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<document id="BBE62851C88D2CD03E6536DBB54A81D7" ID-DOI="10.5281/zenodo.13245646" ID-ISSN="2345-7600" ID-Zenodo-Dep="13245646" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970853461" checkinUser="felipe" docAuthor="Ng, Oliver K. S. Chia Peter K. L." docDate="2006" docId="03DF1C3DFFADFFA77C99FD84FE04FC11" docLanguage="en" docName="RafflesBZool.54.2.381-428.pdf" docOrigin="Raffles Bulletin of Zoology 54 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Parathelphusa ceophallus Ng 1993" docType="treatment" docVersion="2" lastPageNumber="393" masterDocId="FFE66445FFA7FFAB7F27FF80FFFFFF89" masterDocTitle="The Freshwater Crabs Of Sulawesi, With Descriptions Of Two New Genera And Four New Species (Crustacea: Decapoda: Brachyura: Parathelphusidae)" masterLastPageNumber="428" masterPageNumber="381" pageNumber="391" updateTime="1722977516455" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6DC2494FD968EE35BCC74D0ADBF56820">The Freshwater Crabs Of Sulawesi, With Descriptions Of Two New Genera And Four New Species (Crustacea: Decapoda: Brachyura: Parathelphusidae)</mods:title>
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<mods:namePart id="12F23F084C0E3C59F7794D5454E4F07C">Ng, Oliver K. S. Chia Peter K. L.</mods:namePart>
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<mods:title id="492FC132188CF9D4DD2A3D119D878BB8">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="EC84CA3587562B13170513F1E0E03BD6">2006</mods:date>
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<mods:identifier id="AE8C7CB6AF96E2F4A3A886F64FA37EE3" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C997E94BE2179D5FB300A7B10D006EA5" type="Zenodo-Dep">13245646</mods:identifier>
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<treatment id="03DF1C3DFFADFFA77C99FD84FE04FC11" LSID="urn:lsid:plazi:treatment:03DF1C3DFFADFFA77C99FD84FE04FC11" httpUri="http://treatment.plazi.org/id/03DF1C3DFFADFFA77C99FD84FE04FC11" lastPageId="12" lastPageNumber="393" pageId="10" pageNumber="391">
<subSubSection id="C36CFEA0FFADFFA17C99FD84FAB5FD95" box="[958,1354,516,540]" pageId="10" pageNumber="391" type="nomenclature">
<paragraph id="8BC9AD2BFFADFFA17C99FD84FAB5FD95" blockId="10.[958,1354,516,572]" box="[958,1354,516,540]" pageId="10" pageNumber="391">
<heading id="D0811A47FFADFFA17C99FD84FAB5FD95" bold="true" box="[958,1354,516,540]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="391" reason="2">
<taxonomicName id="4C76D6A8FFADFFA17C99FD84FAB5FD95" ID-CoL="934Y5" authority="Ng, 1993" authorityName="Ng" authorityYear="1993" box="[958,1354,516,540]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFADFFA17C99FD84FAB5FD95" bold="true" box="[958,1354,516,540]" pageId="10" pageNumber="391">
<emphasis id="B9027139FFADFFA17C99FD84FB20FD95" bold="true" box="[958,1247,516,540]" italics="true" pageId="10" pageNumber="391">Parathelphusa ceophallus</emphasis>
Ng, 1993
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36CFEA0FFADFFA17B7BFDA5FB52FDB5" box="[1116,1197,549,572]" pageId="10" pageNumber="391" type="description">
<paragraph id="8BC9AD2BFFADFFA17B7BFDA5FB52FDB5" blockId="10.[958,1354,516,572]" box="[1116,1197,549,572]" pageId="10" pageNumber="391">
(
<figureCitation id="134DB1AEFFADFFA17B43FDA5FB5AFDB5" box="[1124,1189,549,572]" captionStart="Fig" captionStartId="11.[112,147,2007,2028]" captionTargetBox="[166,1409,188,1947]" captionTargetId="figure-9@11.[94,1438,127,1989]" captionTargetPageId="11" captionText="Fig. 7. Parathelphusa ceophallus Ng, 1993, holotype male (23.1 x 18.1mm) (RMNH D 42418): A, right dorsal view of carapace; B, frontal view of carapace; C, frontal median triangle; D, anterior thoracic sternum; E, male abdominal segmenst 4-6 and telson; F, left third maxilliped; G, right fourth ambulatory leg; H, left G1 ventral view; I, left G1 dorsal view; J, left G1 distal part, ventral view; K, left G1 distal part, dorsal view; L, left G2 (after Ng, 1993: Figs. 1, 2). Scale bars: A-G = 1.0 mm; H, I, L = 0.5 mm; J, K = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13245660" httpUri="https://zenodo.org/record/13245660/files/figure.png" pageId="10" pageNumber="391">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36CFEA0FFADFFA17C62FDE4FAB1FDF0" box="[837,1358,612,633]" pageId="10" pageNumber="391" type="reference_group">
<paragraph id="8BC9AD2BFFADFFA17C62FDE4FAB1FDF0" blockId="10.[837,1358,612,633]" box="[837,1358,612,633]" pageId="10" pageNumber="391">
<taxonomicName id="4C76D6A8FFADFFA17C62FDE4FB22FDF0" ID-CoL="934Y5" authority="Ng, 1993: 179" authorityName="Ng" authorityPageNumber="179" authorityYear="1993" box="[837,1245,612,633]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFADFFA17C62FDE4FBBCFDF0" box="[837,1091,612,633]" italics="true" pageId="10" pageNumber="391">Parathelphusa ceophallus</emphasis>
Ng, 1993: 179
</taxonomicName>
,
<figureCitation id="134DB1AEFFADFFA17BCEFDE4FAD2FDF0" box="[1257,1325,612,633]" captionStart="Fig" captionStartId="4.[136,171,2065,2086]" captionTargetBox="[136,1476,474,2047]" captionTargetId="figure-161@4.[136,1476,474,2047]" captionTargetPageId="4" captionText="Fig. 1. Parathelphusa pantherina (Schenkel, 1902), paralectotype male (42.4 x 31.2 mm, NMB 21A): A, dorsal view; B, frontal view; C, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13245648" httpUri="https://zenodo.org/record/13245648/files/figure.png" pageId="10" pageNumber="391">Figs. 1</figureCitation>
,
<figureCitation id="134DB1AEFFADFFA17A1BFDE4FAB5FDF0" box="[1340,1354,612,633]" captionStart="Fig" captionStartId="6.[835,869,2037,2058]" captionTargetBox="[835,1475,1200,2020]" captionTargetId="figure-795@6.[835,1475,1200,2020]" captionTargetPageId="6" captionText="Fig. 2. Parathelphusa sarasinorum (Schenkel, 1902), lectotype male (38.9 x 31.5 mm, NMB 22B): A, dorsal view; B, frontal view; C, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13245650" httpUri="https://zenodo.org/record/13245650/files/figure.png" pageId="10" pageNumber="391">2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36CFEA0FFADFFA17C62FD1DFA3AFCCA" pageId="10" pageNumber="391" type="materials_examined">
<paragraph id="8BC9AD2BFFADFFA17C62FD1DFA3AFCCA" blockId="10.[837,1477,669,835]" pageId="10" pageNumber="391">
<emphasis id="B9027139FFADFFA17C62FD1DFBFCFD3B" bold="true" box="[837,1027,669,690]" italics="true" pageId="10" pageNumber="391">Material examined</emphasis>
.
<emphasis id="B9027139FFADFFA17B36FD1DFBE2FD3B" bold="true" box="[1041,1053,669,690]" pageId="10" pageNumber="391"></emphasis>
<materialsCitation id="3B1EA776FFADFFA17B01FD1EFA3AFCCA" collectingDate="1989-11-12" collectingDateMax="1989-11-18" collectingDateMin="1989-11-12" collectionCode="RMNH" collectorName="J. van Tol" county="Jismil Camp" elevation="150" latitude="-4.4416666" location="Sungai" longLatPrecision="922" longitude="122.98333" municipality="Labuhan Tobela" pageId="10" pageNumber="391" specimenCode="D 42418" specimenCount="1" stateProvince="Sulawesi Tenggara" typeStatus="holotype">
<typeStatus id="54CD1389FFADFFA17B01FD1EFB7CFD3A" box="[1062,1155,670,691]" pageId="10" pageNumber="391" type="holotype">Holotype</typeStatus>
: male (23.1 x
<quantity id="4C8E00CEFFADFFA17A33FD1EFA9BFD3A" box="[1300,1380,670,691]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="10" pageNumber="391" unit="mm" value="18.1">18.1mm</quantity>
) (
<collectionCode id="ED6735EEFFADFFA17A5DFD1EFA3BFD3A" box="[1402,1476,670,691]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="391" type="Museum">RMNH</collectionCode>
<specimenCode id="DBD00550FFADFFA17C62FD3AFC60FD46" box="[837,927,698,719]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="391" type="Museum">D 42418</specimenCode>
),
<collectingCounty id="62A8D5A7FFADFFA17C96FD3AFBD0FD46" box="[945,1071,698,719]" pageId="10" pageNumber="391">Jismil Camp</collectingCounty>
, a few kilometres from
<collectingMunicipality id="6BAD3751FFADFFA17A38FD3AFA3FFD46" box="[1311,1472,698,719]" pageId="10" pageNumber="391">Labuhan Tobela</collectingMunicipality>
, along
<location id="8EA9FBF0FFADFFA17CA0FD57FC33FD65" LSID="urn:lsid:plazi:treatment:03DF1C3DFFADFFA77C99FD84FE04FC11:8EA9FBF0FFADFFA17CA0FD57FC33FD65" box="[903,972,727,748]" county="Jismil Camp" latitude="-4.4416666" longLatPrecision="922" longitude="122.98333" municipality="Labuhan Tobela" name="Sungai" pageId="10" pageNumber="391" stateProvince="Sulawesi Tenggara">Sungai</location>
(River)
<location id="8EA9FBF0FFADFFA17B0EFD57FB2DFD65" LSID="urn:lsid:plazi:treatment:03DF1C3DFFADFFA77C99FD84FE04FC11:8EA9FBF0FFADFFA17B0EFD57FB2DFD65" box="[1065,1234,727,748]" county="Jismil Camp" latitude="-4.4416666" longLatPrecision="922" longitude="122.98333" municipality="Labuhan Tobela" name="Labuhan Tobela" pageId="10" pageNumber="391" stateProvince="Sulawesi Tenggara">Labuhan Tobela</location>
, rivulet through hardly disturbed primary evergreen rainforest, boulder covered with calcareous deposits,
<quantity id="4C8E00CEFFADFFA17B3BFC91FB9EFCAF" box="[1052,1121,785,806]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="391" unit="m" value="150.0">
<elevation id="005B4A18FFADFFA17B3BFC91FB9EFCAF" box="[1052,1121,785,806]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="391" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<location id="8EA9FBF0FFADFFA17B48FC91FAAEFCAF" LSID="urn:lsid:plazi:treatment:03DF1C3DFFADFFA77C99FD84FE04FC11:8EA9FBF0FFADFFA17B48FC91FAAEFCAF" box="[1135,1361,785,806]" county="Jismil Camp" latitude="-4.4416666" longLatPrecision="922" longitude="122.98333" municipality="Labuhan Tobela" name="northern Pulau Buton" pageId="10" pageNumber="391" stateProvince="Sulawesi Tenggara">northern Pulau Buton</location>
,
<geoCoordinate id="EE42CBECFFADFFA17A79FC91FA3CFCAF" box="[1374,1475,783,806]" degrees="4" direction="south" minutes="26" orientation="latitude" pageId="10" pageNumber="391" precision="15" seconds="30" value="-4.4416666">
4
<emphasis id="B9027139FFADFFA17A4CFC8FFA8AFCAF" bold="true" box="[1387,1397,783,806]" pageId="10" pageNumber="391">°</emphasis>
26'30&quot;S
</geoCoordinate>
<geoCoordinate id="EE42CBECFFADFFA17C62FCAEFC5FFCCA" box="[837,928,812,835]" degrees="122" direction="east" minutes="59" orientation="longitude" pageId="10" pageNumber="391" precision="925" value="122.98333">
122
<emphasis id="B9027139FFADFFA17C4EFCACFC8CFCCA" bold="true" box="[873,883,812,835]" pageId="10" pageNumber="391">°</emphasis>
59'E
</geoCoordinate>
,
<collectingRegion id="49B263C9FFADFFA17C8BFCAEFB93FCCA" box="[940,1132,814,835]" country="Indonesia" name="Sulawesi Tenggara" pageId="10" pageNumber="391">Southeast Sulawesi</collectingRegion>
, coll.
<collectorName id="2683C8FDFFADFFA17B8CFCAEFAEDFCCA" box="[1195,1298,814,835]" pageId="10" pageNumber="391">J. van Tol</collectorName>
,
<date id="FFC88BEBFFADFFA17A39FCAEFA3EFCCA" box="[1310,1473,814,835]" pageId="10" pageNumber="391" value="1989-11-12" valueMax="1989-11-18" valueMin="1989-11-12">
<collectingDate id="EF8C7203FFADFFA17A39FCAEFA3EFCCA" box="[1310,1473,814,835]" pageId="10" pageNumber="391" value="1989-11-12" valueMax="1989-11-18" valueMin="1989-11-12">12-18 Nov.1989</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36CFEA0FFADFFA17C62FCEBFBA5F8AB" pageId="10" pageNumber="391" type="diagnosis">
<paragraph id="8BC9AD2BFFADFFA17C62FCEBFBA5F8AB" blockId="10.[837,1477,875,1826]" pageId="10" pageNumber="391">
<emphasis id="B9027139FFADFFA17C62FCEBFC48FC0A" bold="true" box="[837,951,875,899]" italics="true" pageId="10" pageNumber="391">Diagnosis</emphasis>
.
<emphasis id="B9027139FFADFFA17CEEFCEBFC29FC0A" bold="true" box="[969,982,875,899]" pageId="10" pageNumber="391"></emphasis>
Carapace dorsal surfaces smooth; cervical groove shallow, narrow; H-shaped central depression pronounced; branchial regions slightly inflated. Epigastric and postorbital cristae continuous, sharp, subparallel to frontal margin, with slight cleft at beginning of cervical groove; outer edge of cristae not reaching anterolateral margin, stopping before very shallow cleft separating first and second epibranchial teeth. Front slightly sinuous, entire; frontal median triangle distinct, but upper ridged margin does not join lateral margins, separated by small but distinct gap. Anterolateral margin arcuate, weakly cristae, external orbital angle broadly triangular, outer margin almost straight, about 3 times length of inner margin, first and second epibranchial teeth very low, lobiform, barely discernible. Posterolateral regions lined with distinct oblique striae. Suture between thoracic sternites 2 and 3 incomplete, gently concave towards buccal cavity. Palm of chelipeds slightly inflated, fingers longer than palm, pollex not pigmented black but dactylus pigmented black throughout length. Ambulatory legs relatively short, second pair longest. Dorsal margin of merus slightly cristate, gently serrated, without distinct subterminal spine. Male abdomen reaching imaginary line joining anterior edges of cheliped bases, T-shaped; segment 6 distinctly longer than segment 7; lateral margins of segment 7 gently concave; lateral margins of segment 6 subparallel, gently sinuous. G1 relatively straight, directed upwards, outer margin gently sinuous along distal two-thirds, proximal part with very deep V-shaped cleft; distal part tapering. G2 with well developed distal segment, about 0.6 times length of basal segment. (Adapted from Ng, 1993)
</paragraph>
</subSubSection>
<subSubSection id="C36CFEA0FFADFFA17C62F8CBFA9EF8EB" box="[837,1377,1867,1891]" pageId="10" pageNumber="391" type="distribution">
<paragraph id="8BC9AD2BFFADFFA17C62F8CBFA9EF8EB" blockId="10.[837,1377,1867,1891]" box="[837,1377,1867,1891]" pageId="10" pageNumber="391">
<emphasis id="B9027139FFADFFA17C62F8CBFC35F8EA" bold="true" box="[837,970,1867,1891]" italics="true" pageId="10" pageNumber="391">Distribution</emphasis>
.
<emphasis id="B9027139FFADFFA17CFDF8CBFC18F8EA" bold="true" box="[986,999,1867,1891]" pageId="10" pageNumber="391"></emphasis>
Pulau Buton,
<collectingRegion id="49B263C9FFADFFA17BAFF8CBFAA2F8EB" box="[1160,1373,1867,1890]" country="Indonesia" name="Sulawesi Tenggara" pageId="10" pageNumber="391">Southeast Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C36CFEA0FFADFFA77C62F80BFE04FC11" lastPageId="12" lastPageNumber="393" pageId="10" pageNumber="391" type="discussion">
<paragraph id="8BC9AD2BFFADFFA77C62F80BFD06FEF1" blockId="10.[837,1477,1931,2114]" lastBlockId="12.[136,775,193,376]" lastPageId="12" lastPageNumber="393" pageId="10" pageNumber="391">
<emphasis id="B9027139FFADFFA17C62F80BFC52F82B" box="[837,941,1931,1955]" italics="true" pageId="10" pageNumber="391">
<emphasis id="B9027139FFADFFA17C62F80BFC59F82A" bold="true" box="[837,934,1931,1955]" italics="true" pageId="10" pageNumber="391">Remarks</emphasis>
.
</emphasis>
<emphasis id="B9027139FFADFFA17C96F80BFC41F82A" bold="true" box="[945,958,1931,1955]" pageId="10" pageNumber="391"></emphasis>
Ng (1993), commented that
<taxonomicName id="4C76D6A8FFADFFA17BC0F80BFA8AF82B" authorityName="Ng" authorityYear="1993" box="[1255,1397,1931,1954]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFADFFA17BC0F80BFA8AF82B" box="[1255,1397,1931,1954]" italics="true" pageId="10" pageNumber="391">P. ceophallus</emphasis>
</taxonomicName>
is most similar to species of the
<emphasis id="B9027139FFADFFA17B6BF82BFAA5F84B" box="[1100,1370,1963,1986]" italics="true" pageId="10" pageNumber="391">
<taxonomicName id="4C76D6A8FFADFFA17B6BF82BFB26F84B" authorityName="De Man" authorityYear="1892" box="[1100,1241,1963,1986]" class="Malacostraca" family="Gecarcinucidae" genus="Telphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="celebensis">P. celebensis</taxonomicName>
-
<taxonomicName id="4C76D6A8FFADFFA17BD7F82BFAA5F84B" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[1264,1370,1963,1986]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="pallida">P. pallida</taxonomicName>
</emphasis>
complex. However, it can be distinguished from the
<taxonomicName id="4C76D6A8FFADFFA17A16F84BFA3BF86B" authorityName="De Man" authorityYear="1892" box="[1329,1476,1995,2018]" class="Malacostraca" family="Gecarcinucidae" genus="Telphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="391" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="B9027139FFADFFA17A16F84BFA3BF86B" box="[1329,1476,1995,2018]" italics="true" pageId="10" pageNumber="391">P. celebensis</emphasis>
</taxonomicName>
complex in that it possess a cleft on the postorbital cristae where the cervical groove begins (
<figureCitation id="134DB1AEFFADFFA17B9CF78BFAEFF7AB" box="[1211,1296,2059,2082]" captionStart="Fig" captionStartId="11.[112,147,2007,2028]" captionTargetBox="[166,1409,188,1947]" captionTargetId="figure-9@11.[94,1438,127,1989]" captionTargetPageId="11" captionText="Fig. 7. Parathelphusa ceophallus Ng, 1993, holotype male (23.1 x 18.1mm) (RMNH D 42418): A, right dorsal view of carapace; B, frontal view of carapace; C, frontal median triangle; D, anterior thoracic sternum; E, male abdominal segmenst 4-6 and telson; F, left third maxilliped; G, right fourth ambulatory leg; H, left G1 ventral view; I, left G1 dorsal view; J, left G1 distal part, ventral view; K, left G1 distal part, dorsal view; L, left G2 (after Ng, 1993: Figs. 1, 2). Scale bars: A-G = 1.0 mm; H, I, L = 0.5 mm; J, K = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13245660" httpUri="https://zenodo.org/record/13245660/files/figure.png" pageId="10" pageNumber="391">Fig. 7A</figureCitation>
) versus the cleft being absent in other species of the complex. The dorsal margin of the frontal median triangle of
<taxonomicName id="4C76D6A8FFABFFA77D09FF41FD40FF51" authorityName="Ng" authorityYear="1993" box="[558,703,193,216]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFABFFA77D09FF41FD40FF51" box="[558,703,193,216]" italics="true" pageId="12" pageNumber="393">P. ceophallus</emphasis>
</taxonomicName>
is also separated from the lateral margins by a distinct gap (
<figureCitation id="134DB1AEFFABFFA77D8FFF61FD05FF71" box="[680,762,225,248]" captionStart="Fig" captionStartId="11.[112,147,2007,2028]" captionTargetBox="[166,1409,188,1947]" captionTargetId="figure-9@11.[94,1438,127,1989]" captionTargetPageId="11" captionText="Fig. 7. Parathelphusa ceophallus Ng, 1993, holotype male (23.1 x 18.1mm) (RMNH D 42418): A, right dorsal view of carapace; B, frontal view of carapace; C, frontal median triangle; D, anterior thoracic sternum; E, male abdominal segmenst 4-6 and telson; F, left third maxilliped; G, right fourth ambulatory leg; H, left G1 ventral view; I, left G1 dorsal view; J, left G1 distal part, ventral view; K, left G1 distal part, dorsal view; L, left G2 (after Ng, 1993: Figs. 1, 2). Scale bars: A-G = 1.0 mm; H, I, L = 0.5 mm; J, K = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13245660" httpUri="https://zenodo.org/record/13245660/files/figure.png" pageId="12" pageNumber="393">Fig. 7C</figureCitation>
).
<taxonomicName id="4C76D6A8FFABFFA77FAFFE81FE61FE91" authorityName="Ng" authorityYear="1993" box="[136,414,257,280]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFABFFA77FAFFE81FE61FE91" box="[136,414,257,280]" italics="true" pageId="12" pageNumber="393">Parathelphusa ceophallus</emphasis>
</taxonomicName>
also possess a very distinct, broad and deep V-shaped cleft along the proximal outer margin of its G1, against a shallow cleft or the margin appearing entire that the other species of the complex possess (
<figureCitation id="134DB1AEFFABFFA77D58FEE1FD13FEF1" box="[639,748,353,376]" captionStart="Fig" captionStartId="11.[112,147,2007,2028]" captionTargetBox="[166,1409,188,1947]" captionTargetId="figure-9@11.[94,1438,127,1989]" captionTargetPageId="11" captionText="Fig. 7. Parathelphusa ceophallus Ng, 1993, holotype male (23.1 x 18.1mm) (RMNH D 42418): A, right dorsal view of carapace; B, frontal view of carapace; C, frontal median triangle; D, anterior thoracic sternum; E, male abdominal segmenst 4-6 and telson; F, left third maxilliped; G, right fourth ambulatory leg; H, left G1 ventral view; I, left G1 dorsal view; J, left G1 distal part, ventral view; K, left G1 distal part, dorsal view; L, left G2 (after Ng, 1993: Figs. 1, 2). Scale bars: A-G = 1.0 mm; H, I, L = 0.5 mm; J, K = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13245660" httpUri="https://zenodo.org/record/13245660/files/figure.png" pageId="12" pageNumber="393">Fig. 7H, I</figureCitation>
).
</paragraph>
<caption id="DF09FDA3FFACFFA07F57F857FB67F7CA" ID-DOI="http://doi.org/10.5281/zenodo.13245660" ID-Zenodo-Dep="13245660" httpUri="https://zenodo.org/record/13245660/files/figure.png" pageId="11" pageNumber="392" startId="11.[112,147,2007,2028]" targetBox="[166,1409,188,1947]" targetPageId="11" targetType="figure">
<paragraph id="8BC9AD2BFFACFFA07F57F857FB67F7CA" blockId="11.[112,1452,2007,2115]" pageId="11" pageNumber="392">
Fig. 7.
<taxonomicName id="4C76D6A8FFACFFA07F93F857FDEEF865" authority="Ng, 1993" authorityName="Ng" authorityYear="1993" box="[180,529,2007,2028]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="392" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFACFFA07F93F857FE4FF865" box="[180,432,2007,2028]" italics="true" pageId="11" pageNumber="392">Parathelphusa ceophallus</emphasis>
Ng, 1993
</taxonomicName>
, holotype male (23.1 x 18.1mm) (RMNH D 42418): A, right dorsal view of carapace; B, frontal view of carapace; C, frontal median triangle; D, anterior thoracic sternum; E, male abdominal segmenst 4-6 and telson; F, left third maxilliped; G, right fourth ambulatory leg; H, left G1 ventral view; I, left G1 dorsal view; J, left G1 distal part, ventral view; K, left G1 distal part, dorsal view; L, left G2 (after Ng, 1993: Figs. 1, 2). Scale bars: A-G = 1.0 mm; H, I, L = 0.5 mm; J, K = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BC9AD2BFFABFFA77FAFFE21FE04FC11" blockId="12.[136,775,417,920]" pageId="12" pageNumber="393">
The other two characters that Ng (1993) used to differentiate
<taxonomicName id="4C76D6A8FFABFFA77FAFFE41FEE4FE51" authorityName="Ng" authorityYear="1993" box="[136,283,449,472]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFABFFA77FAFFE41FEE4FE51" box="[136,283,449,472]" italics="true" pageId="12" pageNumber="393">P. ceophallus</emphasis>
</taxonomicName>
from the rest of the complex, namely a low, less distinct epibranchial teeth and the outer edge of the postorbital cristae stopping well before the anterolateral margin are no longer applicable as several poorly known taxa in this group are now better understood. While they work for
<taxonomicName id="4C76D6A8FFABFFA77FAFFDE1FF0DFDF1" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[136,242,609,632]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B9027139FFABFFA77FAFFDE1FF0DFDF1" box="[136,242,609,632]" italics="true" pageId="12" pageNumber="393">P. pallida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C76D6A8FFABFFA77E00FDE1FE4CFDF1" authorityName="De Man" authorityYear="1892" box="[295,435,609,632]" class="Malacostraca" family="Gecarcinucidae" genus="Telphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="B9027139FFABFFA77E00FDE1FE4CFDF1" box="[295,435,609,632]" italics="true" pageId="12" pageNumber="393">P. celebensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A231CC42FFABFFA77E9DFDE1FE09FDF1" box="[442,502,609,632]" pageId="12" pageNumber="393" sensu="stricto">s. str.</taxonomicNameLabel>
,
<taxonomicName id="4C76D6A8FFABFFA77D26FDE1FD6EFDF1" baseAuthorityName="Roux" baseAuthorityYear="1904" box="[513,657,609,632]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="possoensis">
<emphasis id="B9027139FFABFFA77D26FDE1FD6EFDF1" box="[513,657,609,632]" italics="true" pageId="12" pageNumber="393">P. possoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C76D6A8FFABFFA77D87FDE1FCF8FDF1" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[672,775,609,632]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="crocea">
<emphasis id="B9027139FFABFFA77D87FDE1FCF8FDF1" box="[672,775,609,632]" italics="true" pageId="12" pageNumber="393">P. crocea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C76D6A8FFABFFA77F90FD01FEBCFD11" baseAuthorityName="Roux" baseAuthorityYear="1904" box="[183,323,641,664]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="linduensis">
<emphasis id="B9027139FFABFFA77F90FD01FEBCFD11" box="[183,323,641,664]" italics="true" pageId="12" pageNumber="393">P. linduensis</emphasis>
</taxonomicName>
all have very low epibranchial teeth that are almost continuous with the anterolateral margin like
<taxonomicName id="4C76D6A8FFABFFA77DD7FD21FF06FD51" authorityName="Ng" authorityYear="1993" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFABFFA77DD7FD21FF06FD51" italics="true" pageId="12" pageNumber="393">P. ceophallus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C76D6A8FFABFFA77E23FD41FDF2FD51" baseAuthorityName="Roux" baseAuthorityYear="1904" box="[260,525,705,728]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="linduensis">
<emphasis id="B9027139FFABFFA77E23FD41FDF2FD51" box="[260,525,705,728]" italics="true" pageId="12" pageNumber="393">Parathelphusa linduensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C76D6A8FFABFFA77D1AFD41FD5EFD51" authorityName="Chia &amp; Ng" authorityYear="2006" box="[573,673,705,728]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="sorella">
<emphasis id="B9027139FFABFFA77D1AFD41FD5EFD51" box="[573,673,705,728]" italics="true" pageId="12" pageNumber="393">P. sorella</emphasis>
</taxonomicName>
also have its outer edge of the postorbital cristae stopping well before the anterolateral margin. However,
<taxonomicName id="4C76D6A8FFABFFA77D2AFC81FD5FFC91" authorityName="Ng" authorityYear="1993" box="[525,672,769,792]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="393" phylum="Arthropoda" rank="species" species="ceophallus">
<emphasis id="B9027139FFABFFA77D2AFC81FD5FFC91" box="[525,672,769,792]" italics="true" pageId="12" pageNumber="393">P. ceophallus</emphasis>
</taxonomicName>
, can still be distinguished from these on the basis of the form of its cervical groove, frontal median triangle and G1 (see also Ng, 1993). It is also the only member of the species complex known from Pulau Buton thus far.
</paragraph>
</subSubSection>
</treatment>
</document>