177 lines
22 KiB
XML
177 lines
22 KiB
XML
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<mods:title id="B037E06150F668FA32F60A82D708F1EF">Late Oligocene decapod crustaceans from the Trbovlje Formation of Slovenia, with a description of two new species of hymenosomatid crabs</mods:title>
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<mods:namePart id="1ABC4A3C9FA4FCAF822672D2521B4664">Gašparič, Rok</mods:namePart>
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<mods:namePart id="EACC1DC57A4E24C9A6B5984B64F2A300">Hyžný, Matúš</mods:namePart>
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<mods:namePart id="8D8BC15A815EC58F2E95CC374CA3D416">Hitij, Tomaž</mods:namePart>
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<mods:namePart id="1649F46F9CBB88465457B6CAD2C10598">Šoster, Aleš</mods:namePart>
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<mods:title id="68221583633952C0E0EFC2C179EAE93C">Palaeontologia Electronica</mods:title>
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<mods:date id="367D8D3C21D3FDAE1E0D88456166E387">2024</mods:date>
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<mods:title id="D89EED1D230D4BE116755DF85B6139A9">a 4</mods:title>
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<treatment id="03FF1D38366DCE3513FBEC3D3D41EB6A" ID-DOI="http://doi.org/10.5281/zenodo.11033758" ID-Zenodo-Dep="11033758" LSID="urn:lsid:plazi:treatment:03FF1D38366DCE3513FBEC3D3D41EB6A" httpUri="http://treatment.plazi.org/id/03FF1D38366DCE3513FBEC3D3D41EB6A" lastPageNumber="11" pageId="10" pageNumber="11">
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<subSubSection id="C34CFFA5366DCE3513FBEC3D3B77EA32" box="[199,784,1590,1615]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BE9AC2E366DCE3513FBEC3D3B77EA32" blockId="10.[199,784,1590,1647]" box="[199,784,1590,1615]" pageId="10" pageNumber="11">
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<heading id="D0A11B42366DCE3513FBEC3D3B77EA32" box="[199,784,1590,1615]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="11" reason="4">
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<taxonomicName id="4C56D7AD366DCE3513FBEC3D3B77EA32" ID-CoL="BSB7K" authority="A. Milne-Edwards, 1860" authorityName="A. Milne-Edwards" authorityYear="1860" box="[199,784,1590,1615]" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="michelini">
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<emphasis id="B922703C366DCE3513FBEC3D393AEA32" box="[199,349,1590,1615]" italics="true" pageId="10" pageNumber="11">Necronectes</emphasis>
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cf.
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<emphasis id="B922703C366DCE3512BBEC3D3988EA32" box="[391,495,1590,1615]" italics="true" pageId="10" pageNumber="11">michelini</emphasis>
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A. Milne-Edwards, 1860
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34CFFA5366DCE351298EC5D3A53EA12" box="[420,564,1622,1647]" pageId="10" pageNumber="11" type="description">
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<paragraph id="8BE9AC2E366DCE351298EC5D3A53EA12" blockId="10.[199,784,1590,1647]" box="[420,564,1622,1647]" pageId="10" pageNumber="11">
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<figureCitation id="136DB0AB366DCE351298EC5D3A62EA12" box="[420,517,1622,1647]" captionStart="FIGURE 8" captionStartId="11.[188,279,1460,1483]" captionTargetBox="[187,1435,167,1440]" captionTargetId="figure-114@11.[187,1435,167,1440]" captionTargetPageId="11" captionText="FIGURE 8. Necronectes cf. michelini Milne-Edwards, 1860 (A–D) and Necronectes michelini Milne-Edwards, 1860 (E, F): A. RGA/SMNH 5009, imprint of the outer side of chela; B. RGA/SMNH 5010, imprint of the inner side of chela; C. RGA/SMNH 5009, whitened latex cast; D. RGA/SMNH 5010, whitened latex cast; E. MNHN.F.R03775, (paralectotype), outer side of chela; F. MNHN.F.R03775, (paralectotype), inner side of chela. All scale bars equal 10 mm. Photos A and D: RECOLNAT (ANR-11-INBS-0004) - Jocelyn FALCONNET – 2017." figureDoi="http://doi.org/10.5281/zenodo.10962389" httpUri="https://zenodo.org/record/10962389/files/figure.png" pageId="10" pageNumber="11">Figure 8</figureCitation>
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A-D
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</paragraph>
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</subSubSection>
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<subSubSection id="C34CFFA5366DCE3513FCEC8A3957EB67" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8BE9AC2E366DCE3513FCEC8A3957EB67" blockId="10.[192,792,1664,1818]" pageId="10" pageNumber="11">
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<emphasis id="B922703C366DCE3513FCEC8A394BEAE7" bold="true" box="[192,300,1665,1690]" pageId="10" pageNumber="11">Material.</emphasis>
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A single “crusher” chela (
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<collectionCode id="ED4734EB366DCE3511BAEC8B3AA7EAE7" box="[646,704,1664,1690]" pageId="10" pageNumber="11">RGA</collectionCode>
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/
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<collectionCode id="ED4734EB366DCE3511F4EC8B3B7FEAE7" box="[712,792,1664,1690]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="10" pageNumber="11">SMNH</collectionCode>
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5009-5010) preserved as an imprint. Measurements: length of propodus =
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<quantity id="4CAE01CB366DCE35111AECCA3AF2EAA7" box="[550,661,1729,1754]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="10" pageNumber="11" unit="mm" value="53.6">53.6 mm</quantity>
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; maximum height of manus =
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<quantity id="4CAE01CB366DCE351296ECEA3A7EEA87" box="[426,537,1761,1786]" metricMagnitude="-2" metricUnit="m" metricValue="3.19" pageId="10" pageNumber="11" unit="mm" value="31.9">31.9 mm</quantity>
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; length of dactylus = 31.0 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34CFFA5366DCE351074EAAD3BF4EE22" pageId="10" pageNumber="11" type="description">
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<paragraph id="8BE9AC2E366DCE351074EAAD3BF4EE22" blockId="10.[840,1441,166,1816]" pageId="10" pageNumber="11">
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<emphasis id="B922703C366DCE351074EAAD3B85ECC2" bold="true" box="[840,994,166,191]" pageId="10" pageNumber="11">Description.</emphasis>
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Chela robust.
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<collectingRegion id="499262CC366DCE3517A3EAAD3C88ECC2" box="[1183,1263,166,191]" country="Papua New Guinea" name="Manus" pageId="10" pageNumber="11">Manus</collectingRegion>
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trapezoidal to suboval in outline; upper and lower margin arched and converging proximally; surface appears smooth, without indication of spines on distal margin at dactylus articulation. Fingers long and stout; as long as manus. Fixed finger robust; occlusal margin with four large molariform teeth, followed by several smaller conical teeth distally; first and second proximal teeth joined; fingertip curved upwards. Dactylus stout, curved forward; with rounded upper margin; strong proximal knob-like flattened molariform tooth followed by six rounded conical teeth of different sizes; strongly curved fingertip.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34CFFA5366DCE351074E8653D41EB6A" pageId="10" pageNumber="11" type="discussion">
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<paragraph id="8BE9AC2E366DCE351074E8653CADE99A" blockId="10.[840,1441,166,1816]" pageId="10" pageNumber="11">
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<emphasis id="B922703C366DCE351074E8653BA7EEFA" bold="true" box="[840,960,622,647]" pageId="10" pageNumber="11">Remarks.</emphasis>
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The original cuticle has been dissolved and the massive fingers are preserved as hollow imprints. Therefore, some details cannot be observed in the studied specimen. However, after preparing the latex cast, the preserved features of the chela are sufficient to assign the material to
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<taxonomicName id="4C56D7AD366DCE351074E9253D7DEF3A" authority="A. Milne-Edwards, 1881" authorityName="A. Milne-Edwards" authorityYear="1881" box="[840,1306,814,839]" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B922703C366DCE351074E9253BB9EF3A" box="[840,990,814,839]" italics="true" pageId="10" pageNumber="11">Necronectes</emphasis>
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A.
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<bibRefCitation id="EFC7D1DF366DCE351729E9253D7DEF3A" author="Milne-Edwards, A." box="[1045,1306,814,839]" pageId="10" pageNumber="11" refId="ref8139" refString="Milne-Edwards, A. 1881. Note sur quelques Crustaces fossiles des environs de Biarritz. Annales des Sciences geologiques, 11 (2): 1 - 8." year="1881">Milne-Edwards, 1881</bibRefCitation>
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</taxonomicName>
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. Observed important taxonomic characters include lack of distal spines on manus at position of dactylus articulation and rounded upper margin of manus, which are the key characters for an assignment to
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<taxonomicName id="4C56D7AD366DCE351074E9C53BB9EF9A" authorityName="A. Milne-Edwards" authorityYear="1881" box="[840,990,974,999]" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B922703C366DCE351074E9C53BB9EF9A" box="[840,990,974,999]" italics="true" pageId="10" pageNumber="11">Necronectes</emphasis>
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</taxonomicName>
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, and differentiate it from closely related
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<taxonomicName id="4C56D7AD366DCE351098E9E53CC9E87A" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[932,1198,1006,1032]" class="Malacostraca" family="Portunidae" genus="Scylla" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B922703C366DCE351098E9E53B8CE875" box="[932,1003,1006,1032]" italics="true" pageId="10" pageNumber="11">Scylla</emphasis>
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<bibRefCitation id="EFC7D1DF366DCE3510C9E9E53CC9E87A" author="de Haan, W." box="[1013,1198,1006,1031]" pageId="10" pageNumber="11" refId="ref6985" refString="de Haan, W. 1833 - 1850. Crustacea. In Von Siebold, P. F. (ed.), Fauna Japonica. Muller, Amsterdam, pp. xvii, xxxi, 244 pp." year="1833">De Haan, 1833</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation id="EFC7D1DF366DCE351783E9E53BB2E85A" author="Osso, A. & Gagnaison, C. & France" pageId="10" pageNumber="11" pagination="367 - 383" refId="ref8425" refString="Osso, A. and Gagnaison, C. 2019. An appraisal of the Middle-Late Miocene fossil decapod crustaceans of the ' Fa-luns' (Anjou-Touraine, France). Geodiversitas, 41 (9): 367 - 383. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 9" year="2019">Ossó and Gagnaison, 2019</bibRefCitation>
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). The herein presented specimen strongly resembles
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<taxonomicName id="4C56D7AD366DCE351701EE253C66E81A" authority="Milne-Edwards, 1860" authorityName="A. Milne-Edwards" authorityYear="1860" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="michelini">
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<emphasis id="B922703C366DCE351701EE253D2EE83A" box="[1085,1353,1070,1095]" italics="true" pageId="10" pageNumber="11">Necronectes michelini</emphasis>
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Milne-Edwards, 1860
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</taxonomicName>
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, from the Miocene of
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<collectingCountry id="F341ECBE366DCE351630EE453D38E81A" box="[1292,1375,1102,1127]" name="France" pageId="10" pageNumber="11">France</collectingCountry>
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(
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<figureCitation id="136DB0AB366DCE35164CEE453BEAE8FA" captionStart="FIGURE 8" captionStartId="11.[188,279,1460,1483]" captionTargetBox="[187,1435,167,1440]" captionTargetId="figure-114@11.[187,1435,167,1440]" captionTargetPageId="11" captionText="FIGURE 8. Necronectes cf. michelini Milne-Edwards, 1860 (A–D) and Necronectes michelini Milne-Edwards, 1860 (E, F): A. RGA/SMNH 5009, imprint of the outer side of chela; B. RGA/SMNH 5010, imprint of the inner side of chela; C. RGA/SMNH 5009, whitened latex cast; D. RGA/SMNH 5010, whitened latex cast; E. MNHN.F.R03775, (paralectotype), outer side of chela; F. MNHN.F.R03775, (paralectotype), inner side of chela. All scale bars equal 10 mm. Photos A and D: RECOLNAT (ANR-11-INBS-0004) - Jocelyn FALCONNET – 2017." figureDoi="http://doi.org/10.5281/zenodo.10962389" httpUri="https://zenodo.org/record/10962389/files/figure.png" pageId="10" pageNumber="11">Figure 8</figureCitation>
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E-F), somewhat differing in well-developed proximal molariform teeth on fixed finger and dactylus. Given similarities in preserved specimen, the stratigraphic span and geographic distribution of
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<taxonomicName id="4C56D7AD366DCE3516B9EEC53BD7E97A" authorityName="A. Milne-Edwards" authorityYear="1860" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="michelini">
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<emphasis id="B922703C366DCE3516B9EEC53BD7E97A" italics="true" pageId="10" pageNumber="11">N. michelini</emphasis>
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</taxonomicName>
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, it is also possible that the occurrence reported here, from the upper Oligocene of
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<collectingCountry id="F341ECBE366DCE351668EF053B08E93A" name="Slovenia" pageId="10" pageNumber="11">Slovenia</collectingCountry>
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, belongs to this species. Nevertheless, we refrain to assign it to
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<taxonomicName id="4C56D7AD366DCE35177EEF453CABE91A" authorityName="A. Milne-Edwards" authorityYear="1860" box="[1090,1228,1358,1383]" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="michelini">
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<emphasis id="B922703C366DCE35177EEF453CABE91A" box="[1090,1228,1358,1383]" italics="true" pageId="10" pageNumber="11">N. michelini</emphasis>
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</taxonomicName>
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because of insufficient preservation of diagnostic features on the manus and absence preserved carapace remains. Due to the fragmentary preservation we leave the specimen in open nomenclature.
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</paragraph>
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<paragraph id="8BE9AC2E366DCE351074EFFD3C87EA6D" blockId="10.[840,1441,166,1816]" box="[840,1248,1526,1552]" pageId="10" pageNumber="11">
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<emphasis id="B922703C366DCE351074EFFD3BC6EA72" bold="true" box="[840,929,1526,1551]" pageId="10" pageNumber="11">Range.</emphasis>
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Late Oligocene (Chattian).
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</paragraph>
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<paragraph id="8BE9AC2E366DCE351074EC153D41EB6A" blockId="10.[840,1441,166,1816]" pageId="10" pageNumber="11">
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||
<emphasis id="B922703C366DCE351074EC153B84EA45" bold="true" box="[840,995,1566,1592]" pageId="10" pageNumber="11">Occurrence.</emphasis>
|
||
<taxonomicName id="4C56D7AD366DCE3510D3EC153CE3EA4A" authorityName="A. Milne-Edwards" authorityYear="1881" box="[1007,1156,1566,1591]" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B922703C366DCE3510D3EC153CE3EA4A" box="[1007,1156,1566,1591]" italics="true" pageId="10" pageNumber="11">Necronectes</emphasis>
|
||
</taxonomicName>
|
||
is a well-known genus known from a number of Oligocene and Miocene occurrences (
|
||
<bibRefCitation id="EFC7D1DF366DCE3510D4EC553C98EA0A" author="Schweitzer, C. E. & Iturralde-Vinent, M. & Hetler, J. H. & Velez-Juarbe, J." box="[1000,1279,1630,1656]" pageId="10" pageNumber="11" pagination="111 - 136" refId="ref8672" refString="Schweitzer, C. E., Iturralde-Vinent, M., Hetler, J. H., and Velez-Juarbe, J. 2006. Oligocene and Miocene decapods (Thalassinidea and Brachyura) from the Caribbean. Annals of Carnegie Museum, 75 (2): 111 - 136. https: // doi. org / 10.2992 / 0097 - 4463 (2006) 75 [111: OAMDTA] 2.0. CO; 2" year="2006">Schweitzer et al., 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC7D1DF366DCE351637EC553BE0EAEA" author="Luque, J. & Schweitzer, C. E. & Santana, W. & Portell, R. W. & Vega, F. J. & Klompmaker, A. A. & Brachyura" pageId="10" pageNumber="11" pagination="1 - 85" refId="ref7930" refString="Luque, J., Schweitzer, C. E., Santana, W., Portell, R. W., Vega, F. J., and Klompmaker, A. A. 2017. Checklist of fossil decapod crustaceans from tropical America. part I: Anomura and Brachyura. Nauplius, 25 (0): 1 - 85. https: // doi. org / 10.1590 / 2358 - 2936 e 2017025" year="2017">Luque et al., 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC7D1DF366DCE3510AEEC753CC5EAEA" author="Hyzny, M. & Dulai, A." box="[914,1186,1662,1688]" pageId="10" pageNumber="11" refId="ref7300" refString="Hyzny, M. and Dulai, A. 2021. Badenian Decapods of Hungary. GeoLitera, Institute of Geosciences, University of Szeged, Hungary, 291 pp." year="2021">Hyžný and Dulai, 2021</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C56D7AD366DCE351788EC753CD7EACA" authority="A. Milne-Edwards, 1860" authorityName="A. Milne-Edwards" authorityYear="1860" class="Malacostraca" family="Portunidae" genus="Necronectes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="michelini">
|
||
<emphasis id="B922703C366DCE351788EC753B0EEACA" italics="true" pageId="10" pageNumber="11">Necronectes michelini</emphasis>
|
||
A. Milne-Edwards, 1860
|
||
</taxonomicName>
|
||
, has so far been reported from the middle Miocene of
|
||
<collectingCountry id="F341ECBE366DCE3517C2ECB53D36EAAA" box="[1278,1361,1726,1751]" name="France" pageId="10" pageNumber="11">France</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC7D1DF366DCE351663ECB53C29EA8A" author="Osso, A. & Gagnaison, C. & France" pageId="10" pageNumber="11" pagination="367 - 383" refId="ref8425" refString="Osso, A. and Gagnaison, C. 2019. An appraisal of the Middle-Late Miocene fossil decapod crustaceans of the ' Fa-luns' (Anjou-Touraine, France). Geodiversitas, 41 (9): 367 - 383. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 9" year="2019">Ossó and Gagnaison, 2019</bibRefCitation>
|
||
) and the middle Miocene of
|
||
<collectingCountry id="F341ECBE366DCE351074ECF53BCAEB6A" box="[840,941,1790,1815]" name="Hungary" pageId="10" pageNumber="11">Hungary</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC7D1DF366DCE351086ECF53D7CEB6A" author="Lorenthey, E. & Beurlen, K." box="[954,1307,1790,1816]" pageId="10" pageNumber="11" refId="ref7842" refString="Lorenthey, E. and Beurlen, K. 1929. Die fossilen Decapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica, 3: 1 - 420." year="1929">Lőrenthey and Beurlen, 1929</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |