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<document id="29861258EDDC9B69EAA809D7811C1186" ID-CLB-Dataset="33559" ID-DOI="http://dx.doi.org/10.3897/zookeys.444.7537" ID-GBIF-Dataset="e53fb8f2-2a76-4856-b154-1faef6ee81fd" ID-PMC="PMC4205731" ID-Pensoft-Pub="1313-2970-444-1" ID-PubMed="25349499" ID-ZooBank="FB83BDD3958A456DBFEA9C6C28D3E4D5" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-444-1" ModsDocOrigin="ZooKeys 444" ModsDocTitle="Shallow-water zoantharians (Cnidaria, Hexacorallia) from the Central Indo-Pacific" checkinTime="1451245266514" checkinUser="pensoft" docAuthor="Reimer, James D., Poliseno, Angelo &amp; Hoeksema, Bert W." docDate="2014" docId="0889229B068BCE3E8604ED4865987E75" docLanguage="en" docName="ZooKeys 444: 1-57" docOrigin="ZooKeys 444" docSource="http://dx.doi.org/10.3897/zookeys.444.7537" docTitle="Acrozoanthus australiae Saville-Kent 1893" docType="treatment" docVersion="5" lastPageNumber="6" masterDocId="FFCCA07FFFC4625F1C6AFFB4EC3BFFB7" masterDocTitle="Shallow-water zoantharians (Cnidaria, Hexacorallia) from the Central Indo-Pacific" masterLastPageNumber="57" masterPageNumber="1" pageNumber="6" updateTime="1701385215740" updateUser="ExternalLinkService">
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<mods:title id="3168A058F288499FD1755022C0480D05">Shallow-water zoantharians (Cnidaria, Hexacorallia) from the Central Indo-Pacific</mods:title>
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<mods:namePart id="CD6460370F13F3A3A7597AC9C4230A23">Reimer, James D.</mods:namePart>
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<mods:namePart id="5F7848BDA21AEAAF71FA78A128674300">Poliseno, Angelo</mods:namePart>
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<mods:namePart id="28339E5F6A07DF20F783D12A315C2ECE">Hoeksema, Bert W.</mods:namePart>
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<treatment id="0889229B068BCE3E8604ED4865987E75" ID-GBIF-Taxon="152055734" LSID="urn:lsid:plazi:treatment:0889229B068BCE3E8604ED4865987E75" httpUri="http://treatment.plazi.org/id/0889229B068BCE3E8604ED4865987E75" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="84992BDAD3FCD9E890064A97831BDFBC" pageId="5" pageNumber="6" type="multiple">
<paragraph id="A0F760297036DCBFDDB7261EB528BE61" pageId="5" pageNumber="6">Taxon classification Animalia Zoantharia Zoanthidae</paragraph>
</subSubSection>
<subSubSection id="033F75F8BC3763C5BAE1311D55E29EE3" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="A0252B4DE2DC1CE2E19E48711F995A44" pageId="5" pageNumber="6">
1.
<taxonomicName id="54C879DAC586827B87428C5FE7013F9C" ID-CoL="9RR2" authority="Saville-Kent, 1893" authorityName="Saville-Kent" authorityYear="1893" class="Anthozoa" family="Zoanthidae" genus="Acrozoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrozoanthus australiae" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="species" species="australiae">
Acrozoanthus australiae
<normalizedToken id="ACEC9BC5E233456B69D978CB819C9906" originalValue="SavilleKent">Saville-Kent</normalizedToken>
, 1893
</taxonomicName>
Figures 2A, 3
</paragraph>
</subSubSection>
<subSubSection id="83313AFE506ABF47803ED752C8FC5C3A" pageId="5" pageNumber="6" type="specimens examined">
<paragraph id="1599523DE988568F08547B4339074DAA" pageId="5" pageNumber="6">Specimens examined</paragraph>
