treatments-xml/data/26/28/70/2628702E7FCCF34D222FA914F9A40801.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.794.28214" ID-GBIF-Dataset="875bbc72-e40b-42b3-95ef-ea83589cdc5e" ID-PMC="PMC6224371" ID-Pensoft-Pub="1313-2970-794-45" ID-PubMed="30416340" ID-ZBK="8B5352CFCFBE4A39AFBBE1430EC9E6E6" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-794-45" ModsDocOrigin="ZooKeys 794" ModsDocTitle="Taxonomy of Baetis Leach in Israel (Ephemeroptera, Baetidae)" checkinTime="1541169075417" checkinUser="pensoft" docAuthor="Yanai, Zohar, Gattolliat, Jean-Luc &amp; Dorchin, Netta" docDate="2018" docId="2628702E7FCCF34D222FA914F9A40801" docLanguage="en" docName="ZooKeys 794: 45-84" docOrigin="ZooKeys 794" docSource="http://dx.doi.org/10.3897/zookeys.794.28214" docTitle="Baetis golanensis Yanai &amp; Gattolliat, sp. n." docType="treatment" docUuid="08E663AD-EAF1-4DE4-B70C-CA52AA808552" docUuidSource="ZooBank" docVersion="5" lastPageNumber="45" masterDocId="D139FFC3FFFAFFBACD1F5768FFEF2B60" masterDocTitle="Taxonomy of Baetis Leach in Israel (Ephemeroptera, Baetidae)" masterLastPageNumber="84" masterPageNumber="45" pageNumber="45" updateTime="1668166366486" updateUser="ExternalLinkService">
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<mods:title>Taxonomy of Baetis Leach in Israel (Ephemeroptera, Baetidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yanai, Zohar</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dorchin, Netta</mods:namePart>
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<mods:date>2018</mods:date>
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<mods:number>794</mods:number>
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<mods:start>45</mods:start>
<mods:end>84</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.794.28214</mods:url>
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<treatment ID-GBIF-Taxon="149052375" LSID="urn:lsid:zoobank.org:act:08E663AD-EAF1-4DE4-B70C-CA52AA808552" httpUri="http://treatment.plazi.org/id/2628702E7FCCF34D222FA914F9A40801" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="http://zoobank.org/08E663AD-EAF1-4DE4-B70C-CA52AA808552" authority="Yanai &amp; Gattolliat" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis golanensis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="golanensis">Baetis golanensis Yanai &amp; Gattolliat</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. n.</taxonomicNameLabel>
Figures 4B, 7, 8
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="reference_group">
<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Baetis</taxonomicName>
A12: 
<bibRefCitation author="Samocha, M" journalOrPublisher="MSc thesis, Tel Aviv University, Tel Aviv, Israel" pageId="0" pageNumber="45" title="Ephemeroptera of Israel." year="1972">Samocha 1972</bibRefCitation>
: 44-45, pl XVIII, figs 7-9; pl XIX, figs 1-9.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis lutheri" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="lutheri">Baetis lutheri</taxonomicName>
: 
<bibRefCitation author="Koch, S" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="45" pagination="89 - 112" title="Mayflies of the Northern Levant (Insecta: Ephemeroptera)." url="https://doi.org/10.1080/09397140.1988.10637565" volume="2" year="1988">Koch 1988</bibRefCitation>
: 93.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="differential diagnosis">
<paragraph pageId="0" pageNumber="45">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="45">
Among the species with well-developed median caudal filaments in the 
<taxonomicName lsidName="B. lutheri" pageId="0" pageNumber="45" rank="species" species="lutheri">B. lutheri</taxonomicName>
species group, 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
is the only one with a combination of the following characters: right mandibular incisors of similar size, irregular blunt quadrangular spines on the distal margin of terga, and non-sclerotized protuberances next to the coxae. This is the only species in Israel with a developed median caudal filament that also has ventral protuberances near coxa bases and subapical setae on the claws.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">Length (mature nymphs). Female (n = 11): 5.1-6.3 mm; cerci 2.2-2.9 mm; median caudal filament 1.4-2.4 mm. Male (n = 9): 4.6-6.1 mm; cerci 2.2-3.0 mm; median caudal filament 1.7-2.4 mm.</paragraph>
<paragraph pageId="0" pageNumber="45">Colouration. General colour brown (Figure 4B). Head brown, antennae ecru, area around compound eyes and ocelli ecru. Turbinate eyes in male nymph amber. Thorax dark brown. Legs ivory to ecru, claws, apex of femora and tibiae brown. Abdominal terga uniformly brown, terga V, IX and X sometimes distally lighter. Abdominal sterna ivory to beige. Gills almost transparent, tracheation light brown. Cerci ecru to brown, without bands or pattern, median caudal filament sometimes lighter.</paragraph>
