treatments-xml/data/24/4B/88/244B882F23207622FF7121DEFAD7A38C.xml
2024-06-21 12:31:35 +02:00

200 lines
23 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="4EA1FCB830026ABCD900B15FD814AD71" ID-CLB-Dataset="39115" ID-DOI="10.11646/zootaxa.4109.2.3" ID-GBIF-Dataset="60a1e58f-6277-414a-a302-7fd2a6b2da1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="271419" ID-ZooBank="3DF4E74C-F422-48AD-AFEE-F3B0FEA8F443" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462515728922" checkinUser="plazi" docAuthor="Mens, Lotte P., Schütte, Kai, Stokvis, Frank R. &amp; Dijkstra, Klaas-Douwe B." docDate="2016" docId="244B882F23207622FF7121DEFAD7A38C" docLanguage="en" docName="zootaxa.4109.2.3.pdf.imf" docOrigin="Zootaxa 4109 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acisoma ascalaphoides Rambur 1842" docType="treatment" docVersion="9" lastPageNumber="167" masterDocId="D872F057232E762CFFE62149FFCFA027" masterDocTitle="Six, not two, species of Acisoma pintail dragonfly (Odonata: Libellulidae)" masterLastPageNumber="172" masterPageNumber="153" pageNumber="167" updateTime="1698662321709" updateUser="ExternalLinkService">
<mods:mods id="E67BCBD018626297C6B5DB9C9BAFB34E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="62D7B162C08734EBCCB4E1F85E9386F1">
<mods:title id="E589D07D1DBF39AE263571A14ED539EE">Six, not two, species of Acisoma pintail dragonfly (Odonata: Libellulidae)</mods:title>
</mods:titleInfo>
<mods:name id="CB7CC589118CFF44B60947A4E73566CB" type="personal">
<mods:role id="F9D456E5790830AEA32D4B51C6413AEA">
<mods:roleTerm id="9403A1F4664ECC527112545967871004">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="802B2EBD405F8738A5A546E58900FE29">Mens, Lotte P.</mods:namePart>
</mods:name>
<mods:name id="A27DB679DB4D050E86647625700D1FA8" type="personal">
<mods:role id="6382D3B4578DA13F95080C2292995878">
<mods:roleTerm id="E87B44E8BCC1E1EE3B8A8BC3624252D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6033E4ADD16651B78696605D189A0B5C">Schütte, Kai</mods:namePart>
</mods:name>
<mods:name id="E9FBFCDFA624696DE4ACD4426CEBB69F" type="personal">
<mods:role id="E7BB2467A8BE67439F81B6140CFB0330">
<mods:roleTerm id="FF346979D7D16D1D2C666ADF8BCABA96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9EC247BDDEE03E87FC1293C3A4F8C78">Stokvis, Frank R.</mods:namePart>
</mods:name>
<mods:name id="F66471E56FB32E4146DB9FFD07955F36" type="personal">
<mods:role id="088CFBC945F288C84A9D592816D1D98B">
<mods:roleTerm id="FA8E6DE3720E92E444F3C03F334DF906">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FA71A94698171ED921EAD116AB3F7172">Dijkstra, Klaas-Douwe B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E4298F22CF561473FCD51AC663E64179">text</mods:typeOfResource>
<mods:relatedItem id="23090428874125A728F7F0632A8A6113" type="host">
<mods:titleInfo id="58C4A084D41ED0E73C8D579A5F3CAE9D">
<mods:title id="23595A82F3C011186432835F175D479C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5EA45C45B01C67603BEF6F31BF684A89">
<mods:date id="69731F3582BD7F330A847DC914729734">2016</mods:date>
<mods:detail id="494DDA02013A895982A7D417075F838F" type="volume">
<mods:number id="035A4856283D8EE91D5887EDBCC4A2F1">4109</mods:number>
</mods:detail>
<mods:detail id="2407D5AF8FC5694EB77E9FC265ECAE31" type="issue">
<mods:number id="3A623EE770AFC0EA5C624F64208A530F">2</mods:number>
</mods:detail>
<mods:extent id="FF4FD065A8A76FF00DD6D38663F731E0" unit="page">
