659 lines
81 KiB
XML
659 lines
81 KiB
XML
<document id="C1EA22EF922701AE07E4CF4FC6594933" ID-CLB-Dataset="8721" ID-DOI="10.11646/zootaxa.4823.1.1" ID-GBIF-Dataset="a43d0118-9df5-4970-a316-0dae5c9caebf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401916" ID-ZooBank="3DF06F26-FD35-452E-9056-1D05B49946EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609340622775" checkinUser="plazi" docAuthor="Dombrow, Holger E. & Colville, Jonathan F." docDate="2020" docId="03DFBE05E023FFB7FF64D0E9FD3BFC28" docLanguage="en" docName="zootaxa.4823.1.1.pdf" docOrigin="Zootaxa 4823" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Beckhoplia occidentalis Dombrow 2005" docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="FFE6C67DE01CFFF5FFF3D41BFFBDFF8D" masterDocTitle="Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="64" updateTime="1698849823920" updateUser="ExternalLinkService">
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<mods:title id="BB165174BE6FCF3B967331511A994240">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
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<mods:namePart id="49B20590817BE3FEF6D971CAF7BDE68A">Dombrow, Holger E.</mods:namePart>
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<mods:namePart id="A34328DAC82EA2E1DFE693979CE8ECE3">Colville, Jonathan F.</mods:namePart>
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<mods:nameIdentifier id="DA56E4B668C34A6EC0EF23139F7C00A3" type="email">jonathan.colville@gmail.com</mods:nameIdentifier>
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<mods:title id="C350D9296E9754C269B4446F15CE116B">Zootaxa</mods:title>
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<mods:date id="6A53CB9299E6C55B57D672451AF550DF">2020</mods:date>
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<mods:number id="86BBBF4C8E2BC78F4A273E3AC576FF5E">2020-08-10</mods:number>
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<treatment id="03DFBE05E023FFB7FF64D0E9FD3BFC28" ID-DOI="http://doi.org/10.5281/zenodo.4455307" ID-GBIF-Taxon="177013526" ID-Zenodo-Dep="4455307" LSID="urn:lsid:plazi:treatment:03DFBE05E023FFB7FF64D0E9FD3BFC28" httpUri="http://treatment.plazi.org/id/03DFBE05E023FFB7FF64D0E9FD3BFC28" lastPageId="66" lastPageNumber="67" pageId="63" pageNumber="64">
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<taxonomicName id="4C767490E023FFCAFF64D0E9FD3EFA81" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[151,643,1266,1293]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="occidentalis">
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<emphasis id="B902D301E023FFCAFF64D0E9FD3EFA81" bold="true" box="[151,643,1266,1293]" pageId="63" pageNumber="64">
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<emphasis id="B902D301E023FFCAFF64D0E9FE0BFA80" bold="true" box="[151,438,1266,1293]" italics="true" pageId="63" pageNumber="64">Beckhoplia occidentalis</emphasis>
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<bibRefCitation id="EFE772E2E023FFCAFE4DD0E9FD3EFA81" author="Dombrow, H." box="[446,643,1266,1292]" pageId="63" pageNumber="64" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
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<paragraph id="8BC90F13E023FFCAFF64D10DFE63FABC" blockId="63.[151,858,1266,1364]" box="[151,478,1302,1329]" pageId="63" pageNumber="64">
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(
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<figureCitation id="134D1396E023FFCAFF6CD10DFF40FABD" box="[159,253,1302,1328]" captionStart="FIGURE 31" captionStartId="17.[151,248,683,707]" captionTargetBox="[426,1161,185,655]" captionTargetId="figure-17@17.[422,1165,181,659]" captionTargetPageId="17" captionText="FIGURE 31. General drawing of the outline of Beckhoplia Dombrow, 2005 species, ♂♂. a, ventrites; b, elytra; c, metathoracic pleurite I; d, metathoracic pleurite II; e, mesothoracic pleurite; f, pronotum; g, prothoracic thorax; h, mesothoracic thorax; j, mesothoracic coxa; k, mesothoracic trochanter; l, metathoracic thorax; m, metathoracic coxa; n, metathoracic trochanter. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401936" httpUri="https://zenodo.org/record/4401936/files/figure.png" pageId="63" pageNumber="64">Figs. 31</figureCitation>
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,
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<figureCitation id="134D1396E023FFCAFEFAD10DFE9AFABD" box="[265,295,1302,1328]" captionStart="FIGURES 39–40" captionStartId="23.[151,264,749,773]" captionTargetBox="[432,1159,536,689]" captionTargetId="figure-79@23.[423,1164,532,700]" captionTargetPageId="23" captionText="FIGURES 39–40. General drawings of the tip of pygidium of Beckhoplia Dombrow, 2005 species in posterior view ♂♂. 39, tip linear; 40, tip rounded. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401942" httpUri="https://zenodo.org/record/4401942/files/figure.png" pageId="63" pageNumber="64">39</figureCitation>
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,
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<figureCitation id="134D1396E023FFCAFEC0D10DFEECFABC" box="[307,337,1302,1329]" captionStart="FIGURES 55–58" captionStartId="24.[151,264,1387,1411]" captionTargetBox="[155,1432,874,1323]" captionTargetId="figure-95@24.[151,1436,870,1327]" captionTargetPageId="24" captionText="FIGURES 55–58. General drawing of the types of mesothoracic pretarsomere of Beckhoplia Dombrow, 2005 species, in lateral view showing the distinctly thickened, roundish cutting missing ventrally, ♂♂, otherwise indicated. 55, with a shortened smooth ridge, with roundish cutting; 56, with a shortened granulate ridge; 57, slender, with a full undulate ridge, ♀♀; 58, with a shortened smooth ridge. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401950" httpUri="https://zenodo.org/record/4401950/files/figure.png" pageId="63" pageNumber="64">58</figureCitation>
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,
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<figureCitation id="134D1396E023FFCAFEAED10DFE1DFABD" box="[349,416,1302,1328]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="63" pageNumber="64">92-93</figureCitation>
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<figureCitation id="134D1396E023FFCAFE5FD10DFE6BFABC" box="[428,470,1302,1329]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="63" pageNumber="64">115</figureCitation>
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)
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<emphasis id="B902D301E023FFCAFF64D19BFE34FA15" box="[151,393,1408,1432]" italics="true" pageId="63" pageNumber="64">Beckhoplia occidentalis</emphasis>
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<bibRefCitation id="EFE772E2E023FFCAFE63D19BFDD6FA14" author="Dombrow, H." box="[400,619,1408,1433]" pageId="63" pageNumber="64" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005: 127</bibRefCitation>
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.
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locality.
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[
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,
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<collectionCode id="ED6797D6E023FFCAFE79D1DDFE7AFA6C" box="[394,455,1478,1505]" country="United Kingdom" name="National Collection of Passiflora" pageId="63" pageNumber="64" type="Herbarium">NCP</collectionCode>
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]: Nieuwoudtville
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E
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.
