasis bold="true" box="[1047,1437,1304,1326]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[1047,1294,1304,1326]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus sylvaticus</emphasis>
(
<bibRefCitation author="Lewis, G." box="[1307,1429,1304,1326]" pageId="8" pageNumber="9" pagination="22 - 41" refId="ref64859" refString="Lewis, G. (1896) On the Coccinellidae of Japan. Annals and Magazine of Natural History, Series 6, 17 (97), 22 - 41. https: // doi. org / 10.1080 / 00222939608680319" type="journal article" year="1896">Lewis, 1896</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1333,1355]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1333,1355]" pageId="8" pageNumber="9">7. Pronotum entirely black................................................................................ 8</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1437,1362,1385]" pageId="8" pageNumber="9">
<keyLead box="[151,1437,1362,1385]" pageId="8" pageNumber="9">
- Pronotum yellowish brown with a black spot at base (
<figureCitation box="[742,810,1363,1385]" captionStart="FIGURE 9" captionStartId="16.[151,250,1058,1082]" captionTargetBox="[151,1436,345,1035]" captionTargetId="figure-78@16.[151,1436,345,1035]" captionTargetPageId="16" captionText="FIGURE 9. S. ohtai (Yang &amp; Wu, 1972): a dorsal view, paratype, b ventral view, paratype, c lateral view, paratype, d frontal view, paratype, e abdomen, holotype, f penis, holotype, g tegmen, ventral view, holotype, h labels, paratype." figureDoi="http://doi.org/10.5281/zenodo.7328104" httpUri="https://zenodo.org/record/7328104/files/figure.png" pageId="8" pageNumber="9">Fig. 9a</figureCitation>
).................
<taxonomicName authority="(Yang &amp; Wu, 1972)" baseAuthorityName="Yang &amp; Wu" baseAuthorityYear="1972" box="[1032,1437,1362,1384]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ohtai">
<emphasis bold="true" box="[1032,1437,1362,1384]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[1032,1234,1362,1384]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus ohtai</emphasis>
(
<bibRefCitation author="Yang, C. T. &amp; Wu, R. H." box="[1246,1429,1362,1384]" pageId="8" pageNumber="9" pagination="115 - 128" refId="ref66088" refString="Yang, C. T. &amp; Wu, R. H. (1972) Notes on some Coccinellidae of Taiwan. Journal of Agriculture and Forestry, 21, 115 - 128. https: // doi. org / 10.2337 / diab. 21.2.128 b" type="journal article" year="1972">Yang &amp; Wu, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1392,1414]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1392,1414]" pageId="8" pageNumber="9">8. Parameres not dilated.................................................................................. 9</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1421,1443]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1421,1443]" pageId="8" pageNumber="9">
- Parameres distinctly dilated in lateral view (
<figureCitation box="[649,727,1421,1443]" captionStart="FIGURE 10" captionStartId="17.[151,250,1275,1299]" captionTargetBox="[162,1426,184,1251]" captionTargetId="figure-18@17.[151,1436,181,1252]" captionTargetPageId="17" captionText="FIGURE 10. S. subulatus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f tegmen, lateral view, g tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–g: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328108" httpUri="https://zenodo.org/record/7328108/files/figure.png" pageId="8" pageNumber="9">Fig. 10f</figureCitation>
)..................
<emphasis bold="true" box="[957,1436,1421,1443]" pageId="8" pageNumber="9">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[957,1353,1421,1443]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subulatus" status="sp. nov.">
<emphasis bold="true" box="[957,1205,1421,1443]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus subulatus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1422,1443]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1451,1473]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1451,1473]" pageId="8" pageNumber="9">9. Penis guide widest at base in ventral view................................................................. 10</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1480,1502]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1480,1502]" pageId="8" pageNumber="9">
- Penis guide approximately widest at 1/2 length in ventral view (
<figureCitation box="[840,921,1480,1502]" captionStart="FIGURE 11" captionStartId="18.[151,250,1508,1532]" captionTargetBox="[154,1425,418,1484]" captionTargetId="figure-103@18.[151,1436,417,1485]" captionTargetPageId="18" captionText="FIGURE 11. S. lishanicus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328110" httpUri="https://zenodo.org/record/7328110/files/figure.png" pageId="8" pageNumber="9">Fig. 11h</figureCitation>
)..
<emphasis bold="true" box="[952,1436,1480,1502]" pageId="8" pageNumber="9">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[952,1353,1480,1502]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lishanicus" status="sp. nov.">
<emphasis bold="true" box="[952,1204,1480,1502]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus lishanicus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1480,1501]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1509,1531]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1509,1531]" pageId="8" pageNumber="9">10. Parameres about 3/4 as long as penis guide................................................................ 11</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1538,1561]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1538,1561]" pageId="8" pageNumber="9">
- Parameres about 1/2 as long as penis guide......................................
<taxonomicName authority="Kitano, 2012" authorityName="Kitano" authorityYear="2012" box="[1099,1436,1538,1560]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="jenai">
<emphasis bold="true" box="[1099,1436,1538,1560]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[1099,1299,1539,1560]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus jenai</emphasis>
<bibRefCitation author="Kitano, T." box="[1306,1436,1538,1560]" pageId="8" pageNumber="9" pagination="221 - 229" refId="ref64661" refString="Kitano, T. (2012) Five new species of Scymnini (Coleoptera, Coccinellidae) from East Asia. Elytra, 2 (2), 221 - 229." type="journal article" year="2012">Kitano, 2012</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="8" pageNumber="9">
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1437,1568,1590]" pageId="8" pageNumber="9">
<keyLead box="[151,1437,1568,1590]" pageId="8" pageNumber="9">
11. Penis guide apically pointed in ventral view (Fig.
<quantity box="[705,742,1568,1589]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="8" pageNumber="9" unit="g" value="12.0">12g</quantity>
)..........................
<taxonomicName authority="(Araki, 1963)" baseAuthorityName="Araki" baseAuthorityYear="1963" box="[1063,1437,1568,1590]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nakanei">
<emphasis bold="true" box="[1063,1437,1568,1590]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[1063,1294,1568,1590]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus nakanei</emphasis>
(
<bibRefCitation author="Araki, M." box="[1307,1429,1568,1590]" pageId="8" pageNumber="9" pagination="251 - 255" refId="ref63508" refString="Araki, M. (1963) Descriptions of four new species of the genus Scymnus from Japan (Coleoptera: Coccinellidae). The Scientific Reports of Kyoto Prefectural University. Natural science and living science, 3 (5), 251 - 255." type="journal article" year="1963">Araki, 1963</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="8.[151,1437,981,1619]" box="[151,1436,1597,1619]" pageId="8" pageNumber="9">
<keyLead box="[151,1436,1597,1619]" pageId="8" pageNumber="9">
- Penis guide apically rounded in ventral view.................................
<taxonomicName authority="Kitano, 2012" authorityName="Kitano" authorityYear="2012" box="[1052,1436,1597,1619]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hosonaga">
<emphasis bold="true" box="[1052,1436,1597,1619]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[1052,1300,1597,1619]" italics="true" pageId="8" pageNumber="9">Sasajiscymnus hosonaga</emphasis>
<bibRefCitation author="Kitano, T." box="[1306,1436,1597,1619]" pageId="8" pageNumber="9" pagination="221 - 229" refId="ref64661" refString="Kitano, T. (2012) Five new species of Scymnini (Coleoptera, Coccinellidae) from East Asia. Elytra, 2 (2), 221 - 229." type="journal article" year="2012">Kitano, 2012</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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