<mods:start id="FBFA81029717EB74EA0BCC7DC412986D">153</mods:start>
<mods:end id="C6434C1BC2B9E0DC465AFEA9FB269976">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="54F3575EB4CD0A2C35A1ED453D64F84E">journal article</mods:classification>
<mods:identifier id="8100A3415C2E57D377DDDAA6D1D1007B" type="CLB-Dataset">39115</mods:identifier>
<mods:identifier id="84353F3F97C4F2FA002E5734CDC3C95B" type="DOI">10.11646/zootaxa.4109.2.3</mods:identifier>
<mods:identifier id="28F23D3E3D0447781570AE8799469629" type="GBIF-Dataset">60a1e58f-6277-414a-a302-7fd2a6b2da1a</mods:identifier>
<mods:identifier id="69EDE9C40C7838E8910B308393FCFAF0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="741920D48B99E290E6B18A6ACB652F3D" type="Zenodo-Dep">271419</mods:identifier>
<mods:identifier id="5829F19FA85EC6056F25221F348C7FA6" type="ZooBank">3DF4E74C-F422-48AD-AFEE-F3B0FEA8F443</mods:identifier>
</mods:mods>
<treatment id="244B882F23207622FF7121DEFAD7A38C" ID-DOI="http://doi.org/10.5281/zenodo.5673009" ID-GBIF-Taxon="120674213" ID-Zenodo-Dep="5673009" LSID="urn:lsid:plazi:treatment:244B882F23207622FF7121DEFAD7A38C" httpUri="http://treatment.plazi.org/id/244B882F23207622FF7121DEFAD7A38C" lastPageNumber="167" pageId="14" pageNumber="167">
<subSubSection id="E4F86AB223207622FF7121DEFE22A0F4" pageId="14" pageNumber="167" type="nomenclature">
<paragraph id="AC5D393923207622FF7121DEFCFDA095" blockId="14.[151,818,151,211]" box="[151,818,151,178]" pageId="14" pageNumber="167">
<heading id="F7158E5523207622FF7121DEFCFDA095" bold="true" box="[151,818,151,178]" fontSize="11" level="1" pageId="14" pageNumber="167" reason="1">
<emphasis id="9E96E52B23207622FF7121DEFCFDA095" bold="true" box="[151,818,151,178]" pageId="14" pageNumber="167">
<taxonomicName id="6BE242BA23207622FF7121DEFD93A095" ID-CoL="9WT33" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[151,604,151,178]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="167" phylum="Arthropoda" rank="species" species="ascalaphoides">
<emphasis id="9E96E52B23207622FF7121DEFE66A096" bold="true" box="[151,425,151,177]" italics="true" pageId="14" pageNumber="167">Acisoma ascalaphoides</emphasis>
<bibRefCitation id="C87344C823207622FE5621D1FD93A095" author="Rambur" box="[432,604,152,178]" pageId="14" pageNumber="167" refString="Rambur, J. P. (1842) Histoire naturelle des insectes: Nevropteres. Librairie Encyclopedique de Roret, Paris." type="book" year="1842">Rambur, 1842</bibRefCitation>
</taxonomicName>
—Littoral Pintail
</emphasis>
</heading>
</paragraph>
<paragraph id="AC5D393923207622FF7121F3FE22A0F4" blockId="14.[151,818,151,211]" box="[151,493,186,211]" pageId="14" pageNumber="167">
(
<figureCitation id="34D925BC23207622FF7921F3FF25A0F4" box="[159,234,186,211]" captionStart="FIGURE 4" captionStartId="7.[151,250,705,727]" captionTargetBox="[223,1345,203,654]" captionTargetId="figure@7.[216,1367,190,668]" captionTargetPageId="7" captionText="FIGURE 4. Head of Acisoma species in rostral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides (typical), D: A. panorpoides (pale), E: A. attenboroughi, F: A. ascalaphoides, G: A. trifidum." httpUri="https://zenodo.org/record/271423/files/figure.png" pageId="14" pageNumber="167">Figs. 4</figureCitation>
F, 5H, 6G, 7E, 8E, 10E)
</paragraph>
</subSubSection>
<subSubSection id="E4F86AB223207622FF7121B6FB03A1CF" pageId="14" pageNumber="167" type="materials_examined">
<paragraph id="AC5D393923207622FF7121B6FB03A1CF" blockId="14.[151,1437,255,939]" pageId="14" pageNumber="167">
<emphasis id="9E96E52B23207622FF7121B6FECBA13F" bold="true" box="[151,260,255,280]" pageId="14" pageNumber="167">Material.</emphasis>
<collectingCountry id="D4F579A923207622FEE821B6FE58A13F" box="[270,407,255,280]" name="Madagascar" pageId="14" pageNumber="167">Madagascar</collectingCountry>
:
<specimenCount id="BAE4F2B023207622FE432049FE3AA13F" box="[421,501,255,280]" pageId="14" pageNumber="167" type="male">3 male</specimenCount>
, Maroantsetra, Voloina,
<date id="D85C1FF923207622FCEA21B6FC5FA13F" box="[780,912,255,280]" pageId="14" pageNumber="167" value="1971-12-30">30-xii-1971</date>
, P. &amp; J. Minet (
<collectionCode id="CAF3A1FC23207622FBAA2049FB68A13F" LSID="urn:lsid:biocol.org:col:34988" box="[1100,1191,256,280]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="167">MNHN</collectionCode>
);
<specimenCount id="BAE4F2B023207622FB5B2049FAD6A13F" box="[1213,1305,255,280]" pageId="14" pageNumber="167" type="male">2 males</specimenCount>
, Tolagnaro, Mandena,
<date id="D85C1FF923207622FEEB206BFE4EA11C" box="[269,385,290,315]" pageId="14" pageNumber="167" value="2004-02-12">12-ii-2004</date>
/
<date id="D85C1FF923207622FE6C206BFDC9A11C" box="[394,518,290,315]" pageId="14" pageNumber="167" value="2004-03-19">19-iii-2004</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23207622FD7D206AFD3BA11C" LSID="urn:lsid:biocol.org:col:34992" box="[667,756,291,315]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="167">RMNH</collectionCode>
);
<specimenCount id="BAE4F2B023207622FCEF206AFCACA11C" box="[777,867,290,315]" pageId="14" pageNumber="167" type="male">4 males</specimenCount>
, Tolagnaro, Sainte Luce, Akaifira,
<date id="D85C1FF923207622FB16206BFABCA11C" box="[1264,1395,290,315]" pageId="14" pageNumber="167" value="2006-12-02">02-xii-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23207622FF1D200FFE9DA179" box="[251,338,326,350]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="167">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023207622FE8E200FFE78A179" box="[360,439,325,350]" pageId="14" pageNumber="167" type="male">1 male</specimenCount>
, Tolagnaro, Sainte Luce, S8, forest, marecage,
<date id="D85C1FF923207622FC36200CFB8AA179" box="[976,1093,325,350]" pageId="14" pageNumber="167" value="2006-10-18">18-x-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23207622FB3A200FFAFCA179" box="[1244,1331,326,350]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="167">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023207622FAAF200FFA57A179" box="[1353,1432,325,350]" pageId="14" pageNumber="167" type="male">1 male</specimenCount>
, Tolagnaro, Sainte Luce, S9, marecage west,
<date id="D85C1FF923207622FD6A202EFCC7A1A7" box="[652,776,359,384]" pageId="14" pageNumber="167" value="2004-04-01">01-iv-2004</date>
, P. Razafindraibe (
<collectionCode id="CAF3A1FC23207622FC3B2021FBFBA1A7" box="[989,1076,360,384]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="167">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023207622FBAF2021FB6CA1A7" box="[1097,1187,359,384]" pageId="14" pageNumber="167" type="male">2 males</specimenCount>
, Tolagnaro, Mandena, bridge from M15 to M16,
<date id="D85C1FF923207622FE5D20C3FDFFA184" box="[443,560,394,419]" pageId="14" pageNumber="167" value="2006-10-13">13-x-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23207622FD5920C2FCD9A184" box="[703,790,395,419]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="167">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023207622FCCC20C2FCB8A184" box="[810,887,394,419]" pageId="14" pageNumber="167" type="male">1 male</specimenCount>
, Tolagnaro, Mandena (Citronelle),
<date id="D85C1FF923207622FAE720C3FAB9A184" box="[1281,1398,394,419]" pageId="14" pageNumber="167" value="2006-10-13">13-x-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23207622FF1A20E7FE9CA1E1" box="[252,339,430,454]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="167">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023207622FE8D20E4FE09A1E1" box="[363,454,429,454]" pageId="14" pageNumber="167" type="male">5 males</specimenCount>
, Tolagnaro, Mandena,
<date id="D85C1FF923207622FD3620E4FC86A1E1" box="[720,841,429,454]" pageId="14" pageNumber="167" value="2004-03-17">17-iii-2004</date>
/
<date id="D85C1FF923207622FCB420E4FC03A1E1" box="[850,972,429,454]" pageId="14" pageNumber="167" value="2004-03-18">18-iii-2004</date>
/
<date id="D85C1FF923207622FC0620E4FB95A1E1" box="[992,1114,429,454]" pageId="14" pageNumber="167" value="2004-03-19">19-iii-2004</date>
/
<date id="D85C1FF923207622FB8820E4FB27A1E1" box="[1134,1256,429,454]" pageId="14" pageNumber="167" value="2004-03-22">22-iii-2004</date>
/
<date id="D85C1FF923207622FB1A20E4FAB5A1E1" box="[1276,1402,429,454]" pageId="14" pageNumber="167" value="2004-03-27">27-iii-2004</date>
, P. Razafindraibe (
<collectionCode id="CAF3A1FC23207622FED92099FE59A1CF" box="[319,406,464,488]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="167">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023207622FE4F2099FE3AA1CF" box="[425,501,463,488]" pageId="14" pageNumber="167" type="male">1 male</specimenCount>
, Tolagnaro, Mandena, marecage,
<date id="D85C1FF923207622FC802086FC13A1CF" box="[870,988,463,488]" pageId="14" pageNumber="167" value="2004-02-12">12-ii-2004</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23207622FB8F2099FB0FA1CF" box="[1129,1216,464,488]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="167">ZMUH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="E4F86AB223207622FF2120BBFAD7A38C" pageId="14" pageNumber="167" type="description">
<paragraph id="AC5D393923207622FF2120BBFBB3A2D9" blockId="14.[151,1437,255,939]" pageId="14" pageNumber="167">
<emphasis id="9E96E52B23207622FF2120BBFEB3A22C" bold="true" box="[199,380,498,523]" pageId="14" pageNumber="167">Male diagnosis.</emphasis>
Distinctive species confined to the east coast of
<collectingCountry id="D4F579A923207622FC6C20BBFBC0A22C" box="[906,1039,498,523]" name="Madagascar" pageId="14" pageNumber="167">Madagascar</collectingCountry>
that appears somewhat intermediate between
<taxonomicName id="6BE242BA23207622FF1F235FFEA0A20A" box="[249,367,533,557]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="167" phylum="Arthropoda" rank="species" species="trifidum">
<emphasis id="9E96E52B23207622FF1F235FFEA0A20A" box="[249,367,533,557]" italics="true" pageId="14" pageNumber="167">A. trifidum</emphasis>
</taxonomicName>
and the species formerly treated under
<taxonomicName id="6BE242BA23207622FCFA235FFC0CA20A" box="[796,963,533,557]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="167" phylum="Arthropoda" rank="species" species="panorpoides">
<emphasis id="9E96E52B23207622FCFA235FFC0CA20A" box="[796,963,533,557]" italics="true" pageId="14" pageNumber="167">A. panorpoides</emphasis>
</taxonomicName>
. Its unique characters are the (1) uniformly brown antefrons, clypeus, labrum and labium (
<figureCitation id="34D925BC23207622FD4F237EFD22A277" box="[681,749,567,592]" captionStart="FIGURE 4" captionStartId="7.[151,250,705,727]" captionTargetBox="[223,1345,203,654]" captionTargetId="figure@7.[216,1367,190,668]" captionTargetPageId="7" captionText="FIGURE 4. Head of Acisoma species in rostral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides (typical), D: A. panorpoides (pale), E: A. attenboroughi, F: A. ascalaphoides, G: A. trifidum." httpUri="https://zenodo.org/record/271423/files/figure.png" pageId="14" pageNumber="167">Fig. 4</figureCitation>
F, 5H); (2) largely brown thorax with faint and limited paler markings (
<figureCitation id="34D925BC23207622FEEA2313FE82A254" box="[268,333,602,627]" captionStart="FIGURE 6" captionStartId="7.[151,250,1802,1824]" captionTargetBox="[196,1382,1322,1745]" captionTargetId="figure@7.[193,1384,1321,1753]" captionTargetPageId="7" captionText="FIGURE 6. Thorax of Acisoma species in lateral view. A: A. variegatum, B: A. inflatum (typical), C: A. inflatum (dark), D: A. panorpoides, E: A. attenboroughi (typical), F: A. attenboroughi (dark), G: A. ascalaphoides, H: A. trifidum." httpUri="https://zenodo.org/record/271425/files/figure.png" pageId="14" pageNumber="167">Fig. 6</figureCitation>
G); (3) mostly 2 cells in Fw triangle, although 37% of examined wings with 1 cell; (4) abdomen that narrows most on S56 while S710 are slender (
<figureCitation id="34D925BC23207622FD522334FD39A2B1" box="[692,758,637,662]" captionStart="FIGURE 7. S 2 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="14" pageNumber="167">Fig. 7</figureCitation>
E); (5) ventrally entirely dark abdomen with extensive dorsal white markings only on S36, which are solid and clean-cut but not confluent across dorsal carina (
<figureCitation id="34D925BC23207622FAEB23D6FA9EA29F" box="[1293,1361,671,696]" captionStart="FIGURE 7. S 2 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="14" pageNumber="167">Fig. 7</figureCitation>
E); (6) narrow lobe of hamule with notched tip (
<figureCitation id="34D925BC23207622FDBD238BFD54A2FC" box="[603,667,706,731]" captionStart="FIGURE 8" captionStartId="9.[151,250,911,933]" captionTargetBox="[157,1430,195,882]" captionTargetId="figure@9.[151,1435,193,890]" captionTargetPageId="9" captionText="FIGURE 8. Secondary genitalia of Acisoma species in ventral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides, D: A. attenboroughi, E: A. ascalaphoides, F: A. trifidum. Structure of hamule indicated." httpUri="https://zenodo.org/record/271427/files/figure.png" pageId="14" pageNumber="167">Fig. 8</figureCitation>
E); and (7) black dorsum of cerci, not largely white (
<figureCitation id="34D925BC23207622FB04238BFAEBA2FC" box="[1250,1316,706,731]" captionStart="FIGURE 7. S 2 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="14" pageNumber="167">Fig. 7</figureCitation>
E). Similar only to
<taxonomicName id="6BE242BA23207622FF0C23AFFEAFA2DA" box="[234,352,741,765]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="167" phylum="Arthropoda" rank="species" species="trifidum">
<emphasis id="9E96E52B23207622FF0C23AFFEAFA2DA" box="[234,352,741,765]" italics="true" pageId="14" pageNumber="167">A. trifidum</emphasis>
</taxonomicName>
are the (8) more numerous Fw Ax, usually 8½ but occasionally 8 or 9½.
</paragraph>
<paragraph id="AC5D393923207622FF21224EFAD7A38C" blockId="14.[151,1437,255,939]" pageId="14" pageNumber="167">
<emphasis id="9E96E52B23207622FF21224EFE7BA307" bold="true" box="[199,436,775,800]" pageId="14" pageNumber="167">Range and ecology.</emphasis>
Known only from littoral forest fragments at the southern (Tolagnaro, formerly Fort Dauphin) and northern (Voloina) ends of Madagascars east coast (
<figureCitation id="34D925BC23207622FC912263FC76A364" box="[887,953,810,835]" captionStart="FIGURE 9" captionStartId="11.[151,250,1877,1899]" captionTargetBox="[418,1156,196,1812]" captionTargetId="figure@11.[400,1172,193,1823]" captionTargetPageId="11" captionText="FIGURE 9. Distribution of Afrotropical Acisoma species based on examined specimens and verified records in Odonata Database of Africa (see supplementary materials): A: A. inflatum (circles), A. attenboroughi (triangles), unverified records of A. inflatum or A. variegatum (grey dots); B: A. variegatum (circles), A. ascalaphoides (squares), A. trifidum (crosses), unverified records of A. inflatum or A. variegatum (grey dots). All records are provided in Suppl. Tab. 1 available from https: // science. naturalis. nl / dijkstra." httpUri="https://zenodo.org/record/271428/files/figure.png" pageId="14" pageNumber="167">Fig. 9</figureCitation>
B). Only about 10% of the original cover of these forests remains, all in small patches of which only 13% are protected (
<bibRefCitation id="C87344C823207622FC042204FB1FA341" author="Consiglio" box="[994,1232,845,870]" pageId="14" pageNumber="167" refString="Consiglio, T., Schatz, G. E., McPherson, G., Lowry II, P. P., Rabenantoandro, J., Rogers, Z. S., Rabevohitra, R. &amp; Rabehevitra D. (2006) Deforestation and plant diversity of Madagascar's littoral forests. Conservation Biology, 20, 1799 - 1803. http: // dx. doi. org / 10.1111 / j. 1523 - 1739.2006.00562. x" type="journal article" year="2006">
Consiglio
<emphasis id="9E96E52B23207622FBB02207FB41A342" box="[1110,1166,845,869]" italics="true" pageId="14" pageNumber="167">et al.</emphasis>
2006
</bibRefCitation>
). The species may occur elsewhere along the coast, but as it seems restricted to this habitat, could well be under threat (
<bibRefCitation id="C87344C823207622FACE2226FEBDA38C" author="Schutte" pageId="14" pageNumber="167" refString="Schutte, K. &amp; Razafindraibe, P. (2007) Checklist of dragonflies of the littoral forests near Tolagnaro (Fort Dauphin). In: Ganzhorn J. U., Goodman, S. M., Vincelette, M. (Eds.), Biodiversity, ecology and conservation of littoral ecosystems in southeastern Madagascar, Tolagnaro (Fort Dauphin). Smithsonian Institution, Monitoring and Assessment of Biodiversity Series 11, Washington, pp. 163 - 165." type="book chapter" year="2007">Schütte &amp; Razafindraibe 2007</bibRefCitation>
). The larvae are possibly adapted to more acidic water than
<taxonomicName id="6BE242BA23207622FBE722DAFB76A38C" box="[1025,1209,915,939]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="167" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="9E96E52B23207622FBE722DAFB76A38C" box="[1025,1209,915,939]" italics="true" pageId="14" pageNumber="167">A. attenboroughi</emphasis>
</taxonomicName>
<emphasis id="9E96E52B23207622FB2622DBFAD7A38C" bold="true" box="[1216,1304,914,939]" pageId="14" pageNumber="167">
<taxonomicNameLabel id="85A5585023207622FB2622DBFAD7A38C" box="[1216,1304,914,939]" pageId="14" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>