<paragraph pageId="0" pageNumber="45">Head. No carina between antennae; antennae with few fine setae, pedicel without distal lobe (Figure 7A). Labrum (Figure 7B): dorsal surface with scattered, fine, short setae and seta bases, with one median pair of long setae (sometimes with one additional pair of shorter setae); lateral margin with distal arc of 3-4 long, simple, stout setae and 3-8 lateral setae; ventral surface with five small, stout setae laterodistally; distal margin with row of 30-40 fine, long, feathered setae. Hypopharynx (Figure 7D): lingua tri-lobed; surface of lingua and superlingua densely covered with thin setae; lateral margin of superlingua serrated proximally. Right mandible (Figure 7C) with incisors composed of two sets of four and three denticles, all denticles nearly of same size; prostheca with small pointed denticles; space between prostheca and mola without setae or crenulation, mola apex with tuft of setae; base of mandible with many short, thin setae and seta bases. Left mandible (Figure 7E) with incisors composed of six denticles; prostheca with four broad denticles and comb-shaped structure; space between prostheca and mola without setae or crenulation; base of mola without spines; base of mandible with many setae and seta bases. Maxillae (Figure 7F) with four broad teeth; lacinia with one row of small setae and long, serrated setae; base of lacinia with row of five long, stout setae; one seta perpendicular to lacinia margin; palpus 2-segmented, segments I and II subequal; segment II with few minute, thin setae apically, apical nipple with single scale. Labium (Figure 7G): glossa thin, slightly shorter than paraglossa; glossa inner margin and apex with long, stout setae; paraglossa thick, not curved, with 4-5 setae medio-dorsally; paraglossa with apical stout, blunt seta and distal third with three rows of long, feathered setae; palpus 3-segmented; segment I same length as segments II and III combined; segment II with distolateral protuberance slightly developed and dorsal row of 4-6 long, pointed setae; segment III symmetrical, conical, with distinct nipple apically, covered with thick setae. Mentum and palpi covered with many short fine setae ventrally.</paragraph>
<caption pageId="0" pageNumber="45">
<paragraph pageId="0" pageNumber="45">
Figure 7. 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis golanensis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="golanensis">Baetis golanensis</taxonomicName>
sp. n., nymph. A antennal pedicel and first flagellomeres B labrum (left, ventral; right, dorsal) C right mandible D hypopharynx E left mandible F maxilla G labium.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">Thorax. Forelegs (Figure 8A): coxae and trochanters with scattered, minute, stout setae. Femora with dorsal row of 30-40 long, blunt setae, sometimes with few submarginal setae, approximately 1/3 of femur width, with dense group of 5-10 sharper setae proximally; ventral roughly arranged row of 10-20 minute, stout setae; dorsoapical setal patch composed of few thin setae; surface with scattered seta bases and few feathery, brush-like scales, more evident near femur margins. Tibiae with dorsal row of thin, short setae; ventral margin with two rows of sparse minute, stout setae and brush of thin setae apically; tibiopatellar suture present. Tarsi with 5-12 pointed setae ventrally; dorsal margin with row of short, thin setae; one pointed seta, shorter than most ventral setae, at ventral tarsus-claw meeting point. Tarsal claws (Figure 8B) hooked, with row of 8-10 acute teeth and two subapical thin setae (sometimes hardly visible). Mid- and hindlegs similar to forelegs. Round, unsclerotized protuberances, sometimes hardly seen, adjacent to coxa bases (Figure 9).</paragraph>
<paragraph pageId="0" pageNumber="45">Abdomen. Terga shagreened, with few thin setae and many seta bases, without scales or scale bases; distal margin of all terga with irregular row of blunt, quadrangular spines, with occasional fine seta (Figure 8C), spines absent or rare on terga I and II, more abundant and developed on following terga. Gill I slightly reduced, without visible tracheation (Figure 8D); gills II to VII oval, without marginal spines, main tracheation branched and pigmented (Figure 8E). Paraproct (Figure 8F) surface with few hair-like setae and seta bases; margin with 13-15 short, regular, triangular spines; postero-lateral extension with few seta bases, margin with 18-21 small, triangular spines, sometimes with few submarginal, spatulated scales.</paragraph>
<caption pageId="0" pageNumber="45">
<paragraph pageId="0" pageNumber="45">
Figure 8. 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis golanensis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="golanensis">Baetis golanensis</taxonomicName>
sp. n., nymph. A foreleg B tarsal claw C abdominal tergum V, surface and distal margin D gill I E gill IV F paraproct plate.
</paragraph>
</caption>
<caption pageId="0" pageNumber="45">
<paragraph pageId="0" pageNumber="45">
Figure 9. 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis pacis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pacis">Baetis pacis</taxonomicName>
sp. n., nymph, ventral, showing protuberances next to coxae (arrows), which are typical to nymphs of 
<taxonomicName lsidName="B. pacis" pageId="0" pageNumber="45" rank="species" species="pacis">B. pacis</taxonomicName>
and 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="affinities">
<paragraph pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="45">
The new species clearly belongs to the 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis lutheri" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="lutheri">Baetis lutheri</taxonomicName>
species group sensu 
<bibRefCitation author="Mueller-Liebenau, I" journalOrPublisher="Gewaesser und Abwaesser" pageId="0" pageNumber="45" pagination="1 - 214" title="Revision der europaeischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera)." volume="49" year="1969">
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
(1969)
</bibRefCitation>
based on the reduction of the median caudal filament, blunt spines on distal margin of terga, two subapical setae on the claws, and single scale on the apical nipple of the maxillary palp (
<bibRefCitation author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="45" title="The mayflies of Europe (Ephemeroptera)." year="2012">
Bauernfeind and 
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
2012
</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis lutheri" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="lutheri">Baetis lutheri</taxonomicName>
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
, 1967 appears to be the closest species to 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
, based on its morphology, yet differs from it by the incisors of the right mandible, which is variable in size, the arrangement and shape of the setae along the dorsal margin of femur, and the more regular, blunt teeth along the distal margin of the terga (
<bibRefCitation pageId="0" pageNumber="45">
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1967
</bibRefCitation>
, 
<bibRefCitation author="Mueller-Liebenau, I" journalOrPublisher="Gewaesser und Abwaesser" pageId="0" pageNumber="45" pagination="1 - 214" title="Revision der europaeischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera)." volume="49" year="1969">1969</bibRefCitation>
). In 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis vardarensis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="vardarensis">Baetis vardarensis</taxonomicName>
Ikonomov, 1962 the ventral protuberances next to the coxae are pointed and more sclerotized than in 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
(
<bibRefCitation pageId="0" pageNumber="45">
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1974
</bibRefCitation>
: fig. 4i; 
<bibRefCitation author="Eiseler, B" journalOrPublisher="Lauterbornia" pageId="0" pageNumber="45" pagination="1 - 112" title="Identification key to the mayfly larvae of the German Highlands and Lowlands." url="https://www.zobodat.at/pdf/Lauterbornia_2005_53_0001-0112.pdf" volume="53" year="2005">Eiseler 2005</bibRefCitation>
: figs 76-77), and 
<taxonomicName lsidName="B. vardarensis" pageId="0" pageNumber="45" rank="species" species="vardarensis">B. vardarensis</taxonomicName>
has several rows of a smaller number of teeth along the paraproct margin (
<bibRefCitation author="Ikonomov, P" journalOrPublisher="Faculte des Sciences Naturelles de l'universite de Skopja, Biologie" pageId="0" pageNumber="45" pagination="83 - 140" title="Baetidae (Ephemeroptera) macedoniens." volume="13" year="1962">Ikonomov 1962</bibRefCitation>
, 
<bibRefCitation author="Mueller-Liebenau, I" journalOrPublisher="Gewaesser und Abwaesser" pageId="0" pageNumber="45" pagination="1 - 214" title="Revision der europaeischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera)." volume="49" year="1969">
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1969
</bibRefCitation>
). 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis estrelensis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="estrelensis">Baetis estrelensis</taxonomicName>
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
, 1974, 
<taxonomicName lsidName="B. mirkae" pageId="0" pageNumber="45" rank="species" species="mirkae">B. mirkae</taxonomicName>
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
&amp; Godunko, 2008, 
<taxonomicName lsidName="B. nigrescens" pageId="0" pageNumber="45" rank="species" species="nigrescens">B. nigrescens</taxonomicName>
<normalizedToken originalValue="Navás">Navas</normalizedToken>
, 1932 and 
<taxonomicName lsidName="B. pacis" pageId="0" pageNumber="45" rank="species" species="pacis">B. pacis</taxonomicName>
sp. n. can easily be separated from 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
by their extremely reduced median caudal filament (Figure 4B vs. Figure 4E). Moreover, 
<taxonomicName lsidName="B. mirkae" pageId="0" pageNumber="45" rank="species" species="mirkae">B. mirkae</taxonomicName>
from Cyprus has many spatulas on body surface, and an additional row of long setae on the proximal part of the dorsal margin of femora, which distinguish it from 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
. It also differs from 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
and 
<taxonomicName lsidName="B. lutheri" pageId="0" pageNumber="45" rank="species" species="lutheri">B. lutheri</taxonomicName>
by lacking the brush scales on femora (
<bibRefCitation author="Soldan, T" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="45" pagination="79 - 104" title="Two new species of the genus Baetis Leach, 1815 (Ephemeroptera: Baetidae) from Cyprus." url="https://doi.org/10.3161/067.058.0104" volume="58" year="2008">
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
and Godunko 2008
</bibRefCitation>
). 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis meridionalis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="meridionalis">Baetis meridionalis</taxonomicName>
Ikonomov, 1954, which was assigned to the 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis lutheri" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="lutheri">Baetis lutheri</taxonomicName>
species group by 
<bibRefCitation author="Mueller-Liebenau, I" journalOrPublisher="Gewaesser und Abwaesser" pageId="0" pageNumber="45" pagination="1 - 214" title="Revision der europaeischen Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera)." volume="49" year="1969">
<normalizedToken originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
(1969)
</bibRefCitation>
, differs from 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
by the lack of scale at the apex of its maxillary palpus, short setae on margin of femora and shape of spines on the distal margin of terga. It is noteworthy that the description of 
<taxonomicName lsidName="B. meridionalis" pageId="0" pageNumber="45" rank="species" species="meridionalis">B. meridionalis</taxonomicName>
is superficial, it has no type material that could be examined, and its assignment to this species group is uncertain (
<bibRefCitation author="Soldan, T" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="45" pagination="79 - 104" title="Two new species of the genus Baetis Leach, 1815 (Ephemeroptera: Baetidae) from Cyprus." url="https://doi.org/10.3161/067.058.0104" volume="58" year="2008">
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
and Godunko 2008
</bibRefCitation>
; 
<bibRefCitation author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="45" title="The mayflies of Europe (Ephemeroptera)." year="2012">
Bauernfeind and 
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
2012
</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis lutheri" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="lutheri">Baetis lutheri</taxonomicName>
has a wide distribution in the West Palearctic (
<bibRefCitation author="Bauernfeind, E" journalOrPublisher="Apollo Books, Ollerup" pageId="0" pageNumber="45" title="The mayflies of Europe (Ephemeroptera)." year="2012">
Bauernfeind and 
<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
2012
</bibRefCitation>
) but recent molecular analysis has shown that it may constitute a complex of closely related species (
<bibRefCitation author="Tenchini, R" journalOrPublisher="The European Zoological Journal" pageId="0" pageNumber="45" pagination="254 - 267" title="DNA barcoding and faunistic criteria for a revised taxonomy of Italian Ephemeroptera." url="https://doi.org/10.1080/24750263.2018.1480732" volume="85" year="2018">Tenchini et al. 2018</bibRefCitation>
). 