<paragraph id="28FA2D8A5256AB4B76BB621C02500E39" pageId="5" pageNumber="6">
(n=16). RMNH Coel 23405, Tg. Bengteng (=Galghoek), Ambon, Moluccas, depth = 3 to 4 m, collected November 10, 1990 by J.C. den Hartog; RMNH Coel 23406, outer bay, Ruhmatiga, Hitu, Ambon, Moluccas, depth = approx. 3 m, collected December 3, 1990 by J.C. den Hartog; RMNH Coel 23407, station 17, southeast side of Pombo Island, Ambon, Moluccas, depth = 6 m, collected November 17, 1994 by J.C. den Hartog; RMNH Coel 23408, west-northwest of Barrang Lompo, Spermonde Archipelago, South Sulawesi, depth = 1.5 to 4 m, collected December 23, 1994 by J.C. den Hartog; RMNH Coel 23409, entrance of harbor near light beacon, northwest of Gusung, Spermonde Archipelago, South Sulawesi, depth = 5 to 7 m, collected October 7, 1990 by J.C. den Hartog; RMNH Coel 23410, 7.5 km west of Makassar, Spermonde Archipelago, South Sulawesi (
<geoCoordinate id="4E481490E26AD6E97AACC93AC60E6C8F" direction="south" orientation="latitude" precision="925" value="-5.116667">05°07'S</geoCoordinate>
,
<geoCoordinate id="FCA9E53918FFD2F5E7A6D8917552216F" direction="east" orientation="longitude" precision="925" value="119.333336">119°20'E</geoCoordinate>
), depth = NA, collected May 31, 1994 by J.C. den Hartog; RMNH Coel 23411, west of Gusung (=
<normalizedToken id="D41B2A0106EA83F81C49D4DDC4D3BD38" originalValue="LaeLae">Lae-Lae</normalizedToken>
Keke) (=1 km northwest of Makassar), Spermonde Archipelago, South Sulawesi (
<geoCoordinate id="2F418A546C9641CF47AFE49E184BC891" direction="south" orientation="latitude" precision="92" value="-5.125">05°07.5'S</geoCoordinate>
,
<geoCoordinate id="61766676342986446842F1715CA94B98" direction="east" orientation="longitude" precision="925" value="119.38333">119°23'E</geoCoordinate>
), depth = NA, collected May 31, 1994 by J.C. den Hartog; RMNH Coel 24100, station MAL04, south coast northeast of Cape Hahurong, Ambon, Moluccas (
<geoCoordinate id="5BDF2A362F694EC9B1B3CC3A566EFE6E" direction="south" orientation="latitude" precision="925" value="-3.7833333">03°47'S</geoCoordinate>
,
<geoCoordinate id="2C244249FA53D2D9FCD0AEBCB2E1DC8A" direction="east" orientation="longitude" precision="925" value="128.1">128°06'E</geoCoordinate>
), depth = 2 to 28 m, collected June 6, 1996 by J.C. den Hartog; RMNH Coel 40361,
<normalizedToken id="DD8047A43E095CB1F227E149792571C4" originalValue="NNMLIPIWWF">NNM-LIPI-WWF</normalizedToken>
Expedition station BAL.16, southeast side of Pulau Serangan, Bali (
<geoCoordinate id="15934F78D0B8636943A7D305C5EAE7CB" direction="south" orientation="latitude" precision="15" value="-8.746667">08°44'48&quot;S</geoCoordinate>
,
<geoCoordinate id="0F69C56986BD3B6C628DB487F6F235EA" direction="east" orientation="longitude" precision="15" value="115.240555">115°14'26&quot;E</geoCoordinate>
), depth = to 10 m, collected April 6, 2001 by J. Goud; RMNH Coel 40549,
<normalizedToken id="A14FC81CABE5C61C30F25DA184B306CA" originalValue="SnelliusII">Snellius-II</normalizedToken>
Expedition station 4.011, reef edge west of Mai, Maisel Islands, Banda Sea (
<geoCoordinate id="A5CC0D3DDB49BD6D5E52FB7A5225797D" direction="south" orientation="latitude" precision="925" value="-5.4666667">05°28'S</geoCoordinate>
,
<geoCoordinate id="527EC900896E8B3EE4AF4F9A433A138D" direction="east" orientation="longitude" precision="925" value="127.51667">127°31'E</geoCoordinate>
