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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.803.28903" ID-GBIF-Dataset="6c6c0b59-636f-4502-8b48-642ba231eb16" ID-PMC="PMC6390078" ID-Pensoft-Pub="1313-2970-803-7" ID-PubMed="30814899" ID-ZBK="936CFD88F297440EA9BE4C258AE9BD09" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-803-7" ModsDocOrigin="ZooKeys 803" ModsDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1545315537716" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo &amp; Balke, Michael" docDate="2018" docId="0B5149F72140AA3F08E731D92770F49B" docLanguage="en" docName="ZooKeys 803: 7-70" docOrigin="ZooKeys 803" docSource="http://dx.doi.org/10.3897/zookeys.803.28903" docTitle="Exocelina yoginofi Shaverdo &amp; Balke, sp. n." docType="treatment" docUuid="1F4C342B-17BB-4331-83CF-5AC202F7A881" docUuidSource="ZooBank" docVersion="7" lastPageNumber="43" masterDocId="5133FFF6125A1065FF83FF96FFF3FFD4" masterDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="70" masterPageNumber="7" pageNumber="40" updateTime="1668166558239" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sagata, Katayo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>803</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>7</mods:start>
<mods:end>70</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.803.28903</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.803.28903</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-803-7</mods:identifier>
<mods:identifier type="ZBK">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
<mods:identifier type="ZooBank">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154125613" LSID="urn:lsid:zoobank.org:act:1F4C342B-17BB-4331-83CF-5AC202F7A881" httpUri="http://treatment.plazi.org/id/0B5149F72140AA3F08E731D92770F49B" lastPageId="36" lastPageNumber="43" pageId="33" pageNumber="40">
<subSubSection pageId="33" pageNumber="40" type="nomenclature">
<paragraph pageId="33" pageNumber="40">
24.
<taxonomicName LSID="http://zoobank.org/1F4C342B-17BB-4331-83CF-5AC202F7A881" authority="Shaverdo &amp; Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina yoginofi" order="Coleoptera" pageId="33" pageNumber="40" phylum="Arthropoda" rank="species" species="yoginofi">Exocelina yoginofi Shaverdo &amp; Balke</taxonomicName>
<taxonomicNameLabel pageId="33" pageNumber="40">sp. n.</taxonomicNameLabel>
Figs 22, 46
</paragraph>
</subSubSection>
<subSubSection lastPageId="35" lastPageNumber="42" pageId="34" pageNumber="41" type="reference_group">
<paragraph pageId="34" pageNumber="41">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="34" pageNumber="41" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="34" pageNumber="41" start="start">Exocelina</pageBreakToken>
</taxonomicName>
undescribed sp. MB1302:
<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/ncomms5001" year="2014">Toussaint et al. 2014</bibRefCitation>
: supplementary figs 1-4, table 2;
<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
: supplementary figs S1-S2, table S3.