<bibRefCitation author="Koch, S" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="45" pagination="89 - 112" title="Mayflies of the Northern Levant (Insecta: Ephemeroptera)." url="https://doi.org/10.1080/09397140.1988.10637565" volume="2" year="1988">Koch (1988)</bibRefCitation>
reported it from Lebanon and Syria, but these reports may refer to 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
. 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis golanensis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="golanensis">Baetis golanensis</taxonomicName>
also corresponds to the undescribed &quot; 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Baetis</taxonomicName>
A12&quot; mentioned by Samocha, also from Israel (1972: 44, pls 
<normalizedToken originalValue="XVIII–XIX">XVIII-XIX</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">The species name reflects its presently known distribution range, the Golan Heights.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution and ecology.</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis golanensis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="golanensis">Baetis golanensis</taxonomicName>
is a rare species in Israel. It was reported by Samocha (as &quot; 
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis" order="Ephemeroptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">Baetis</taxonomicName>
A12&quot;) from springs and streams in the western Golan and a few localities along the upper Jordan River. This restricted distribution is confirmed in the present study. Two distinct habitats are typical for 
<taxonomicName lsidName="B. golanensis" pageId="0" pageNumber="45" rank="species" species="golanensis">B. golanensis</taxonomicName>
in Israel: streams with large water discharge and rapid currents (upper Jordan River; Figure 2D), and small tributaries with shallow, quiet water running over basaltic pebble substrate (small streams and spring-fed brooks in the Golan Heights; Figure 2E). In all cases, the water is pristine, well oxygenized (&gt;85% dissolved oxygen) and contains aquatic vegetation. In all collecting events of the species, temperatures ranged between 19-25°C and salinity levels between 160-170 ppm. In Israel, mature nymphs were collected mainly in spring and early summer (
<normalizedToken originalValue="May–July">May-July</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="materials_examined">
<paragraph pageId="0" pageNumber="45">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="45">
Holotype. ISRAEL: 1N, Divsha Spring, 
<geoCoordinate direction="north" orientation="latitude" precision="5" value="33.0901">33.0901°N</geoCoordinate>
, 
<geoCoordinate direction="east" orientation="longitude" precision="5" value="35.6483">35.6483°E</geoCoordinate>
, ca. 150 m a.s.l., 11.v.2016, Z. Yanai &amp; N. Dorchin, SMNHTAU292000. Paratypes. ISRAEL: 14N (3N on slides), Divsha Spring, 10.vi.2014, Z. Yanai; 77N, Divsha Spring, 06.xi.2015, Z. Yanai &amp; S. Cohen; 74N, Divsha Spring, 11.v.2016, Z. Yanai &amp; N. Dorchin; 19N, Divsha Spring, 17.v.2016, Z. Yanai; 3N, Divsha Spring, 02.x.2016, Z. Yanai &amp; N. Truskanov; 16N (1N on slide), Tina Spring, 10.iii.2017, Z. Yanai &amp; J.-L. Gattolliat; 4N, Divsha Spring, 11.iii.2017, Z. Yanai &amp; J.-L. Gattolliat; 44N, Yehudiyya Stream, 11.iii.2017, Z. Yanai &amp; J.-L. Gattolliat. Other material. ISRAEL: 1N, Maymon Spring, 22.vi.2014, Z. Yanai; 6N (1N on slide), Gilbon Stream (downstream Gilbon Spring), 15.vii.2014, Z. Yanai; 4N, Divsha Spring, 11.v.2015, Y. Hershkovitz &amp; T. Eshcoly; 1N, Orevim Stream, 11.v.2015, Y. Hershkovitz; 14N, Samakh Stream, 13.v.2015, Y. Hershkovitz &amp; T. Eshcoly; 11N, Jordan River (Ateret Fortress), 29.x.2015, Z. Yanai &amp; Y. Brenner; 7N, Senir Stream (Bet Hillel), 05.xi.2015, Z. Yanai &amp; S. Cohen; 16N, Tina Spring, 06.xi.2015, Z. Yanai &amp; S. Cohen; 2N, Gilbon Stream (upstream Devora Waterfall), 11.v.2016, Z. Yanai &amp; N. Dorchin; 2N, Tina Spring, 16.v.2016, Z. Yanai &amp; A. Charvet; 162N, Jordan River (Ateret Fortress), 16.v.2016, Z. Yanai &amp; A. Charvet; 1N, Senir Stream (Beth Hillel), 17.v.2016, Z. Yanai &amp; A. Charvet; 3N, Jordan River (Neot Mordekhay), 01.vi.2016, Y. Hershkovitz; 8N (2N on slides), Jordan River (Ateret Fortress), 16.x.2016, Z. Yanai.
</paragraph>
</subSubSection>
</treatment>
</document>