), depth = 1 to 30 m, collected September 7, 1984; RMNH Coel 40550,
<normalizedToken id="68A28F238C8E12EEB76B3BEA76CB1C58" originalValue="SnelliusII">Snellius-II</normalizedToken>
Expedition station 4.001, near Tawiri, Ambon Bay, Moluccas (
<geoCoordinate id="A24014ECBDBF24B7FBE08F32BEA1FAC5" direction="south" orientation="latitude" precision="925" value="-3.7">03°42'S</geoCoordinate>
,
<geoCoordinate id="BBEBE385D49FFA91167927865E2C052E" direction="east" orientation="longitude" precision="925" value="128.11667">128°07'E</geoCoordinate>
), depth = approx. 1.5 to 8 m, collected September 4, 1984; RMNH Coel 40554,
<normalizedToken id="7EFE6B22B74F9BED7D5874CE6338D8B5" originalValue="SnelliusII">Snellius-II</normalizedToken>
Expedition station 4.006, near Eri, Ambon Bay, Moluccas (
<geoCoordinate id="1E3AD1848682224900698B7F362B6188" direction="south" orientation="latitude" precision="925" value="-3.75">03°45'S</geoCoordinate>
,
<geoCoordinate id="A94F0E96E72318235A3B05C267FA4D2B" direction="east" orientation="longitude" precision="925" value="128.13333">128°8'E</geoCoordinate>
), depth = approx. 1.5 to 5 m, collected September 4, 1984; RMNH Coel 40556,
<normalizedToken id="6D8E926D3312E5E4BF6E8519790C893F" originalValue="SnelliusII">Snellius-II</normalizedToken>
Expedition station 4.006, near Eri, Ambon Bay, Moluccas (
<geoCoordinate id="48E137F6DCAA3C3B977C6304B702538B" direction="south" orientation="latitude" precision="925" value="-3.75">03°45'S</geoCoordinate>
,
<geoCoordinate id="D42A41D1F3D0F7A9B56431C5A538A440" direction="east" orientation="longitude" precision="925" value="128.13333">128°8'E</geoCoordinate>
), depth = approx. 1.5 to 5 m, collected August 29, 1984; RMNH Coel 40558,
<normalizedToken id="B50C204C00CB70AF4C80919319DB7AE3" originalValue="SnelliusII">Snellius-II</normalizedToken>
Expedition station 4.030, west coast of Binongko, Tukang Besi Islands, Banda Sea (
<geoCoordinate id="F881CC84B5E8B6BB1EA89451DC004CAE" direction="south" orientation="latitude" precision="925" value="-5.9166665">05°55'S</geoCoordinate>
,
<geoCoordinate id="43A882AEE5ED25467188E21EE691623D" direction="east" orientation="longitude" precision="925" value="123.98333">123°59'E</geoCoordinate>
), depth = approx. 3 to 4 m, collected September 10, 1984 by M. Slierings; RMNH Coel 40566, west side of Pulau Samalona, 7.5 km west of Makassar, Spermonde Archipelago, South Sulawesi (
<geoCoordinate id="A2D65D1449725A041D09957B82C66A23" direction="south" orientation="latitude" precision="925" value="-5.116667">05°07'S</geoCoordinate>
,
<geoCoordinate id="EB6AE95BE298DE090A6B428F0D688BC4" direction="east" orientation="longitude" precision="925" value="119.333336">119°20'E</geoCoordinate>
), depth = NA, collected February 18, 1994 by B.W. Hoeksema; RMNH Coel 40569, Fauna Malesiana Marine Sulawesi Expedition station SUL.06, Pantai Parigi, Pulau Lembeh, Selat Lembeh, North Sulawesi (
<geoCoordinate id="425AF091BFEBBD54C7CE94B7CEE55AA7" direction="north" orientation="latitude" precision="925" value="1.4666667">01°28'N</geoCoordinate>
,
<geoCoordinate id="77511A92C7340D63E90C8F292D844500" direction="east" orientation="longitude" precision="925" value="125.23333">125°14'E</geoCoordinate>
), depth = 0 to 6 m, collected October 15, 1994 by M. Slierings.
</paragraph>
</subSubSection>
<subSubSection id="F186E3DA130FDE714EA566CF60846184" pageId="5" pageNumber="6" type="photographic records">
<paragraph id="AB92DCD21E25E5E10A36BF1AB7BE9223" pageId="5" pageNumber="6">Photographic records</paragraph>
<paragraph id="27DB2D5ACA5C4DC04409CC867DFC800A" pageId="5" pageNumber="6">
(n=6). West side of Pulau
<normalizedToken id="1B632EDE80C79C57A8D36E2C76691720" originalValue="LaeLae">Lae-Lae</normalizedToken>
(
<geoCoordinate id="A8886B3D204C96EE93C78DF60A07F3C3" direction="south" orientation="latitude" precision="15" value="-5.134722">05°08'05&quot;S</geoCoordinate>
,
<geoCoordinate id="C447B67924CF8FA46BADC460CBBF2961" direction="east" orientation="longitude" precision="15" value="119.3875">119°23'15&quot;E</geoCoordinate>
), South Sulawesi, May 24, 1997; station MAL.19 (
<geoCoordinate id="BD2A6815F9D3910D53EB357CE11DEBFE" direction="south" orientation="latitude" precision="925" value="-3.7166667">03°43'S</geoCoordinate>
,
<geoCoordinate id="860A3C77388D3972BADD642EB9AA0855" direction="east" orientation="longitude" precision="925" value="128.05">128°03'E</geoCoordinate>
), Tanjune Batu Dua, east of Hatu, north coast of Ambon Bay, Moluccas, November 19, 1996; station MAL.22 (
<geoCoordinate id="2AA4C7224AE823E364E537797E833A1B" direction="south" orientation="latitude" precision="925" value="-3.8">03°48'S</geoCoordinate>
,
<geoCoordinate id="99F4E40E61142448EB84D0240E398704" direction="east" orientation="longitude" precision="925" value="128.1">128°06'E</geoCoordinate>
), southwest coast, east of Tunjung Nusanive, Ambon Bay, Moluccas, November 21, 1996; Nusa Penida, Lombok Strait, east Bali, May 26, 1998 (
<geoCoordinate id="F73B70D72507F48F659C4CA9012F9953" direction="south" orientation="latitude" precision="15" value="-8.682222">08°40'56&quot;S</geoCoordinate>
,
<geoCoordinate id="86C7EAB990F4BFE9EDECB88ABB6D2706" direction="east" orientation="longitude" precision="15" value="115.48222">115°28'56&quot;E</geoCoordinate>
); northwest Pulau Samalona, Spermonde Archipelago, South Sulawesi (
<geoCoordinate id="8B62CE6C8DB0391AFCBFA3E7D0DBB28E" direction="south" orientation="latitude" precision="15" value="-5.1236115">05°07'25&quot;S</geoCoordinate>
,
<geoCoordinate id="A979D6C33E1328E1C9CC207840FF3220" direction="east" orientation="longitude" precision="15" value="119.33611">119°20'10&quot;E</geoCoordinate>
), January 12, 1997; western slope of Bone Lola shoal, Spermonde Archipelago, South Sulawesi (
<geoCoordinate id="A4FF77341C49A16F2B4E20CC4F286A9C" direction="south" orientation="latitude" precision="15" value="-5.054167">05°03'15&quot;S</geoCoordinate>
,
<geoCoordinate id="8EF82F214E1543FAA74F76ECFC0A0AFF" direction="east" orientation="longitude" precision="15" value="119.354164">119°21'15&quot;E</geoCoordinate>
), April 22, 1998.
</paragraph>
</subSubSection>
<subSubSection id="921FE6C1EDF22545B1CA1B0D0D24FE07" pageId="5" pageNumber="6" type="description">
<paragraph id="350D201E103250179AF3F7E64E8AFF1B" pageId="5" pageNumber="6">Description.</paragraph>
<paragraph id="4F1E6ACDF54FA7A1B77118440049F77A" pageId="5" pageNumber="6">
Non-incrusted zooxanthellate zoantharian that inhabits the outside of eunicid worm tubes (
<bibRefCitation id="14937A5E6867DA9A498A12E256C67E88" author="Haddon, AC" journalOrPublisher="Scientific Proceedings of the Royal Dublin Society" pageId="25" pageNumber="26" pagination="344 - 346" title="Branched worm-tubes and Acrozoanthus." volume="VIII" year="1895">Haddon 1895</bibRefCitation>
), with a unique asexual form of
<normalizedToken id="E2036ACD17F75146FCAF0577F5D192A6" originalValue="“budding”">&quot;budding&quot;</normalizedToken>
(
<bibRefCitation id="1E10A94A5E5CB399B65B1AB019BB2065" pageId="5" pageNumber="6">Ryland 1997</bibRefCitation>
). Easily recognizable as it is an epibiont on outside surface of eunicid worm tube, and has a reduced stoloniferous coenenchyme, long pale yellow-green or pale purple tentacles (n=approx. 40-50) with occasional fluorescent green markings and black tips, and light brown/purple to white ectoderm with similarly colored oral disks (Figure 2A). Preserved specimens in this study had polyps of average 6.0 mm in height (range 2.5-14 mm), 3.2 mm in width (range 2-5 mm) (n=8 specimens examined [RMNH Coel 40361, 40549, 40550, 40554, 40556, 40558, 40566, 40569], 10 polyps/specimen), and oral disks approximately 6 mm in diameter when expanded in situ (partially adapted from
<bibRefCitation id="CA5EBD9D0D9111389DE0C53E5BCB6A7B" author="Reimer, JD" journalOrPublisher="Marine Biodiversity" pageId="29" pageNumber="30" pagination="313 - 323" title="New records and molecular characterization of Acrozoanthus (Cnidaria: Anthozoa: Zoanthidae) from Taiwan." url="10.1007/s12526-010-0069-5" volume="41" year="2011 b">Reimer et al. 2011b</bibRefCitation>
). Specimen RMNH Coel 40566 had much larger polyps than other specimens (average height 10.6 mm, average width 4.3 mm), but this may be due to preservation in formalin as opposed to ethanol than to any phenotypic difference.
</paragraph>
<caption id="D9E0606CA6956A472F1E8A867D8772A0" pageId="5" pageNumber="6">
<paragraph id="18C5A523D2C60B172D90FAFB40EBA5C4" pageId="5" pageNumber="6">
Figure 2. Images of
<taxonomicName id="2CE53DA55DE878D85CBCE8F9F24EEB3F" class="Anthozoa" family="Zoanthidae" genus="Acrozoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrozoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Acrozoanthus</taxonomicName>
and
<taxonomicName id="61B0647EFA21F30994AA9851F7F37A4B" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Zoanthus</taxonomicName>
species from photographic records in this study. A
<taxonomicName id="60D76A77A754F29D73DCB9097B3F577B" class="Anthozoa" family="Zoanthidae" genus="Acrozoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrozoanthus australiae" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="species" species="australiae">Acrozoanthus australiae</taxonomicName>
at Nusa Penida, Lombok Strait, east Bali, May 26, 1998 B
<taxonomicName id="F08AF48577BE65A84697E1D3F083F889" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus sansibaricus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="species" species="sansibaricus">Zoanthus sansibaricus</taxonomicName>
at Station BER.26, northeast Buliulin (south of Samama Island), East Kalimantan, Berau Islands, October 15, 2003 C
<taxonomicName id="3206B015404D0EC96CBE9C318196237C" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Zoanthus</taxonomicName>
sp. at west side of Pulau Samalona, Spermonde Archipelago, South Sulawesi, September 16, 1997; and D
<taxonomicName id="534FF72EFE1D83DAA0B6758DA5E1AD69" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Zoanthus</taxonomicName>
sp. west of Gusung (=Pulau
<normalizedToken id="14C4C7AC9E1D74897DA6311361266D93" originalValue="LaeLae">Lae-Lae</normalizedToken>
Keke), Spermonde Archipelago, South Sulawesi, October 11 1997.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0CAD22DD03E40543F3718018C7AF02B1" pageId="5" pageNumber="6" type="distribution">
<paragraph id="9FF7021112D08D86C25DB71677CE99E3" pageId="5" pageNumber="6">Distribution.</paragraph>
<paragraph id="A45DB530340FFDFA38AA0C9576361461" pageId="5" pageNumber="6">Regions recorded in this study (Figure 3). East coast of Bali (5), Spermonde Archipelago (9), Tukang Besi Islands (12), Maisel Islands (13), Moluccas (14), Lembeh Strait (17).</paragraph>
<caption id="145E631D2CD6506215CC3619CB53FD5A" pageId="5" pageNumber="6">
<paragraph id="DB24511F8636C75232DC81ADC339784C" pageId="5" pageNumber="6">
Figure 3. Distribution of
<taxonomicName id="5EC0F0BADCD58B0D5A0DD489E42FD148" class="Anthozoa" family="Zoanthidae" genus="Acrozoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrozoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Acrozoanthus</taxonomicName>
and
<taxonomicName id="23992E04C20096FEEE47110ABBE8B312" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Zoanthus</taxonomicName>
species from specimens and photographic records from this study.
<taxonomicName id="8CCAAC03D020D8164D5DAC4A32201216" class="Anthozoa" family="Zoanthidae" genus="Acrozoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrozoanthus australiae" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="species" species="australiae">Acrozoanthus australiae</taxonomicName>
specimens in red,
<taxonomicName id="ED83FFBD62F8D2E50A29D9A59E12CE52" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus sansibaricus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="species" species="sansibaricus">Zoanthus sansibaricus</taxonomicName>
in green, and
<taxonomicName id="550EB789490E7426E801D86035E66A29" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Zoanthus</taxonomicName>
sp. in blue. Region numbers correspond to locations given in
<normalizedToken id="132E3141BC05868F28450525F98332DD" originalValue="species">species'</normalizedToken>
information. Boxes indicate presence of specimens (with or without photographic records), while circles indicate only photographic records. Overlapping symbols indicate the same region.
</paragraph>
</caption>
<paragraph id="6B3586271D8A958EBD01470CBF03EABB" pageId="5" pageNumber="6">
Previous records. Originally described from Australia, where it has been reported from both the coast of northern Queensland, and the region around Darwin in the Northern Territory. Subsequent records reported from North Sulawesi, Indonesia (
<bibRefCitation id="6F09F30A7F2C766B7166E86D6403E235" author="Sinniger, F" journalOrPublisher="Marine Biology" pageId="29" pageNumber="30" pagination="1121 - 1128" title="Phylogeny of the order Zoantharia (Anthozoa, Hexacorallia) based on the mitochondrial ribosomal genes." url="10.1007/s00227-005-0016-3" volume="147" year="2005">Sinniger et al. 2005</bibRefCitation>
,
<bibRefCitation id="CAFAAE9729AA16386C39055FB8ECD49D" author="Reimer, JD" journalOrPublisher="Marine Biodiversity" pageId="29" pageNumber="30" pagination="313 - 323" title="New records and molecular characterization of Acrozoanthus (Cnidaria: Anthozoa: Zoanthidae) from Taiwan." url="10.1007/s12526-010-0069-5" volume="41" year="2011 b">Reimer et al. 2011b</bibRefCitation>
), and photographic records from Mactan Island, Philippines (
<bibRefCitation id="971090A7EE5EB99B6DBCD00B568F3374" author="Reimer, JD" journalOrPublisher="Marine Biodiversity" pageId="29" pageNumber="30" pagination="313 - 323" title="New records and molecular characterization of Acrozoanthus (Cnidaria: Anthozoa: Zoanthidae) from Taiwan." url="10.1007/s12526-010-0069-5" volume="41" year="2011 b">Reimer et al. 2011b</bibRefCitation>
). Also reported from southern Taiwan (
<bibRefCitation id="E99E73BBE6EB3B21D24D68D8464433AD" author="Reimer, JD" journalOrPublisher="Marine Biodiversity" pageId="29" pageNumber="30" pagination="313 - 323" title="New records and molecular characterization of Acrozoanthus (Cnidaria: Anthozoa: Zoanthidae) from Taiwan." url="10.1007/s12526-010-0069-5" volume="41" year="2011 b">Reimer et al. 2011b</bibRefCitation>
) and at Ningaloo, Western Australia (Y. Irei and J.D. Reimer, unpubl. data).
</paragraph>
</subSubSection>
<subSubSection id="4C7628F5EB1E5B723F55F19C312FA7BE" pageId="5" pageNumber="6" type="remarks">
<paragraph id="1ECB89D68AC6B88D9F6225341DEECF53" pageId="5" pageNumber="6">Remarks.</paragraph>
<paragraph id="78C7C4D970895922A27C306441C5A988" pageId="5" pageNumber="6">
This genus is positioned within the genus
<taxonomicName id="65DC41E1529AF8FC37AAC90AAA3496BF" class="Anthozoa" family="Zoanthidae" genus="Zoanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoanthus" order="Zoantharia" pageId="5" pageNumber="6" phylum="Cnidaria" rank="genus">Zoanthus</taxonomicName>
based on phylogenetic analyses (
<bibRefCitation id="8C2B69EC0C946928A0EF8F6104F3B21D" author="Reimer, JD" journalOrPublisher="Marine Biodiversity" pageId="29" pageNumber="30" pagination="313 - 323" title="New records and molecular characterization of Acrozoanthus (Cnidaria: Anthozoa: Zoanthidae) from Taiwan." url="10.1007/s12526-010-0069-5" volume="41" year="2011 b">Reimer et al. 2011b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>