</paragraph>
<paragraph lastPageId="35" lastPageNumber="42" pageId="34" pageNumber="41">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina yoginofi" order="Coleoptera" pageId="34" pageNumber="41" phylum="Arthropoda" rank="species" species="yoginofi">Exocelina yoginofi</taxonomicName>
<pageBreakToken pageId="35" pageNumber="42" start="start">_</pageBreakToken>
New_Guinea_MB1302:
<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
: supplementary information S5-S6.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="42" type="type locality">
<paragraph pageId="35" pageNumber="42">Type locality.</paragraph>
<paragraph pageId="35" pageNumber="42">
Papua New Guinea: Eastern Highlands Province, Kainantu, Yoginofi,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-6.363333">06°21.80'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="145.75766">145°45.46'E</geoCoordinate>
, 1,900 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="42" type="type material">
<paragraph pageId="35" pageNumber="42">Type material.</paragraph>
<paragraph pageId="35" pageNumber="42">Holotype: male &quot;Papua New Guinea: Eastern Highlands, Kainantu, Yoginofi, 1900m, 9.v.1994, 06.21.799S 145.45.463E, Balke &amp; Sagata (PNG 55)&quot; (ZSM). Paratypes: 6 males, 8 females with the same label as the holotype, one male with an additional green label &quot;M.Balke 1302&quot; (NHMW, ZSM). 1 male, 1 female &quot;Papua New Guinea: Eastern Highlands, 37 km S Goroka, Hogave vill., Mt. Michael, 2179-2800m, 9.-15.vii.2009, 06.22.479S 145.15.256E, Sagata (PNG230)&quot; (ZSM).</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="42" type="description">
<paragraph pageId="35" pageNumber="42">Description.</paragraph>
<paragraph pageId="35" pageNumber="42">Body size and form: Beetle medium-sized: TL-H 3.45-4.0 mm, TL 3.85-4.45 mm, MW 1.85-2.15 mm (holotype: TL-H 3.9 mm, TL 4.3 mm, MW 2.0 mm), with oblong-oval habitus.</paragraph>
<paragraph pageId="35" pageNumber="42">Coloration: Dark brown to piceous, with paler pronotum. Head reddish to dark brown, paler anteriorly and posteriorly. Pronotum brown to dark brown, with reddish brown lateral sides and sometimes anteriorly and narrowly posteriorly, darker on disc, sometimes to piceous. Elytra brown to piceous, sometimes with narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish brown (Fig. 22).</paragraph>
<paragraph pageId="35" pageNumber="42">Surface sculpture: Submatt dorsally. Head with rather dense punctation (spaces between punctures 1-2 times size of punctures), finer and sparser anteriorly; diameter of punctures smaller than or equal to diameter of cells of microreticulation. Pronotum and elytra with sparser and finer punctation than on head. Head, pronotum and elytra with rather strongly impressed microreticulation. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine, sparse but distinct punctation.</paragraph>
<paragraph pageId="35" pageNumber="42">Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 rounded.</paragraph>
<paragraph pageId="35" pageNumber="42">
Male: Antennomere simple (Fig. 22). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with narrow anterior band of ca. 40 and posterior row of 14 relatively long setae (Fig. 46D). Median lobe in lateral view slightly curved, with apex dully pointed, slightly bent downwards; in ventral view, broadened subdistally, with broad, rounded apex. Paramere very slightly concave on dorsal side and with dense setae on subdistal part; proximal setae inconspicuous (Fig. 46
<normalizedToken originalValue="AC">A-C</normalizedToken>
). Abdominal ventrite 6 with 6-8 lateral striae on each side.
</paragraph>
<paragraph pageId="35" pageNumber="42">Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="42" type="affinities">
<paragraph pageId="35" pageNumber="42">Affinities.</paragraph>
<paragraph pageId="35" pageNumber="42">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina yoginofi" order="Coleoptera" pageId="35" pageNumber="42" phylum="Arthropoda" rank="species" species="yoginofi">Exocelina yoginofi</taxonomicName>
sp. n. is similar to
<taxonomicName lsidName="E. simbaiensis" pageId="35" pageNumber="42" rank="species" species="simbaiensis">E. simbaiensis</taxonomicName>
sp. n. in body form, size, and coloration, but differs from it in distinctly finer punctation and microreticulation and in having the apex of the median lobe not thickened. Also, see under
<taxonomicName lsidName="E. okapa" pageId="35" pageNumber="42" rank="species" species="okapa">E. okapa</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="43" type="distribution">
<paragraph pageId="36" pageNumber="43">
<pageBreakToken pageId="36" pageNumber="43" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="36" pageNumber="43">Papua New Guinea: Eastern Highlands Province (Fig. 50).</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="43" type="etymology">
<paragraph pageId="36" pageNumber="43">Etymology.</paragraph>
<paragraph pageId="36" pageNumber="43">The species is named after Yoginofi Village. The name is a noun in the nominative singular standing in apposition.</paragraph>
</subSubSection>
</treatment>
</document>