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<emphasis id="B902D301E023FFCAFF64D214FEF8F9A5" bold="true" box="[151,325,1550,1577]" pageId="63" pageNumber="64">Type material.</emphasis>
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[1]
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<typeStatus id="54CDB1B1E023FFCAFE81D214FE62F9A4" box="[370,479,1551,1577]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="9D70C49AE023FFCAFE15D214FE43F9A4" box="[486,510,1551,1577]" pageId="63" pageNumber="64" type="male">♂</specimenCount>
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: white label pr «
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<collectingCountry id="F3614F83E023FFCAFD46D214FCC3F9A4" box="[693,894,1551,1577]" name="South Africa" pageId="63" pageNumber="64">SOUTH AFRICA</collectingCountry>
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<collectingRegion id="49B2C1F1E023FFCAFC62D214FB84F9A4" box="[913,1081,1551,1577]" country="South Africa" name="Northern Cape" pageId="63" pageNumber="64">Northern Cape</collectingRegion>
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, Hantam | Nieuwoudtville
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<quantity id="4C8EA2F6E023FFCAFA97D215FA21F9A4" box="[1380,1436,1550,1577]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="64" unit="km" value="5.0">5 km</quantity>
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S at | 31.22.37S–19.09.13E» bs pr «Time 14:30, 17. Sept. 2001 | alt.
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<quantity id="4C8EA2F6E023FFCAFC66D229FC60F9C0" box="[917,989,1586,1613]" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="63" pageNumber="64" unit="m" value="752.0">752 m</quantity>
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,
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<taxonomicName id="4C767490E023FFCAFC14D228FB20F9C0" box="[999,1181,1587,1613]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="family">ASTERACEAE</taxonomicName>
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<taxonomicName id="4C767490E023FFCAFB42D228FADDF9C0" box="[1201,1376,1587,1613]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="acaulis">
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<emphasis id="B902D301E023FFCAFB42D228FADDF9C0" box="[1201,1376,1587,1613]" italics="true" pageId="63" pageNumber="64">Arctotis acaulis</emphasis>
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</taxonomicName>
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| leg. Holger Dombrow # 339» || blue label pr «Bioregion: | Shale Renosterveld | Vegetation Type: FRs2 – | Nieuwoudtville Shale Renosterveld» || yellow label hw «18» pr «.721 | coll. Dombrow» || red label pr «
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<typeStatus id="54CDB1B1E023FFCAFB53D260FAB7F918" box="[1184,1290,1659,1685]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
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<taxonomicName id="4C767490E023FFCAFAEDD260FA21F918" authorityName="Dombrow" authorityYear="2005" box="[1310,1436,1659,1685]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E023FFCAFAEDD260FA21F918" box="[1310,1436,1659,1685]" italics="true" pageId="63" pageNumber="64">Beckhoplia</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C767490E023FFCAFF57D284FE96F934" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[164,299,1695,1721]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
|
||
<emphasis id="B902D301E023FFCAFF57D284FE96F934" box="[164,299,1695,1721]" italics="true" pageId="63" pageNumber="64">occidentalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE023FFCAFEC1D284FEDAF934" box="[306,359,1695,1721]" pageId="63" pageNumber="64" rank="species">sp.n.</taxonomicNameLabel>
|
||
| det.: H. Dombrow 2003» (
|
||
<collectionCode id="ED6797D6E023FFCAFD51D284FD5CF934" box="[674,737,1695,1721]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="63" pageNumber="64" type="Museum">ZSM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E023FFCAFF64D2FDFEF2F8C4" blockId="63.[151,1436,1766,1865]" pageId="63" pageNumber="64">
|
||
<emphasis id="B902D301E023FFCAFF64D2FDFDB5F88D" bold="true" box="[151,520,1766,1792]" pageId="63" pageNumber="64">Additional material examined.</emphasis>
|
||
<materialsCitation id="3B1E054EE023FFCAFDE2D2FCFCC3F8A8" ID-GBIF-Occurrence="3022772329" collectingDate="2000-07-09" collectorName="Ingrid Nanni" country="South Africa" latitude="-31.24" location="Northern Cape" longLatPrecision="731" longitude="19.08" pageId="63" pageNumber="64" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="Northern Cape">
|
||
<specimenCount id="9D70C49AE023FFCAFDE2D2FCFDE9F88C" box="[529,596,1767,1793]" pageId="63" pageNumber="64" type="male">7 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D70C49AE023FFCAFD91D2FDFD18F88C" box="[610,677,1766,1793]" pageId="63" pageNumber="64" type="female">6 ♀♀</specimenCount>
|
||
: white label pr «
|
||
<collectingCountry id="F3614F83E023FFCAFC95D2FCFB34F88C" box="[870,1161,1767,1793]" name="South Africa" pageId="63" pageNumber="64">Republic of South Africa</collectingCountry>
|
||
, |
|
||
<collectingRegion id="49B2C1F1E023FFCAFB56D2FCFAF2F88C" box="[1189,1359,1767,1793]" country="South Africa" name="Northern Cape" pageId="63" pageNumber="64">Northern Cape</collectingRegion>
|
||
, Nieuwoudtville, |
|
||
<geoCoordinate id="EE4269D4E023FFCAFEDDD310FE30F8A8" box="[302,397,1803,1829]" degrees="31.24" direction="south" orientation="latitude" pageId="63" pageNumber="64" precision="555" value="-31.24">31.24S–</geoCoordinate>
|
||
<geoCoordinate id="EE4269D4E023FFCAFE7ED310FE63F8A8" box="[397,478,1803,1829]" degrees="19.08" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="555" value="19.08">19.08E</geoCoordinate>
|
||
, 04.–
|
||
<date id="FFC829D3E023FFCAFDEAD310FD20F8A8" box="[537,669,1803,1829]" pageId="63" pageNumber="64" value="2000-07-09">
|
||
<collectingDate id="EF8CD03BE023FFCAFDEAD310FD20F8A8" box="[537,669,1803,1829]" pageId="63" pageNumber="64" value="2000-07-09">07.09.2000</collectingDate>
|
||
</date>
|
||
, | leg.
|
||
<collectorName id="26836AC5E023FFCAFD18D310FCC3F8A8" box="[747,894,1803,1829]" pageId="63" pageNumber="64">Ingrid Nänni</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1E054EE023FFCAFC8CD310FEF6F8C4" ID-GBIF-Occurrence="3022772324" collectionCode="ESDW" collectorName="Dombrow" country="South Africa" pageId="63" pageNumber="64" specimenCount="1" stateProvince="Northern Cape">
|
||
» || yellow label hw «55» pr «.721 | coll.
|
||
<collectorName id="26836AC5E023FFCAFAA8D310FF6EF8C4" pageId="63" pageNumber="64">Dombrow</collectorName>
|
||
» (
|
||
<collectionCode id="ED6797D6E023FFCAFF1DD334FEF9F8C4" box="[238,324,1839,1865]" pageId="63" pageNumber="64">ESDW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E023FFB5FF64D36DFEAAFF58" blockId="63.[151,1436,1910,2009]" lastBlockId="64.[151,1437,150,213]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
|
||
<emphasis id="B902D301E023FFCAFF64D36DFEE3F81C" bold="true" box="[151,350,1910,1937]" pageId="63" pageNumber="64">Taxonomic note.</emphasis>
|
||
The following specimens from the original
|
||
<typeStatus id="54CDB1B1E023FFCAFCA8D36CFC36F81C" box="[859,907,1911,1937]" pageId="63" pageNumber="64">type</typeStatus>
|
||
series are transferred to several other species, because of the distinct morphological differences see descriptions of the target species: The
|
||
<specimenCount id="9D70C49AE023FFCAFB6FD381FB66F838" box="[1180,1243,1946,1973]" pageId="63" pageNumber="64" type="female">6 ♀♀</specimenCount>
|
||
from the locality Botterkloof Pass are transferred to
|
||
<taxonomicName id="4C767490E023FFCAFDD1D3A4FD0BF854" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[546,694,1983,2009]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="caliginosa" status="sp. nov.">
|
||
<emphasis id="B902D301E023FFCAFDD1D3A4FD0BF854" box="[546,694,1983,2009]" italics="true" pageId="63" pageNumber="64">B. caliginosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE023FFCAFD4DD3A4FD48F854" box="[702,757,1983,2009]" pageId="63" pageNumber="64" rank="species">sp.n.</taxonomicNameLabel>
|
||
; the
|
||
<specimenCount id="9D70C49AE023FFCAFCDFD3A4FCD1F854" box="[812,876,1983,2009]" pageId="63" pageNumber="64" type="male">3 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D70C49AE023FFCAFC50D3A4FC4CF854" box="[931,1009,1983,2009]" pageId="63" pageNumber="64" type="female">10 ♀♀</specimenCount>
|
||
from the locality Farm Nuwedam are transferred to
|
||
<taxonomicName id="4C767490E05CFFB5FECBD48CFE12FF3C" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[312,431,151,177]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="pumilla" status="sp. nov.">
|
||
<emphasis id="B902D301E05CFFB5FECBD48CFE12FF3C" box="[312,431,151,177]" italics="true" pageId="64" pageNumber="65">B. pumilla</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE05CFFB5FE4BD48CFE50FF3C" box="[440,493,151,177]" pageId="64" pageNumber="65" rank="species">sp.n.</taxonomicNameLabel>
|
||
The locality has to be adjusted because the coordinate is located
|
||
<quantity id="4C8EA2F6E05CFFB5FB2DD48DFAA4FF3C" box="[1246,1305,150,177]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="64" pageNumber="65" unit="km" value="5.0">5 km</quantity>
|
||
E of Nieuwoudtville.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05CFFB5FF64D519FB7EFE90" blockId="64.[151,1219,258,285]" box="[151,1219,258,285]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF64D519FE1FFE90" bold="true" box="[151,418,258,285]" pageId="64" pageNumber="65">
|
||
Condition of
|
||
<typeStatus id="54CDB1B1E05CFFB5FEC7D518FE20FE90" box="[308,413,259,285]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Very good: Glued on a card. Dissected, parameres glued behind beetle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E05CFFB5FF64D551FC6BFDB1" pageId="64" pageNumber="65" type="diagnosis">
|
||
<paragraph id="8BC90F13E05CFFB5FF64D551FC6BFDB1" blockId="64.[151,1437,330,572]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF64D551FE1CFEE9" bold="true" box="[151,417,330,356]" pageId="64" pageNumber="65">Differential diagnosis.</emphasis>
|
||
Differs from
|
||
<taxonomicName id="4C767490E05CFFB5FDB0D550FCD9FEE8" authority=": Elytra" authorityName="Elytra" box="[579,868,331,357]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="dolichiocnemis">
|
||
<emphasis id="B902D301E05CFFB5FDB0D550FCB2FEE8" box="[579,783,331,357]" italics="true" pageId="64" pageNumber="65">B. dolichiocnemis</emphasis>
|
||
: Elytra
|
||
</taxonomicName>
|
||
of
|
||
<taxonomicName id="4C767490E05CFFB5FC63D550FB84FEE8" authorityName="Dombrow" authorityYear="2020" box="[912,1081,331,357]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B902D301E05CFFB5FC63D550FB84FEE8" box="[912,1081,331,357]" italics="true" pageId="64" pageNumber="65">B. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
without a longitudinal depression along suture. Elytra of
|
||
<taxonomicName id="4C767490E05CFFB5FE3ED574FD2BFE04" box="[461,662,367,393]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="dolichiocnemis">
|
||
<emphasis id="B902D301E05CFFB5FE3ED574FD2BFE04" box="[461,662,367,393]" italics="true" pageId="64" pageNumber="65">B. dolichiocnemis</emphasis>
|
||
</taxonomicName>
|
||
with a longitudinal depression along suture. Elytra of
|
||
<taxonomicName id="4C767490E05CFFB5FB05D574FA21FE04" authorityName="Dombrow" authorityYear="2020" box="[1270,1436,367,393]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B902D301E05CFFB5FB05D574FA21FE04" box="[1270,1436,367,393]" italics="true" pageId="64" pageNumber="65">B. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
partly covering propygidium. Elytra of
|
||
<taxonomicName id="4C767490E05CFFB5FDA0D588FCA1FE20" box="[595,796,403,429]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="dolichiocnemis">
|
||
<emphasis id="B902D301E05CFFB5FDA0D588FCA1FE20" box="[595,796,403,429]" italics="true" pageId="64" pageNumber="65">B. dolichiocnemis</emphasis>
|
||
</taxonomicName>
|
||
protruding the propygidium. Interno-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E05CFFB5FEC3D5ACFE6AFE5D" authorityName="Dombrow" authorityYear="2020" box="[304,471,438,464]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B902D301E05CFFB5FEC3D5ACFE6AFE5D" box="[304,471,438,464]" italics="true" pageId="64" pageNumber="65">B. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
curly bracket-shaped in dorsal view (
|
||
<figureCitation id="134D1396E05CFFB5FC8DD5ADFC60FE5C" box="[894,989,438,465]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="64" pageNumber="65">Figs. 92</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E05CFFB5FC1BD5ADFBA8FE5C" box="[1000,1045,438,465]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="64" pageNumber="65">115</figureCitation>
|
||
). Interno-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E05CFFB5FEF3D5C0FE74FE79" box="[256,457,474,500]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="dolichiocnemis">
|
||
<emphasis id="B902D301E05CFFB5FEF3D5C0FE74FE79" box="[256,457,474,500]" italics="true" pageId="64" pageNumber="65">B. dolichiocnemis</emphasis>
|
||
</taxonomicName>
|
||
parabola-shaped in dorsal view (
|
||
<figureCitation id="134D1396E05CFFB5FCCED5C1FC26FE79" box="[829,923,474,500]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="64" pageNumber="65">Figs. 96</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E05CFFB5FC54D5C1FC69FE79" box="[935,980,474,500]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="64" pageNumber="65">117</figureCitation>
|
||
). Externo-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E05CFFB5FF45D5E4FEE3FD95" authorityName="Dombrow" authorityYear="2020" box="[182,350,510,536]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B902D301E05CFFB5FF45D5E4FEE3FD95" box="[182,350,510,536]" italics="true" pageId="64" pageNumber="65">B. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
curly bracket-shaped, convex medially in dorsal view (
|
||
<figureCitation id="134D1396E05CFFB5FC20D5E5FB8EFD95" box="[979,1075,510,536]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="64" pageNumber="65">Figs. 92</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E05CFFB5FBCCD5E5FBD1FD94" box="[1087,1132,510,537]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="64" pageNumber="65">115</figureCitation>
|
||
). Externo-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E05CFFB5FEBDD639FDAAFDB1" box="[334,535,546,572]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="dolichiocnemis">
|
||
<emphasis id="B902D301E05CFFB5FEBDD639FDAAFDB1" box="[334,535,546,572]" italics="true" pageId="64" pageNumber="65">B. dolichiocnemis</emphasis>
|
||
</taxonomicName>
|
||
undulate in dorsal view (
|
||
<figureCitation id="134D1396E05CFFB5FCC0D639FC2DFDB6" box="[819,912,545,572]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="64" pageNumber="65">Figs. 96</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E05CFFB5FC6FD639FC74FDB1" box="[924,969,546,572]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="64" pageNumber="65">117</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E05CFFB7FF64D672FCADFD41" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65" type="description">
|
||
<paragraph id="8BC90F13E05CFFB5FF64D672FE80FD0E" blockId="64.[151,1437,617,2038]" box="[151,317,617,643]" pageId="64" pageNumber="65">
|
||
<heading id="D081B87FE05CFFB5FF64D672FE80FD0E" box="[151,317,617,643]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF64D672FE80FD0E" bold="true" box="[151,317,617,643]" pageId="64" pageNumber="65">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05CFFB5FF34D696FC83FD2A" blockId="64.[151,1437,617,2038]" box="[199,830,653,680]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF34D696FE3DFD2A" bold="true" box="[199,384,653,679]" pageId="64" pageNumber="65">Measurements:</emphasis>
|
||
length
|
||
<quantity id="4C8EA2F6E05CFFB5FE27D696FDE3FD2A" box="[468,606,653,679]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="8.8" metricValueMin="7.6" pageId="64" pageNumber="65" unit="mm" value="8.2" valueMax="8.8" valueMin="7.6">7.6–8.8 mm</quantity>
|
||
; width 4.0–
|
||
<quantity id="4C8EA2F6E05CFFB5FD11D696FC87FD2A" box="[738,826,653,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="64" pageNumber="65" unit="mm" value="4.5">4.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05CFFB5FF34D6AAFABBFD47" blockId="64.[151,1437,617,2038]" box="[199,1286,688,715]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF34D6AAFEE6FD46" bold="true" box="[199,347,689,715]" pageId="64" pageNumber="65">Integument:</emphasis>
|
||
Black; funiculum brown annulated; elytra transparently brown, margins blackened.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05CFFB5FF34D6CFFAFEF977" blockId="64.[151,1437,617,2038]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF34D6CFFEFAFD63" bold="true" box="[199,327,724,750]" pageId="64" pageNumber="65">Tegument:</emphasis>
|
||
<emphasis id="B902D301E05CFFB5FEBCD6CFFE14FD63" box="[335,425,724,750]" italics="true" pageId="64" pageNumber="65">Clypeus</emphasis>
|
||
bare; inner side of the reflexed anterior and lateral margins loose setose with short, erect, black setae.
|
||
<emphasis id="B902D301E05CFFB5FED0D6E3FEDDFC9F" box="[291,352,760,786]" italics="true" pageId="64" pageNumber="65">Head</emphasis>
|
||
loose setose with long, erect, black setae.
|
||
<emphasis id="B902D301E05CFFB5FCB1D6E2FC76FC9F" box="[834,971,760,786]" italics="true" pageId="64" pageNumber="65">Eye canthus</emphasis>
|
||
with a loose setose ciliary of long, black setae.
|
||
<emphasis id="B902D301E05CFFB5FF2ED707FE91FCBB" box="[221,300,796,822]" italics="true" pageId="64" pageNumber="65">Scapus</emphasis>
|
||
loose setose with long, black setae, whitish ventrally, sparse and medium-long on
|
||
<emphasis id="B902D301E05CFFB5FB3FD707FAFAFCBB" box="[1228,1351,796,822]" italics="true" pageId="64" pageNumber="65">pedicellus;</emphasis>
|
||
funicle I–II bare,
|
||
<emphasis id="B902D301E05CFFB5FEF4D724FE28FCD4" box="[263,405,831,857]" italics="true" pageId="64" pageNumber="65">funicle II–IV</emphasis>
|
||
with few very short and short, erect, brown setae;
|
||
<emphasis id="B902D301E05CFFB5FC3DD724FBABFCD4" box="[974,1046,831,857]" italics="true" pageId="64" pageNumber="65">clavus</emphasis>
|
||
sparse setose with short, reclinate, black and very short, semi-erect, pale-brown setae.
|
||
<emphasis id="B902D301E05CFFB5FD1ED77FFCE1FCF0" box="[749,860,868,893]" italics="true" pageId="64" pageNumber="65">Pronotum</emphasis>
|
||
loose setose with long, erect, black setae, shorter towards posterior margin; with a loose setose ciliary of distinctly long, black setae on lateral margin.
|
||
<emphasis id="B902D301E05CFFB5FADDD79CFA21FC2C" box="[1326,1436,903,929]" italics="true" pageId="64" pageNumber="65">Scutellum</emphasis>
|
||
loose setose with short and medium-long, semi-erect, brown setae.
|
||
<emphasis id="B902D301E05CFFB5FC77D7B0FC77FC48" box="[900,970,939,965]" italics="true" pageId="64" pageNumber="65">Elytra</emphasis>
|
||
without pattern. Sparse setose with short, reclinate and semi-erect, black setae; with two loose, longitudinal rows of medium-long, semi-erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus; a juxta-humeral row from anterior margin between humerus and scutellar torus towards apical torus. Suture with a loose row of short and long, robust, erect, black setae.
|
||
<emphasis id="B902D301E05CFFB5FE4CD021FDADFBD9" box="[447,528,1082,1108]" italics="true" pageId="64" pageNumber="65">Thorax</emphasis>
|
||
and
|
||
<emphasis id="B902D301E05CFFB5FDBAD021FD16FBD9" box="[585,683,1082,1108]" italics="true" pageId="64" pageNumber="65">pleurites</emphasis>
|
||
loose pilose with long, black hairs.
|
||
<emphasis id="B902D301E05CFFB5FBBAD021FB6CFBDE" box="[1097,1233,1082,1107]" italics="true" pageId="64" pageNumber="65">Ventrite I–V</emphasis>
|
||
loose setose with short, reclinate, black setae; medium-long and long, semi-erect, black setae interspersed.
|
||
<emphasis id="B902D301E05CFFB5FB8DD046FAB2FBFA" box="[1150,1295,1117,1143]" italics="true" pageId="64" pageNumber="65">Propygidium</emphasis>
|
||
loose setose with long, reclinate, black setae, erect on
|
||
<emphasis id="B902D301E05CFFB5FD86D09AFD60FB16" box="[629,733,1153,1179]" italics="true" pageId="64" pageNumber="65">pygidium</emphasis>
|
||
.
|
||
<emphasis id="B902D301E05CFFB5FD1ED099FC1DFB16" box="[749,928,1153,1179]" italics="true" pageId="64" pageNumber="65">Tip of pygidium</emphasis>
|
||
with a sparse setose ciliary of medium-long and long, black setae.
|
||
<emphasis id="B902D301E05CFFB5FE7DD0BEFDE1FB32" box="[398,604,1189,1215]" italics="true" pageId="64" pageNumber="65">Prothoracic femur</emphasis>
|
||
loose pilose with long, black hairs; bare internally.
|
||
<emphasis id="B902D301E05CFFB5FB68D0BEFAE7FB32" box="[1179,1370,1189,1215]" italics="true" pageId="64" pageNumber="65">Prothoracic tibia</emphasis>
|
||
loose setose with short, reclinate and semi-erect, black setae; erect and medium-long ventrally; short, semi-erect, white setae interspersed dorsally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; with a loose setose ciliary of medium-long, black setae on internal ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth.
|
||
<emphasis id="B902D301E05CFFB5FD72D14CFCD9FAFF" box="[641,868,1367,1394]" italics="true" pageId="64" pageNumber="65">Mesothoracic femur</emphasis>
|
||
loose pilose with long, black hairs; distinctly long ventrally; bare internally.
|
||
<emphasis id="B902D301E05CFFB5FE32D160FD24FA18" box="[449,665,1403,1429]" italics="true" pageId="64" pageNumber="65">Mesothoracic tibia</emphasis>
|
||
loose setose with short and medium-long, reclinate, whitish setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of short, black bristles; lengthening dorsally; missing internally.
|
||
<emphasis id="B902D301E05CFFB5FB4FD1D8FA21FA50" box="[1212,1436,1475,1501]" italics="true" pageId="64" pageNumber="65">Metathoracic femur</emphasis>
|
||
loose setose with short and long, semi-erect, black, setae; bare internally.
|
||
<emphasis id="B902D301E05CFFB5FC36D1FDFB28F98D" box="[965,1173,1510,1536]" italics="true" pageId="64" pageNumber="65">Metathoracic tibia</emphasis>
|
||
sparse setose with very short and short, reclinate, brown setae; medium-long, semi-erect, robust, black setae interspersed; denser, long and finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally.
|
||
<emphasis id="B902D301E05CFFB5FD5AD249FC5BF9E6" box="[681,998,1618,1644]" italics="true" pageId="64" pageNumber="65">Prothoracic tarsomere I–IV</emphasis>
|
||
bare.
|
||
<emphasis id="B902D301E05CFFB5FBC2D249FB76F9E1" box="[1073,1227,1618,1644]" italics="true" pageId="64" pageNumber="65">Mesothoracic</emphasis>
|
||
and
|
||
<emphasis id="B902D301E05CFFB5FAFBD249FEFCF902" italics="true" pageId="64" pageNumber="65">metathoracic tarsomere I–IV</emphasis>
|
||
sparse setose with very short, reclinate, pale-brown setae.
|
||
<emphasis id="B902D301E05CFFB5FC29D26DFB34F902" box="[986,1161,1654,1679]" italics="true" pageId="64" pageNumber="65">Tarsomere I–IV</emphasis>
|
||
of all thoracic legs with a setaceous, apical, marginal ciliary of short, dark red-brown setae, lengthening towards
|
||
<emphasis id="B902D301E05CFFB5FB8DD282FA28F93E" box="[1150,1429,1689,1715]" italics="true" pageId="64" pageNumber="65">metathoracic tarsomeres</emphasis>
|
||
, fine dorsally and more robust ventrally.
|
||
<emphasis id="B902D301E05CFFB5FD92D2A5FD44F95A" box="[609,761,1726,1751]" italics="true" pageId="64" pageNumber="65">Pretarsomere</emphasis>
|
||
of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; few, long, semi-erect, black setae at apical margin, shorter and dark red-brown ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05CFFB5FF34D31FFD2CF8EB" blockId="64.[151,1437,617,2038]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF34D31FFEADF893" bold="true" box="[199,272,1796,1822]" pageId="64" pageNumber="65">Head:</emphasis>
|
||
Suture between head and clypeus linear, crenated. Head dense punctate with small, shallow, roundish, not merging punctures. Funicle I–II cylindrical; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05CFFB5FF34D36BFD20F87B" blockId="64.[151,1437,617,2038]" pageId="64" pageNumber="65">
|
||
<emphasis id="B902D301E05CFFB5FF34D36BFEF4F807" bold="true" box="[199,329,1904,1930]" pageId="64" pageNumber="65">Pronotum:</emphasis>
|
||
Widest medially; dense punctate with shallow, very small and small, roundish punctures, denser within medial furrow.Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly in anterior quarter and slightly convex attenuate posteriorly. With an indistinct, shortened, shallow medial furrow from posterior margin terminating on disk. Posterior corner rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D48DFBB8FF59" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D48DFEF5FF3D" bold="true" box="[199,328,150,176]" pageId="65" pageNumber="66">Scutellum:</emphasis>
|
||
Parabola-shaped; slightly longer than broad; dense punctate with shallow, small to large, roundish punctures, micropunctate within. Lateral margin slightly convex. Tip rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D4C5FB17FE20" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D4C5FEA0FF75" bold="true" box="[199,285,222,248]" pageId="65" pageNumber="66">Elytra:</emphasis>
|
||
Ovate; longer than broad; widest in anterior third; covering propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate along scutellum and suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; with two sparse, longitudinal rows of small, deep, roundish punctures: a juxta-sutural row from anterior margin crossing scutellar torus to apical torus; a supra-humeral row from anterior margin between humerus and scutellar torus to apical torus.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D5ADFB16FD95" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D5ADFEF8FE5D" bold="true" box="[199,325,438,464]" pageId="65" pageNumber="66">Abdomen:</emphasis>
|
||
Ventrite I distinctly shortened medially; ventrite V slightly constricted ventrally. Propygidium with an indistinct stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin moderately convex; lateral margin subparallel in upper fifth, linearly attenuate apically; tip linear.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D639FBACFD7D" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D639FE3BFDB1" bold="true" box="[199,390,546,572]" pageId="65" pageNumber="66">Prothoracic leg:</emphasis>
|
||
Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between apical and median tooth double in width than between median and basal tooth. Shape of interspace between teeth concave. Tarsomere I–IV combined extremely short; equal in length; slightly more than
|
||
<superScript id="7C03A25BE05DFFB4FA77D671FA31FDF5" attach="right" box="[1412,1420,618,632]" fontSize="6" pageId="65" pageNumber="66">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FA67D667FF10FD25" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="65" pageNumber="66" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05DFFB4FA67D667FA21FD07" attach="none" box="[1428,1436,636,650]" fontSize="6" pageId="65" pageNumber="66">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Tarsomere IV with a single ventral extension. Pretarsomere distinctly thickened; smooth ventrally; slightly less than
|
||
<superScript id="7C03A25BE05DFFB4FEA4D6A9FEE2FD4D" attach="right" box="[343,351,690,704]" fontSize="6" pageId="65" pageNumber="66">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E05DFFB4FE94D6DEFECAFD5E" attach="left" box="[359,375,709,723]" fontSize="6" pageId="65" pageNumber="66">10</subScript>
|
||
more in length of tarsomere I–IV combined. Internal claw slightly more than
|
||
<superScript id="7C03A25BE05DFFB4FB26D6A9FB60FD4D" attach="right" box="[1237,1245,690,704]" fontSize="6" pageId="65" pageNumber="66">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E05DFFB4FB16D6DEFB48FD5E" attach="left" box="[1253,1269,709,723]" fontSize="6" pageId="65" pageNumber="66">10</subScript>
|
||
more in length of pretarsomere. External claw slightly more than
|
||
<superScript id="7C03A25BE05DFFB4FD39D6CDFD6FFD69" attach="right" box="[714,722,726,740]" fontSize="6" pageId="65" pageNumber="66">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FD29D6F2FCBAFD7D" box="[730,775,726,759]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="65" pageNumber="66" unit="in" value="10.0">
|
||
<subScript id="17F20D56E05DFFB4FD29D6F2FD57FD7A" attach="left" box="[730,746,745,759]" fontSize="6" pageId="65" pageNumber="66">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of internal claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D6E1FE67FC0D" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D6E1FE26FC99" bold="true" box="[199,411,762,788]" pageId="65" pageNumber="66">Mesothoracic leg:</emphasis>
|
||
Tibia slightly less than
|
||
<superScript id="7C03A25BE05DFFB4FD55D6E1FD13FC85" attach="right" box="[678,686,762,776]" fontSize="6" pageId="65" pageNumber="66">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FD45D717FD66FC99" box="[694,731,762,794]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="65" pageNumber="66" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05DFFB4FD45D717FD03FC97" attach="left" box="[694,702,780,794]" fontSize="6" pageId="65" pageNumber="66">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined very short; slightly less than
|
||
<superScript id="7C03A25BE05DFFB4FEF1D705FEB7FCA1" attach="right" box="[258,266,798,812]" fontSize="6" pageId="65" pageNumber="66">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FEE1D72BFE85FCB5" box="[274,312,798,830]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="65" pageNumber="66" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05DFFB4FEE1D72BFEA7FCB3" attach="left" box="[274,282,816,830]" fontSize="6" pageId="65" pageNumber="66">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere with a shortened smooth ridge ventrally, roundish cutting missing; as long as tarsomere I–IV combined. External claw slightly shorter than pretarsomere. Internal claw slightly less than
|
||
<superScript id="7C03A25BE05DFFB4FF64D77DFF22FCF9" attach="none" box="[151,159,870,884]" fontSize="6" pageId="65" pageNumber="66">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FF54D763FF71FC0D" box="[167,204,870,902]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="65" pageNumber="66" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05DFFB4FF54D763FF12FC0B" attach="left" box="[167,175,888,902]" fontSize="6" pageId="65" pageNumber="66">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D791FDABFB9D" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D791FE26FC29" bold="true" box="[199,411,906,932]" pageId="65" pageNumber="66">Metathoracic leg:</emphasis>
|
||
Tibia slightly longer than femur. Tarsomere I–IV combined distinctly short; slightly more than
|
||
<superScript id="7C03A25BE05DFFB4FF3CD7B5FF6AFC31" attach="right" box="[207,215,942,956]" fontSize="6" pageId="65" pageNumber="66">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FF2CD7DAFEB0FC45" box="[223,269,942,975]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="65" pageNumber="66" unit="in" value="10.0">
|
||
<subScript id="17F20D56E05DFFB4FF2CD7DAFF52FC42" attach="left" box="[223,239,961,975]" fontSize="6" pageId="65" pageNumber="66">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly more than
|
||
<superScript id="7C03A25BE05DFFB4FA77D7B5FA31FC31" attach="right" box="[1412,1420,942,956]" fontSize="6" pageId="65" pageNumber="66">1</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FA67D7DAFF10FC61" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="65" pageNumber="66" unit="in" value="2.0">
|
||
<subScript id="17F20D56E05DFFB4FA67D7DAFA21FC42" attach="none" box="[1428,1436,961,975]" fontSize="6" pageId="65" pageNumber="66">2</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw slightly more than
|
||
<superScript id="7C03A25BE05DFFB4FC4AD7C9FC7CFC6D" attach="right" box="[953,961,978,992]" fontSize="6" pageId="65" pageNumber="66">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E05DFFB4FC3AD7FEFC64FC7E" attach="left" box="[969,985,997,1011]" fontSize="6" pageId="65" pageNumber="66">10</subScript>
|
||
more in length of pretarsomere. Internal claw
|
||
<superScript id="7C03A25BE05DFFB4FF20D7EDFF66FB89" attach="right" box="[211,219,1014,1028]" fontSize="6" pageId="65" pageNumber="66">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05DFFB4FF10D013FEB5FB9D" box="[227,264,1014,1046]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="65" pageNumber="66" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05DFFB4FF10D013FF56FB9B" attach="left" box="[227,235,1032,1046]" fontSize="6" pageId="65" pageNumber="66">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D001FD90FBB8" blockId="65.[151,1437,150,2013]" box="[199,557,1050,1077]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D001FEECFBB9" bold="true" box="[199,337,1050,1076]" pageId="65" pageNumber="66">Parameres:</emphasis>
|
||
(
|
||
<figureCitation id="134D1396E05DFFB4FE93D001FE55FBB9" box="[352,488,1050,1076]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="65" pageNumber="66">Figs. 92–93</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E05DFFB4FE07D001FD9CFBB8" box="[500,545,1050,1077]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="65" pageNumber="66">115</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D05BFCCBFBD4" blockId="65.[151,1437,150,2013]" box="[199,886,1086,1115]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D05BFE05FBD5" bold="true" box="[199,440,1086,1115]" pageId="65" pageNumber="66">
|
||
<specimenCount id="9D70C49AE05DFFB4FF34D05BFF4FFBD6" box="[199,242,1088,1115]" pageId="65" pageNumber="66" type="female">♀♀</specimenCount>
|
||
. Measurements:
|
||
</emphasis>
|
||
length
|
||
<quantity id="4C8EA2F6E05DFFB4FDFFD025FD2BFBD4" box="[524,662,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="9.2" metricValueMin="7.4" pageId="65" pageNumber="66" unit="mm" value="8.3" valueMax="9.2" valueMin="7.4">7.4–9.2 mm</quantity>
|
||
; width
|
||
<quantity id="4C8EA2F6E05DFFB4FD1BD025FCCFFBD4" box="[744,882,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.4" metricValueMin="4.4" pageId="65" pageNumber="66" unit="mm" value="4.9" valueMax="5.4" valueMin="4.4">4.4–5.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D079FF5BFB2D" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D079FEE6FBF1" bold="true" box="[199,347,1122,1148]" pageId="65" pageNumber="66">Integument:</emphasis>
|
||
Only lateral margin of elytra blackened; mesothoracic, metathoracic tibia and all tarsi pale redbrown.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D0B1FCADF889" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D0B1FEFAFB49" bold="true" box="[199,327,1194,1220]" pageId="65" pageNumber="66">Tegument:</emphasis>
|
||
<emphasis id="B902D301E05DFFB4FEBDD0B1FE15FB49" box="[334,424,1194,1220]" italics="true" pageId="65" pageNumber="66">Clypeus</emphasis>
|
||
sparse setose with short, erect, pale-brown setae. Colour of setae on
|
||
<emphasis id="B902D301E05DFFB4FB5FD0B1FB5FFB49" box="[1196,1250,1194,1220]" italics="true" pageId="65" pageNumber="66">head</emphasis>
|
||
and
|
||
<emphasis id="B902D301E05DFFB4FAEAD0B0FA21FB49" box="[1305,1436,1194,1220]" italics="true" pageId="65" pageNumber="66">eye canthus</emphasis>
|
||
pale-brown.
|
||
<emphasis id="B902D301E05DFFB4FED5D0D5FEC1FB65" box="[294,380,1230,1256]" italics="true" pageId="65" pageNumber="66">Funicle</emphasis>
|
||
with few very short semi-erect, whitish setae; denser on
|
||
<emphasis id="B902D301E05DFFB4FBF1D0D5FBF7FB65" box="[1026,1098,1230,1256]" italics="true" pageId="65" pageNumber="66">clavus</emphasis>
|
||
. Colour of setae on
|
||
<emphasis id="B902D301E05DFFB4FAC3D0D4FA21FB65" box="[1328,1436,1231,1256]" italics="true" pageId="65" pageNumber="66">pronotum</emphasis>
|
||
brown.
|
||
<emphasis id="B902D301E05DFFB4FF1DD0E9FEE1FA81" box="[238,348,1266,1292]" italics="true" pageId="65" pageNumber="66">Scutellum</emphasis>
|
||
loose setose with very short, semi-erect, brown setae; extremely short to very short, semi-erect, whitish setae interspersed medially and towards tip. Colour of setae on
|
||
<emphasis id="B902D301E05DFFB4FC47D10DFC48FABD" box="[948,1013,1302,1328]" italics="true" pageId="65" pageNumber="66">elytra</emphasis>
|
||
brown. Colour of hairs on
|
||
<emphasis id="B902D301E05DFFB4FAD7D10DFAD0FABD" box="[1316,1389,1302,1328]" italics="true" pageId="65" pageNumber="66">thorax</emphasis>
|
||
and
|
||
<emphasis id="B902D301E05DFFB4FF64D121FF44FAD9" box="[151,249,1338,1364]" italics="true" pageId="65" pageNumber="66">pleurites</emphasis>
|
||
pale-brown. Colour of setae on
|
||
<emphasis id="B902D301E05DFFB4FD97D120FD79FAD9" box="[612,708,1339,1364]" italics="true" pageId="65" pageNumber="66">ventrites</emphasis>
|
||
yellowish; interspersed setae long and pale-brown.
|
||
<emphasis id="B902D301E05DFFB4FAF8D121FA21FAD9" box="[1291,1436,1338,1364]" italics="true" pageId="65" pageNumber="66">Propygidium</emphasis>
|
||
and
|
||
<emphasis id="B902D301E05DFFB4FF34D145FE92FAF5" box="[199,303,1374,1400]" italics="true" pageId="65" pageNumber="66">pygidium</emphasis>
|
||
loose setose with short, semi-erect, yellowish setae; long, semi-erect, pale-brown setae interspersed.
|
||
<emphasis id="B902D301E05DFFB4FF64D198FEF8FA11" box="[151,325,1410,1436]" italics="true" pageId="65" pageNumber="66">Tip of pygidium</emphasis>
|
||
with a loose setose ciliary of medium-long, brown setae. Colour of hairs on
|
||
<emphasis id="B902D301E05DFFB4FB67D199FAE3FA11" box="[1172,1374,1410,1436]" italics="true" pageId="65" pageNumber="66">prothoracic femur</emphasis>
|
||
palebrown; medium-long, erect, brown setae interspersed ventrally; bare internally. Colour of hairs on
|
||
<emphasis id="B902D301E05DFFB4FAF6D1BDFF65FA69" italics="true" pageId="65" pageNumber="66">mesothoracic femur</emphasis>
|
||
pale-brown to brown. Interspersed setae on
|
||
<emphasis id="B902D301E05DFFB4FD3FD1D1FC23FA69" box="[716,926,1482,1508]" italics="true" pageId="65" pageNumber="66">mesothoracic tibia</emphasis>
|
||
red-brown; additionally with few semi-erect, medium-long, transparently red-brown bristles dorsally. Posterior margin with a bristled ciliary of medium-long, transparently red-brown bristles; not lengthening dorsally; missing internally.
|
||
<emphasis id="B902D301E05DFFB4FBE8D209FB40F9A1" box="[1051,1277,1554,1580]" italics="true" pageId="65" pageNumber="66">Metathoracic femur</emphasis>
|
||
sparse setose with medium-long, erect, whitish, setae; long, erect, pale-brown setae interspersed; bare internally. Colour of setae on
|
||
<emphasis id="B902D301E05DFFB4FF4ED241FE30F9F9" box="[189,397,1626,1652]" italics="true" pageId="65" pageNumber="66">metathoracic tibia</emphasis>
|
||
whitish; interspersed setae transparently red-brown.
|
||
<emphasis id="B902D301E05DFFB4FC1CD241FB7EF9F9" box="[1007,1219,1626,1652]" italics="true" pageId="65" pageNumber="66">Metathoracic tibia</emphasis>
|
||
sparse setose with very short and short, reclinate, brown setae; medium-long, semi-erect, robust, black setae interspersed; denser, long and finer internally. Colour of bristles on ciliary transparently red-brown.
|
||
<emphasis id="B902D301E05DFFB4FC23D2B8FA8EF931" box="[976,1331,1698,1724]" italics="true" pageId="65" pageNumber="66">Tarsomere I–IV of thoracic legs</emphasis>
|
||
with few very short, reclinate, whitish setae; colour of bristles on ciliary transparently red-brown. Colour of setae on
|
||
<emphasis id="B902D301E05DFFB4FABDD2DCFF5AF889" italics="true" pageId="65" pageNumber="66">pretarsomere</emphasis>
|
||
of all thoracic legs transparently pale red-brown.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D315FBA6F819" blockId="65.[151,1437,150,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D315FEADF8A5" bold="true" box="[199,272,1806,1832]" pageId="65" pageNumber="66">Head:</emphasis>
|
||
Clypeus very slightly depressed laterally; dense punctate with large, deep, roundish punctures. Anterior margin of clypeus slightly vertically reflexed. Lateral margin slightly reflexed. Suture between head and clypeus undulate. Head dense punctate with very large, deep, roundish punctures. Eye below head level. Funicle I–II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D385FA28F835" blockId="65.[151,1437,150,2013]" box="[199,1429,1950,1977]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D385FEF4F835" bold="true" box="[199,329,1950,1976]" pageId="65" pageNumber="66">Pronotum:</emphasis>
|
||
Lateral margin linearly attenuate in anterior two fifth, convex posteriorly. Medial furrow missing.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05DFFB4FF34D3D9FDAFF851" blockId="65.[151,1437,150,2013]" box="[199,530,1986,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B902D301E05DFFB4FF34D3D9FEF5F851" bold="true" box="[199,328,1986,2012]" pageId="65" pageNumber="66">Scutellum:</emphasis>
|
||
As broad as long.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05EFFB7FF34D48DFD8AFF3D" blockId="66.[151,1437,150,722]" box="[199,567,150,176]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF34D48DFEA0FF3D" bold="true" box="[199,285,150,176]" pageId="66" pageNumber="67">Elytra:</emphasis>
|
||
Lateral trench indistinct.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05EFFB7FF34D4A1FB84FF75" blockId="66.[151,1437,150,722]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF34D4A1FEF8FF59" bold="true" box="[199,325,186,212]" pageId="66" pageNumber="67">Abdomen:</emphasis>
|
||
Distinctly pulvinate; ventrite II–V broadened medially; ventrite I–IV broadest. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly declined posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05EFFB7FF34D519FE93FE5D" blockId="66.[151,1437,150,722]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF34D519FE3BFE91" bold="true" box="[199,390,258,284]" pageId="66" pageNumber="67">Prothoracic leg:</emphasis>
|
||
Femur and tibia robust. Median tooth broad triangular-shaped. Basal tooth very small. Tarsomere I–IV combined distinctly short; slightly less than
|
||
<superScript id="7C03A25BE05EFFB7FD0FD53DFCB9FEB9" attach="right" box="[764,772,294,308]" fontSize="6" pageId="66" pageNumber="67">3</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FCFFD523FC8CFECD" box="[780,817,294,326]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="66" pageNumber="67" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05EFFB7FCFFD523FCA9FECB" attach="left" box="[780,788,312,326]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia; tarsomere IV without ventral extension. Pretarsomere slightly thickened; with a shortened smooth ridge ventrally, roundish cutting missing; slightly less than
|
||
<superScript id="7C03A25BE05EFFB7FF3CD575FF6AFEF1" attach="right" box="[207,215,366,380]" fontSize="6" pageId="66" pageNumber="67">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FF2CD59AFEB0FE05" box="[223,269,366,399]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="66" pageNumber="67" unit="in" value="10.0">
|
||
<subScript id="17F20D56E05EFFB7FF2CD59AFF52FE02" attach="left" box="[223,239,385,399]" fontSize="6" pageId="66" pageNumber="67">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. Internal claw slender; not broadened medially; as long as pretarsomere. External claw slightly less than
|
||
<superScript id="7C03A25BE05EFFB7FDB1D589FDF7FE2D" attach="right" box="[578,586,402,416]" fontSize="6" pageId="66" pageNumber="67">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FDA1D5BFFDCAFE21" box="[594,631,402,434]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="66" pageNumber="67" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05EFFB7FDA1D5BFFDE7FE3F" attach="left" box="[594,602,420,434]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of internal claw. External claw slightly more than
|
||
<superScript id="7C03A25BE05EFFB7FB0FD589FAB9FE2D" attach="right" box="[1276,1284,402,416]" fontSize="6" pageId="66" pageNumber="67">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FAFFD5BFFA8CFE21" box="[1292,1329,402,434]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="66" pageNumber="67" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05EFFB7FAFFD5BFFAA9FE3F" attach="left" box="[1292,1300,420,434]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of internal claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05EFFB7FF34D5C1FAC9FDB1" blockId="66.[151,1437,150,722]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF34D5C1FE21FE79" bold="true" box="[199,412,474,500]" pageId="66" pageNumber="67">Mesothoracic leg:</emphasis>
|
||
Tibia slightly more than
|
||
<superScript id="7C03A25BE05EFFB7FD48D5C1FD7EFE65" attach="right" box="[699,707,474,488]" fontSize="6" pageId="66" pageNumber="67">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FD38D5F6FD44FE79" box="[715,761,474,507]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="66" pageNumber="67" unit="in" value="10.0">
|
||
<subScript id="17F20D56E05EFFB7FD38D5F6FD66FE76" attach="left" box="[715,731,493,507]" fontSize="6" pageId="66" pageNumber="67">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined distinctly short. Pretarsomere slightly thickened; slightly more than
|
||
<superScript id="7C03A25BE05EFFB7FD20D5E5FD66FD81" attach="right" box="[723,731,510,524]" fontSize="6" pageId="66" pageNumber="67">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FD11D60AFCB3FD95" box="[738,782,510,543]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="66" pageNumber="67" unit="in" value="10.0">
|
||
<subScript id="17F20D56E05EFFB7FD11D60AFD4FFD92" attach="left" box="[738,754,529,543]" fontSize="6" pageId="66" pageNumber="67">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw slender; not broadened medially; as long as pretarsomere. Internal claw slightly more than
|
||
<superScript id="7C03A25BE05EFFB7FBC2D639FB84FDBD" attach="right" box="[1073,1081,546,560]" fontSize="6" pageId="66" pageNumber="67">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FBB3D62FFBDBFDB1" box="[1088,1126,546,578]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="66" pageNumber="67" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05EFFB7FBB3D62FFBF5FDCF" attach="left" box="[1088,1096,564,578]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05EFFB7FF34D65DFCADFD41" blockId="66.[151,1437,150,722]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF34D65DFE24FDED" bold="true" box="[199,409,582,608]" pageId="66" pageNumber="67">Metathoracic leg:</emphasis>
|
||
Femur slightly thickened. Tibia cornet-shaped;
|
||
<superScript id="7C03A25BE05EFFB7FC43D65DFC05FDD9" attach="right" box="[944,952,582,596]" fontSize="6" pageId="66" pageNumber="67">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FC33D642FC50FDED" box="[960,1005,582,615]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="66" pageNumber="67" unit="in" value="10.0">
|
||
<subScript id="17F20D56E05EFFB7FC33D642FC6DFDEA" attach="left" box="[960,976,601,615]" fontSize="6" pageId="66" pageNumber="67">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined very short; slightly less than
|
||
<superScript id="7C03A25BE05EFFB7FDD7D671FD91FDF5" attach="right" box="[548,556,618,632]" fontSize="6" pageId="66" pageNumber="67">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FDC7D667FDE7FD09" box="[564,602,618,650]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="66" pageNumber="67" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05EFFB7FDC7D667FD81FD07" attach="left" box="[564,572,636,650]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slender; with a shortened smooth ridge ventrally; slightly less than
|
||
<superScript id="7C03A25BE05EFFB7FE56D695FE10FD11" attach="right" box="[421,429,654,668]" fontSize="6" pageId="66" pageNumber="67">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FE47D6BBFE67FD25" box="[436,474,654,686]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="66" pageNumber="67" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05EFFB7FE47D6BBFE01FD23" attach="left" box="[436,444,672,686]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw
|
||
<superScript id="7C03A25BE05EFFB7FBE8D695FB9EFD11" attach="right" box="[1051,1059,654,668]" fontSize="6" pageId="66" pageNumber="67">2</superScript>
|
||
/
|
||
<subScript id="17F20D56E05EFFB7FBD8D6BBFB8EFD23" attach="left" box="[1067,1075,672,686]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
more in length of pretarsomere. Internal claw slightly more
|
||
<superScript id="7C03A25BE05EFFB7FE3ED6A9FE68FD4D" attach="right" box="[461,469,690,704]" fontSize="6" pageId="66" pageNumber="67">3</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E05EFFB7FE2ED6DFFDBFFD41" box="[477,514,690,722]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="66" pageNumber="67" unit="in" value="5.0">
|
||
<subScript id="17F20D56E05EFFB7FE2ED6DFFE58FD5F" attach="left" box="[477,485,708,722]" fontSize="6" pageId="66" pageNumber="67">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E05EFFB7FF64D6E1FCE8FCD0" pageId="66" pageNumber="67" type="distribution">
|
||
<paragraph id="8BC90F13E05EFFB7FF64D6E1FBEDFC99" blockId="66.[151,1104,762,789]" box="[151,1104,762,789]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF64D6E1FE8CFC99" bold="true" box="[151,305,762,788]" pageId="66" pageNumber="67">Distribution.</emphasis>
|
||
<collectingRegion id="49B2C1F1E05EFFB7FEC4D6E1FDF5FC99" box="[311,584,762,788]" country="South Africa" name="Northern Cape" pageId="66" pageNumber="67">Northern Cape Province</collectingRegion>
|
||
(
|
||
<figureCitation id="134D1396E05EFFB7FDA4D6E1FD05FC99" box="[599,696,762,788]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="66" pageNumber="67">Fig. 140</figureCitation>
|
||
), only known from the
|
||
<typeStatus id="54CDB1B1E05EFFB7FC4CD6E0FC52FC98" box="[959,1007,763,789]" pageId="66" pageNumber="67">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E05EFFB7FF64D759FCE8FCD0" blockId="66.[151,853,834,861]" box="[151,853,834,861]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF64D759FEE3FCD1" bold="true" box="[151,350,834,860]" pageId="66" pageNumber="67">
|
||
Vegetation
|
||
<typeStatus id="54CDB1B1E05EFFB7FEEFD759FEE4FCD1" box="[284,345,834,860]" pageId="66" pageNumber="67">Type</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Nieuwoudtville Shale Renosterveld [FRs 2].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E05EFFB7FF64D791FD3BFC28" box="[151,646,906,933]" pageId="66" pageNumber="67" type="biology_ecology">
|
||
<paragraph id="8BC90F13E05EFFB7FF64D791FD3BFC28" blockId="66.[151,646,906,933]" box="[151,646,906,933]" pageId="66" pageNumber="67">
|
||
<emphasis id="B902D301E05EFFB7FF64D791FE95FC29" bold="true" box="[151,296,906,932]" pageId="66" pageNumber="67">Host Plants.</emphasis>
|
||
<taxonomicName id="4C767490E05EFFB7FEDED791FE11FC29" box="[301,428,906,932]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C767490E05EFFB7FE44D790FDDAFC29" box="[439,615,906,933]" class="Magnoliopsida" family="Asteraceae" genus="Arctotis" kingdom="Plantae" order="Asterales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="acaulis">
|
||
<emphasis id="B902D301E05EFFB7FE44D790FDDAFC29" box="[439,615,906,933]" italics="true" pageId="66" pageNumber="67">Arctotis acaulis</emphasis>
|
||
</taxonomicName>
|
||
L.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |