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<mods:title id="A618D29239449A67DFC5E0A51117E55A">Oysters (Bivalvia: Ostreidae And Gryphaeidae) Recorded From Malaysia And Singapore</mods:title>
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<heading id="D0A13638FFF3FFF9FC51AF0CDB08FAA6" bold="true" box="[963,1341,1312,1336]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="483" reason="2">
<taxonomicName id="4C56FAD7FFF3FFF9FC51AF0CDB08FAA6" ID-CoL="6WYJ6" authority="(Gould, 1850)" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[963,1341,1312,1336]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="mordax">
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<emphasis id="B9225D46FFF3FFF9FC51AF0CDAA3FAA6" bold="true" box="[963,1174,1312,1336]" italics="true" pageId="2" pageNumber="483">Saccostrea mordax</emphasis>
(
<bibRefCitation id="EFC7FCA5FFF3FFF9FB35AF0CDB03FAA6" author="Gould, A" box="[1191,1334,1312,1336]" pageId="2" pageNumber="483" pagination="343 - 348" refId="ref7883" refString="Gould, A. A., 1850. Shells of the United States Exploring Expedition. Proceedings of the Boston Society of Natural History, 3: 343 - 348." type="journal article" year="1850">Gould, 1850</bibRefCitation>
)
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<paragraph id="8BE98154FFF3FFF9FC11AF6DDB48FAC6" blockId="2.[899,1405,1312,1368]" box="[899,1405,1345,1368]" pageId="2" pageNumber="483">
Common name: Crenulate rock oyster (
<figureCitation id="136D9DD1FFF3FFF9FAA1AF6DDB40FAC6" box="[1331,1397,1345,1368]" captionStart="Fig" captionStartId="4.[134,165,2065,2084]" captionTargetBox="[136,1470,1433,2040]" captionTargetId="figure-616@4.[132,1474,1429,2040]" captionTargetPageId="4" captionText="Fig. 2. Saccostrea mordax (Gould, 1850) from Pulau Langkawi, Malaysia (NHM20090289). A, external view of the right shell valve, B, internal view of the right shell valve and C, internal view of the left shell valve." figureDoi="http://doi.org/10.5281/zenodo.5342236" httpUri="https://zenodo.org/record/5342236/files/figure.png" pageId="2" pageNumber="483">Fig. 2</figureCitation>
)
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<paragraph id="8BE98154FFF3FFF9FCD3AF53DB54FA0A" blockId="2.[833,1472,1407,1745]" box="[833,1377,1407,1428]" pageId="2" pageNumber="483">
<treatmentCitationGroup id="AB46A67AFFF3FFF9FCD3AF53DB54FA0A" box="[833,1377,1407,1428]" pageId="2" pageNumber="483">
<treatmentCitation id="0AF7A745FFF3FFF9FCD3AF53DA85FA0A" author="Lamarck, J" box="[833,1200,1407,1428]" page="200" pageId="2" pageNumber="483" year="1819">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3AF53DA85FA0A" ID-CoL="75CTN" authority="Lamarck, 1819: 200" authorityName="Lamarck" authorityPageNumber="200" authorityYear="1819" box="[833,1200,1407,1428]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="cucullata">
<emphasis id="B9225D46FFF3FFF9FCD3AF53DDD7FA0A" box="[833,994,1407,1428]" italics="true" pageId="2" pageNumber="483">Ostrea cucullata</emphasis>
<bibRefCitation id="EFC7FCA5FFF3FFF9FC78AF53DA85FA0A" author="Lamarck, J" box="[1002,1200,1407,1428]" pageId="2" pageNumber="483" refId="ref8436" refString="Lamarck, J. B. P. de M., 1819. Histoire naturelle des animaux sans vertebres. Suite des conchiferes. Chez L'Auteur, Paris. 258 pp." type="book" year="1819">Lamarck,1819: 200</bibRefCitation>
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[non
<bibRefCitation id="EFC7FCA5FFF3FFF9FB79AF53DB6EFA0A" author="Born, I" box="[1259,1371,1407,1428]" pageId="2" pageNumber="483" refId="ref7511" refString="Born, I., 1778. Index Rerum Naturalium Musei Caesarei Vindobonensis, pars prima, Testacea. Ex Officina Krausiana, Vindobonae, Vienna. 442 pp." type="book" year="1778">Born, 1778</bibRefCitation>
]
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<treatmentCitation id="0AF7A745FFF3FFF9FCD3AFB0DAA8FA2F" author="Gould, A" box="[833,1181,1436,1457]" page="346" pageId="2" pageNumber="483" year="1850">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3AFB0DAA8FA2F" authority="Gould, 1850: 346" authorityName="Gould" authorityPageNumber="346" authorityYear="1850" box="[833,1181,1436,1457]" class="Bivalvia" family="Ostreidae" genus="Ostraea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="mordax">
<emphasis id="B9225D46FFF3FFF9FCD3AFB0DDD4FA2F" box="[833,993,1436,1457]" italics="true" pageId="2" pageNumber="483">Ostraea mordax</emphasis>
<bibRefCitation id="EFC7FCA5FFF3FFF9FC78AFB0DAA8FA2F" author="Gould, A" box="[1002,1181,1436,1457]" pageId="2" pageNumber="483" pagination="343 - 348" refId="ref7883" refString="Gould, A. A., 1850. Shells of the United States Exploring Expedition. Proceedings of the Boston Society of Natural History, 3: 343 - 348." type="journal article" year="1850">Gould, 1850: 346</bibRefCitation>
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;
<bibRefCitation id="EFC7FCA5FFF3FFF9FB38AFB0DB74FA2F" author="Sowerby, G" box="[1194,1345,1436,1457]" pageId="2" pageNumber="483" refId="ref9218" refString="Sowerby, G. B., 1871. Conchologia Iconica. Volume 18. Ostrea. L. Reeve &amp; Co, London." type="book" year="1871">Sowerby, 1871</bibRefCitation>
, Pl. 15, Fig. 31a, b.
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<paragraph id="8BE98154FFF3FFF9FCD3AFFADB33FA75" blockId="2.[833,1472,1407,1745]" box="[833,1286,1494,1515]" pageId="2" pageNumber="483">
<treatmentCitationGroup id="AB46A67AFFF3FFF9FCD3AFFADB33FA75" box="[833,1286,1494,1515]" pageId="2" pageNumber="483">
<treatmentCitation id="0AF7A745FFF3FFF9FCD3AFFADB37FA75" author="Lamy, E" box="[833,1282,1494,1515]" page="159" pageId="2" pageNumber="483" year="1929">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3AFFADB37FA75" ID-CoL="6TD6R" authority="Lamy, 1929: 159" authorityName="Lamy" authorityPageNumber="159" authorityYear="1929" box="[833,1282,1494,1515]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="variety" species="forskali" variety="mordax">
<emphasis id="B9225D46FFF3FFF9FCD3AFFADDE7FA75" box="[833,978,1494,1515]" italics="true" pageId="2" pageNumber="483">Ostrea forskali</emphasis>
var.
<emphasis id="B9225D46FFF3FFF9FB95AFFADA7AFA75" box="[1031,1103,1494,1515]" italics="true" pageId="2" pageNumber="483">mordax</emphasis>
<bibRefCitation id="EFC7FCA5FFF3FFF9FBC5AFFADB37FA75" author="Lamy, E" box="[1111,1282,1494,1515]" pageId="2" pageNumber="483" pagination="233 - 275" refId="ref8492" refString="Lamy, E., 1929. Revision des Ostrea vivants du Museum National d'Histoire Naturelle de Paris. Journal de Conchyliologie, 7: 1 - 46, 71 - 108, 133 - 168, 233 - 275." type="journal article" year="1929">Lamy, 1929: 159</bibRefCitation>
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.
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<paragraph id="8BE98154FFF3FFF9FCD3AFDFDB2DF999" blockId="2.[833,1472,1407,1745]" box="[833,1304,1522,1544]" pageId="2" pageNumber="483">
<treatmentCitationGroup id="AB46A67AFFF3FFF9FCD3AFDFDB2DF999" box="[833,1304,1522,1544]" pageId="2" pageNumber="483">
<treatmentCitation id="0AF7A745FFF3FFF9FCD3AFDFDAB1F999" author="Iredale, T" box="[833,1156,1522,1544]" page="399" pageId="2" pageNumber="483" year="1939">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3AFDFDAB1F999" ID-CoL="6TD4X" authority="Iredale, 1939: 399" authorityName="Iredale" authorityPageNumber="399" authorityYear="1939" box="[833,1156,1522,1544]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="amasa">
<emphasis id="B9225D46FFF3FFF9FCD3AFDFDDFDF996" box="[833,968,1523,1544]" italics="true" pageId="2" pageNumber="483">Ostrea amasa</emphasis>
<bibRefCitation id="EFC7FCA5FFF3FFF9FC5DAFDEDAB1F999" author="Iredale, T" box="[975,1156,1522,1543]" pageId="2" pageNumber="483" pagination="211 - 425" refId="ref8146" refString="Iredale, T., 1939. Mollusca. In: Iredale, T. (ed.), Great Barrier Reef Expedition 1928 - 1929, Scientific Reports 5 (6), Part I. British Museum (Natural History), London. Pp. 211 - 425." type="book chapter" year="1939">Iredale, 1939: 399</bibRefCitation>
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, Pl. 17, Fig. 8.
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<paragraph id="8BE98154FFF3FFF9FCD3AC23DB4FF9BA" blockId="2.[833,1472,1407,1745]" box="[833,1402,1551,1572]" pageId="2" pageNumber="483">
<treatmentCitationGroup id="AB46A67AFFF3FFF9FCD3AC23DB4FF9BA" box="[833,1402,1551,1572]" pageId="2" pageNumber="483">
<treatmentCitation id="0AF7A745FFF3FFF9FCD3AC23DAFAF9BA" author="Thomson, J" box="[833,1231,1551,1572]" page="154" pageId="2" pageNumber="483" year="1954">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3AC23DAFAF9BA" ID-CoL="6BBHR" authority="Thomson, 1954: 154" authorityName="Thomson" authorityPageNumber="154" authorityYear="1954" box="[833,1231,1551,1572]" class="Bivalvia" family="Ostreidae" genus="Crassostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="amasa">
<emphasis id="B9225D46FFF3FFF9FCD3AC23DDCEF9BA" box="[833,1019,1551,1572]" italics="true" pageId="2" pageNumber="483">Crassostrea amasa</emphasis>
<bibRefCitation id="EFC7FCA5FFF3FFF9FB90AC23DAFAF9BA" author="Thomson, J" box="[1026,1231,1551,1572]" pageId="2" pageNumber="483" pagination="132 - 168" refId="ref9494" refString="Thomson, J. M., 1954. The genera of oysters and the Australian species. Australian Journal of Marine and Freshwater Research, 5: 132 - 168." type="journal article" year="1954">Thomson, 1954: 154</bibRefCitation>
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, Pl. 7,
<figureCitation id="136D9DD1FFF3FFF9FA85AC23DB6FF9BA" box="[1303,1370,1551,1572]" captionStart="Fig" captionStartId="3.[111,142,2039,2058]" captionTargetBox="[155,1408,188,2011]" captionTargetId="figure-64@3.[152,1411,187,2014]" captionTargetPageId="3" captionText="Fig. 1. Saccostrea cuccullata (Born, 1778) from Pulau Langkawi, Malaysia, showing external and internal views of the right and left shell valves. AD, ecotype occurring on mangrove trees (NHM20090286), EH, ecotype occurring on low intertidal beach rocks (NHM20090287), IL, ecotype occurring on intertidal rocks (NHM20090288)." figureDoi="http://doi.org/10.5281/zenodo.5342234" httpUri="https://zenodo.org/record/5342234/files/figure.png" pageId="2" pageNumber="483">Figs. 1</figureCitation>
,
<figureCitation id="136D9DD1FFF3FFF9FAF5AC23DB40F9BA" box="[1383,1397,1551,1572]" captionStart="Fig" captionStartId="4.[134,165,2065,2084]" captionTargetBox="[136,1470,1433,2040]" captionTargetId="figure-616@4.[132,1474,1429,2040]" captionTargetPageId="4" captionText="Fig. 2. Saccostrea mordax (Gould, 1850) from Pulau Langkawi, Malaysia (NHM20090289). A, external view of the right shell valve, B, internal view of the right shell valve and C, internal view of the left shell valve." figureDoi="http://doi.org/10.5281/zenodo.5342236" httpUri="https://zenodo.org/record/5342236/files/figure.png" pageId="2" pageNumber="483">2</figureCitation>
.
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<treatmentCitationGroup id="AB46A67AFFF3FFF9FCD3AC00DBB0F9DF" box="[833,1413,1580,1601]" pageId="2" pageNumber="483">
<treatmentCitation id="0AF7A745FFF3FFF9FCD3AC00DACBF9DF" author="Thomson, J" box="[833,1278,1580,1601]" page="157" pageId="2" pageNumber="483" year="1954">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3AC00DACBF9DF" authority="Thomson, 1954: 157" authorityName="Thomson" authorityPageNumber="157" authorityYear="1954" box="[833,1278,1580,1601]" class="Bivalvia" family="Ostreidae" genus="Crassostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B9225D46FFF3FFF9FCD3AC00DA1FF9DF" box="[833,1066,1580,1601]" italics="true" pageId="2" pageNumber="483">Crassostrea tuberculata</emphasis>
<bibRefCitation id="EFC7FCA5FFF3FFF9FBA3AC00DACBF9DF" author="Thomson, J" box="[1073,1278,1580,1601]" pageId="2" pageNumber="483" pagination="132 - 168" refId="ref9494" refString="Thomson, J. M., 1954. The genera of oysters and the Australian species. Australian Journal of Marine and Freshwater Research, 5: 132 - 168." type="journal article" year="1954">Thomson, 1954: 157</bibRefCitation>
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, Pl. 8,
<figureCitation id="136D9DD1FFF3FFF9FAD4AC00DBB4F9DF" box="[1350,1409,1580,1601]" captionStart="Fig" captionStartId="5.[111,142,2065,2084]" captionTargetBox="[195,1361,187,2040]" captionTargetId="figure-61@5.[195,1362,187,2040]" captionTargetPageId="5" captionText="Fig. 3. Crassostrea gigas (Thünberg, 1793) from Johor Strait, Malaysia (NHM20090290). A, external view of the right shell valve, B, external view of the left shell valve. C, internal view of the right shell valve D, internal view of the left shell valve." figureDoi="http://doi.org/10.5281/zenodo.5342238" httpUri="https://zenodo.org/record/5342238/files/figure.png" pageId="2" pageNumber="483">Fig. 3</figureCitation>
.
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</paragraph>
<paragraph id="8BE98154FFF3FFF9FCD3AC65DB2AF9C0" blockId="2.[833,1472,1407,1745]" box="[833,1311,1609,1630]" pageId="2" pageNumber="483">
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<taxonomicName id="4C56FAD7FFF3FFF9FCD3AC65DDCCF9C0" ID-CoL="6WYJ6" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[833,1017,1609,1630]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="mordax">
<emphasis id="B9225D46FFF3FFF9FCD3AC65DDCCF9C0" box="[833,1017,1609,1630]" italics="true" pageId="2" pageNumber="483">Saccostrea mordax</emphasis>
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<bibRefCitation id="EFC7FCA5FFF3FFF9FB87AC65DAE6F9C0" author="Torigoe, K" box="[1045,1235,1609,1630]" pageId="2" pageNumber="483" pagination="291 - 481" refId="ref9595" refString="Torigoe, K., 1981. Oysters in Japan. Journal of Science-Hiroshima University Series B Division 1, 29: 291 - 481." type="journal article" year="1981">Torigoe, 1981: 328</bibRefCitation>
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, Pl. 17.
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</paragraph>
<paragraph id="8BE98154FFF3FFF9FCD3AC4ADA72F909" blockId="2.[833,1472,1407,1745]" pageId="2" pageNumber="483">
<emphasis id="B9225D46FFF3FFF9FCD3AC4ADA3CF9E5" box="[833,1033,1638,1659]" italics="true" pageId="2" pageNumber="483">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3AC4ADDC2F9E5" ID-CoL="6WYJ6" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[833,1015,1638,1659]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="mordax">Saccostrea mordax</taxonomicName>
</emphasis>
Qi &amp; Choe, 2000: 138139,
<figureCitation id="136D9DD1FFF3FFF9FA8DAC4ADBB1F9E5" box="[1311,1412,1638,1659]" captionStart="Fig" captionStartId="5.[111,142,2065,2084]" captionTargetBox="[195,1361,187,2040]" captionTargetId="figure-61@5.[195,1362,187,2040]" captionTargetPageId="5" captionText="Fig. 3. Crassostrea gigas (Thünberg, 1793) from Johor Strait, Malaysia (NHM20090290). A, external view of the right shell valve, B, external view of the left shell valve. C, internal view of the right shell valve D, internal view of the left shell valve." figureDoi="http://doi.org/10.5281/zenodo.5342238" httpUri="https://zenodo.org/record/5342238/files/figure.png" pageId="2" pageNumber="483">Fig. 3CE</figureCitation>
; Lam, 2003: 112113, Pl. 13.
</paragraph>
<paragraph id="8BE98154FFF3FFF9FCD3ACB3DDA9F94F" blockId="2.[833,1472,1407,1745]" pageId="2" pageNumber="483">
<treatmentCitationGroup id="AB46A67AFFF3FFF9FCD3ACB3DDA9F94F" pageId="2" pageNumber="483">
<taxonomicName id="4C56FAD7FFF3FFF9FCD3ACB3DA3EF92A" ID-CoL="75CTN" baseAuthorityName="Born" baseAuthorityYear="1778" box="[833,1035,1695,1716]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="2" pageNumber="483" phylum="Mollusca" rank="species" species="cucullata">
<emphasis id="B9225D46FFF3FFF9FCD3ACB3DA3EF92A" box="[833,1035,1695,1716]" italics="true" pageId="2" pageNumber="483">Saccostrea cucullata</emphasis>
</taxonomicName>
<treatmentCitation id="0AF7A745FFF3FFF9FBBBACB3DB3AF92A" author="Morris, S" box="[1065,1295,1695,1716]" page="125" pageId="2" pageNumber="483" year="1985">
<bibRefCitation id="EFC7FCA5FFF3FFF9FBBBACB3DB3AF92A" author="Morris, S" box="[1065,1295,1695,1716]" pageId="2" pageNumber="483" pagination="119 - 138" refId="ref8723" refString="Morris, S., 1985. Preliminary guide to the oysters of Hong Kong. Asian Marine Biology, 2: 119 - 138." type="journal article" year="1985">Morris, 1985: 125128</bibRefCitation>
</treatmentCitation>
, Pl. 3, Fig. A, B, C, D.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34CD2DFFFF3FFF9FCD3ACD9DAB4F829" pageId="2" pageNumber="483" type="materials_examined">
<paragraph id="8BE98154FFF3FFF9FCD3ACD9DD8FF8DA" blockId="2.[833,1472,1781,1860]" pageId="2" pageNumber="483">
<emphasis id="B9225D46FFF3FFF9FCD3ACD9DA00F894" bold="true" box="[833,1077,1781,1802]" italics="true" pageId="2" pageNumber="483">Material examined. </emphasis>
<materialsCitation id="3B3E8B09FFF3FFF9FBD3ACDADD83F8DA" collectionCode="USNM" country="Fiji" location="Feejee Islands'" municipality="Exploring Expedition" pageId="2" pageNumber="483" specimenCount="1" typeStatus="syntype">
<typeStatus id="54ED3FF6FFF3FFF9FBD3ACDADA9DF895" box="[1089,1192,1782,1803]" pageId="2" pageNumber="483" type="syntype">Syntypes</typeStatus>
: [
<collectionCode id="ED471991FFF3FFF9FB2FACDADB38F895" box="[1213,1293,1782,1803]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="483" type="Museum">USNM</collectionCode>
] 5958,
<location id="8E89D78FFFF3FFF9FAE3ACDADDA5F8B9" LSID="urn:lsid:plazi:treatment:03FF3042FFF3FFFFFC51AF0CDB26FC0B:8E89D78FFFF3FFF9FAE3ACDADDA5F8B9" country="Fiji" municipality="Exploring Expedition" name="Feejee Islands'" pageId="2" pageNumber="483">Feejee Islands</location>
(
<collectingCountry id="F341C1C4FFF3FFF9FC08AD3EDDF4F8B9" box="[922,961,1810,1831]" name="Fiji" pageId="2" pageNumber="483">Fiji</collectingCountry>
), coll. United States
<collectingMunicipality id="6B8D1B2EFFF3FFF9FB00AD3EDB53F8B9" box="[1170,1382,1810,1831]" pageId="2" pageNumber="483">Exploring Expedition</collectingMunicipality>
, no date, 3 dry shells
</materialsCitation>
.
</paragraph>
<paragraph id="8BE98154FFF3FFF9FCD3AD45DAB4F829" blockId="2.[833,1472,1897,1975]" pageId="2" pageNumber="483">
<materialsCitation id="3B3E8B09FFF3FFF9FCD3AD45DA48F829" collectingDate="2004-02-04" collectionCode="NHM" collectorName="B. Morton" country="Malaysia" location="Malaysia" pageId="2" pageNumber="483" specimenCount="1">
Voucher: [
<collectionCode id="ED471991FFF3FFF9FC3AAD45DDD0F8E0" box="[936,997,1897,1918]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="483" type="Herbarium">NHM</collectionCode>
] 20090289, Pelangi Beach rocks, low intertidal, Pulau Langkawi,
<collectingCountry id="F341C1C4FFF3FFF9FC6AADAADA63F805" box="[1016,1110,1926,1947]" name="Malaysia" pageId="2" pageNumber="483">Malaysia</collectingCountry>
, coll.
<collectorName id="26A3E482FFF3FFF9FB08ADAADB3CF805" box="[1178,1289,1926,1947]" pageId="2" pageNumber="483">B. Morton</collectorName>
,
<date id="FFE8A794FFF3FFF9FA84ADAADBB9F805" box="[1302,1420,1926,1947]" pageId="2" pageNumber="483" value="2004-02-04">
<collectingDate id="EFAC5E7CFFF3FFF9FA84ADAADBB9F805" box="[1302,1420,1926,1947]" pageId="2" pageNumber="483" value="2004-02-04">4 Feb.2004</collectingDate>
</date>
, two individuals preserved in alcohol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34CD2DFFFF3FFFFFCD3ADCCDFBAFACB" lastPageId="4" lastPageNumber="485" pageId="2" pageNumber="483" type="description">
<paragraph id="8BE98154FFF3FFFFFCD3ADCCDF7CFDEB" blockId="2.[833,1472,2016,2104]" lastBlockId="4.[133,775,190,629]" lastPageId="4" lastPageNumber="485" pageId="2" pageNumber="483">
<emphasis id="B9225D46FFF3FFF9FCD3ADCCDDEAF866" bold="true" box="[833,991,2016,2040]" italics="true" pageId="2" pageNumber="483">Description. </emphasis>
Shell medium, up to
<quantity id="4CAE2CB1FFF3FFF9FB4AADCDDB12F866" box="[1240,1319,2017,2040]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="483" unit="mm" value="47.0">47 mm</quantity>
in height,
<quantity id="4CAE2CB1FFF3FFF9FA37ADCDDD5FF786" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="483" unit="mm" value="30.0">30 mm</quantity>
in length and
<quantity id="4CAE2CB1FFF3FFF9FB98A22DDA7CF786" box="[1034,1097,2049,2072]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="483" unit="mm" value="6.0">6 mm</quantity>
in width. Outline either triangular or an elongate D-shaped depending upon the substratum. Left valve flat and attached completely.The dorsal margin very thin and easily broken when collected. Margin of left valve built up steeply along the anterior and posterior sides. Margins thick, with packed layers of growth squamae running along the margin. Closely and evenly spaced ribs perpendicular to these end as marginal crenulations. In most cases, where the oyster grows on unlimited flat rock surfaces, the anterior margin is reduced so that the right valve is in touch with the substratum. When viewed along the mouth-anus axis, the shape of the oyster is almost a right-angled triangle with height given by the height of the posterior margin and base given by the left valve. Hinge line straight and short. Ligament area elongated to form an obvious left beak.
</paragraph>
<caption id="DF29D1DCFFF2FFF8FFFDADDBDF85F7A3" ID-DOI="http://doi.org/10.5281/zenodo.5342234" ID-Zenodo-Dep="5342234" httpUri="https://zenodo.org/record/5342234/files/figure.png" pageId="3" pageNumber="484" startId="3.[111,142,2039,2058]" targetBox="[155,1408,188,2011]" targetPageId="3">
<paragraph id="8BE98154FFF2FFF8FFFDADDBDF85F7A3" blockId="3.[111,1450,2039,2109]" pageId="3" pageNumber="484">
Fig. 1.
<taxonomicName id="4C56FAD7FFF2FFF8FF3DADDBDFD1F794" authority="(Born, 1778)" baseAuthorityName="Born" baseAuthorityYear="1778" box="[175,484,2039,2058]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="3" pageNumber="484" phylum="Mollusca" rank="species" species="cuccullata">
<emphasis id="B9225D46FFF2FFF8FF3DADDBDF59F794" box="[175,364,2039,2058]" italics="true" pageId="3" pageNumber="484">Saccostrea cuccullata</emphasis>
(
<bibRefCitation id="EFC7FCA5FFF2FFF8FEE8ADDBDFEBF794" author="Born, I" box="[378,478,2039,2058]" pageId="3" pageNumber="484" refId="ref7511" refString="Born, I., 1778. Index Rerum Naturalium Musei Caesarei Vindobonensis, pars prima, Testacea. Ex Officina Krausiana, Vindobonae, Vienna. 442 pp." type="book" year="1778">Born, 1778</bibRefCitation>
)
</taxonomicName>
from Pulau Langkawi, Malaysia, showing external and internal views of the right and left shell valves. AD, ecotype occurring on mangrove trees (NHM20090286), EH, ecotype occurring on low intertidal beach rocks (NHM20090287), IL, ecotype occurring on intertidal rocks (NHM20090288).
</paragraph>
</caption>
<paragraph id="8BE98154FFF5FFFFFF17A8B2DC08FC0B" blockId="4.[133,773,670,917]" pageId="4" pageNumber="485">Right valve convex, thicker than the left valve and white with purple patches on the less eroded ventral area. Shell usually eroded with exposed conchiolin scales near the dorsal end. Parallel grooves extend from half way along the dorso-ventral axis to the ventral shell margin. These grooves reveal uneroded growth lines on the right valve. Evenly-spaced crenulations around the shell margin are more obvious anteriorly and posteriorly.</paragraph>
<paragraph id="8BE98154FFF5FFFFFF17A992DFBAFACB" blockId="4.[133,773,958,1365]" pageId="4" pageNumber="485">Interior of the shell white with a shiny, pearly appearance. A band of dark green conchiolin usually occur near the ventral margin of the right valve. This band may be absent in some specimens. It appears that the conchiolin and prismatic calcite layers are deposited alternately at the right, inner, ventral shell surface. Adductor muscle scar relatively large and positioned in the posterior ventral half of the pallial area, circular to elongate-oval according to the growth form of the shell and mostly white, slightly stained with growth bands or purple. Chomata white, short, rod-shaped and arranged in a single line around the inner margin. They are more prominent dorsally and fade as they extend towards the ventral shell margin.</paragraph>
</subSubSection>
<subSubSection id="C34CD2DFFFF5FFFFFCD3AA92DDB3FE4B" pageId="4" pageNumber="485" type="distribution">
<paragraph id="8BE98154FFF5FFFFFCD3AA92DDB3FE4B" blockId="4.[833,1472,190,469]" pageId="4" pageNumber="485">
<emphasis id="B9225D46FFF5FFFFFCD3AA92DB0EFF48" box="[833,1339,190,214]" italics="true" pageId="4" pageNumber="485">
<emphasis id="B9225D46FFF5FFFFFCD3AA92DA52FF48" bold="true" box="[833,1127,190,214]" italics="true" pageId="4" pageNumber="485">Distribution and habitat. </emphasis>
<taxonomicName id="4C56FAD7FFF5FFFFFBFDAA93DB0EFF48" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[1135,1339,191,214]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="species" species="mordax">Saccostrea mordax</taxonomicName>
</emphasis>
occurs only on oceanic, exposed rocky shores as individuals among the
<taxonomicName id="4C56FAD7FFF5FFFFFCD3AAD3DDA1FE88" authorityName="Recluz" authorityYear="1848" box="[833,916,255,278]" class="Bivalvia" family="Mytilidae" genus="Septifer" kingdom="Animalia" order="Mytilida" pageId="4" pageNumber="485" phylum="Mollusca" rank="genus">
<emphasis id="B9225D46FFF5FFFFFCD3AAD3DDA1FE88" box="[833,916,255,278]" italics="true" pageId="4" pageNumber="485">Septifer</emphasis>
</taxonomicName>
and
<taxonomicName id="4C56FAD7FFF5FFFFFC5AAAD3DA1AFE88" box="[968,1071,255,278]" class="Maxillopoda" family="Tetraclitidae" genus="Tetraclita" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="485" phylum="Arthropoda" rank="genus">
<emphasis id="B9225D46FFF5FFFFFC5AAAD3DA1AFE88" box="[968,1071,255,278]" italics="true" pageId="4" pageNumber="485">Tetraclita</emphasis>
</taxonomicName>
zone on slopes or in pools frequently refreshed by strong waves (Lam &amp; Morton, 2004). This species is widely distributed in the Indo-West Pacific, for example,
<collectingCountry id="F341C1C4FFF5FFFFFC39AB72DDDFFEEB" box="[939,1002,350,373]" name="Japan" pageId="4" pageNumber="485">Japan</collectingCountry>
,
<collectingCountry id="F341C1C4FFF5FFFFFC6AAB72DA09FEEB" box="[1016,1084,350,373]" name="South Korea" pageId="4" pageNumber="485">Korea</collectingCountry>
,
<collectingCountry id="F341C1C4FFF5FFFFFBDBAB72DAA9FEEB" box="[1097,1180,350,373]" name="Taiwan" pageId="4" pageNumber="485">Taiwan</collectingCountry>
,
<collectingCountry id="F341C1C4FFF5FFFFFB3BAB72DB1EFEEB" box="[1193,1323,350,373]" name="Hong Kong" pageId="4" pageNumber="485">Hong Kong</collectingCountry>
, South
<collectingCountry id="F341C1C4FFF5FFFFFA12AB72DBF5FEEB" box="[1408,1472,350,373]" name="China" pageId="4" pageNumber="485">China</collectingCountry>
Sea, peninsular
<collectingCountry id="F341C1C4FFF5FFFFFB90AB52DA5AFE0B" box="[1026,1135,382,405]" name="Malaysia" pageId="4" pageNumber="485">Malaysia</collectingCountry>
,
<collectingCountry id="F341C1C4FFF5FFFFFBEDAB52DAC2FE0B" box="[1151,1271,382,405]" name="Singapore" pageId="4" pageNumber="485">Singapore</collectingCountry>
,
<collectingCountry id="F341C1C4FFF5FFFFFA95AB52DB4EFE0B" box="[1287,1403,382,405]" name="Indonesia" pageId="4" pageNumber="485">Indonesia</collectingCountry>
,
<collectingCountry id="F341C1C4FFF5FFFFFA19AB52DD91FE2B" name="Vanuatu" pageId="4" pageNumber="485">New Hebrides</collectingCountry>
[now
<collectingCountry id="F341C1C4FFF5FFFFFC63ABB2DA64FE2B" box="[1009,1105,414,437]" name="Vanuatu" pageId="4" pageNumber="485">Vanuatu</collectingCountry>
] and
<collectingCountry id="F341C1C4FFF5FFFFFB01ABB2DAC2FE2B" box="[1171,1271,414,437]" name="Australia" pageId="4" pageNumber="485">Australia</collectingCountry>
(Lam &amp; Morton, 2006).
</paragraph>
</subSubSection>
<subSubSection id="C34CD2DFFFF5FFFFFCD3ABD2DB26FC0B" pageId="4" pageNumber="485" type="discussion">
<paragraph id="8BE98154FFF5FFFFFCD3ABD2DB26FC0B" blockId="4.[833,1471,510,917]" pageId="4" pageNumber="485">
<emphasis id="B9225D46FFF5FFFFFCD3ABD2DAF0FD88" box="[833,1221,510,534]" italics="true" pageId="4" pageNumber="485">
<emphasis id="B9225D46FFF5FFFFFCD3ABD2DDFAFD88" bold="true" box="[833,975,510,534]" italics="true" pageId="4" pageNumber="485">Remarks. </emphasis>
<taxonomicName id="4C56FAD7FFF5FFFFFC4EABD3DAF0FD88" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[988,1221,511,534]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="species" species="mordax">Saccostrea mordax</taxonomicName>
</emphasis>
has been identified previously as
<taxonomicName id="4C56FAD7FFF5FFFFFC77A833DA78FDA8" authorityName="Thomson" authorityYear="1954" box="[997,1101,543,566]" class="Bivalvia" family="Ostreidae" genus="Crassostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="species" species="amasa">
<emphasis id="B9225D46FFF5FFFFFC77A833DA78FDA8" box="[997,1101,543,566]" italics="true" pageId="4" pageNumber="485">S. amasa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C56FAD7FFF5FFFFFB1EA833DDB1FDCB" authority="(Thompson, 1954)" baseAuthorityName="Thompson" baseAuthorityYear="1954" class="Bivalvia" family="Ostreidae" genus="Crassostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B9225D46FFF5FFFFFB1EA833DB18FDA8" box="[1164,1325,543,566]" italics="true" pageId="4" pageNumber="485">S. tuberculata</emphasis>
(Thompson, 1954)
</taxonomicName>
. This species is often confused with
<taxonomicName id="4C56FAD7FFF5FFFFFABCA813DB8AFDC8" baseAuthorityName="Born" baseAuthorityYear="1778" box="[1326,1471,575,598]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="species" species="cuccullata">
<emphasis id="B9225D46FFF5FFFFFABCA813DB8AFDC8" box="[1326,1471,575,598]" italics="true" pageId="4" pageNumber="485">S. cuccullata</emphasis>
</taxonomicName>
because of similar shell characters. Lam &amp; Morton (2006) suggest that
<taxonomicName id="4C56FAD7FFF5FFFFFC41A853DA73FD08" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[979,1094,639,662]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="species" species="mordax">
<emphasis id="B9225D46FFF5FFFFFC41A853DA73FD08" box="[979,1094,639,662]" italics="true" pageId="4" pageNumber="485">S. mordax</emphasis>
</taxonomicName>
occurs only on exposed, wholly marine rocky shores whereas the other
<taxonomicName id="4C56FAD7FFF5FFFFFB7EA8B3DB55FD28" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[1260,1376,671,694]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="genus">
<emphasis id="B9225D46FFF5FFFFFB7EA8B3DB55FD28" box="[1260,1376,671,694]" italics="true" pageId="4" pageNumber="485">Saccostrea</emphasis>
</taxonomicName>
lineages occupy a wide range of habitats from brackish mangroves to somewhat less exposed marine shores. Un-eroded
<taxonomicName id="4C56FAD7FFF5FFFFFCD3A8D3DA28FC88" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[833,1053,767,790]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="species" species="mordax">
<emphasis id="B9225D46FFF5FFFFFCD3A8D3DA28FC88" box="[833,1053,767,790]" italics="true" pageId="4" pageNumber="485">Saccostrea mordax</emphasis>
</taxonomicName>
individuals are distinct from other species of
<taxonomicName id="4C56FAD7FFF5FFFFFC52A933DA0EFCA8" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[960,1083,799,822]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="4" pageNumber="485" phylum="Mollusca" rank="genus">
<emphasis id="B9225D46FFF5FFFFFC52A933DA0EFCA8" box="[960,1083,799,822]" italics="true" pageId="4" pageNumber="485">Saccostrea</emphasis>
</taxonomicName>
in terms of the regularly-spaced grooves radiating from the umbone to the ventral margin of the right valve, its triangular shell shape and finely, regularly, m-shaped plicated valve margin.
</paragraph>
</subSubSection>
</treatment>
</document>