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<mods:title id="1478F859842DF3B13B79C2478BEF8495">A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895)</mods:title>
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<taxonomicName id="4C644D4079109369FF3ADEB149A1F95F" box="[151,467,1668,1694]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="klakahensis" status="sp. nov.">
<emphasis id="B910EAD179109369FF3ADEB149A1F95F" bold="true" box="[151,467,1668,1694]" italics="true" pageId="1" pageNumber="553">Cyrtodactylus klakahensis</emphasis>
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<paragraph id="8BDB36C379109369FF3ADE9349DDF8C5" blockId="1.[151,1086,1668,1796]" pageId="1" pageNumber="553">
English: Klakah bent-toed gecko; 
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: Klakah Bogenfinger-Gecko Chresonym: 
<taxonomicName id="4C644D4079109369FE8ADEFC4A54F920" box="[295,550,1737,1762]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="553" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD179109369FE8ADEFC4A54F920" box="[295,550,1737,1762]" italics="true" pageId="1" pageNumber="553">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
— 
<bibRefCitation id="EFF54B3279109369FDEEDEFF4A92F920" author="Mertens" box="[579,736,1737,1761]" pageId="1" pageNumber="567" refString="Mertens, R. (1934) Die Amphibien und Reptilien der Deutschen Limnologischen Sunda-Expedition. In: Thienemann- Festband, A. (Ed.), Tropische Binnengewasser. Archiv fur Hydrobiologie, 40, pp. 677 - 701." type="journal article" year="1934">Mertens 1934</bibRefCitation>
, 
<emphasis id="B910EAD179109369FD46DEFC4C74F923" box="[747,1030,1737,1762]" italics="true" pageId="1" pageNumber="553">Archiv für Hydrobiologie</emphasis>
: 689 
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–3; 
<tableCitation id="C6E6037879109369FE86DEDE490FF8C5" box="[299,381,1771,1796]" captionStart="TABLE 2" captionStartId="3.[151,244,151,176]" captionTargetBox="[151,1436,239,1258]" captionTargetPageId="3" captionText="TABLE 2. Metric (in mm) and meristic data for the type series of Cyrtodactylus klakahensis sp. nov. Abbreviations are defined in Table 1." httpUri="http://table.plazi.org/id/DF1B664B7912936BFF3AD8A24A45FF15" pageId="1" pageNumber="553" tableUuid="DF1B664B7912936BFF3AD8A24A45FF15">Table 2</tableCitation>
&amp; 
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.
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22476 (
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&amp; 3A–C; 
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), an adult male, collected in 1928 or 1929 by members of the 
<emphasis id="B910EAD179109369FF3ADF674A09F8AA" box="[151,635,1874,1899]" italics="true" pageId="1" pageNumber="553">Deutsche Limnologische Sunda-Expedition</emphasis>
at Klakah, Lumajang Regency, Jawa Timur Province, Java, 
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.
</paragraph>
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.
</emphasis>
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22477 (
<figureCitation id="135F2A4679109369FE73DF404A4DF84F" box="[478,575,1909,1934]" captionStart="FIGURE 2" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[303,1286,221,1902]" captionTargetId="figure@8.[285,1298,193,1927]" captionTargetPageId="8" captionText="FIGURE 2. Paratype series of Cyrtodactylus klakahensis sp. nov. in dorsal and ventral view. (A) SMF 22477, an adult male. (B) SMF 22478, an adult female. (C) SMF 22479, an unsexed juvenile specimen. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267814/files/figure.png" pageId="1" pageNumber="553">Figure 2</figureCitation>
A; 
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), an adult male; 
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22478 (
<figureCitation id="135F2A4679109369FBADDF404C13F84F" box="[1024,1121,1909,1934]" captionStart="FIGURE 2" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[303,1286,221,1902]" captionTargetId="figure@8.[285,1298,193,1927]" captionTargetPageId="8" captionText="FIGURE 2. Paratype series of Cyrtodactylus klakahensis sp. nov. in dorsal and ventral view. (A) SMF 22477, an adult male. (B) SMF 22478, an adult female. (C) SMF 22479, an unsexed juvenile specimen. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267814/files/figure.png" pageId="1" pageNumber="553">Figure 2</figureCitation>
B; 
<tableCitation id="C6E6037879109369FB2DDF404CA6F84F" box="[1152,1236,1909,1934]" captionStart="TABLE 2" captionStartId="3.[151,244,151,176]" captionTargetBox="[151,1436,239,1258]" captionTargetPageId="3" captionText="TABLE 2. Metric (in mm) and meristic data for the type series of Cyrtodactylus klakahensis sp. nov. Abbreviations are defined in Table 1." httpUri="http://table.plazi.org/id/DF1B664B7912936BFF3AD8A24A45FF15" pageId="1" pageNumber="553" tableUuid="DF1B664B7912936BFF3AD8A24A45FF15">Table 2</tableCitation>
), an adult female; and 
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22479 (
<figureCitation id="135F2A4679109369FEF6DFA249CEF871" box="[347,444,1943,1968]" captionStart="FIGURE 2" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[303,1286,221,1902]" captionTargetId="figure@8.[285,1298,193,1927]" captionTargetPageId="8" captionText="FIGURE 2. Paratype series of Cyrtodactylus klakahensis sp. nov. in dorsal and ventral view. (A) SMF 22477, an adult male. (B) SMF 22478, an adult female. (C) SMF 22479, an unsexed juvenile specimen. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267814/files/figure.png" pageId="1" pageNumber="553">Figure 2</figureCitation>
C; 
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), an unsexed juvenile. All specimens have the same collection information as the 
<typeStatus id="54DF886179109369FF6DDF8F4954F812" box="[192,294,1978,2003]" pageId="1" pageNumber="553" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF1B664B7913936AFF3AD8AD4B94FF6E" box="[151,998,152,175]" pageId="2" pageNumber="554">
<paragraph id="8BDB36C37913936AFF3AD8AD4B94FF6E" blockId="2.[151,998,152,175]" box="[151,998,152,175]" pageId="2" pageNumber="554">
<emphasis id="B910EAD17913936AFF3AD8AD4978FF6E" bold="true" box="[151,266,152,175]" pageId="2" pageNumber="554">TABLE 1.</emphasis>
Metric and meristic characters with abbreviations used in this study.
</paragraph>
</caption>
<paragraph id="8BDB36C37913936AFF0CD8F84B61FF22" blockId="2.[161,1424,205,1879]" box="[161,787,205,227]" pageId="2" pageNumber="554">Character Abbreviation Definition</paragraph>
<paragraph id="8BDB36C37913936AFF0CD8C94BC5FED3" blockId="2.[161,1424,205,1879]" box="[161,951,252,274]" pageId="2" pageNumber="554">Snout-vent length SVL from tip of snout to cloaca</paragraph>
<paragraph id="8BDB36C37913936AFF0CD91F4B03FE81" blockId="2.[161,1424,205,1879]" box="[161,881,298,320]" pageId="2" pageNumber="554">Axial length AxialL from axilla to groin</paragraph>
<paragraph id="8BDB36C37913936AFF0CD96C4CF1FEAE" blockId="2.[161,1424,205,1879]" box="[161,1155,345,367]" pageId="2" pageNumber="554">Tail length TailL length of original tail, from cloaca to tip of tail</paragraph>
<paragraph id="8BDB36C37913936AFF0CD9BD4B8DFDEB" blockId="2.[161,1424,205,1879]" pageId="2" pageNumber="554">Arm length ArmL from insertion of antebrachium with body wall to claw of longest finger Leg length LegL from insertion of femur with body wall to claw of longest toe Head length HeadL from tip of snout to articulation of quadrate bone Head width HeadW measured at level of ear openings</paragraph>
<paragraph id="8BDB36C37913936AFF0CDA774B87FD99" blockId="2.[161,1424,205,1879]" box="[161,1013,578,600]" pageId="2" pageNumber="554">Head height HeadH measured at level of ear opening</paragraph>
<paragraph id="8BDB36C37913936AFF0CDA444C16FD46" blockId="2.[161,1424,205,1879]" box="[161,1124,625,647]" pageId="2" pageNumber="554">Snout length SnoutL from tip of snout to anterior margin of orbit</paragraph>
<paragraph id="8BDB36C37913936AFF0CDA954C39FD25" blockId="2.[161,1424,205,1879]" pageId="2" pageNumber="554">Orbit-ear length OrbEarL from posterior margin of orbit to anterior margin of ear opening Orbital diameter OrbD from anterior to posterior margin of orbit</paragraph>
<paragraph id="8BDB36C37913936AFF0CDACB4C72FC83" blockId="2.[161,1424,205,1879]" pageId="2" pageNumber="554">Ear length EarL from anterior to posterior margin of ear opening Mental length MentalL maximum length of mental shield</paragraph>
<paragraph id="8BDB36C37913936AFF0CDB6F4B8EFCB1" blockId="2.[161,1424,205,1879]" box="[161,1020,858,880]" pageId="2" pageNumber="554">Mental width MentalW maximum width of mental shield</paragraph>
<paragraph id="8BDB36C37913936AFF0CDBBC4D29FC5E" blockId="2.[161,1424,205,1879]" box="[161,1371,905,927]" pageId="2" pageNumber="554">Dorsal tubercles DTR number of tubercle rows on dorsum at midbody, counted in one row</paragraph>
<paragraph id="8BDB36C37913936AFD06DB9D4B0CFC7F" blockId="2.[161,1424,205,1879]" box="[683,894,936,958]" pageId="2" pageNumber="554">between lateral folds</paragraph>
<paragraph id="8BDB36C37913936AFF0CDBE34D13FC2D" blockId="2.[161,1424,205,1879]" box="[161,1377,982,1004]" pageId="2" pageNumber="554">Paravertebral tubercles PVT number of tubercles counted in a longitudinal row between posterior</paragraph>
<paragraph id="8BDB36C37913936AFD06DBC04C97FBCA" blockId="2.[161,1424,205,1879]" box="[683,1253,1013,1035]" pageId="2" pageNumber="554">insertion of fore limb and anterior insertion of hind limb</paragraph>
<paragraph id="8BDB36C37913936AFF0CDC114DF8FBFB" blockId="2.[161,1424,205,1879]" box="[161,1418,1060,1082]" pageId="2" pageNumber="554">Ventral scales VS number of ventral scales at midbody, counted in one row between lateral</paragraph>
<paragraph id="8BDB36C37913936AFD06DC774AACFB99" blockId="2.[161,1424,205,1879]" box="[683,734,1090,1112]" pageId="2" pageNumber="554">folds</paragraph>
<paragraph id="8BDB36C37913936AFF0CDC444DE2FB46" blockId="2.[161,1424,205,1879]" box="[161,1424,1137,1159]" pageId="2" pageNumber="554">Precloacofemoral scales PFS number of enlarged precloacofemoral scales, counted along lowest, pore-</paragraph>
<paragraph id="8BDB36C37913936AFD06DCA54B56FB67" blockId="2.[161,1424,205,1879]" box="[683,804,1168,1190]" pageId="2" pageNumber="554">bearing row</paragraph>
<paragraph id="8BDB36C37913936AFF0CDC8B4C7AFB15" blockId="2.[161,1424,205,1879]" box="[161,1032,1214,1236]" pageId="2" pageNumber="554">Precloacofemoral pores PFP number of precloacofemoral pores</paragraph>
<paragraph id="8BDB36C37913936AFF0CDCD84B9FFAC2" blockId="2.[161,1424,205,1879]" box="[161,1005,1261,1283]" pageId="2" pageNumber="554">Postcloacal tubercles PCT number of postcloacal tubercles</paragraph>
<paragraph id="8BDB36C37913936AFF0CDD2B4D1EF9DD" blockId="2.[161,1424,205,1879]" pageId="2" pageNumber="554">Subdigital lamellae under 4th LT 4 subdigital scales under 4th toe, counted from first enlarged scale (true toe lamellae) on lower side of toe to scale proximal to apical scale Supralabial scales 1 SupraLab1 labial scales of upper jaw, beginning with first enlarged scale bordering rostral shield, ending with last enlarged scale bordering labial angle Supralabial scales 2 SupraLab2 labial scales of upper jaw, beginning with first enlarged scale bordering rostral shield, ending with enlarged scale below anterior margin of orbit Infralabial scales InfraLab labial scales of lower jaw, beginning with first scale bordering mental</paragraph>
<paragraph id="8BDB36C37913936AFD06DE104D09F959" blockId="2.[161,1424,205,1879]" pageId="2" pageNumber="554">shield, ending with last enlarged scale bordering labial angle Internasal scales InterNas number of scales between rostronasals, bordering rostral shield Supraciliar scales SC number of enlarged scales extending from anterior-ventral to posterior-</paragraph>
<paragraph id="8BDB36C37913936AFD06DE944B0AF976" blockId="2.[161,1424,205,1879]" box="[683,888,1697,1719]" pageId="2" pageNumber="554">medial edge of orbit</paragraph>
<paragraph id="8BDB36C37913936AFF0CDEFA4D13F924" blockId="2.[161,1424,205,1879]" box="[161,1377,1743,1765]" pageId="2" pageNumber="554">
Interorbital scales 
<collectionCode id="ED75AE067913936AFE7CDEFA4988F924" box="[465,506,1743,1765]" httpUri="http://grbio.org/cool/dcki-wrnq" name="Institute of Oceanographic Sciences" pageId="2" pageNumber="554">IOS</collectionCode>
number of scales counted in one row between medial edges of orbits
</paragraph>
<paragraph id="8BDB36C37913936AFD06DEDA4B4EF8C5" blockId="2.[161,1424,205,1879]" box="[683,828,1774,1796]" pageId="2" pageNumber="554">across occiput</paragraph>
<paragraph id="8BDB36C37913936AFF0CDF284D23F8F4" blockId="2.[161,1424,205,1879]" box="[161,1361,1821,1845]" pageId="2" pageNumber="554">Gular scales GulS number of gular scales bordering pair of 1st postmentals (excluding</paragraph>
<paragraph id="8BDB36C37913936AFD06DF744B85F896" blockId="2.[161,1424,205,1879]" box="[683,1015,1853,1879]" pageId="2" pageNumber="554">enlarged second 2nd postmentals)</paragraph>
<caption id="DF1B664B7912936BFF3AD8A24A45FF15" ID-Table-UUID="DF1B664B7912936BFF3AD8A24A45FF15" httpUri="http://table.plazi.org/id/DF1B664B7912936BFF3AD8A24A45FF15" pageId="3" pageNumber="555" targetBox="[151,1436,239,1258]" targetIsTable="true" targetPageId="3">
<paragraph id="8BDB36C37912936BFF3AD8A24A45FF15" blockId="3.[151,1436,151,213]" pageId="3" pageNumber="555">
<emphasis id="B910EAD17912936BFF3AD8A2496BFF71" bold="true" box="[151,281,151,176]" pageId="3" pageNumber="555">TABLE 2.</emphasis>
Metric (in mm) and meristic data for the type series of 
<taxonomicName id="4C644D407912936BFC55D8A24D5BFF71" box="[1016,1321,151,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="klakahensis" status="sp. nov.">
<emphasis id="B910EAD17912936BFC55D8A24D5BFF71" box="[1016,1321,151,176]" italics="true" pageId="3" pageNumber="555">Cyrtodactylus klakahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357AA7912936BFA94D8A24DEEFF71" box="[1337,1436,151,176]" pageId="3" pageNumber="555" rank="species">
<emphasis id="B910EAD17912936BFA94D8A24DEEFF71" bold="true" box="[1337,1436,151,176]" pageId="3" pageNumber="555">sp. nov.</emphasis>
</taxonomicNameLabel>
Abbreviations are defined in Table 1.
</paragraph>
</caption>
<paragraph id="8BDB36C37912936BFE03D8DA4CC6FB2B" pageId="3" pageNumber="555">
<table id="F964C46379126C97FF3AD8DA4DEEFB2B" box="[151,1436,239,1258]" gridcols="5" gridrows="16" pageId="3" pageNumber="555">
<tr id="3554348179126C97FF3AD8DA4DEEFEE2" box="[151,1436,239,291]" gridrow="0" pageId="3" pageNumber="555" rowspan-0="1">
<th id="76855DFD79126C97FE03D8DA4A54FEE2" box="[430,550,239,291]" gridcol="1" gridrow="0" pageId="3" pageNumber="555">Holotype SMF 22476</th>
<th id="76855DFD79126C97FD01D8DA4B56FEE2" box="[684,804,239,291]" gridcol="2" gridrow="0" pageId="3" pageNumber="555">Paratype SMF 22477</th>
<th id="76855DFD79126C97FC07D8DA4C51FEE2" box="[938,1059,239,291]" gridcol="3" gridrow="0" pageId="3" pageNumber="555">Paratype SMF 22478</th>
<th id="76855DFD79126C97FB0AD8DA4DEEFEE2" box="[1191,1436,239,291]" gridcol="4" gridrow="0" pageId="3" pageNumber="555">Paratype SMF 22479</th>
</tr>
<tr id="3554348179126C97FF3AD9004DEEFE8A" box="[151,1436,309,331]" gridrow="1" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3AD9004975FE8A" box="[151,263,309,331]" gridcol="0" gridrow="1" pageId="3" pageNumber="555">Sex</th>
<td id="76855DFD79126C97FE03D9004A54FE8A" box="[430,550,309,331]" gridcol="1" gridrow="1" pageId="3" pageNumber="555">male</td>
<td id="76855DFD79126C97FD01D9004B56FE8A" box="[684,804,309,331]" gridcol="2" gridrow="1" pageId="3" pageNumber="555">male</td>
<td id="76855DFD79126C97FC07D9004C51FE8A" box="[938,1059,309,331]" gridcol="3" gridrow="1" pageId="3" pageNumber="555">female</td>
<td id="76855DFD79126C97FB0AD9004DEEFE8A" box="[1191,1436,309,331]" gridcol="4" gridrow="1" pageId="3" pageNumber="555">unsexed juvenile</td>
</tr>
<tr id="3554348179126C97FF3AD9684DEEFEB2" box="[151,1436,349,371]" gridrow="2" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3AD9684975FEB2" box="[151,263,349,371]" gridcol="0" gridrow="2" pageId="3" pageNumber="555">SVL</th>
<td id="76855DFD79126C97FE03D9684A54FEB2" box="[430,550,349,371]" gridcol="1" gridrow="2" pageId="3" pageNumber="555">68.1</td>
<td id="76855DFD79126C97FD01D9684B56FEB2" box="[684,804,349,371]" gridcol="2" gridrow="2" pageId="3" pageNumber="555">61.1</td>
<td id="76855DFD79126C97FC07D9684C51FEB2" box="[938,1059,349,371]" gridcol="3" gridrow="2" pageId="3" pageNumber="555">67.2</td>
<td id="76855DFD79126C97FB0AD9684DEEFEB2" box="[1191,1436,349,371]" gridcol="4" gridrow="2" pageId="3" pageNumber="555">37.2</td>
</tr>
<tr id="3554348179126C97FF3AD9B04DEEFE03" box="[151,1436,389,450]" gridrow="3" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3AD9B04975FE03" box="[151,263,389,450]" gridcol="0" gridrow="3" pageId="3" pageNumber="555">AxialL ArmL</th>
<td id="76855DFD79126C97FE03D9B04A54FE03" box="[430,550,389,450]" gridcol="1" gridrow="3" pageId="3" pageNumber="555">31.9 25.9</td>
<td id="76855DFD79126C97FD01D9B04B56FE03" box="[684,804,389,450]" gridcol="2" gridrow="3" pageId="3" pageNumber="555">27.3 21.7</td>
<td id="76855DFD79126C97FC07D9B04C51FE03" box="[938,1059,389,450]" gridcol="3" gridrow="3" pageId="3" pageNumber="555">28.7 24.1</td>
<td id="76855DFD79126C97FB0AD9B04DEEFE03" box="[1191,1436,389,450]" gridcol="4" gridrow="3" pageId="3" pageNumber="555">14.0 13.6</td>
</tr>
<tr id="3554348179126C97FF3AD9E14DEEFE2B" box="[151,1436,468,490]" gridrow="4" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3AD9E14975FE2B" box="[151,263,468,490]" gridcol="0" gridrow="4" pageId="3" pageNumber="555">LegL</th>
<td id="76855DFD79126C97FE03D9E14A54FE2B" box="[430,550,468,490]" gridcol="1" gridrow="4" pageId="3" pageNumber="555">36.3</td>
<td id="76855DFD79126C97FD01D9E14B56FE2B" box="[684,804,468,490]" gridcol="2" gridrow="4" pageId="3" pageNumber="555">28.9</td>
<td id="76855DFD79126C97FC07D9E14C51FE2B" box="[938,1059,468,490]" gridcol="3" gridrow="4" pageId="3" pageNumber="555">34.0</td>
<td id="76855DFD79126C97FB0AD9E14DEEFE2B" box="[1191,1436,468,490]" gridcol="4" gridrow="4" pageId="3" pageNumber="555">16.8</td>
</tr>
<tr id="3554348179126C97FF3AD9C94DEEFDD3" box="[151,1436,508,530]" gridrow="5" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3AD9C94975FDD3" box="[151,263,508,530]" gridcol="0" gridrow="5" pageId="3" pageNumber="555">HeadL</th>
<td id="76855DFD79126C97FE03D9C94A54FDD3" box="[430,550,508,530]" gridcol="1" gridrow="5" pageId="3" pageNumber="555">18.1</td>
<td id="76855DFD79126C97FD01D9C94B56FDD3" box="[684,804,508,530]" gridcol="2" gridrow="5" pageId="3" pageNumber="555">16.8</td>
<td id="76855DFD79126C97FC07D9C94C51FDD3" box="[938,1059,508,530]" gridcol="3" gridrow="5" pageId="3" pageNumber="555">16.6</td>
<td id="76855DFD79126C97FB0AD9C94DEEFDD3" box="[1191,1436,508,530]" gridcol="4" gridrow="5" pageId="3" pageNumber="555">10.4</td>
</tr>
<tr id="3554348179126C97FF3ADA114DEEFDA0" box="[151,1436,548,609]" gridrow="6" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADA114975FDA0" box="[151,263,548,609]" gridcol="0" gridrow="6" pageId="3" pageNumber="555">HeadW HeadH</th>
<td id="76855DFD79126C97FE03DA114A54FDA0" box="[430,550,548,609]" gridcol="1" gridrow="6" pageId="3" pageNumber="555">10.6 7.0</td>
<td id="76855DFD79126C97FD01DA114B56FDA0" box="[684,804,548,609]" gridcol="2" gridrow="6" pageId="3" pageNumber="555">10.4 7.2</td>
<td id="76855DFD79126C97FC07DA114C51FDA0" box="[938,1059,548,609]" gridcol="3" gridrow="6" pageId="3" pageNumber="555">10.5 8.3</td>
<td id="76855DFD79126C97FB0ADA114DEEFDA0" box="[1191,1436,548,609]" gridcol="4" gridrow="6" pageId="3" pageNumber="555">6.1 4.2</td>
</tr>
<tr id="3554348179126C97FF3ADA464DEEFD48" box="[151,1436,627,649]" gridrow="7" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADA464975FD48" box="[151,263,627,649]" gridcol="0" gridrow="7" pageId="3" pageNumber="555">SnoutL</th>
<td id="76855DFD79126C97FE03DA464A54FD48" box="[430,550,627,649]" gridcol="1" gridrow="7" pageId="3" pageNumber="555">7.6</td>
<td id="76855DFD79126C97FD01DA464B56FD48" box="[684,804,627,649]" gridcol="2" gridrow="7" pageId="3" pageNumber="555">7.0</td>
<td id="76855DFD79126C97FC07DA464C51FD48" box="[938,1059,627,649]" gridcol="3" gridrow="7" pageId="3" pageNumber="555">7.9</td>
<td id="76855DFD79126C97FB0ADA464DEEFD48" box="[1191,1436,627,649]" gridcol="4" gridrow="7" pageId="3" pageNumber="555">4.2</td>
</tr>
<tr id="3554348179126C97FF3ADAAE4DEEFD70" box="[151,1436,667,689]" gridrow="8" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADAAE4975FD70" box="[151,263,667,689]" gridcol="0" gridrow="8" pageId="3" pageNumber="555">OrbEarL</th>
<td id="76855DFD79126C97FE03DAAE4A54FD70" box="[430,550,667,689]" gridcol="1" gridrow="8" pageId="3" pageNumber="555">5.9</td>
<td id="76855DFD79126C97FD01DAAE4B56FD70" box="[684,804,667,689]" gridcol="2" gridrow="8" pageId="3" pageNumber="555">5.6</td>
<td id="76855DFD79126C97FC07DAAE4C51FD70" box="[938,1059,667,689]" gridcol="3" gridrow="8" pageId="3" pageNumber="555">6.7</td>
<td id="76855DFD79126C97FB0ADAAE4DEEFD70" box="[1191,1436,667,689]" gridcol="4" gridrow="8" pageId="3" pageNumber="555">2.9</td>
</tr>
<tr id="3554348179126C97FF3ADAF64DEEFCC1" box="[151,1436,707,768]" gridrow="9" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADAF64975FCC1" box="[151,263,707,768]" gridcol="0" gridrow="9" pageId="3" pageNumber="555">OrbD EarL</th>
<td id="76855DFD79126C97FE03DAF64A54FCC1" box="[430,550,707,768]" gridcol="1" gridrow="9" pageId="3" pageNumber="555">3.7 1.3</td>
<td id="76855DFD79126C97FD01DAF64B56FCC1" box="[684,804,707,768]" gridcol="2" gridrow="9" pageId="3" pageNumber="555">3.4 1.3</td>
<td id="76855DFD79126C97FC07DAF64C51FCC1" box="[938,1059,707,768]" gridcol="3" gridrow="9" pageId="3" pageNumber="555">3.4 1.4</td>
<td id="76855DFD79126C97FB0ADAF64DEEFCC1" box="[1191,1436,707,768]" gridcol="4" gridrow="9" pageId="3" pageNumber="555">1.9 0.5</td>
</tr>
<tr id="3554348179126C97FF3ADB274DEEFCE9" box="[151,1436,786,808]" gridrow="10" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADB274975FCE9" box="[151,263,786,808]" gridcol="0" gridrow="10" pageId="3" pageNumber="555">DTR</th>
<td id="76855DFD79126C97FE03DB274A54FCE9" box="[430,550,786,808]" gridcol="1" gridrow="10" pageId="3" pageNumber="555">19</td>
<td id="76855DFD79126C97FD01DB274B56FCE9" box="[684,804,786,808]" gridcol="2" gridrow="10" pageId="3" pageNumber="555">17</td>
<td id="76855DFD79126C97FC07DB274C51FCE9" box="[938,1059,786,808]" gridcol="3" gridrow="10" pageId="3" pageNumber="555">17</td>
<td id="76855DFD79126C97FB0ADB274DEEFCE9" box="[1191,1436,786,808]" gridcol="4" gridrow="10" pageId="3" pageNumber="555">15</td>
</tr>
<tr id="3554348179126C97FF3ADB0F4DEEFC91" box="[151,1436,826,848]" gridrow="11" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADB0F4975FC91" box="[151,263,826,848]" gridcol="0" gridrow="11" pageId="3" pageNumber="555">PVT</th>
<td id="76855DFD79126C97FE03DB0F4A54FC91" box="[430,550,826,848]" gridcol="1" gridrow="11" pageId="3" pageNumber="555">26</td>
<td id="76855DFD79126C97FD01DB0F4B56FC91" box="[684,804,826,848]" gridcol="2" gridrow="11" pageId="3" pageNumber="555">29</td>
<td id="76855DFD79126C97FC07DB0F4C51FC91" box="[938,1059,826,848]" gridcol="3" gridrow="11" pageId="3" pageNumber="555">33</td>
<td id="76855DFD79126C97FB0ADB0F4DEEFC91" box="[1191,1436,826,848]" gridcol="4" gridrow="11" pageId="3" pageNumber="555">26</td>
</tr>
<tr id="3554348179126C97FF3ADB574DEEFC5E" box="[151,1436,866,927]" gridrow="12" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADB574975FC5E" box="[151,263,866,927]" gridcol="0" gridrow="12" pageId="3" pageNumber="555">VS PFS</th>
<td id="76855DFD79126C97FE03DB574A54FC5E" box="[430,550,866,927]" gridcol="1" gridrow="12" pageId="3" pageNumber="555">38 40</td>
<td id="76855DFD79126C97FD01DB574B56FC5E" box="[684,804,866,927]" gridcol="2" gridrow="12" pageId="3" pageNumber="555">36 38</td>
<td id="76855DFD79126C97FC07DB574C51FC5E" box="[938,1059,866,927]" gridcol="3" gridrow="12" pageId="3" pageNumber="555">36 38</td>
<td id="76855DFD79126C97FB0ADB574DEEFC5E" box="[1191,1436,866,927]" gridcol="4" gridrow="12" pageId="3" pageNumber="555">35 38</td>
</tr>
<tr id="3554348179126C97FF3ADB844DEEFC06" box="[151,1436,945,967]" gridrow="13" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADB844975FC06" box="[151,263,945,967]" gridcol="0" gridrow="13" pageId="3" pageNumber="555">PFP</th>
<td id="76855DFD79126C97FE03DB844A54FC06" box="[430,550,945,967]" gridcol="1" gridrow="13" pageId="3" pageNumber="555">37</td>
<td id="76855DFD79126C97FD01DB844B56FC06" box="[684,804,945,967]" gridcol="2" gridrow="13" pageId="3" pageNumber="555">38</td>
<td id="76855DFD79126C97FC07DB844C51FC06" box="[938,1059,945,967]" gridcol="3" gridrow="13" pageId="3" pageNumber="555">0</td>
<td id="76855DFD79126C97FB0ADB844DEEFC06" box="[1191,1436,945,967]" gridcol="4" gridrow="13" pageId="3" pageNumber="555">0</td>
</tr>
<tr id="3554348179126C97FF3ADBEC4DEEFB03" box="[151,1436,985,1218]" gridrow="14" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADBEC4975FB03" box="[151,263,985,1218]" gridcol="0" gridrow="14" pageId="3" pageNumber="555">LT 4 (proximal) LT 4 (distal) LT 4 SupraLab1 SupraLab2 InfraLab</th>
<td id="76855DFD79126C97FE03DBEC4A54FB03" box="[430,550,985,1218]" gridcol="1" gridrow="14" pageId="3" pageNumber="555">9 11 20 R10 L10 R6 L5 R9 L9</td>
<td id="76855DFD79126C97FD01DBEC4B56FB03" box="[684,804,985,1218]" gridcol="2" gridrow="14" pageId="3" pageNumber="555">7 10 17 R11 L10 R5 L5 R9 L10</td>
<td id="76855DFD79126C97FC07DBEC4C51FB03" box="[938,1059,985,1218]" gridcol="3" gridrow="14" pageId="3" pageNumber="555">7 10 17 R10 L11 R5 L6 R8 L10</td>
<td id="76855DFD79126C97FB0ADBEC4DEEFB03" box="[1191,1436,985,1218]" gridcol="4" gridrow="14" pageId="3" pageNumber="555">8 9 17 R11 L10 R5 L6 R9 L8</td>
</tr>
<tr id="3554348179126C97FF3ADCE14DEEFB2B" box="[151,1436,1236,1258]" gridrow="15" pageId="3" pageNumber="555">
<th id="76855DFD79126C97FF3ADCE14975FB2B" box="[151,263,1236,1258]" gridcol="0" gridrow="15" pageId="3" pageNumber="555">GulS</th>
<td id="76855DFD79126C97FE03DCE14A54FB2B" box="[430,550,1236,1258]" gridcol="1" gridrow="15" pageId="3" pageNumber="555">4</td>
<td id="76855DFD79126C97FD01DCE14B56FB2B" box="[684,804,1236,1258]" gridcol="2" gridrow="15" pageId="3" pageNumber="555">4</td>
<td id="76855DFD79126C97FC07DCE14C51FB2B" box="[938,1059,1236,1258]" gridcol="3" gridrow="15" pageId="3" pageNumber="555">4</td>
<td id="76855DFD79126C97FB0ADCE14DEEFB2B" box="[1191,1436,1236,1258]" gridcol="4" gridrow="15" pageId="3" pageNumber="555">4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDB36C37912936BFF6ADD1F4B36FA38" blockId="3.[151,1436,1322,2033]" pageId="3" pageNumber="555">
<emphasis id="B910EAD17912936BFF6ADD1F4934FA82" bold="true" box="[199,326,1322,1347]" pageId="3" pageNumber="555">Definition.</emphasis>
<taxonomicName id="4C644D407912936BFEE3DD1F4A0AFA82" box="[334,632,1322,1347]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="klakahensis">
<emphasis id="B910EAD17912936BFEE3DD1F4A0AFA82" box="[334,632,1322,1347]" italics="true" pageId="3" pageNumber="555">Cyrtodactylus klakahensis</emphasis>
</taxonomicName>
can be distinguished from all other congeners occuring in the Greater Sunda Islands (including Sulawesi) and Lesser Sunda Islands by the following combination of characters: (1) a deep precloacal groove in males, (2) three rows of enlarged precloacofemoral scales, of which the third row bears 37–38 pores in males, (3) three or four rows of posterior precloacal scales, (4) raised and strongly keeled dorsal tubercles in 15–19 rows at midbody, (5) an indistinct lateral fold, (6) 17–20 subdigital lamellae under the 4th toe, and (7) subcaudal scales which are not transversely enlarged.
</paragraph>
</subSubSection>
<subSubSection id="C37E654879129362FF6ADE31489DF8FE" lastPageId="10" lastPageNumber="562" pageId="3" pageNumber="555" type="description">
<paragraph id="8BDB36C37912936BFF6ADE314A5AF8FC" blockId="3.[151,1436,1322,2033]" pageId="3" pageNumber="555">
<emphasis id="B910EAD17912936BFF6ADE314A4DF9DC" bold="true" box="[199,575,1540,1565]" pageId="3" pageNumber="555">Comparison with other species.</emphasis>
The new species can be readily distinguished from the Greater Sunda Island congeners 
<taxonomicName id="4C644D407912936BFEBEDE1D4A82F980" authority="Iskandar et al., 2011" authorityName="Iskandar et al." authorityYear="2011" box="[275,752,1576,1601]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="batik">
<emphasis id="B910EAD17912936BFEBEDE1D4981F980" box="[275,499,1576,1601]" italics="true" pageId="3" pageNumber="555">Cyrtodactylus batik</emphasis>
<bibRefCitation id="EFF54B327912936BFE50DE1D4A82F980" author="Iskandar" box="[509,752,1576,1601]" pageId="3" pageNumber="567" refString="Iskandar, D. T., Rachmansah, A. &amp; Umilaela (2011) A new bent-toed gecko of the genus Cyrtodactylus Gray, 1827 (Reptilia, Gekkonidae) from Mount Tompotika, eastern peninsula of Sulawesi, Indonesia. Zootaxa, 2838, 65 - 78." type="journal article" year="2011">
Iskandar 
<emphasis id="B910EAD17912936BFDC5DE1F4AEFF980" box="[616,669,1576,1601]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
., 2011
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFD53DE1C4C28F980" authority="Peters, 1871" authorityName="Peters" authorityYear="1871" box="[766,1114,1576,1601]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="B910EAD17912936BFD53DE1C4B63F981" box="[766,785,1577,1600]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFC8EDE1D4BDEF980" box="[803,940,1576,1601]" italics="true" pageId="3" pageNumber="555">consobrinus</emphasis>
(
<bibRefCitation id="EFF54B327912936BFC13DE1C4C23F980" author="Peters" box="[958,1105,1577,1601]" pageId="3" pageNumber="567" refString="Peters, W. (1871) Uber neue Reptilien aus Ostafrika und Sarawak (Borneo), vorzuglich aus der Sammlung des Hrn. Marquis J. Doria zu Genua. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften, Berlin, Germany, 566 - 581." type="book chapter" year="1871">Peters, 1871</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFBC4DE1C4D06F980" authority="Hikida, 1990" authorityName="Hikida" authorityYear="1990" box="[1129,1396,1576,1601]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="B910EAD17912936BFBC4DE1C4C0EF981" box="[1129,1148,1577,1600]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFB20DE1F4CA2F980" box="[1165,1232,1578,1601]" italics="true" pageId="3" pageNumber="555">ingeri</emphasis>
<bibRefCitation id="EFF54B327912936BFB76DE1D4D06F980" author="Hikida" box="[1243,1396,1576,1601]" pageId="3" pageNumber="567" refString="Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology, 13 (3), 91 - 107." type="journal article" year="1990">Hikida, 1990</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFA2FDE1C49A4F9A4" authority="Boulenger, 1897" authorityName="Boulenger" authorityYear="1897" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="jellesmae">
<emphasis id="B910EAD17912936BFA2FDE1C4DE7F981" box="[1410,1429,1577,1600]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFF3ADE794973F9A4" box="[151,257,1612,1637]" italics="true" pageId="3" pageNumber="555">jellesmae</emphasis>
(
<bibRefCitation id="EFF54B327912936BFEA2DE7949BDF9A4" author="Boulenger" box="[271,463,1612,1637]" pageId="3" pageNumber="566" refString="Boulenger, G. A. (1897) A catalogue of the reptiles and batrachians of Celebes with special reference to the collections made by Drs P &amp; F Sarasin in 1893 - 1896. Proceedings of the Royal Society of London, 1897, 193 - 237." type="proceedings paper" year="1897">Boulenger, 1897</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFE4FDE784B66F9A4" authority="Werner, 1896" authorityName="Werner" authorityYear="1896" box="[482,788,1612,1637]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B910EAD17912936BFE4FDE784987F9A5" box="[482,501,1613,1636]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFDAEDE794A12F9A4" box="[515,608,1612,1637]" italics="true" pageId="3" pageNumber="555">lateralis</emphasis>
(
<bibRefCitation id="EFF54B327912936BFDC2DE784B7EF9A4" author="Werner" box="[623,780,1612,1637]" pageId="3" pageNumber="568" refString="Werner, F. (1896) Zweiter Beitrag zur Herpetologie der indo-orientalischen Region. Verhandlungen der Zoologisch- Botanischen Gesellschaft in Osterreich, 46, 6 - 24." type="journal article" year="1896">Werner, 1896</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFCB2DE784BCBF9A4" box="[799,953,1612,1637]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B910EAD17912936BFCB2DE784B40F9A5" box="[799,818,1613,1636]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFCEDDE794BCBF9A4" box="[832,953,1612,1637]" italics="true" pageId="3" pageNumber="555">malayanus</emphasis>
</taxonomicName>
(de 
<bibRefCitation id="EFF54B327912936BFC41DE794C01F9A4" author="Rooij" box="[1004,1139,1612,1637]" pageId="3" pageNumber="566" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Vol. I: Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, 384 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">Rooij, 1915</bibRefCitation>
), 
<taxonomicName id="4C644D407912936BFB25DE784DEAF9A4" authority="Hikida, 1990" authorityName="Hikida" authorityYear="1990" box="[1160,1432,1612,1637]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="matsuii">
<emphasis id="B910EAD17912936BFB25DE784CE9F9A5" box="[1160,1179,1613,1636]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFB04DE784C8EF9A5" box="[1193,1276,1613,1636]" italics="true" pageId="3" pageNumber="555">matsuii</emphasis>
<bibRefCitation id="EFF54B327912936BFAAEDE794DEAF9A4" author="Hikida" box="[1283,1432,1612,1637]" pageId="3" pageNumber="567" refString="Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology, 13 (3), 91 - 107." type="journal article" year="1990">Hikida, 1990</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFF3ADE444A7AF948" authority="Riyanto et al., 2014" authorityName="Riyanto et al." authorityYear="2014" box="[151,520,1648,1673]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="semiadii">
<emphasis id="B910EAD17912936BFF3ADE4448D8F949" box="[151,170,1649,1672]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFF17DE45496BF948" box="[186,281,1648,1673]" italics="true" pageId="3" pageNumber="555">semiadii</emphasis>
<bibRefCitation id="EFF54B327912936BFE8CDE454A7AF948" author="Riyanto" box="[289,520,1648,1673]" pageId="3" pageNumber="567" refString="Riyanto, A., Bauer, A. M. &amp; Yudha, D. S. (2014) A new small karst-dwelling species of Cyrtodactylus from Java, Indonesia. Zootaxa, 3785 (4), 589 - 599. http: // dx. doi. org / 10.11646 / zootaxa. 3785.4.7" type="journal article" year="2014">
Riyanto 
<emphasis id="B910EAD17912936BFE29DE4749C4F948" box="[388,438,1648,1673]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
., 2014
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFDBBDE444B08F948" authority="Taylor, 1962" authorityName="Taylor" authorityYear="1962" box="[534,890,1648,1673]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="B910EAD17912936BFDBBDE444A5BF949" box="[534,553,1649,1672]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFD94DE454AACF948" box="[569,734,1648,1673]" italics="true" pageId="3" pageNumber="555">quadrivirgatus</emphasis>
<bibRefCitation id="EFF54B327912936BFD4ADE454B08F948" author="Taylor" box="[743,890,1648,1673]" pageId="3" pageNumber="568" refString="Taylor, E. H. (1962) New oriental reptiles. The University of Kansas Science Bulletin, 43, 209 - 263." type="journal article" year="1962">Taylor, 1962</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFC2ADE444C85F948" authority="Hayden et al., 2008" authorityName="Hayden et al." authorityYear="2008" box="[903,1271,1648,1673]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="wallacei">
<emphasis id="B910EAD17912936BFC2ADE444BE8F949" box="[903,922,1649,1672]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFC07DE454C7BF948" box="[938,1033,1648,1673]" italics="true" pageId="3" pageNumber="555">wallacei</emphasis>
<bibRefCitation id="EFF54B327912936BFBBFDE454C85F948" author="Hayden" box="[1042,1271,1648,1673]" pageId="3" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Hayden 
<emphasis id="B910EAD17912936BFBDFDE474CD7F948" box="[1138,1189,1648,1673]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
., 2008
</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C644D407912936BFA9BDE44495FF96C" authority="Hikida, 1990" authorityName="Hikida" authorityYear="1990" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="yoshii">
<emphasis id="B910EAD17912936BFA9BDE444D3BF949" box="[1334,1353,1649,1672]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFAF4DE454DEEF948" box="[1369,1436,1648,1673]" italics="true" pageId="3" pageNumber="555">yoshii</emphasis>
<bibRefCitation id="EFF54B327912936BFF3ADEA1495FF96C" author="Hikida" box="[151,301,1684,1709]" pageId="3" pageNumber="567" refString="Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology, 13 (3), 91 - 107." type="journal article" year="1990">Hikida, 1990</bibRefCitation>
</taxonomicName>
, and from the Lesser Sunda Island species 
<taxonomicName id="4C644D407912936BFC8DDEA04CFCF96C" authority="Weber, 1890" authorityName="Weber" authorityYear="1890" box="[800,1166,1684,1709]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="darmandvillei">
<emphasis id="B910EAD17912936BFC8DDEA04B41F96D" box="[800,819,1685,1708]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFCEEDEA14B92F96C" box="[835,992,1684,1709]" italics="true" pageId="3" pageNumber="555">darmandvillei</emphasis>
(
<bibRefCitation id="EFF54B327912936BFC5CDEA14CF4F96C" author="Weber" box="[1009,1158,1684,1709]" pageId="3" pageNumber="568" refString="Weber, M. (1890) Reptilia from the Malay Archipelago. I. Sauria, Crocodylidae, Chelonia. In: Weber, M. (Ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, Vol. I. E. J. Brill. Leiden, pp. 158 - 177." type="book chapter" year="1890">Weber, 1890</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFB36DEA048A8F910" authority="Das, 1993" authorityName="Das" authorityYear="1993" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="gordongekkoi">
<emphasis id="B910EAD17912936BFB36DEA04CDCF96D" box="[1179,1198,1685,1708]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFB13DEA14D25F96C" box="[1214,1367,1684,1709]" italics="true" pageId="3" pageNumber="555">gordongekkoi</emphasis>
(
<bibRefCitation id="EFF54B327912936BFAC5DEA048A0F910" author="Das" pageId="3" pageNumber="566" refString="Das, I. (1993) Cnemaspis gordongekkoi, a new gecko from Lombok, Indonesia, and the biogeography of oriental species of Cnemaspis (Squamata: Sauria: Gekkonidae). Hamadryad, 18, 1 - 9." type="journal article" year="1993">Das, 1993</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFF4BDE8C4A41F910" authority="Darevsky, 1964" authorityName="Darevsky" authorityYear="1964" box="[230,563,1720,1745]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="laevigatus">
<emphasis id="B910EAD17912936BFF4BDE8C488BF911" box="[230,249,1721,1744]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFEABDE8D490BF910" box="[262,377,1720,1745]" italics="true" pageId="3" pageNumber="555">laevigatus</emphasis>
Darevsky, 1964
</taxonomicName>
, and 
<taxonomicName id="4C644D407912936BFDC0DE8C4BD9F910" authority="Dunn, 1927" authorityName="Dunn" authorityYear="1927" box="[621,939,1721,1745]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="wetariensis">
<emphasis id="B910EAD17912936BFDC0DE8C4AF2F911" box="[621,640,1721,1744]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFD23DE8F4B7CF910" box="[654,782,1722,1745]" italics="true" pageId="3" pageNumber="555">wetariensis</emphasis>
(
<bibRefCitation id="EFF54B327912936BFCB1DE8C4BD1F910" author="Dunn" box="[796,931,1721,1745]" pageId="3" pageNumber="566" refString="Dunn, E. (1927) Results of the Douglas Burden Expedition to the island of Komodo - III, Lizards from the East Indies. American Museum Novitates, 288, 2 - 3." type="journal article" year="1927">Dunn, 1927</bibRefCitation>
)
</taxonomicName>
by the presence of a deep precloacal groove in males, bearing five pores. Data in 
<tableCitation id="C6E603787912936BFDF2DEE94ACAF934" box="[607,696,1756,1781]" captionStart="TABLE 3" captionStartId="6.[125,203,673,693]" captionTargetBox="[125,1910,727,914]" captionTargetPageId="6" captionText="TABLE 3 (continued)" httpUri="http://table.plazi.org/id/DF1B664B7917936EFFD0DA944939FD74" pageId="3" pageNumber="555" tableUuid="DF1B664B7917936EFFD0DA944939FD74">Table 3</tableCitation>
allow a detailed comparison of 
<taxonomicName id="4C644D407912936BFBE9DEE84C9FF934" box="[1092,1261,1756,1781]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="klakahensis">
<emphasis id="B910EAD17912936BFBE9DEE84C9FF934" box="[1092,1261,1756,1781]" italics="true" pageId="3" pageNumber="555">C. klakahensis</emphasis>
</taxonomicName>
with all other Sundanese bent-toed geckos, and additional characters to distinguish the new species from taxa without a precloacal groove are listed therein.
</paragraph>
<paragraph id="8BDB36C37912936CFF6ADF7C4C0BFA48" blockId="3.[151,1436,1322,2033]" lastBlockId="4.[151,1437,151,1417]" lastPageId="4" lastPageNumber="556" pageId="3" pageNumber="555">
In the following comparisons with species that also possess a precloacal groove, including 
<taxonomicName id="4C644D407912936BFB52DF7D49A2F844" authority="Bleeker, 1860" authorityName="Bleeker" authorityYear="1860" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="agamensis">
<emphasis id="B910EAD17912936BFB52DF7D497FF844" italics="true" pageId="3" pageNumber="555">Cyrtodactylus agamensis</emphasis>
(
<bibRefCitation id="EFF54B327912936BFE8CDF5949BAF844" author="Bleeker" box="[289,456,1900,1925]" pageId="3" pageNumber="566" refString="Bleeker, P. (1860) Reptilien van Agam. Natuurkundig Tijdschrift voor Nederlandsch Indie, 20, 325 - 329." type="journal article" year="1860">Bleeker, 1860</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFE4CDF584B23F844" authority="Mocquard, 1890" authorityName="Mocquard" authorityYear="1890" box="[481,849,1900,1925]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B910EAD17912936BFE4CDF584986F845" box="[481,500,1901,1924]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFDAADF594A03F844" box="[519,625,1900,1925]" italics="true" pageId="3" pageNumber="555">baluensis</emphasis>
(
<bibRefCitation id="EFF54B327912936BFD28DF594B38F844" author="Mocquard" box="[645,842,1900,1925]" pageId="3" pageNumber="567" refString="Mocquard, F. (1890) Diagnoses d'especes nouvelles de reptiles et de batraciens des iles Borneo et Palawan. Le Naturaliste, Paris, France, 144 - 145." type="book chapter" year="1890">Mocquard, 1890</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFCCEDF584D67F844" authority="Inger &amp; King, 1961" authorityName="Inger &amp; King" authorityYear="1961" box="[867,1301,1900,1925]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="cavernicolus">
<emphasis id="B910EAD17912936BFCCEDF584B04F845" box="[867,886,1901,1924]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFC24DF594C6AF844" box="[905,1048,1900,1925]" italics="true" pageId="3" pageNumber="555">cavernicolus</emphasis>
<bibRefCitation id="EFF54B327912936BFB88DF584D67F844" author="Inger" box="[1061,1301,1900,1925]" pageId="3" pageNumber="567" refString="Inger, R. F. &amp; King, W. (1961) A new cave-dwelling lizard of the genus Cyrtodactylus from Niah. Sarawak Museum Journal, 11, 274 - 276." type="journal article" year="1961">Inger &amp; King, 1961</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFA8BDF5849FDF868" authority="Kathriner et al., 2014" authorityName="Kathriner et al." authorityYear="2014" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="celatus">
<emphasis id="B910EAD17912936BFA8BDF584D4BF845" box="[1318,1337,1901,1924]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFAE0DF594DEEF844" box="[1357,1436,1900,1925]" italics="true" pageId="3" pageNumber="555">celatus</emphasis>
<bibRefCitation id="EFF54B327912936BFF3ADFA549FDF868" author="Kathriner" box="[151,399,1936,1961]" pageId="3" pageNumber="567" refString="Kathriner, A., Bauer, A. M., O'Shea, M., Sanchez, C. &amp; Kaiser, H. (2014) Hiding in plain sight: a new species of bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from West Timor, collected by Malcolm Smith in 1924. Zootaxa, 3900 (4), 555 - 568. http: // dx. doi. org / 10.11646 / zootaxa. 3900.4.6" type="journal article" year="2014">
Kathriner 
<emphasis id="B910EAD17912936BFEA6DFA7494FF868" box="[267,317,1936,1961]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
., 2014
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFE31DFA44A69F868" box="[412,539,1937,1961]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17912936BFE31DFA449DDF869" box="[412,431,1937,1960]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFE12DFA44A69F868" box="[447,539,1937,1961]" italics="true" pageId="3" pageNumber="555">fumosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFD87DFA44AA7F868" box="[554,725,1937,1961]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17912936BFD87DFA44A4FF869" box="[554,573,1937,1960]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFDE1DFA74AA7F868" box="[588,725,1938,1961]" italics="true" pageId="3" pageNumber="555">marmoratus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFD48DFA44C33F868" authority="Harvey et al. 2015" authorityName="Harvey et al." authorityYear="2015" box="[741,1089,1936,1961]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="psarops">
<emphasis id="B910EAD17912936BFD48DFA44A8AF869" box="[741,760,1937,1960]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFCA5DFA74B12F868" box="[776,864,1938,1961]" italics="true" pageId="3" pageNumber="555">psarops</emphasis>
<bibRefCitation id="EFF54B327912936BFCC5DFA44C33F868" author="Harvey" box="[872,1089,1936,1961]" pageId="3" pageNumber="566" refString="Harvey, M. B., O'Connell, K. A., Barraza, G., Riyanto, A., Kurniawan, N. &amp; Smith, E. N. (2015) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the southern Bukit Barisan range of Sumatra and an estimation of their phylogeny. Zootaxa, 4020 (3), 495 - 516. http: // dx. doi. org / 10.11646 / zootaxa. 4020.3.5" type="journal article" year="2015">
Harvey 
<emphasis id="B910EAD17912936BFC6EDFA74B84F868" box="[963,1014,1936,1961]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
. 2015
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFBE3DFA44D07F868" authority="Inger, 1958" authorityName="Inger" authorityYear="1958" box="[1102,1397,1936,1961]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="pubisulcus">
<emphasis id="B910EAD17912936BFBE3DFA44C13F869" box="[1102,1121,1937,1960]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFBDDDFA54C9AF868" box="[1136,1256,1936,1961]" italics="true" pageId="3" pageNumber="555">pubisulcus</emphasis>
<bibRefCitation id="EFF54B327912936BFB5CDFA44D07F868" author="Inger" box="[1265,1397,1936,1961]" pageId="3" pageNumber="567" refString="Inger, R. F. (1958) A new gecko of the genus Cyrtodactylus, with a key to the species from Borneo and the Philippine Islands. Sarawak Museum Journal, 8, 261 - 264." type="journal article" year="1958">Inger, 1958</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C644D407912936BFA2FDFA44981F80C" authority="Harvey et al. 2015" authorityName="Harvey et al." authorityYear="2015" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="semicinctus">
<emphasis id="B910EAD17912936BFA2FDFA44DE7F869" box="[1410,1429,1937,1960]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFF3ADF804968F80D" box="[151,282,1973,1996]" italics="true" pageId="3" pageNumber="555">semicinctus</emphasis>
<bibRefCitation id="EFF54B327912936BFE8DDF804981F80C" author="Harvey" box="[288,499,1972,1997]" pageId="3" pageNumber="566" refString="Harvey, M. B., O'Connell, K. A., Barraza, G., Riyanto, A., Kurniawan, N. &amp; Smith, E. N. (2015) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the southern Bukit Barisan range of Sumatra and an estimation of their phylogeny. Zootaxa, 4020 (3), 495 - 516. http: // dx. doi. org / 10.11646 / zootaxa. 4020.3.5" type="journal article" year="2015">
Harvey 
<emphasis id="B910EAD17912936BFED4DF8349D8F80C" box="[377,426,1972,1997]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
. 2015
</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C644D407912936BFD81DF804BE0F80C" authority="Linkem et al., 2008" authorityName="Linkem et al." authorityYear="2008" box="[556,914,1972,1997]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B910EAD17912936BFD81DF804A4DF80D" box="[556,575,1973,1996]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFDE0DF804ADFF80D" box="[589,685,1973,1996]" italics="true" pageId="3" pageNumber="555">spinosus</emphasis>
<bibRefCitation id="EFF54B327912936BFD19DF814BE0F80C" author="Linkem" box="[692,914,1972,1997]" pageId="3" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Linkem 
<emphasis id="B910EAD17912936BFCBFDF834B31F80C" box="[786,835,1972,1997]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
., 2008
</bibRefCitation>
</taxonomicName>
, the characters for 
<taxonomicName id="4C644D407912936BFBCBDF804D7BF80C" box="[1126,1289,1972,1997]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="klakahensis">
<emphasis id="B910EAD17912936BFBCBDF804C0BF80D" box="[1126,1145,1973,1996]" italics="true" pageId="3" pageNumber="555">C</emphasis>
. 
<emphasis id="B910EAD17912936BFB2BDF814D7BF80C" box="[1158,1289,1972,1997]" italics="true" pageId="3" pageNumber="555">klakahensis</emphasis>
</taxonomicName>
are provided in parentheses. 
<taxonomicName id="4C644D407912936BFEE5DFED4A16F830" box="[328,612,2008,2033]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="555" phylum="Chordata" rank="species" species="agamensis">
<emphasis id="B910EAD17912936BFEE5DFED4A16F830" box="[328,612,2008,2033]" italics="true" pageId="3" pageNumber="555">Cyrtodactylus agamensis</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="EFF54B327912936BFD43DFED4BCDF830" author="Rosler" box="[750,959,2008,2033]" pageId="3" pageNumber="567" refString="Rosler, H., Richards, S. J. &amp; Gunther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae). Salamandra, 43 (4), 193 - 230." type="journal article" year="2007">
Rösler 
<emphasis id="B910EAD17912936BFCECDFEF4B01F830" box="[833,883,2008,2033]" italics="true" pageId="3" pageNumber="555">et al</emphasis>
. 2007
</bibRefCitation>
, based on the single known specimen, an adult female) possesses 67 (35–38) VS; 26 (17–20) LT4; 54 enlarged PFS (38–40); and a single enlarged scale in the precloacal groove (five scales in the precloacal groove). 
<taxonomicName id="4C644D407915936CFCEAD88B4C25FF16" box="[839,1111,190,215]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B910EAD17915936CFCEAD88B4C25FF16" box="[839,1111,190,215]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus baluensis</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="EFF54B327915936CFB73D88B4D19FF16" author="Hikida" box="[1246,1387,190,215]" pageId="4" pageNumber="567" refString="Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology, 13 (3), 91 - 107." type="journal article" year="1990">Hikida 1990</bibRefCitation>
and obtained from specimens listed in the Appendix) possesses precloacal scales that are separated from the femoral scales (enlarged PFS in both sexes); 4–10 precloacal- and 9–11 femoral pores in males (37–38 PFP in a continuous series in males); no posterior precloacal scales (posterior precloacal scales present); and enlarged subcaudals (enlarged subcaudals absent). 
<taxonomicName id="4C644D407915936CFE59D97B4B59FEA6" box="[500,811,334,359]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="cavernicolus">
<emphasis id="B910EAD17915936CFE59D97B4B59FEA6" box="[500,811,334,359]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus cavernicolus</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="EFF54B327915936CFC17D97B4CBCFEA6" author="Grismer" box="[954,1230,334,360]" pageId="4" pageNumber="566" refString="Grismer, L. L. &amp; Leong, T. M. (2005) New species of Cyrtodactylus (Squamata: Gekkonidae) from southern Peninsular Malaysia. Journal of Herpetology, 39 (4), 584 - 591. http: // dx. doi. org / 10.1670 / 43 - 05 A. 1" type="journal article" year="2005">Grismer &amp; Leong 2005</bibRefCitation>
) possesses 51–58 (35–38) VS; 22–26 (17–20) LT4; no enlarged femoral scales (enlarged PFS in three rows present in both sexes); no femoral pores (pores, including those on the femur, present in males); and dorsal bands (dorsal blotches). 
<taxonomicName id="4C644D407915936CFF3AD98849FEFE17" box="[151,396,445,470]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="celatus">
<emphasis id="B910EAD17915936CFF3AD98849FEFE17" box="[151,396,445,470]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus celatus</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="EFF54B327915936CFDB8D9884B75FE17" author="Kathriner" box="[533,775,445,470]" pageId="4" pageNumber="567" refString="Kathriner, A., Bauer, A. M., O'Shea, M., Sanchez, C. &amp; Kaiser, H. (2014) Hiding in plain sight: a new species of bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from West Timor, collected by Malcolm Smith in 1924. Zootaxa, 3900 (4), 555 - 568. http: // dx. doi. org / 10.11646 / zootaxa. 3900.4.6" type="journal article" year="2014">
Kathriner 
<emphasis id="B910EAD17915936CFD24D98B4ACEFE17" box="[649,700,445,470]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. 2014
</bibRefCitation>
; Rösler &amp; Kaiser, in press.; and obtained from specimens listed in the Appendix) is a small-sized species with adult SVL of 
<quantity id="4C9C9B267915936CFCDAD9D44C69FE38" box="[887,1051,481,506]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="4.36" metricValueMin="3.84" pageId="4" pageNumber="556" unit="mm" value="41.0" valueMax="43.6" valueMin="38.4">38.4−43.6 mm</quantity>
(
<quantity id="4C9C9B267915936CFB84D9D44CBDFE3B" box="[1065,1231,481,506]" metricMagnitude="-2" metricUnit="m" metricValue="6.459999999999999" metricValueMax="6.81" metricValueMin="6.11" pageId="4" pageNumber="556" unit="mm" value="64.6" valueMax="68.1" valueMin="61.1">61.1−68.1 mm</quantity>
) that possesses no femoral pores (pores, including those on the femur, present in males). 
<taxonomicName id="4C644D407915936CFC19DA304CC7FDDF" box="[948,1205,517,542]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17915936CFC19DA304CC7FDDF" box="[948,1205,517,542]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
(data obtained from specimens listed in the Appendix) possesses widely scattered, roundish, flat, and smooth dorsal tubercles in 4–7 rows at midbody (closely arranged, trihedral, raised, and strongly keeled dorsal tubercles in 15–19 rows at midbody); no or a low number of scattered tubercles on the limbs (limbs strongly tuberculated); a total number of 16 pores, ten of which are precloacal pores, separated from three femoral pores by ten enlarged pore-less scales 
<quantity id="4C9C9B267915936CFADADAA14DEEFD70" box="[1399,1436,660,689]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="556" unit="in" value="1.0">1 in</quantity>
males (37–38 PFP in a continuous series in males); and an ear opening forming a horizontal cleft2 (ear opening vertically elongated). 
<taxonomicName id="4C644D407915936CFE21DAD64AC8FD3A" box="[396,698,739,764]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17915936CFE21DAD64AC8FD3A" box="[396,698,739,764]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus marmoratus</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="EFF54B327915936CFC93DAD64C74FD3A" author="Rosler" box="[830,1030,739,764]" pageId="4" pageNumber="567" refString="Rosler, H., Richards, S. J. &amp; Gunther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae). Salamandra, 43 (4), 193 - 230." type="journal article" year="2007">
Rösler 
<emphasis id="B910EAD17915936CFC22DAD14BB2FD3D" box="[911,960,739,764]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. 2007
</bibRefCitation>
and obtained from specimens listed in the Appendix) possesses 38–47 (35–38) VS; 48–56 3 (38–40) enlarged PFS; 45–53 4 (37–38) PFP in males; and pores present in females (no pores present in females). Based on our examinations, 
<taxonomicName id="4C644D407915936CFBDFDB1B4D6DFC87" box="[1138,1311,814,838]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17915936CFBDFDB1B4CF7FC84" box="[1138,1157,814,837]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFB3BDB1A4D6DFC87" box="[1174,1311,815,838]" italics="true" pageId="4" pageNumber="556">marmoratus</emphasis>
</taxonomicName>
also lacks posterior precloacal scales (posterior precloacal scales present), possesses a different arrangement of postmental and gular scales, and has differently shaped postcloacal tubercles. The reader is referred to Figure 3 for a comparison of the shape and arrangement of dorsal tubercles at midbody, the precloacal region in males, and the postmental and gular scale pattern between 
<taxonomicName id="4C644D407915936CFDD2DB8B4B51FC17" box="[639,803,957,982]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="klakahensis">
<emphasis id="B910EAD17915936CFDD2DB8B4AE0FC14" box="[639,658,958,981]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFD0DDB884B51FC17" box="[672,803,957,982]" italics="true" pageId="4" pageNumber="556">klakahensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C644D407915936CFC9DDB8B4BDFFC17" box="[816,941,958,982]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17915936CFC9DDB8B4B31FC14" box="[816,835,958,981]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFCFCDB8B4BDFFC17" box="[849,941,958,982]" italics="true" pageId="4" pageNumber="556">fumosus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C644D407915936CFC44DB8B4CE1FC17" box="[1001,1171,958,982]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17915936CFC44DB8B4B8EFC14" box="[1001,1020,958,981]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFBA7DB8A4CE1FC17" box="[1034,1171,959,982]" italics="true" pageId="4" pageNumber="556">marmoratus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C644D407915936CFB0DDB884DEEFC17" box="[1184,1436,957,982]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="psarops">
<emphasis id="B910EAD17915936CFB0DDB884DEEFC17" box="[1184,1436,957,982]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus psarops</emphasis>
</taxonomicName>
possesses weakly keeled dorsal tubercles in 23–26 rows at midbody (strongly keeled tubercles in 15–19 rows at midbody); tubercles often present on the brachium (tubercles on brachium absent); 38–49 (35–38) VS; 28–32 (37– 38) PFP in males; a single, greatly enlarged, pore-bearing scale at the apex of the pore-bearing scale series (apical scale of pore-series not greatly enlarged); no posterior precloacal scales (posterior precloacal scales present); and a single (2–3) PCT. 
<taxonomicName id="4C644D407915936CFED4DC474AEFFB4A" box="[377,669,1138,1163]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="pubisulcus">
<emphasis id="B910EAD17915936CFED4DC474AEFFB4A" box="[377,669,1138,1163]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus pubisulcus</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="EFF54B327915936CFC98DC474BBEFB4B" author="Hikida" box="[821,972,1138,1163]" pageId="4" pageNumber="567" refString="Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology, 13 (3), 91 - 107." type="journal article" year="1990">Hikida 1990</bibRefCitation>
; 
<bibRefCitation id="EFF54B327915936CFC73DC464CDFFB4B" author="Das" box="[990,1197,1138,1163]" pageId="4" pageNumber="566" refString="Das, I. &amp; Jim, L. L. (2000) A new species of Cyrtodactylus (Sauria: Gekkonidae) from Pulau Tioman, Malaysia. The Raffles Bulletin of Zoology, 48 (2), 223 - 231." type="journal article" year="2000">Das &amp; Jim 2000</bibRefCitation>
; and obtained from specimens listed in the Appendix) possesses 37–58 (35–38) VS; no enlarged femoral scales (enlarged PFS present); no femoral pores (pores, including those on the femur, present in males); and no posterior precloacal scales (precloacal scales present). 
<taxonomicName id="4C644D407915936CFE7CDCE84A88FB37" box="[465,762,1245,1270]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="semicinctus">
<emphasis id="B910EAD17915936CFE7CDCE84A88FB37" box="[465,762,1245,1270]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus semicinctus</emphasis>
</taxonomicName>
possesses weakly keeled dorsal tubercles in 24–27 rows at midbody (strongly keeled tubercles in 15–19 rows at midbody); and a single, greatly enlarged, pore-bearing scale at apex of the pore-bearing scale series (apical scale of pore-series not greatly enlarged). 
<taxonomicName id="4C644D407915936CFB38DD104DEEFAFF" box="[1173,1436,1317,1342]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B910EAD17915936CFB38DD104DEEFAFF" box="[1173,1436,1317,1342]" italics="true" pageId="4" pageNumber="556">Cyrtodactylus spinosus</emphasis>
</taxonomicName>
(data from 
<bibRefCitation id="EFF54B327915936CFEB7DD794989FAA4" author="Linkem" box="[282,507,1356,1381]" pageId="4" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Linkem 
<emphasis id="B910EAD17915936CFED6DD7B49DDFAA4" box="[379,431,1356,1381]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. 2008
</bibRefCitation>
) possesses 38–44 5 (35–38) VS; no femoral pores (pores, including those on the femur, present in males); and lateral and caudal spines (lateral and caudal spines absent).
</paragraph>
<paragraph id="8BDB36C37915936CFF3ADDE44AD3F91C" blockId="4.[151,1437,1489,2033]" pageId="4" pageNumber="556">
1. 
<bibRefCitation id="EFF54B327915936CFF6ADDE44909FA26" author="Boulenger" box="[199,379,1489,1511]" pageId="4" pageNumber="566" refString="Boulenger, G. A. (1897) A catalogue of the reptiles and batrachians of Celebes with special reference to the collections made by Drs P &amp; F Sarasin in 1893 - 1896. Proceedings of the Royal Society of London, 1897, 193 - 237." type="proceedings paper" year="1897">Boulenger (1897)</bibRefCitation>
provided a count of 42 PFP for 
<taxonomicName id="4C644D407915936CFD67DDE74B4EFA26" box="[714,828,1489,1511]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17915936CFD67DDE74AA9FA26" box="[714,731,1490,1511]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFD44DDE44B4EFA26" box="[745,828,1489,1511]" italics="true" pageId="4" pageNumber="556">fumosus</emphasis>
</taxonomicName>
. We re-examined the material used by Boulenger, which is clearly conspecific with the 
<typeStatus id="54DF88617915936CFE58DDC44A53F9C7" box="[501,545,1521,1542]" pageId="4" pageNumber="556">type</typeStatus>
specimen housed in NMB, and found that his single adult male specimen (BMNH 1896.12.9.3, from Rurukan, North Sulawesi) does not possess a continuous series of PFP. De 
<bibRefCitation id="EFF54B327915936CFB0DDE3A4D56F9E4" author="Rooij" box="[1184,1316,1551,1573]" pageId="4" pageNumber="566" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Vol. I: Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, 384 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">Rooij (1915)</bibRefCitation>
provided a count of 42–52 PFP, but included data of specimens from Sulawesi (
<taxonomicName id="4C644D407915936CFCDADE1B4B95F982" box="[887,999,1581,1603]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17915936CFCDADE1B4BFAF982" box="[887,904,1582,1603]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFC39DE184B95F982" box="[916,999,1581,1603]" italics="true" pageId="4" pageNumber="556">fumosus</emphasis>
</taxonomicName>
), Halmahera (identified as 
<taxonomicName id="4C644D407915936CFB56DE1B4DEEF982" box="[1275,1436,1581,1603]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="philippinicus">
<emphasis id="B910EAD17915936CFB56DE1B4D7EF982" box="[1275,1292,1582,1603]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFAB5DE184DEEF982" box="[1304,1436,1581,1603]" italics="true" pageId="4" pageNumber="556">philippinicus</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFF54B327915936CFF61DE7949E6F9A3" author="Steindachner" box="[204,404,1612,1634]" pageId="4" pageNumber="568" refString="Steindachner, F. (1867) Reptilien. In: Steindachner, F. (Ed.), Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair, Zoologischer Theil, Vol. 1, part 3. K. Gerold's Sohn / Kaiserlich-Konigliche Hof- und Staatsdruckerei, Vienna, pp. 1 - 98." type="book chapter" year="1867">Steindachner, 1867</bibRefCitation>
] by 
<bibRefCitation id="EFF54B327915936CFE6FDE784A16F9A3" author="Boettger" box="[450,612,1612,1634]" pageId="4" pageNumber="566" refString="Boettger, O. (1900) Die Reptilien und Batrachier. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 25, 325 - 400." type="journal article" year="1900">Boettger [1900]</bibRefCitation>
and subsequently described as a new taxon currently known as 
<taxonomicName id="4C644D407915936CFB59DE784DEEF9A3" box="[1268,1436,1612,1634]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="halmahericus">
<emphasis id="B910EAD17915936CFB59DE784D77F9A3" box="[1268,1285,1613,1634]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFABFDE794DEEF9A3" box="[1298,1436,1612,1634]" italics="true" pageId="4" pageNumber="556">halmahericus</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFF54B327915936CFF60DE59491BF940" author="Mertens" box="[205,361,1643,1665]" pageId="4" pageNumber="567" refString="Mertens, R. (1929) Zwei neue Haftzeher aus dem Indo-Australischen Archipel (Rept.). Senckenbergiana, 11, 237 - 241." type="journal article" year="1929">Mertens, 1929</bibRefCitation>
]), and Java (misidentified 
<taxonomicName id="4C644D407915936CFD2CDE594B6FF940" box="[641,797,1644,1665]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17915936CFD2CDE594AE0F940" box="[641,658,1644,1665]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFD0CDE594B6FF940" box="[673,797,1644,1665]" italics="true" pageId="4" pageNumber="556">marmoratus</emphasis>
</taxonomicName>
) in her definition of 
<taxonomicName id="4C644D407915936CFBADDE594C01F940" box="[1024,1139,1643,1665]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17915936CFBADDE594C63F940" box="[1024,1041,1644,1665]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFB8DDE5E4C01F940" box="[1056,1139,1643,1665]" italics="true" pageId="4" pageNumber="556">fumosus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFF54B327915936CFB02DE594D18F940" author="Brongersma" box="[1199,1386,1643,1665]" pageId="4" pageNumber="566" refString="Brongersma, L. D. (1934) Contributions to Indo-Australian herpetology. Zoologische Mededelingen, 17, 161 - 251." type="journal article" year="1934">Brongersma 1934</bibRefCitation>
). De Rooij’s (1915) count of 42–52 PFP for 
<taxonomicName id="4C644D407915936CFDFDDEBF4ACDF95E" box="[592,703,1673,1695]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17915936CFDFDDEBF4A13F95E" box="[592,609,1674,1695]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFDC1DEBC4ACDF95E" box="[620,703,1673,1695]" italics="true" pageId="4" pageNumber="556">fumosus</emphasis>
</taxonomicName>
is often cited in the literature (e.g., 
<bibRefCitation id="EFF54B327915936CFB88DEBC4CABF95E" author="Oliver" box="[1061,1241,1673,1695]" pageId="4" pageNumber="567" refString="Oliver, P., Edgar, P., Mumpuni, Iskandar, D. T. &amp; Lilley, R. (2009) A new species of bent-toed gecko (Cyrtodactylus: Gekkonidae) from Seram Island, Indonesia. Zootaxa, 2115, 47 - 55." type="journal article" year="2009">
Oliver 
<emphasis id="B910EAD17915936CFBC6DEBF4CE5F95E" box="[1131,1175,1673,1695]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EFF54B327915936CFB4FDEBC488FF97F" author="Chan" pageId="4" pageNumber="566" refString="Chan, K. O. &amp; Norhayati, A. (2010) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia. Zootaxa, 2389, 47 - 56." type="journal article" year="2010">Chan &amp; Norhayati 2010</bibRefCitation>
; 
<bibRefCitation id="EFF54B327915936CFEAADE9D49A3F97F" author="Grismer" box="[263,465,1704,1726]" pageId="4" pageNumber="566" refString="Grismer, L. L., Wood, P. L. Jr. &amp; Lim, K. K. P. (2012) Cyrtodactylus majulah, a new species of bent-toed gecko Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology, 60 (2), 487 - 499." type="journal article" year="2012">
Grismer 
<emphasis id="B910EAD17915936CFECCDE9C49FCF97F" box="[353,398,1704,1726]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. 2012
</bibRefCitation>
), although this count is incorrect as demonstrated by our examination and literature survey (see also Remarks on the taxonomy of 
<taxonomicName id="4C644D407915936CFD8FDEFD4AE1F91C" box="[546,659,1735,1757]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17915936CFD8FDEFD4A41F91C" box="[546,563,1736,1757]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFDEDDEF24AE1F91C" box="[576,659,1735,1757]" italics="true" pageId="4" pageNumber="556">fumosus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BDB36C37915936CFF3ADED04AF3F8F8" blockId="4.[151,1437,1489,2033]" pageId="4" pageNumber="556">
2. De 
<bibRefCitation id="EFF54B327915936CFF46DED0491FF93A" author="Rooij" box="[235,365,1765,1787]" pageId="4" pageNumber="566" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Vol. I: Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, 384 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">Rooij (1915)</bibRefCitation>
and 
<bibRefCitation id="EFF54B327915936CFE32DED34A16F93A" author="Brongersma" box="[415,612,1765,1787]" pageId="4" pageNumber="566" refString="Brongersma, L. D. (1934) Contributions to Indo-Australian herpetology. Zoologische Mededelingen, 17, 161 - 251." type="journal article" year="1934">Brongersma (1934)</bibRefCitation>
attributed the shape of the ear-opening to the state of preservation. We examined well preserved specimens of 
<taxonomicName id="4C644D407915936CFE5FDF304AD1F8DB" box="[498,675,1796,1818]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="subSpecies" species="fumosus" subSpecies="sensu">
<emphasis id="B910EAD17915936CFE5FDF304A71F8DB" box="[498,515,1797,1818]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFDBDDF314AD1F8DB" box="[528,675,1796,1818]" italics="true" pageId="4" pageNumber="556">
fumosus 
<taxonomicNameLabel id="A22357AA7915936CFDC6DF304AD1F8DB" box="[619,675,1797,1818]" pageId="4" pageNumber="556" sensu="stricto">sensu</taxonomicNameLabel>
</emphasis>
</taxonomicName>
stricto that exhibited a horizontal, slit-shaped ear opening, indicating that this character is taxonomically informative.
</paragraph>
<paragraph id="8BDB36C37915936CFF3ADF744AF7F8B7" blockId="4.[151,1437,1489,2033]" pageId="4" pageNumber="556">
3. Although 
<bibRefCitation id="EFF54B327915936CFE83DF744985F896" box="[302,503,1857,1879]" pageId="4" pageNumber="567" refString="Rosler, H., Richards, S. J. &amp; Gunther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae). Salamandra, 43 (4), 193 - 230." type="journal article">
Rösler 
<emphasis id="B910EAD17915936CFED5DF7749D4F896" box="[376,422,1857,1879]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. (2007)
</bibRefCitation>
provided a maximum count of 52 PFS for the 
<typeStatus id="54DF88617915936CFC73DF774C78F896" box="[990,1034,1858,1879]" pageId="4" pageNumber="556">type</typeStatus>
series, the 
<typeStatus id="54DF88617915936CFB2DDF744CAFF896" box="[1152,1245,1857,1879]" pageId="4" pageNumber="556" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="4C644D407915936CFAACDF774DEEF896" box="[1281,1436,1858,1879]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17915936CFAACDF774D60F896" box="[1281,1298,1858,1879]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFA8DDF774DEEF896" box="[1312,1436,1858,1879]" italics="true" pageId="4" pageNumber="556">marmoratus</emphasis>
</taxonomicName>
(RMNH.RENA 2710a.1) possesses 56 PFS.
</paragraph>
<paragraph id="8BDB36C37915936CFF3ADF4A4A1CF872" blockId="4.[151,1437,1489,2033]" pageId="4" pageNumber="556">
4. 
<bibRefCitation id="EFF54B327915936CFF6ADF4A49E2F854" box="[199,400,1919,1941]" pageId="4" pageNumber="567" refString="Rosler, H., Richards, S. J. &amp; Gunther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae). Salamandra, 43 (4), 193 - 230." type="journal article">
Rösler 
<emphasis id="B910EAD17915936CFEBCDFB5494DF854" box="[273,319,1919,1941]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. (2007)
</bibRefCitation>
provided a count of 45–50 PFP for male 
<taxonomicName id="4C644D407915936CFCEDDFB54BA8F854" box="[832,986,1920,1941]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17915936CFCEDDFB54B23F854" box="[832,849,1920,1941]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFCF3DFB54BA8F854" box="[862,986,1920,1941]" italics="true" pageId="4" pageNumber="556">marmoratus</emphasis>
</taxonomicName>
, with the 
<typeStatus id="54DF88617915936CFBEEDF4A4CD2F854" box="[1091,1184,1919,1941]" pageId="4" pageNumber="556" type="lectotype">lectotype</typeStatus>
stated to have 45 pores. However, the 
<typeStatus id="54DF88617915936CFEF9DFA849C3F872" box="[340,433,1949,1971]" pageId="4" pageNumber="556" type="lectotype">lectotype</typeStatus>
possesses 52 PFP.
</paragraph>
<paragraph id="8BDB36C37915936CFF3ADF894B22F830" blockId="4.[151,1437,1489,2033]" pageId="4" pageNumber="556">
5. 
<bibRefCitation id="EFF54B327915936CFF6ADF8949EEF813" author="Linkem" box="[199,412,1980,2002]" pageId="4" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Linkem 
<emphasis id="B910EAD17915936CFEB2DF88493CF813" box="[287,334,1980,2002]" italics="true" pageId="4" pageNumber="556">et al</emphasis>
. (2008
</bibRefCitation>
; 
<tableCitation id="C6E603787915936CFE04DF894987F813" box="[425,501,1980,2002]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Metric and meristic characters with abbreviations used in this study." pageId="4" pageNumber="556">Table 1</tableCitation>
) listed 38−44 VS for 
<taxonomicName id="4C644D407915936CFD4EDF884B28F813" box="[739,858,1981,2002]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="556" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B910EAD17915936CFD4EDF884A86F813" box="[739,756,1981,2002]" italics="true" pageId="4" pageNumber="556">C</emphasis>
. 
<emphasis id="B910EAD17915936CFCAEDF884B28F813" box="[771,858,1981,2002]" italics="true" pageId="4" pageNumber="556">spinosus</emphasis>
</taxonomicName>
(counts listed for individual specimens), while in their comparative table (
<tableCitation id="C6E603787915936CFE25DFEE49A7F830" box="[392,469,2011,2033]" captionStart="TABLE 2" captionStartId="3.[151,244,151,176]" captionTargetBox="[151,1436,239,1258]" captionTargetPageId="3" captionText="TABLE 2. Metric (in mm) and meristic data for the type series of Cyrtodactylus klakahensis sp. nov. Abbreviations are defined in Table 1." httpUri="http://table.plazi.org/id/DF1B664B7912936BFF3AD8A24A45FF15" pageId="4" pageNumber="556" tableUuid="DF1B664B7912936BFF3AD8A24A45FF15">Table 2</tableCitation>
) they provided a range of 40−43 VS.
</paragraph>
<paragraph id="8BDB36C37914936DFFDADA1D4F04FD9E" blockId="5.[119,1910,552,958]" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFFDADA1D48AEFDFD" bold="true" box="[119,220,552,572]" pageId="5" pageNumber="557">
<tableCitation id="C6E603787914936DFFDADA1D48A4FDFD" box="[119,214,552,572]" captionStart="TABLE 3" captionStartId="6.[125,203,673,693]" captionTargetBox="[125,1910,727,914]" captionTargetPageId="6" captionText="TABLE 3 (continued)" httpUri="http://table.plazi.org/id/DF1B664B7917936EFFD0DA944939FD74" pageId="5" pageNumber="557" tableUuid="DF1B664B7917936EFFD0DA944939FD74">TABLE 3</tableCitation>
.
</emphasis>
Characters used to distinguish 
<taxonomicName id="4C644D407914936DFE54DA1C4A95FDFD" box="[505,743,553,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="klakahensis" status="sp. nov.">
<emphasis id="B910EAD17914936DFE54DA1C4A95FDFD" box="[505,743,553,572]" italics="true" pageId="5" pageNumber="557">Cyrtodactylus klakahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357AA7914936DFD40DA1D4B45FDFD" box="[749,823,552,572]" pageId="5" pageNumber="557" rank="species">
<emphasis id="B910EAD17914936DFD40DA1D4B45FDFD" bold="true" box="[749,823,552,572]" pageId="5" pageNumber="557">sp. nov.</emphasis>
</taxonomicNameLabel>
from congeneric species occurring in the Sunda Islands (including Sulawesi). The presence of a diagnostic character is coded as ʻ1ʼ, the absence of a character is coded as ʻ0ʼ. For taxa possessing precloacofemoral scales (= scales in a continuous series; column entitled ‘9’), precloacal- and femoral scales (separated
</paragraph>
<paragraph id="8BDB36C37914936DFF07DA5A4F04FD09" blockId="5.[119,1910,552,958]" pageId="5" pageNumber="557">each other by infrascales) are coded ‘n/a’ (columns entitled ‘7’ and ‘8’). Numbers at the head of the table correspond to characters as follows: 1 = tubercles on forelimbs, 2 = tubercles on hindlimbs, 3 = tubercles on head, 4 = number of ventral scales, 5 = enlarged subcaudals, 6 = number of subdigital lamellae under 4th toe, 7 = enlarged precloacal scales (and number of pores in parentheses if present; if pores are present in one sex only, this is indicated either by ‘♂’ or ʻ♀ʼ), 8 = enlarged femoral scales (and number of pores in parentheses if present; if pores are present in</paragraph>
<paragraph id="8BDB36C37914936DFF33DAEC4F07FD2A" blockId="5.[119,1910,552,958]" box="[158,1909,728,748]" pageId="5" pageNumber="557">sex only, this is indicated either by ‘♂’ or ʻ♀ʼ), 9 = enlarged precloacofemoral scales (and number of pores in parentheses if present; if pores are present in males only this is indicated by ‘♂’),</paragraph>
<paragraph id="8BDB36C37914936DFF39DAC94F07FCF3" blockId="5.[119,1910,552,958]" pageId="5" pageNumber="557">= pores in a continuous series, 11 = precloacal groove present (if a groove is present in males only, this is indicated by ‘♂’), 12 = pattern of dorsum (bd = banded; bl = blotched; mo = mottling; pl patternless; st = striped). If data for a character are not available, this is indicated by a ‘?’. Where derived from the literature (‘Lit.’ column), references are abbreviated by letters as follows: A =</paragraph>
<paragraph id="8BDB36C37914936DFF33DB744B39FC7F" blockId="5.[119,1910,552,958]" pageId="5" pageNumber="557">
publication; B = 
<bibRefCitation id="EFF54B327914936DFE94DB744992FC94" author="Rosler" box="[313,480,833,853]" pageId="5" pageNumber="567" refString="Rosler, H., Richards, S. J. &amp; Gunther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae). Salamandra, 43 (4), 193 - 230." type="journal article" year="2007">
Rösler 
<emphasis id="B910EAD17914936DFED6DB7749D1FC95" box="[379,419,833,852]" italics="true" pageId="5" pageNumber="557">et al</emphasis>
. 2007
</bibRefCitation>
; C = 
<bibRefCitation id="EFF54B327914936DFDBFDB744A99FC94" author="Grismer" box="[530,747,833,853]" pageId="5" pageNumber="566" refString="Grismer, L. L. &amp; Leong, T. M. (2005) New species of Cyrtodactylus (Squamata: Gekkonidae) from southern Peninsular Malaysia. Journal of Herpetology, 39 (4), 584 - 591. http: // dx. doi. org / 10.1670 / 43 - 05 A. 1" type="journal article" year="2005">Grismer &amp; Leong 2005</bibRefCitation>
; D = 
<bibRefCitation id="EFF54B327914936DFCB2DB744C59FC94" author="Manthey" box="[799,1067,833,853]" pageId="5" pageNumber="567" refString="Manthey, U. &amp; Grossmann, W. (1997) Amphibien &amp; Reptilien Sudostasiens. Natur und Tier-Verlag, Munster, Germany, 512 pp." type="book" year="1997">Manthey &amp; Grossmann 1997</bibRefCitation>
; E = 
<bibRefCitation id="EFF54B327914936DFBF0DB744D64FC94" author="Iskandar" box="[1117,1302,833,853]" pageId="5" pageNumber="567" refString="Iskandar, D. T., Rachmansah, A. &amp; Umilaela (2011) A new bent-toed gecko of the genus Cyrtodactylus Gray, 1827 (Reptilia, Gekkonidae) from Mount Tompotika, eastern peninsula of Sulawesi, Indonesia. Zootaxa, 2838, 65 - 78." type="journal article" year="2011">
Iskandar 
<emphasis id="B910EAD17914936DFB1FDB774CA9FC95" box="[1202,1243,833,852]" italics="true" pageId="5" pageNumber="557">et al</emphasis>
. 2011
</bibRefCitation>
; F = 
<bibRefCitation id="EFF54B327914936DFAEADB774D8AFC94" author="Inger" box="[1351,1528,833,853]" pageId="5" pageNumber="567" refString="Inger, R. F. &amp; King, W. (1961) A new cave-dwelling lizard of the genus Cyrtodactylus from Niah. Sarawak Museum Journal, 11, 274 - 276." type="journal article" year="1961">Inger &amp; King 1961</bibRefCitation>
; G = 
<bibRefCitation id="EFF54B327914936DF981DB744E9DFC94" author="Kathriner" box="[1580,1775,833,853]" pageId="5" pageNumber="567" refString="Kathriner, A., Bauer, A. M., O'Shea, M., Sanchez, C. &amp; Kaiser, H. (2014) Hiding in plain sight: a new species of bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from West Timor, collected by Malcolm Smith in 1924. Zootaxa, 3900 (4), 555 - 568. http: // dx. doi. org / 10.11646 / zootaxa. 3900.4.6" type="journal article" year="2014">
Kathriner 
<emphasis id="B910EAD17914936DF927DB774EC1FC95" box="[1674,1715,833,852]" italics="true" pageId="5" pageNumber="557">et al</emphasis>
. 2014
</bibRefCitation>
; H = Rösler &amp; Kaiser, in press.; I = 
<bibRefCitation id="EFF54B327914936DFE95DB5149DEFCB9" author="Hikida" box="[312,428,868,888]" pageId="5" pageNumber="567" refString="Hikida, T. (1990) Bornean gekkonid lizards of the genus Cyrtodactylus (Lacertilia: Gekkonidae) with descriptions of three new species. Japanese Journal of Herpetology, 13 (3), 91 - 107." type="journal article" year="1990">Hikida 1990</bibRefCitation>
; J = de 
<bibRefCitation id="EFF54B327914936DFE58DB514A2FFCB9" author="Rooij" box="[501,605,868,888]" pageId="5" pageNumber="566" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Vol. I: Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, 384 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">Rooij 1915</bibRefCitation>
; K = 
<bibRefCitation id="EFF54B327914936DFD3CDB504B44FCB9" author="Brongersma" box="[657,822,869,888]" pageId="5" pageNumber="566" refString="Brongersma, L. D. (1934) Contributions to Indo-Australian herpetology. Zoologische Mededelingen, 17, 161 - 251." type="journal article" year="1934">Brongersma 1934</bibRefCitation>
; L = 
<bibRefCitation id="EFF54B327914936DFCCADB514B8EFCB9" author="Boulenger" box="[871,1020,868,888]" pageId="5" pageNumber="566" refString="Boulenger, G. A. (1897) A catalogue of the reptiles and batrachians of Celebes with special reference to the collections made by Drs P &amp; F Sarasin in 1893 - 1896. Proceedings of the Royal Society of London, 1897, 193 - 237." type="proceedings paper" year="1897">Boulenger 1897</bibRefCitation>
; M = 
<bibRefCitation id="EFF54B327914936DFB99DB504CFFFCB9" author="Das" box="[1076,1165,869,888]" pageId="5" pageNumber="566" refString="Das, I. (1993) Cnemaspis gordongekkoi, a new gecko from Lombok, Indonesia, and the biogeography of oriental species of Cnemaspis (Squamata: Sauria: Gekkonidae). Hamadryad, 18, 1 - 9." type="journal article" year="1993">Das 1993</bibRefCitation>
; N = 
<bibRefCitation id="EFF54B327914936DFB6CDB504DCAFCB9" author="Youmans" box="[1217,1464,868,888]" pageId="5" pageNumber="568" refString="Youmans, T. M. &amp; Grismer, L. L. (2006) A new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History, 10, 61 - 70." type="journal article" year="2006">Youmans &amp; Grismer 2006</bibRefCitation>
; O = 
<bibRefCitation id="EFF54B327914936DFA40DB514EF9FCB9" author="Auffenberg" box="[1517,1675,868,888]" pageId="5" pageNumber="566" refString="Auffenberg, W. (1980) The herpetofauna of Komodo, with notes on adjacent areas. Bulletin of the Florida State Museum, Biological Science, 25 (2), 63 - 67." type="journal article" year="1980">Auffenberg 1980</bibRefCitation>
; P = Darevsky 1964; Q = 
<bibRefCitation id="EFF54B327914936DFFDADBBD4882FC5A" author="Werner" box="[119,240,903,923]" pageId="5" pageNumber="568" refString="Werner, F. (1896) Zweiter Beitrag zur Herpetologie der indo-orientalischen Region. Verhandlungen der Zoologisch- Botanischen Gesellschaft in Osterreich, 46, 6 - 24." type="journal article" year="1896">Werner 1896</bibRefCitation>
; R = 
<bibRefCitation id="EFF54B327914936DFE8CDBBD490BFC5A" author="Das" box="[289,377,904,923]" pageId="5" pageNumber="566" refString="Das, I. (2010) A Field Guide to the Reptiles of South-east Asia. New Holland Publisher, London, 376 pp." type="book" year="2010">Das 2010</bibRefCitation>
; S = 
<bibRefCitation id="EFF54B327914936DFE0ADBBD4A20FC5A" author="Harvey" box="[423,594,903,923]" pageId="5" pageNumber="566" refString="Harvey, M. B., O'Connell, K. A., Barraza, G., Riyanto, A., Kurniawan, N. &amp; Smith, E. N. (2015) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the southern Bukit Barisan range of Sumatra and an estimation of their phylogeny. Zootaxa, 4020 (3), 495 - 516. http: // dx. doi. org / 10.11646 / zootaxa. 4020.3.5" type="journal article" year="2015">
Harvey 
<emphasis id="B910EAD17914936DFE42DBBD4A65FC5B" box="[495,535,903,922]" italics="true" pageId="5" pageNumber="557">et al</emphasis>
. 2015
</bibRefCitation>
; T = 
<bibRefCitation id="EFF54B327914936DFD2CDBBD4A94FC5A" author="Inger" box="[641,742,903,923]" pageId="5" pageNumber="567" refString="Inger, R. F. (1958) A new gecko of the genus Cyrtodactylus, with a key to the species from Borneo and the Philippine Islands. Sarawak Museum Journal, 8, 261 - 264." type="journal article" year="1958">Inger 1958</bibRefCitation>
; U = 
<bibRefCitation id="EFF54B327914936DFCB5DBB24BBAFC5A" author="Riyanto" box="[792,968,903,923]" pageId="5" pageNumber="567" refString="Riyanto, A., Bauer, A. M. &amp; Yudha, D. S. (2014) A new small karst-dwelling species of Cyrtodactylus from Java, Indonesia. Zootaxa, 3785 (4), 589 - 599. http: // dx. doi. org / 10.11646 / zootaxa. 3785.4.7" type="journal article" year="2014">
Riyanto 
<emphasis id="B910EAD17914936DFCC8DBBD4BFFFC5B" box="[869,909,903,922]" italics="true" pageId="5" pageNumber="557">et al</emphasis>
. 2014
</bibRefCitation>
; V = 
<bibRefCitation id="EFF54B327914936DFC54DBB24CDAFC5A" author="Linkem" box="[1017,1192,903,923]" pageId="5" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Linkem 
<emphasis id="B910EAD17914936DFBE8DBBD4C1FFC5B" box="[1093,1133,903,922]" italics="true" pageId="5" pageNumber="557">et al</emphasis>
. 2008
</bibRefCitation>
; W = 
<bibRefCitation id="EFF54B327914936DFB72DBB24D22FC5A" author="Taylor" box="[1247,1360,903,923]" pageId="5" pageNumber="568" refString="Taylor, E. H. (1962) New oriental reptiles. The University of Kansas Science Bulletin, 43, 209 - 263." type="journal article" year="1962">Taylor 1962</bibRefCitation>
; X = 
<bibRefCitation id="EFF54B327914936DFA2FDBB24E43FC5A" author="Hayden" box="[1410,1585,903,923]" pageId="5" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Hayden 
<emphasis id="B910EAD17914936DFA63DBBD4D84FC5B" box="[1486,1526,903,922]" italics="true" pageId="5" pageNumber="557">et al</emphasis>
. 2008
</bibRefCitation>
; Y = 
<bibRefCitation id="EFF54B327914936DF9CFDBBD4EB8FC5A" author="Dunn" box="[1634,1738,904,923]" pageId="5" pageNumber="566" refString="Dunn, E. (1927) Results of the Douglas Burden Expedition to the island of Komodo - III, Lizards from the East Indies. American Museum Novitates, 288, 2 - 3." type="journal article" year="1927">Dunn 1927</bibRefCitation>
. Under the column heading ʻ 
<emphasis id="B910EAD17914936DFF61DB9E48A5FC7C" box="[204,215,939,957]" italics="true" pageId="5" pageNumber="557">n</emphasis>
ʼ we provide the number of adult specimens we examined personally.
</paragraph>
<paragraph id="8BDB36C37914936DFFDADBD74F3AFB2F" pageId="5" pageNumber="557">
<table id="F964C46379146C97FFDADBD74F07FB2F" box="[119,1909,994,1262]" gridcols="12" gridrows="5" pageId="5" pageNumber="557">
<tr id="3554348179146C97FFDADBD74F07FB90" box="[119,1909,994,1105]" gridrow="0" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADBD749E6FB90" box="[119,404,994,1105]" gridcol="0" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFFDADBD749E6FBEA" bold="true" pageId="5" pageNumber="557">
Taxon SVL 1 in adults 
<taxonomicName id="4C644D407914936DFFDADC234894FBEB" box="[119,230,1046,1066]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="klakahensis">
<emphasis id="B910EAD17914936DFFDADC234894FBEB" bold="true" box="[119,230,1046,1066]" italics="true" pageId="5" pageNumber="557">klakahensis</emphasis>
</taxonomicName>
61–68 1
</emphasis>
<taxonomicName id="4C644D407914936DFFDADC0B48A4FB91" box="[119,214,1086,1104]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="agamensis">
<emphasis id="B910EAD17914936DFFDADC0B48A4FB91" box="[119,214,1086,1104]" italics="true" pageId="5" pageNumber="557">agamensis</emphasis>
</taxonomicName>
50 1
</th>
<th id="76855DFD79146C97FE7DDBD749A9FB90" box="[464,475,994,1105]" gridcol="1" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFE7DDBD749A9FBEA" bold="true" box="[464,475,994,1067]" pageId="5" pageNumber="557">2 1</emphasis>
1
</th>
<th id="76855DFD79146C97FDBBDBD74A53FB90" box="[534,545,994,1105]" gridcol="2" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFDBBDBD74A53FBEA" bold="true" box="[534,545,994,1067]" pageId="5" pageNumber="557">3 1</emphasis>
1
</th>
<th id="76855DFD79146C97FDF0DBD74AE4FB90" box="[605,662,994,1105]" gridcol="3" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFDF0DBD74AE4FBEA" bold="true" box="[605,662,994,1067]" pageId="5" pageNumber="557">4 35–38</emphasis>
67
</th>
<th id="76855DFD79146C97FD65DBD74AA1FB90" box="[712,723,994,1105]" gridcol="4" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFD65DBD74AA1FBEA" bold="true" box="[712,723,994,1067]" pageId="5" pageNumber="557">5 0</emphasis>
0
</th>
<th id="76855DFD79146C97FCA0DBD74B34FB90" box="[781,838,994,1105]" gridcol="5" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFCA0DBD74B34FBEA" bold="true" box="[781,838,994,1067]" pageId="5" pageNumber="557">6 17–20</emphasis>
26
</th>
<th id="76855DFD79146C97FCD4DBD74B90FB90" box="[889,994,994,1105]" gridcol="6" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFCD4DBD74BE5FBEA" bold="true" box="[889,919,994,1067]" pageId="5" pageNumber="557">7 n/a</emphasis>
n/a (? 
<emphasis id="B910EAD17914936DFC06DC0D4BC1FB84" bold="true" box="[939,947,1080,1093]" pageId="5" pageNumber="557">a</emphasis>
)
</th>
<th id="76855DFD79146C97FBC6DBD74CA7FB90" box="[1131,1237,994,1105]" gridcol="7" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFBC6DBD74CF8FBEA" bold="true" box="[1131,1162,994,1067]" pageId="5" pageNumber="557">8 n/a</emphasis>
n/a (? 
<emphasis id="B910EAD17914936DFB33DC0D4CD4FB84" bold="true" box="[1182,1190,1080,1093]" pageId="5" pageNumber="557">a</emphasis>
)
</th>
<th id="76855DFD79146C97FA8DDBD74DB8FB90" box="[1312,1482,994,1105]" gridcol="8" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DFA8DDBD74D05FBEB" bold="true" pageId="5" pageNumber="557">9 10 1 (37–38,</emphasis>
♂
<emphasis id="B910EAD17914936DFA20DC234DCCFBEA" bold="true" box="[1421,1470,1046,1067]" pageId="5" pageNumber="557">) 1</emphasis>
1 (? 
<emphasis id="B910EAD17914936DFAEEDC0D4D39FB84" bold="true" box="[1347,1355,1080,1093]" pageId="5" pageNumber="557">a</emphasis>
)?
</th>
<th id="76855DFD79146C97F9A0DBD74E09FB90" box="[1549,1659,994,1105]" gridcol="9" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DF9A0DBD74E04FBEA" bold="true" pageId="5" pageNumber="557">11 12 1 bl</emphasis>
1 bl
</th>
<th id="76855DFD79146C97F913DBD74F77FB90" box="[1726,1797,994,1105]" gridcol="10" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DF913DBD74EBCFBEA" bold="true" pageId="5" pageNumber="557">Lit. A</emphasis>
B
</th>
<th id="76855DFD79146C97F890DBD74F07FB90" box="[1853,1909,994,1105]" gridcol="11" gridrow="0" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DF890DBD74F3AFBEA" bold="true" box="[1853,1866,994,1067]" pageId="5" pageNumber="557">
<emphasis id="B910EAD17914936DF890DBD74F38FC34" bold="true" box="[1853,1866,994,1013]" italics="true" pageId="5" pageNumber="557">n</emphasis>
4
</emphasis>
-
</th>
</tr>
<tr id="3554348179146C97FFDADC514F07FBB9" box="[119,1909,1124,1144]" gridrow="1" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADC5149E6FBB9" box="[119,404,1124,1144]" gridcol="0" gridrow="1" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADC5048BFFBB9" box="[119,205,1125,1144]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B910EAD17914936DFFDADC5048BFFBB9" box="[119,205,1125,1144]" italics="true" pageId="5" pageNumber="557">baluensis</emphasis>
</taxonomicName>
71–95 1
</th>
<td id="76855DFD79146C97FE7DDC5149A9FBB9" box="[464,475,1124,1144]" gridcol="1" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDC514A53FBB9" box="[534,545,1124,1144]" gridcol="2" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DC514AE4FBB9" box="[605,662,1124,1144]" gridcol="3" gridrow="1" pageId="5" pageNumber="557">36–46</td>
<td id="76855DFD79146C97FD65DC514AA1FBB9" box="[712,723,1124,1144]" gridcol="4" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FCA0DC514B34FBB9" box="[781,838,1124,1144]" gridcol="5" gridrow="1" pageId="5" pageNumber="557">19–23</td>
<td id="76855DFD79146C97FCD4DC514B90FBB9" box="[889,994,1124,1144]" gridcol="6" gridrow="1" pageId="5" pageNumber="557">1 (9–11, ♂)</td>
<td id="76855DFD79146C97FBC6DC514CA7FBB9" box="[1131,1237,1124,1144]" gridcol="7" gridrow="1" pageId="5" pageNumber="557">1 (4–10, ♂)</td>
<td id="76855DFD79146C97FA8DDC514DB8FBB9" box="[1312,1482,1124,1144]" gridcol="8" gridrow="1" pageId="5" pageNumber="557">0 0</td>
<td id="76855DFD79146C97F9A0DC514E09FBB9" box="[1549,1659,1124,1144]" gridcol="9" gridrow="1" pageId="5" pageNumber="557">0 bl</td>
<td id="76855DFD79146C97F913DC514F77FBB9" box="[1726,1797,1124,1144]" gridcol="10" gridrow="1" pageId="5" pageNumber="557">A, C, D</td>
<td id="76855DFD79146C97F890DC514F07FBB9" box="[1853,1909,1124,1144]" gridcol="11" gridrow="1" pageId="5" pageNumber="557">9</td>
</tr>
<tr id="3554348179146C97FFDADCB94F07FB61" box="[119,1909,1164,1184]" gridrow="2" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADCB949E6FB61" box="[119,404,1164,1184]" gridcol="0" gridrow="2" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADCB948D7FB5E" box="[119,165,1164,1183]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="batik">
<emphasis id="B910EAD17914936DFFDADCB948D7FB5E" box="[119,165,1164,1183]" italics="true" pageId="5" pageNumber="557">batik</emphasis>
</taxonomicName>
103–113 1
</th>
<td id="76855DFD79146C97FE7DDCB949A9FB61" box="[464,475,1164,1184]" gridcol="1" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDCB94A53FB61" box="[534,545,1164,1184]" gridcol="2" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DCB94AE4FB61" box="[605,662,1164,1184]" gridcol="3" gridrow="2" pageId="5" pageNumber="557">48–57</td>
<td id="76855DFD79146C97FD65DCB94AA1FB61" box="[712,723,1164,1184]" gridcol="4" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FCA0DCB94B34FB61" box="[781,838,1164,1184]" gridcol="5" gridrow="2" pageId="5" pageNumber="557">24–27</td>
<td id="76855DFD79146C97FCD4DCB94B90FB61" box="[889,994,1164,1184]" gridcol="6" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FBC6DCB94CA7FB61" box="[1131,1237,1164,1184]" gridcol="7" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FA8DDCB94DB8FB61" box="[1312,1482,1164,1184]" gridcol="8" gridrow="2" pageId="5" pageNumber="557">0 0</td>
<td id="76855DFD79146C97F9A0DCB94E09FB61" box="[1549,1659,1164,1184]" gridcol="9" gridrow="2" pageId="5" pageNumber="557">0 bd</td>
<td id="76855DFD79146C97F913DCB94F77FB61" box="[1726,1797,1164,1184]" gridcol="10" gridrow="2" pageId="5" pageNumber="557">E</td>
<td id="76855DFD79146C97F890DCB94F07FB61" box="[1853,1909,1164,1184]" gridcol="11" gridrow="2" pageId="5" pageNumber="557">-</td>
</tr>
<tr id="3554348179146C97FFDADC864F07FB06" box="[119,1909,1203,1223]" gridrow="3" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADC8649E6FB06" box="[119,404,1203,1223]" gridcol="0" gridrow="3" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADC864898FB07" box="[119,234,1203,1222]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="cavernicolus">
<emphasis id="B910EAD17914936DFFDADC864898FB07" box="[119,234,1203,1222]" italics="true" pageId="5" pageNumber="557">cavernicolus</emphasis>
</taxonomicName>
64–81 0
</th>
<td id="76855DFD79146C97FE7DDC8649A9FB06" box="[464,475,1203,1223]" gridcol="1" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDC864A53FB06" box="[534,545,1203,1223]" gridcol="2" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DC864AE4FB06" box="[605,662,1203,1223]" gridcol="3" gridrow="3" pageId="5" pageNumber="557">51–58</td>
<td id="76855DFD79146C97FD65DC864AA1FB06" box="[712,723,1203,1223]" gridcol="4" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FCA0DC864B34FB06" box="[781,838,1203,1223]" gridcol="5" gridrow="3" pageId="5" pageNumber="557">22–26</td>
<td id="76855DFD79146C97FCD4DC864B90FB06" box="[889,994,1203,1223]" gridcol="6" gridrow="3" pageId="5" pageNumber="557">1 (4, ♂)</td>
<td id="76855DFD79146C97FBC6DC864CA7FB06" box="[1131,1237,1203,1223]" gridcol="7" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FA8DDC864DB8FB06" box="[1312,1482,1203,1223]" gridcol="8" gridrow="3" pageId="5" pageNumber="557">0 0</td>
<td id="76855DFD79146C97F9A0DC864E09FB06" box="[1549,1659,1203,1223]" gridcol="9" gridrow="3" pageId="5" pageNumber="557">1 bd</td>
<td id="76855DFD79146C97F913DC864F77FB06" box="[1726,1797,1203,1223]" gridcol="10" gridrow="3" pageId="5" pageNumber="557">C, F</td>
<td id="76855DFD79146C97F890DC864F07FB06" box="[1853,1909,1203,1223]" gridcol="11" gridrow="3" pageId="5" pageNumber="557">-</td>
</tr>
<tr id="3554348179146C97FFDADCEF4F07FB2F" box="[119,1909,1242,1262]" gridrow="4" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADCEF49E6FB2F" box="[119,404,1242,1262]" gridcol="0" gridrow="4" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADCEF48C5FB2C" box="[119,183,1242,1261]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="celatus">
<emphasis id="B910EAD17914936DFFDADCEF48C5FB2C" box="[119,183,1242,1261]" italics="true" pageId="5" pageNumber="557">celatus</emphasis>
</taxonomicName>
38−44 1
</th>
<td id="76855DFD79146C97FE7DDCEF49A9FB2F" box="[464,475,1242,1262]" gridcol="1" gridrow="4" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDCEF4A53FB2F" box="[534,545,1242,1262]" gridcol="2" gridrow="4" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DCEF4AE4FB2F" box="[605,662,1242,1262]" gridcol="3" gridrow="4" pageId="5" pageNumber="557">34–42</td>
<td id="76855DFD79146C97FD65DCEF4AA1FB2F" box="[712,723,1242,1262]" gridcol="4" gridrow="4" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FCA0DCEF4B34FB2F" box="[781,838,1242,1262]" gridcol="5" gridrow="4" pageId="5" pageNumber="557">15–18</td>
<td id="76855DFD79146C97FCD4DCEF4B90FB2F" box="[889,994,1242,1262]" gridcol="6" gridrow="4" pageId="5" pageNumber="557">1 (4, ♂)</td>
<td id="76855DFD79146C97FBC6DCEF4CA7FB2F" box="[1131,1237,1242,1262]" gridcol="7" gridrow="4" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FA8DDCEF4DB8FB2F" box="[1312,1482,1242,1262]" gridcol="8" gridrow="4" pageId="5" pageNumber="557">0 0</td>
<td id="76855DFD79146C97F9A0DCEF4E09FB2F" box="[1549,1659,1242,1262]" gridcol="9" gridrow="4" pageId="5" pageNumber="557">1 (♂) bl</td>
<td id="76855DFD79146C97F913DCEF4F77FB2F" box="[1726,1797,1242,1262]" gridcol="10" gridrow="4" pageId="5" pageNumber="557">A, G, H</td>
<td id="76855DFD79146C97F890DCEF4F07FB2F" box="[1853,1909,1242,1262]" gridcol="11" gridrow="4" pageId="5" pageNumber="557">3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDB36C37914936DFFDADD344F3AFA4B" pageId="5" pageNumber="557">
<table id="F964C46379146C97FFDADD344F07FA4B" box="[119,1909,1281,1418]" gridcols="13" gridrows="4" pageId="5" pageNumber="557">
<tr id="3554348179146C97FFDADD344F07FAD4" box="[119,1909,1281,1301]" gridrow="0" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADD344923FAD4" box="[119,337,1281,1301]" gridcol="0" gridrow="0" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADD344894FAD5" box="[119,230,1281,1300]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="B910EAD17914936DFFDADD344894FAD5" box="[119,230,1281,1300]" italics="true" pageId="5" pageNumber="557">consobrinus</emphasis>
</taxonomicName>
97–125
</th>
<th id="76855DFD79146C97FE24DD3449E6FAD4" box="[393,404,1281,1301]" gridcol="1" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FE7DDD3449A9FAD4" box="[464,475,1281,1301]" gridcol="2" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FDBBDD344A53FAD4" box="[534,545,1281,1301]" gridcol="3" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FDF0DD344AE4FAD4" box="[605,662,1281,1301]" gridcol="4" gridrow="0" pageId="5" pageNumber="557">58–71</th>
<th id="76855DFD79146C97FD65DD344AA1FAD4" box="[712,723,1281,1301]" gridcol="5" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FCA0DD344B35FAD4" box="[781,839,1281,1301]" gridcol="6" gridrow="0" pageId="5" pageNumber="557">22–28</th>
<th id="76855DFD79146C97FCD4DD344B90FAD4" box="[889,994,1281,1301]" gridcol="7" gridrow="0" pageId="5" pageNumber="557">1 (8–11, ♂)</th>
<th id="76855DFD79146C97FBC6DD344CA9FAD4" box="[1131,1243,1281,1301]" gridcol="8" gridrow="0" pageId="5" pageNumber="557">0/1 (0–6, ♂)</th>
<th id="76855DFD79146C97FA8DDD344D59FAD4" box="[1312,1323,1281,1301]" gridcol="9" gridrow="0" pageId="5" pageNumber="557">0</th>
<th id="76855DFD79146C97FA1EDD344DCCFAD4" box="[1459,1470,1281,1301]" gridcol="10" gridrow="0" pageId="5" pageNumber="557">0</th>
<th id="76855DFD79146C97F9A0DD344E09FAD4" box="[1549,1659,1281,1301]" gridcol="11" gridrow="0" pageId="5" pageNumber="557">0 bd</th>
<th id="76855DFD79146C97F913DD344F07FAD4" box="[1726,1909,1281,1301]" gridcol="12" gridrow="0" pageId="5" pageNumber="557">A, C, D, I 2</th>
</tr>
<tr id="3554348179146C97FFDADD1D4F07FAFD" box="[119,1909,1320,1340]" gridrow="1" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADD1D4923FAFD" box="[119,337,1320,1340]" gridcol="0" gridrow="1" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADD1D4884FAFA" box="[119,246,1320,1339]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="darmandvillei">
<emphasis id="B910EAD17914936DFFDADD1D4884FAFA" box="[119,246,1320,1339]" italics="true" pageId="5" pageNumber="557">darmandvillei</emphasis>
</taxonomicName>
80−82
</th>
<td id="76855DFD79146C97FE24DD1D49E6FAFD" box="[393,404,1320,1340]" gridcol="1" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FE7DDD1D49A9FAFD" box="[464,475,1320,1340]" gridcol="2" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDD1D4A53FAFD" box="[534,545,1320,1340]" gridcol="3" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DD1D4AE4FAFD" box="[605,662,1320,1340]" gridcol="4" gridrow="1" pageId="5" pageNumber="557">34–36</td>
<td id="76855DFD79146C97FD65DD1D4AA1FAFD" box="[712,723,1320,1340]" gridcol="5" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FCA0DD1D4B35FAFD" box="[781,839,1320,1340]" gridcol="6" gridrow="1" pageId="5" pageNumber="557">23−24</td>
<td id="76855DFD79146C97FCD4DD1D4B90FAFD" box="[889,994,1320,1340]" gridcol="7" gridrow="1" pageId="5" pageNumber="557">n/a</td>
<td id="76855DFD79146C97FBC6DD1D4CA9FAFD" box="[1131,1243,1320,1340]" gridcol="8" gridrow="1" pageId="5" pageNumber="557">n/a</td>
<td id="76855DFD79146C97FA8DDD1D4D59FAFD" box="[1312,1323,1320,1340]" gridcol="9" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FA1EDD1D4DCCFAFD" box="[1459,1470,1320,1340]" gridcol="10" gridrow="1" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9A0DD1D4E09FAFD" box="[1549,1659,1320,1340]" gridcol="11" gridrow="1" pageId="5" pageNumber="557">0 bl</td>
<td id="76855DFD79146C97F913DD1D4F07FAFD" box="[1726,1909,1320,1340]" gridcol="12" gridrow="1" pageId="5" pageNumber="557">A, J, K 2</td>
</tr>
<tr id="3554348179146C97FFDADD7A4F07FAA2" box="[119,1909,1359,1379]" gridrow="2" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADD7A4923FAA2" box="[119,337,1359,1379]" gridcol="0" gridrow="2" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADD6548B3FAA2" box="[119,193,1360,1379]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD17914936DFFDADD6548B3FAA2" box="[119,193,1360,1379]" italics="true" pageId="5" pageNumber="557">fumosus</emphasis>
</taxonomicName>
57–78
</th>
<td id="76855DFD79146C97FE24DD7A49E6FAA2" box="[393,404,1359,1379]" gridcol="1" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FE7DDD7A49A9FAA2" box="[464,475,1359,1379]" gridcol="2" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDD7A4A53FAA2" box="[534,545,1359,1379]" gridcol="3" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DD7A4AE4FAA2" box="[605,662,1359,1379]" gridcol="4" gridrow="2" pageId="5" pageNumber="557">37–50</td>
<td id="76855DFD79146C97FD65DD7A4AA1FAA2" box="[712,723,1359,1379]" gridcol="5" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FCA0DD7A4B35FAA2" box="[781,839,1359,1379]" gridcol="6" gridrow="2" pageId="5" pageNumber="557">17–23</td>
<td id="76855DFD79146C97FCD4DD7A4B90FAA2" box="[889,994,1359,1379]" gridcol="7" gridrow="2" pageId="5" pageNumber="557">n/a (10, ♂)</td>
<td id="76855DFD79146C97FBC6DD7A4CA9FAA2" box="[1131,1243,1359,1379]" gridcol="8" gridrow="2" pageId="5" pageNumber="557">n/a (3, ♂)</td>
<td id="76855DFD79146C97FA8DDD7A4D59FAA2" box="[1312,1323,1359,1379]" gridcol="9" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FA1EDD7A4DCCFAA2" box="[1459,1470,1359,1379]" gridcol="10" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9A0DD7A4E09FAA2" box="[1549,1659,1359,1379]" gridcol="11" gridrow="2" pageId="5" pageNumber="557">1 (♂) bl</td>
<td id="76855DFD79146C97F913DD7A4F07FAA2" box="[1726,1909,1359,1379]" gridcol="12" gridrow="2" pageId="5" pageNumber="557">A, L 3</td>
</tr>
<tr id="3554348179146C97FFDADD434F07FA4B" box="[119,1909,1398,1418]" gridrow="3" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADD434923FA4B" box="[119,337,1398,1418]" gridcol="0" gridrow="3" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADD424881FA4B" box="[119,243,1399,1418]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="gordongekkoi">
<emphasis id="B910EAD17914936DFFDADD424881FA4B" box="[119,243,1399,1418]" italics="true" pageId="5" pageNumber="557">gordongekkoi</emphasis>
</taxonomicName>
71–73
</th>
<td id="76855DFD79146C97FE24DD4349E6FA4B" box="[393,404,1398,1418]" gridcol="1" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FE7DDD4349A9FA4B" box="[464,475,1398,1418]" gridcol="2" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDD434A53FA4B" box="[534,545,1398,1418]" gridcol="3" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DD434AE4FA4B" box="[605,662,1398,1418]" gridcol="4" gridrow="3" pageId="5" pageNumber="557">30</td>
<td id="76855DFD79146C97FD65DD434AA1FA4B" box="[712,723,1398,1418]" gridcol="5" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FCA0DD434B35FA4B" box="[781,839,1398,1418]" gridcol="6" gridrow="3" pageId="5" pageNumber="557">22–23</td>
<td id="76855DFD79146C97FCD4DD434B90FA4B" box="[889,994,1398,1418]" gridcol="7" gridrow="3" pageId="5" pageNumber="557">n/a</td>
<td id="76855DFD79146C97FBC6DD434CA9FA4B" box="[1131,1243,1398,1418]" gridcol="8" gridrow="3" pageId="5" pageNumber="557">n/a</td>
<td id="76855DFD79146C97FA8DDD434D59FA4B" box="[1312,1323,1398,1418]" gridcol="9" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FA1EDD434DCCFA4B" box="[1459,1470,1398,1418]" gridcol="10" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9A0DD434E09FA4B" box="[1549,1659,1398,1418]" gridcol="11" gridrow="3" pageId="5" pageNumber="557">0 bl</td>
<td id="76855DFD79146C97F913DD434F07FA4B" box="[1726,1909,1398,1418]" gridcol="12" gridrow="3" pageId="5" pageNumber="557">A, M 2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDB36C37914936EFFDADDAB4DD5FAA3" blockId="5.[119,1861,1437,1457]" lastBlockId="6.[125,1866,1357,1569]" lastPageId="6" lastPageNumber="558" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADDAB48DFFA71" box="[119,173,1438,1456]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="B910EAD17914936DFFDADDAB48DFFA71" box="[119,173,1438,1456]" italics="true" pageId="5" pageNumber="557">ingeri</emphasis>
</taxonomicName>
65– 
<date id="FFDA10037914936DFE82DDA84934FA70" box="[303,326,1437,1457]" pageId="5" pageNumber="557" value="1976-01-01">76</date>
1 1 1 40–43 1 
<date id="FFDA10037914936DFCA0DDAB4BF6FA70" box="[781,900,1438,1457]" pageId="5" pageNumber="557" value="1923-01-27">23–27 1</date>
(8, only ♂♂ known) 0 0 0 0 bl I, N - ...continued on next page 
<taxonomicName id="4C644D407917936EFFD0DC7348BEFB9A" box="[125,204,1094,1115]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B910EAD17917936EFFD0DC7348BEFB9A" box="[125,204,1094,1115]" italics="true" pageId="6" pageNumber="558">spinosus</emphasis>
</taxonomicName>
70– 
<date id="FFDA10037917936EFEE9DC724929FB9A" box="[324,347,1095,1115]" pageId="6" pageNumber="558" value="1983-01-01">83</date>
1 1 1 38−44 0 
<date id="FFDA10037917936EFCB3DC724BE5FB9A" box="[798,919,1094,1115]" pageId="6" pageNumber="558" value="1919-01-21">19–21 1</date>
(12–13, ♂) 
<emphasis id="B910EAD17917936EFBAFDC774C78FB8E" bold="true" box="[1026,1034,1090,1103]" pageId="6" pageNumber="558">d</emphasis>
1 (? 
<emphasis id="B910EAD17917936EFB08DC774CDEFB8E" bold="true" box="[1189,1196,1090,1103]" pageId="6" pageNumber="558">e</emphasis>
) 0 0 1 (♂) bd V - 
<taxonomicName id="4C644D407917936EFF27DD784903FAA3" box="[138,369,1357,1378]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="agamensis">
<emphasis id="B910EAD17917936EFF27DD784903FAA3" box="[138,369,1357,1378]" italics="true" pageId="6" pageNumber="558">Cyrtodactylus agamensis</emphasis>
</taxonomicName>
is known from a single female only (see 
<bibRefCitation id="EFF54B327917936EFD41DD7B4BE1FAA3" author="Rosler" box="[748,915,1357,1378]" pageId="6" pageNumber="567" refString="Rosler, H., Richards, S. J. &amp; Gunther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae). Salamandra, 43 (4), 193 - 230." type="journal article" year="2007">
Rösler 
<emphasis id="B910EAD17917936EFC83DD784B24FAA3" box="[814,854,1357,1378]" italics="true" pageId="6" pageNumber="558">et al</emphasis>
. 2007
</bibRefCitation>
). Hence, it is not known at present if males possess pores.
</paragraph>
<paragraph id="8BDB36C37914936DFFDADDF04F37F98F" pageId="5" pageNumber="557">
<table id="F964C46379146C97FFDADDF04F07F98F" box="[119,1909,1477,1614]" gridcols="14" gridrows="4" pageId="5" pageNumber="557">
<tr id="3554348179146C97FFDADDF04F07FA18" box="[119,1909,1477,1497]" gridrow="0" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADDF048ABFA18" box="[119,217,1477,1497]" gridcol="0" gridrow="0" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADDF048BEFA19" box="[119,204,1477,1496]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="jellesmae">
<emphasis id="B910EAD17914936DFFDADDF048BEFA19" box="[119,204,1477,1496]" italics="true" pageId="5" pageNumber="557">jellesmae</emphasis>
</taxonomicName>
</th>
<th id="76855DFD79146C97FEA0DDF04934FA18" box="[269,326,1477,1497]" gridcol="1" gridrow="0" pageId="5" pageNumber="557">42–64</th>
<th id="76855DFD79146C97FE24DDF049E6FA18" box="[393,404,1477,1497]" gridcol="2" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FE7DDDF049A9FA18" box="[464,475,1477,1497]" gridcol="3" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FDBBDDF04A53FA18" box="[534,545,1477,1497]" gridcol="4" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FDF0DDF04AE5FA18" box="[605,663,1477,1497]" gridcol="5" gridrow="0" pageId="5" pageNumber="557">40−47</th>
<th id="76855DFD79146C97FD65DDF04AA1FA18" box="[712,723,1477,1497]" gridcol="6" gridrow="0" pageId="5" pageNumber="557">0</th>
<th id="76855DFD79146C97FCA0DDF04B34FA18" box="[781,838,1477,1497]" gridcol="7" gridrow="0" pageId="5" pageNumber="557">16–19</th>
<th id="76855DFD79146C97FCD4DDF04BB9FA18" box="[889,971,1477,1497]" gridcol="8" gridrow="0" pageId="5" pageNumber="557">1</th>
<th id="76855DFD79146C97FBC6DDF04CFAFA18" box="[1131,1160,1477,1497]" gridcol="9" gridrow="0" pageId="5" pageNumber="557">0</th>
<th id="76855DFD79146C97FA8DDDF04D59FA18" box="[1312,1323,1477,1497]" gridcol="10" gridrow="0" pageId="5" pageNumber="557">0</th>
<th id="76855DFD79146C97FA1EDDF04DCCFA18" box="[1459,1470,1477,1497]" gridcol="11" gridrow="0" pageId="5" pageNumber="557">0</th>
<th id="76855DFD79146C97F9A0DDF04E6AFA18" box="[1549,1560,1477,1497]" gridcol="12" gridrow="0" pageId="5" pageNumber="557">0</th>
<th id="76855DFD79146C97F9C9DDF04F07FA18" box="[1636,1909,1477,1497]" gridcol="13" gridrow="0" pageId="5" pageNumber="557">bl A, L 3</th>
</tr>
<tr id="3554348179146C97FFDADDD94F07F9C1" box="[119,1909,1516,1536]" gridrow="1" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADDD948ABF9C1" box="[119,217,1516,1536]" gridcol="0" gridrow="1" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADDD948A6FA3E" box="[119,212,1516,1535]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="laevigatus">
<emphasis id="B910EAD17914936DFFDADDD948A6FA3E" box="[119,212,1516,1535]" italics="true" pageId="5" pageNumber="557">laevigatus</emphasis>
</taxonomicName>
</th>
<td id="76855DFD79146C97FEA0DDD94934F9C1" box="[269,326,1516,1536]" gridcol="1" gridrow="1" pageId="5" pageNumber="557">38–47</td>
<td id="76855DFD79146C97FE24DDD949E6F9C1" box="[393,404,1516,1536]" gridcol="2" gridrow="1" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FE7DDDD949A9F9C1" box="[464,475,1516,1536]" gridcol="3" gridrow="1" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDDD94A53F9C1" box="[534,545,1516,1536]" gridcol="4" gridrow="1" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FDF0DDD94AE5F9C1" box="[605,663,1516,1536]" gridcol="5" gridrow="1" pageId="5" pageNumber="557">30–34</td>
<td id="76855DFD79146C97FD65DDD94AA1F9C1" box="[712,723,1516,1536]" gridcol="6" gridrow="1" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FCA0DDD94B34F9C1" box="[781,838,1516,1536]" gridcol="7" gridrow="1" pageId="5" pageNumber="557">10–15</td>
<td id="76855DFD79146C97FCD4DDD94BB9F9C1" box="[889,971,1516,1536]" gridcol="8" gridrow="1" pageId="5" pageNumber="557">?</td>
<td id="76855DFD79146C97FBC6DDD94CFAF9C1" box="[1131,1160,1516,1536]" gridcol="9" gridrow="1" pageId="5" pageNumber="557">0/1</td>
<td id="76855DFD79146C97FA8DDDD94D59F9C1" box="[1312,1323,1516,1536]" gridcol="10" gridrow="1" pageId="5" pageNumber="557">?</td>
<td id="76855DFD79146C97FA1EDDD94DCCF9C1" box="[1459,1470,1516,1536]" gridcol="11" gridrow="1" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9A0DDD94E6AF9C1" box="[1549,1560,1516,1536]" gridcol="12" gridrow="1" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9C9DDD94F07F9C1" box="[1636,1909,1516,1536]" gridcol="13" gridrow="1" pageId="5" pageNumber="557">mo, pl A, E, O, P 5</td>
</tr>
<tr id="3554348179146C97FFDADE264F07F9E6" box="[119,1909,1555,1575]" gridrow="2" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADE2648ABF9E6" box="[119,217,1555,1575]" gridcol="0" gridrow="2" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADE2648B1F9E7" box="[119,195,1555,1574]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B910EAD17914936DFFDADE2648B1F9E7" box="[119,195,1555,1574]" italics="true" pageId="5" pageNumber="557">lateralis</emphasis>
</taxonomicName>
</th>
<td id="76855DFD79146C97FEA0DE264934F9E6" box="[269,326,1555,1575]" gridcol="1" gridrow="2" pageId="5" pageNumber="557">65−85</td>
<td id="76855DFD79146C97FE24DE2649E6F9E6" box="[393,404,1555,1575]" gridcol="2" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FE7DDE2649A9F9E6" box="[464,475,1555,1575]" gridcol="3" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDE264A53F9E6" box="[534,545,1555,1575]" gridcol="4" gridrow="2" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DE264AE5F9E6" box="[605,663,1555,1575]" gridcol="5" gridrow="2" pageId="5" pageNumber="557">60–64</td>
<td id="76855DFD79146C97FD65DE264AA1F9E6" box="[712,723,1555,1575]" gridcol="6" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FCA0DE264B34F9E6" box="[781,838,1555,1575]" gridcol="7" gridrow="2" pageId="5" pageNumber="557">21–22</td>
<td id="76855DFD79146C97FCD4DE264BB9F9E6" box="[889,971,1555,1575]" gridcol="8" gridrow="2" pageId="5" pageNumber="557">1 (13, ♂)</td>
<td id="76855DFD79146C97FBC6DE264CFAF9E6" box="[1131,1160,1555,1575]" gridcol="9" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FA8DDE264D59F9E6" box="[1312,1323,1555,1575]" gridcol="10" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FA1EDE264DCCF9E6" box="[1459,1470,1555,1575]" gridcol="11" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9A0DE264E6AF9E6" box="[1549,1560,1555,1575]" gridcol="12" gridrow="2" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9C9DE264F07F9E6" box="[1636,1909,1555,1575]" gridcol="13" gridrow="2" pageId="5" pageNumber="557">bl C, D, Q -</td>
</tr>
<tr id="3554348179146C97FFDADE0F4F07F98F" box="[119,1909,1594,1614]" gridrow="3" pageId="5" pageNumber="557">
<th id="76855DFD79146C97FFDADE0F48ABF98F" box="[119,217,1594,1614]" gridcol="0" gridrow="3" pageId="5" pageNumber="557">
<taxonomicName id="4C644D407914936DFFDADE0F48ABF98C" box="[119,217,1594,1613]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="557" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B910EAD17914936DFFDADE0F48ABF98C" box="[119,217,1594,1613]" italics="true" pageId="5" pageNumber="557">malayanus</emphasis>
</taxonomicName>
</th>
<td id="76855DFD79146C97FEA0DE0F4934F98F" box="[269,326,1594,1614]" gridcol="1" gridrow="3" pageId="5" pageNumber="557">70–83</td>
<td id="76855DFD79146C97FE24DE0F49E6F98F" box="[393,404,1594,1614]" gridcol="2" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FE7DDE0F49A9F98F" box="[464,475,1594,1614]" gridcol="3" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDBBDE0F4A53F98F" box="[534,545,1594,1614]" gridcol="4" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FDF0DE0F4AE5F98F" box="[605,663,1594,1614]" gridcol="5" gridrow="3" pageId="5" pageNumber="557">58–62</td>
<td id="76855DFD79146C97FD65DE0F4AA1F98F" box="[712,723,1594,1614]" gridcol="6" gridrow="3" pageId="5" pageNumber="557">1</td>
<td id="76855DFD79146C97FCA0DE0F4B34F98F" box="[781,838,1594,1614]" gridcol="7" gridrow="3" pageId="5" pageNumber="557">21–23</td>
<td id="76855DFD79146C97FCD4DE0F4BB9F98F" box="[889,971,1594,1614]" gridcol="8" gridrow="3" pageId="5" pageNumber="557">1 (8–10)</td>
<td id="76855DFD79146C97FBC6DE0F4CFAF98F" box="[1131,1160,1594,1614]" gridcol="9" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FA8DDE0F4D59F98F" box="[1312,1323,1594,1614]" gridcol="10" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97FA1EDE0F4DCCF98F" box="[1459,1470,1594,1614]" gridcol="11" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9A0DE0F4E6AF98F" box="[1549,1560,1594,1614]" gridcol="12" gridrow="3" pageId="5" pageNumber="557">0</td>
<td id="76855DFD79146C97F9C9DE0F4F07F98F" box="[1636,1909,1594,1614]" gridcol="13" gridrow="3" pageId="5" pageNumber="557">bd C, I, N -</td>
</tr>
</table>
</paragraph>
<caption id="DF1B664B7917936EFFD0DA944939FD74" ID-Table-UUID="DF1B664B7917936EFFD0DA944939FD74" box="[125,331,673,693]" httpUri="http://table.plazi.org/id/DF1B664B7917936EFFD0DA944939FD74" pageId="6" pageNumber="558" targetBox="[125,1910,727,914]" targetIsTable="true" targetPageId="6">
<paragraph id="8BDB36C37917936EFFD0DA944939FD74" blockId="6.[125,331,673,693]" box="[125,331,673,693]" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFFD0DA9448A9FD74" bold="true" box="[125,219,673,693]" pageId="6" pageNumber="558">TABLE 3</emphasis>
(continued)
</paragraph>
</caption>
<paragraph id="8BDB36C37917936EFFD0DAE24F15FCB6" pageId="6" pageNumber="558">
<table id="F964C46379176C97FFD0DAE24F04FC53" box="[125,1910,727,914]" gridcols="13" gridrows="3" pageId="6" pageNumber="558">
<tr id="3554348179176C97FFD0DAE24F04FCE0" box="[125,1910,727,801]" gridrow="0" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DAE2489FFCE0" box="[125,237,727,801]" gridcol="0" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFFD0DAE248CEFD2A" bold="true" box="[125,188,727,747]" pageId="6" pageNumber="558">Taxon</emphasis>
<taxonomicName id="4C644D407917936EFFD0DB39489FFCE0" box="[125,237,780,801]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD17917936EFFD0DB39489FFCE0" box="[125,237,780,801]" italics="true" pageId="6" pageNumber="558">marmoratus</emphasis>
</taxonomicName>
</th>
<th id="76855DFD79176C97FE8CDAE249EDFCE0" box="[289,415,727,801]" gridcol="1" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFE8CDAE24904FCC7" bold="true" pageId="6" pageNumber="558">SVL 1 in adults</emphasis>
45–82 0
</th>
<th id="76855DFD79176C97FE71DAE2499AFCE0" box="[476,488,727,801]" gridcol="2" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFE71DAE24995FD2A" bold="true" box="[476,487,727,747]" pageId="6" pageNumber="558">2</emphasis>
1
</th>
<th id="76855DFD79176C97FD8EDAE24A5DFCE0" box="[547,559,727,801]" gridcol="3" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFD8EDAE24A5CFD2A" bold="true" box="[547,558,727,747]" pageId="6" pageNumber="558">3</emphasis>
1
</th>
<th id="76855DFD79176C97FDC6DAE24AD7FCE0" box="[619,677,727,801]" gridcol="4" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFDC6DAE24A04FD2A" bold="true" box="[619,630,727,747]" pageId="6" pageNumber="558">4</emphasis>
38–47
</th>
<th id="76855DFD79176C97FD75DAE24A96FCE0" box="[728,740,727,801]" gridcol="5" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFD75DAE24A91FD2A" bold="true" box="[728,739,727,747]" pageId="6" pageNumber="558">5</emphasis>
0
</th>
<th id="76855DFD79176C97FCB3DAE24B2AFCE0" box="[798,856,727,801]" gridcol="6" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFCB3DAE24B5BFD2A" bold="true" box="[798,809,727,747]" pageId="6" pageNumber="558">6</emphasis>
19–24
</th>
<th id="76855DFD79176C97FC26DAE24B9EFCE0" box="[907,1004,727,801]" gridcol="7" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFC26DAE24BE4FD2A" bold="true" box="[907,918,727,747]" pageId="6" pageNumber="558">7</emphasis>
1 (16, ♀)
</th>
<th id="76855DFD79176C97FB2FDAE24C91FCE0" box="[1154,1251,727,801]" gridcol="8" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFB2FDAE24CFFFD2A" bold="true" box="[1154,1165,727,747]" pageId="6" pageNumber="558">8</emphasis>
1 (6−7, ♀)
</th>
<th id="76855DFD79176C97FA97DAE24E33FCE0" box="[1338,1601,727,801]" gridcol="9" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFA97DAE24E33FD2A" bold="true" box="[1338,1601,727,747]" pageId="6" pageNumber="558">9 10 11</emphasis>
1 (45–53, ♂) 0 (♀)/1 1
</th>
<th id="76855DFD79176C97F92EDAE24EE8FCE0" box="[1667,1690,727,801]" gridcol="10" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EF92EDAE24EE8FD2A" bold="true" box="[1667,1690,727,747]" pageId="6" pageNumber="558">12</emphasis>
bl
</th>
<th id="76855DFD79176C97F970DAE24F69FCE0" box="[1757,1819,727,801]" gridcol="11" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EF970DAE24F72FD2A" bold="true" box="[1757,1792,727,747]" pageId="6" pageNumber="558">Lit.</emphasis>
A, B
</th>
<th id="76855DFD79176C97F8F2DAE24F04FCE0" box="[1887,1910,727,801]" gridcol="12" gridrow="0" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EF8F2DAE24F1EFD2D" bold="true" box="[1887,1900,727,748]" italics="true" pageId="6" pageNumber="558">n</emphasis>
2
</th>
</tr>
<tr id="3554348179176C97FFD0DB024F04FC8C" box="[125,1910,823,845]" gridrow="1" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DB02489FFC8C" box="[125,237,823,845]" gridcol="0" gridrow="1" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DB0248B3FC8D" box="[125,193,823,844]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="matsuii">
<emphasis id="B910EAD17917936EFFD0DB0248B3FC8D" box="[125,193,823,844]" italics="true" pageId="6" pageNumber="558">matsuii</emphasis>
</taxonomicName>
</th>
<td id="76855DFD79176C97FE8CDB0249EDFC8C" box="[289,415,823,845]" gridcol="1" gridrow="1" pageId="6" pageNumber="558">105 1</td>
<td id="76855DFD79176C97FE71DB02499AFC8C" box="[476,488,823,845]" gridcol="2" gridrow="1" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FD8EDB024A5DFC8C" box="[547,559,823,845]" gridcol="3" gridrow="1" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FDC6DB024AD7FC8C" box="[619,677,823,845]" gridcol="4" gridrow="1" pageId="6" pageNumber="558">48−51</td>
<td id="76855DFD79176C97FD75DB024A96FC8C" box="[728,740,823,845]" gridcol="5" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FCB3DB024B2AFC8C" box="[798,856,823,845]" gridcol="6" gridrow="1" pageId="6" pageNumber="558">22</td>
<td id="76855DFD79176C97FC26DB024B9EFC8C" box="[907,1004,823,845]" gridcol="7" gridrow="1" pageId="6" pageNumber="558">0 (7−8, ♂)</td>
<td id="76855DFD79176C97FB2FDB024C91FC8C" box="[1154,1251,823,845]" gridcol="8" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FA97DB024E33FC8C" box="[1338,1601,823,845]" gridcol="9" gridrow="1" pageId="6" pageNumber="558">0 0 0</td>
<td id="76855DFD79176C97F92EDB024EE8FC8C" box="[1667,1690,823,845]" gridcol="10" gridrow="1" pageId="6" pageNumber="558">bl</td>
<td id="76855DFD79176C97F970DB024F69FC8C" box="[1757,1819,823,845]" gridcol="11" gridrow="1" pageId="6" pageNumber="558">I, N, R</td>
<td id="76855DFD79176C97F8F2DB024F04FC8C" box="[1887,1910,823,845]" gridcol="12" gridrow="1" pageId="6" pageNumber="558">-</td>
</tr>
<tr id="3554348179176C97FFD0DB6B4F04FC53" box="[125,1910,862,914]" gridrow="2" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DB6B489FFC53" box="[125,237,862,914]" gridcol="0" gridrow="2" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DB5648B4FCB9" box="[125,198,867,888]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="psarops">
<emphasis id="B910EAD17917936EFFD0DB5648B4FCB9" box="[125,198,867,888]" italics="true" pageId="6" pageNumber="558">psarops</emphasis>
</taxonomicName>
</th>
<td id="76855DFD79176C97FE8CDB6B49EDFC53" box="[289,415,862,914]" gridcol="1" gridrow="2" pageId="6" pageNumber="558">
? 
<emphasis id="B910EAD17917936EFE86DB6B4941FCAA" bold="true" box="[299,307,862,875]" pageId="6" pageNumber="558">b</emphasis>
1
</td>
<td id="76855DFD79176C97FE71DB6B499AFC53" box="[476,488,862,914]" gridcol="2" gridrow="2" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FD8EDB6B4A5DFC53" box="[547,559,862,914]" gridcol="3" gridrow="2" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FDC6DB6B4AD7FC53" box="[619,677,862,914]" gridcol="4" gridrow="2" pageId="6" pageNumber="558">38–49</td>
<td id="76855DFD79176C97FD75DB6B4A96FC53" box="[728,740,862,914]" gridcol="5" gridrow="2" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FCB3DB6B4B2AFC53" box="[798,856,862,914]" gridcol="6" gridrow="2" pageId="6" pageNumber="558">18–22</td>
<td id="76855DFD79176C97FC26DB6B4B9EFC53" box="[907,1004,862,914]" gridcol="7" gridrow="2" pageId="6" pageNumber="558">
n/a (? 
<emphasis id="B910EAD17917936EFC12DB6B4BB4FCAA" bold="true" box="[959,966,862,875]" pageId="6" pageNumber="558">c</emphasis>
,♀)
</td>
<td id="76855DFD79176C97FB2FDB6B4C91FC53" box="[1154,1251,862,914]" gridcol="8" gridrow="2" pageId="6" pageNumber="558">
n/a (? 
<emphasis id="B910EAD17917936EFB1BDB6B4CCFFCAA" bold="true" box="[1206,1213,862,875]" pageId="6" pageNumber="558">c</emphasis>
,♀)
</td>
<td id="76855DFD79176C97FA97DB6B4E33FC53" box="[1338,1601,862,914]" gridcol="9" gridrow="2" pageId="6" pageNumber="558">1 (28–32, ♂) 0 (♀)/1 1 (0–29,♀)</td>
<td id="76855DFD79176C97F92EDB6B4EE8FC53" box="[1667,1690,862,914]" gridcol="10" gridrow="2" pageId="6" pageNumber="558">bl</td>
<td id="76855DFD79176C97F970DB6B4F69FC53" box="[1757,1819,862,914]" gridcol="11" gridrow="2" pageId="6" pageNumber="558">S</td>
<td id="76855DFD79176C97F8F2DB6B4F04FC53" box="[1887,1910,862,914]" gridcol="12" gridrow="2" pageId="6" pageNumber="558">-</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDB36C37917936EFFD0DB9C4F15FBD4" pageId="6" pageNumber="558">
<table id="F964C46379176C97FFD0DB9C4F04FBEE" box="[125,1910,937,1071]" gridcols="13" gridrows="3" pageId="6" pageNumber="558">
<tr id="3554348179176C97FFD0DB9C4F04FC7F" box="[125,1910,937,958]" gridrow="0" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DB9C489AFC7F" box="[125,232,937,958]" gridcol="0" gridrow="0" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DB9C48ADFC7F" box="[125,223,937,958]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="pubisulcus">
<emphasis id="B910EAD17917936EFFD0DB9C48ADFC7F" box="[125,223,937,958]" italics="true" pageId="6" pageNumber="558">pubisulcus</emphasis>
</taxonomicName>
</th>
<th id="76855DFD79176C97FE8CDB9C4929FC7F" box="[289,347,937,958]" gridcol="1" gridrow="0" pageId="6" pageNumber="558">51–74</th>
<th id="76855DFD79176C97FE3EDB9C499AFC7F" box="[403,488,937,958]" gridcol="2" gridrow="0" pageId="6" pageNumber="558">0/1 1</th>
<th id="76855DFD79176C97FD8EDB9C4A5DFC7F" box="[547,559,937,958]" gridcol="3" gridrow="0" pageId="6" pageNumber="558">1</th>
<th id="76855DFD79176C97FDC6DB9C4AD6FC7F" box="[619,676,937,958]" gridcol="4" gridrow="0" pageId="6" pageNumber="558">37–58</th>
<th id="76855DFD79176C97FD75DB9C4A96FC7F" box="[728,740,937,958]" gridcol="5" gridrow="0" pageId="6" pageNumber="558">0</th>
<th id="76855DFD79176C97FCB3DB9C4B2AFC7F" box="[798,856,937,958]" gridcol="6" gridrow="0" pageId="6" pageNumber="558">16–23</th>
<th id="76855DFD79176C97FC26DB9C4B99FC7F" box="[907,1003,937,958]" gridcol="7" gridrow="0" pageId="6" pageNumber="558">1 (7–9, ♂)</th>
<th id="76855DFD79176C97FB2FDB9C4CECFC7F" box="[1154,1182,937,958]" gridcol="8" gridrow="0" pageId="6" pageNumber="558">0</th>
<th id="76855DFD79176C97FA97DB9C4DA9FC7F" box="[1338,1499,937,958]" gridcol="9" gridrow="0" pageId="6" pageNumber="558">0 0</th>
<th id="76855DFD79176C97F987DB9C4E44FC7F" box="[1578,1590,937,958]" gridcol="10" gridrow="0" pageId="6" pageNumber="558">1</th>
<th id="76855DFD79176C97F92EDB9C4EE7FC7F" box="[1667,1685,937,958]" gridcol="11" gridrow="0" pageId="6" pageNumber="558">bl</th>
<th id="76855DFD79176C97F970DB9C4F04FC7F" box="[1757,1910,937,958]" gridcol="12" gridrow="0" pageId="6" pageNumber="558">A, I, N, T 10</th>
</tr>
<tr id="3554348179176C97FFD0DBE14F04FC28" box="[125,1910,980,1001]" gridrow="1" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DBE1489AFC28" box="[125,232,980,1001]" gridcol="0" gridrow="1" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DBE148B9FC28" box="[125,203,980,1001]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="semiadii">
<emphasis id="B910EAD17917936EFFD0DBE148B9FC28" box="[125,203,980,1001]" italics="true" pageId="6" pageNumber="558">semiadii</emphasis>
</taxonomicName>
</th>
<td id="76855DFD79176C97FE8CDBE14929FC28" box="[289,347,980,1001]" gridcol="1" gridrow="1" pageId="6" pageNumber="558">40–47</td>
<td id="76855DFD79176C97FE3EDBE1499AFC28" box="[403,488,980,1001]" gridcol="2" gridrow="1" pageId="6" pageNumber="558">? 1</td>
<td id="76855DFD79176C97FD8EDBE14A5DFC28" box="[547,559,980,1001]" gridcol="3" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FDC6DBE14AD6FC28" box="[619,676,980,1001]" gridcol="4" gridrow="1" pageId="6" pageNumber="558">35–36</td>
<td id="76855DFD79176C97FD75DBE14A96FC28" box="[728,740,980,1001]" gridcol="5" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FCB3DBE14B2AFC28" box="[798,856,980,1001]" gridcol="6" gridrow="1" pageId="6" pageNumber="558">14–15</td>
<td id="76855DFD79176C97FC26DBE14B99FC28" box="[907,1003,980,1001]" gridcol="7" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FB2FDBE14CECFC28" box="[1154,1182,980,1001]" gridcol="8" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FA97DBE14DA9FC28" box="[1338,1499,980,1001]" gridcol="9" gridrow="1" pageId="6" pageNumber="558">0 0</td>
<td id="76855DFD79176C97F987DBE14E44FC28" box="[1578,1590,980,1001]" gridcol="10" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97F92EDBE14EE7FC28" box="[1667,1685,980,1001]" gridcol="11" gridrow="1" pageId="6" pageNumber="558">bl</td>
<td id="76855DFD79176C97F970DBE14F04FC28" box="[1757,1910,980,1001]" gridcol="12" gridrow="1" pageId="6" pageNumber="558">U -</td>
</tr>
<tr id="3554348179176C97FFD0DBC94F04FBEE" box="[125,1910,1020,1071]" gridrow="2" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DBC9489AFBEE" box="[125,232,1020,1071]" gridcol="0" gridrow="2" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DC35489AFBD4" box="[125,232,1024,1045]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="semicinctus">
<emphasis id="B910EAD17917936EFFD0DC35489AFBD4" box="[125,232,1024,1045]" italics="true" pageId="6" pageNumber="558">semicinctus</emphasis>
</taxonomicName>
</th>
<td id="76855DFD79176C97FE8CDBC94929FBEE" box="[289,347,1020,1071]" gridcol="1" gridrow="2" pageId="6" pageNumber="558">
? 
<emphasis id="B910EAD17917936EFE86DBC94941FBC8" bold="true" box="[299,307,1020,1033]" pageId="6" pageNumber="558">b</emphasis>
</td>
<td id="76855DFD79176C97FE3EDBC9499AFBEE" box="[403,488,1020,1071]" gridcol="2" gridrow="2" pageId="6" pageNumber="558">0 1</td>
<td id="76855DFD79176C97FD8EDBC94A5DFBEE" box="[547,559,1020,1071]" gridcol="3" gridrow="2" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FDC6DBC94AD6FBEE" box="[619,676,1020,1071]" gridcol="4" gridrow="2" pageId="6" pageNumber="558">33–44</td>
<td id="76855DFD79176C97FD75DBC94A96FBEE" box="[728,740,1020,1071]" gridcol="5" gridrow="2" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FCB3DBC94B2AFBEE" box="[798,856,1020,1071]" gridcol="6" gridrow="2" pageId="6" pageNumber="558">19–22</td>
<td id="76855DFD79176C97FC26DBC94B99FBEE" box="[907,1003,1020,1071]" gridcol="7" gridrow="2" pageId="6" pageNumber="558">n/a</td>
<td id="76855DFD79176C97FB2FDBC94CECFBEE" box="[1154,1182,1020,1071]" gridcol="8" gridrow="2" pageId="6" pageNumber="558">n/a</td>
<td id="76855DFD79176C97FA97DBC94DA9FBEE" box="[1338,1499,1020,1071]" gridcol="9" gridrow="2" pageId="6" pageNumber="558">1 (36–38, ♂) 1 (0–19,♀)</td>
<td id="76855DFD79176C97F987DBC94E44FBEE" box="[1578,1590,1020,1071]" gridcol="10" gridrow="2" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97F92EDBC94EE7FBEE" box="[1667,1685,1020,1071]" gridcol="11" gridrow="2" pageId="6" pageNumber="558">bl</td>
<td id="76855DFD79176C97F970DBC94F04FBEE" box="[1757,1910,1020,1071]" gridcol="12" gridrow="2" pageId="6" pageNumber="558">S -</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDB36C37917936EFFD0DC444F15FAC9" pageId="6" pageNumber="558">
<table id="F964C46379176C97FFD0DC444F04FAE2" box="[125,1910,1137,1315]" gridcols="13" gridrows="4" pageId="6" pageNumber="558">
<tr id="3554348179176C97FFD0DC444F04FB47" box="[125,1910,1137,1158]" gridrow="0" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DC4449EDFB47" box="[125,415,1137,1158]" gridcol="0" gridrow="0" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DC444974FB47" box="[125,262,1137,1158]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="B910EAD17917936EFFD0DC444974FB47" box="[125,262,1137,1158]" italics="true" pageId="6" pageNumber="558">quadrivirgatus</emphasis>
</taxonomicName>
51–71 1
</th>
<th id="76855DFD79176C97FE71DC44499AFB47" box="[476,488,1137,1158]" gridcol="1" gridrow="0" pageId="6" pageNumber="558">1</th>
<th id="76855DFD79176C97FD8EDC444A5DFB47" box="[547,559,1137,1158]" gridcol="2" gridrow="0" pageId="6" pageNumber="558">1</th>
<th id="76855DFD79176C97FDC6DC444AD6FB47" box="[619,676,1137,1158]" gridcol="3" gridrow="0" pageId="6" pageNumber="558">34–42</th>
<th id="76855DFD79176C97FD75DC444A96FB47" box="[728,740,1137,1158]" gridcol="4" gridrow="0" pageId="6" pageNumber="558">0</th>
<th id="76855DFD79176C97FCB3DC444B2AFB47" box="[798,856,1137,1158]" gridcol="5" gridrow="0" pageId="6" pageNumber="558">19–20</th>
<th id="76855DFD79176C97FC26DC444C3FFB47" box="[907,1101,1137,1158]" gridcol="6" gridrow="0" pageId="6" pageNumber="558">n/a (3–4, ♂)</th>
<th id="76855DFD79176C97FB2FDC444D7BFB47" box="[1154,1289,1137,1158]" gridcol="7" gridrow="0" pageId="6" pageNumber="558">n/a</th>
<th id="76855DFD79176C97FA97DC444D34FB47" box="[1338,1350,1137,1158]" gridcol="8" gridrow="0" pageId="6" pageNumber="558">1</th>
<th id="76855DFD79176C97FA62DC444DA9FB47" box="[1487,1499,1137,1158]" gridcol="9" gridrow="0" pageId="6" pageNumber="558">0</th>
<th id="76855DFD79176C97F987DC444E44FB47" box="[1578,1590,1137,1158]" gridcol="10" gridrow="0" pageId="6" pageNumber="558">0</th>
<th id="76855DFD79176C97F92EDC444F58FB47" box="[1667,1834,1137,1158]" gridcol="11" gridrow="0" pageId="6" pageNumber="558">bl, st D, N, W</th>
<th id="76855DFD79176C97F8F2DC444F04FB47" box="[1887,1910,1137,1158]" gridcol="12" gridrow="0" pageId="6" pageNumber="558">-</th>
</tr>
<tr id="3554348179176C97FFD0DCA84F04FB73" box="[125,1910,1181,1202]" gridrow="1" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DCA849EDFB73" box="[125,415,1181,1202]" gridcol="0" gridrow="1" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DCA848B9FB73" box="[125,203,1181,1202]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="wallacei">
<emphasis id="B910EAD17917936EFFD0DCA848B9FB73" box="[125,203,1181,1202]" italics="true" pageId="6" pageNumber="558">wallacei</emphasis>
</taxonomicName>
92–114 1
</th>
<td id="76855DFD79176C97FE71DCA8499AFB73" box="[476,488,1181,1202]" gridcol="1" gridrow="1" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FD8EDCA84A5DFB73" box="[547,559,1181,1202]" gridcol="2" gridrow="1" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FDC6DCA84AD6FB73" box="[619,676,1181,1202]" gridcol="3" gridrow="1" pageId="6" pageNumber="558">45–49</td>
<td id="76855DFD79176C97FD75DCA84A96FB73" box="[728,740,1181,1202]" gridcol="4" gridrow="1" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FCB3DCA84B2AFB73" box="[798,856,1181,1202]" gridcol="5" gridrow="1" pageId="6" pageNumber="558">17–25</td>
<td id="76855DFD79176C97FC26DCA84C3FFB73" box="[907,1101,1181,1202]" gridcol="6" gridrow="1" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FB2FDCA84D7BFB73" box="[1154,1289,1181,1202]" gridcol="7" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FA97DCA84D34FB73" box="[1338,1350,1181,1202]" gridcol="8" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FA62DCA84DA9FB73" box="[1487,1499,1181,1202]" gridcol="9" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97F987DCA84E44FB73" box="[1578,1590,1181,1202]" gridcol="10" gridrow="1" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97F92EDCA84F58FB73" box="[1667,1834,1181,1202]" gridcol="11" gridrow="1" pageId="6" pageNumber="558">bd, bl X</td>
<td id="76855DFD79176C97F8F2DCA84F04FB73" box="[1887,1910,1181,1202]" gridcol="12" gridrow="1" pageId="6" pageNumber="558">-</td>
</tr>
<tr id="3554348179176C97FFD0DCFD4F04FB1C" box="[125,1910,1224,1245]" gridrow="2" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DCFD49EDFB1C" box="[125,415,1224,1245]" gridcol="0" gridrow="2" pageId="6" pageNumber="558">
<taxonomicName id="4C644D407917936EFFD0DCFD4894FB1C" box="[125,230,1224,1245]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="wetariensis">
<emphasis id="B910EAD17917936EFFD0DCFD4894FB1C" box="[125,230,1224,1245]" italics="true" pageId="6" pageNumber="558">wetariensis</emphasis>
</taxonomicName>
42–67 1
</th>
<td id="76855DFD79176C97FE71DCFD499AFB1C" box="[476,488,1224,1245]" gridcol="1" gridrow="2" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FD8EDCFD4A5DFB1C" box="[547,559,1224,1245]" gridcol="2" gridrow="2" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FDC6DCFD4AD6FB1C" box="[619,676,1224,1245]" gridcol="3" gridrow="2" pageId="6" pageNumber="558">36–42</td>
<td id="76855DFD79176C97FD75DCFD4A96FB1C" box="[728,740,1224,1245]" gridcol="4" gridrow="2" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FCB3DCFD4B2AFB1C" box="[798,856,1224,1245]" gridcol="5" gridrow="2" pageId="6" pageNumber="558">15–22</td>
<td id="76855DFD79176C97FC26DCFD4C3FFB1C" box="[907,1101,1224,1245]" gridcol="6" gridrow="2" pageId="6" pageNumber="558">n/a (11, ♂)</td>
<td id="76855DFD79176C97FB2FDCFD4D7BFB1C" box="[1154,1289,1224,1245]" gridcol="7" gridrow="2" pageId="6" pageNumber="558">n/a (12–16, ♂)</td>
<td id="76855DFD79176C97FA97DCFD4D34FB1C" box="[1338,1350,1224,1245]" gridcol="8" gridrow="2" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FA62DCFD4DA9FB1C" box="[1487,1499,1224,1245]" gridcol="9" gridrow="2" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97F987DCFD4E44FB1C" box="[1578,1590,1224,1245]" gridcol="10" gridrow="2" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97F92EDCFD4F58FB1C" box="[1667,1834,1224,1245]" gridcol="11" gridrow="2" pageId="6" pageNumber="558">bl A, Y</td>
<td id="76855DFD79176C97F8F2DCFD4F04FB1C" box="[1887,1910,1224,1245]" gridcol="12" gridrow="2" pageId="6" pageNumber="558">4</td>
</tr>
<tr id="3554348179176C97FFD0DCC64F04FAE2" box="[125,1910,1267,1315]" gridrow="3" pageId="6" pageNumber="558">
<th id="76855DFD79176C97FFD0DCC649EDFAE2" box="[125,415,1267,1315]" gridcol="0" gridrow="3" pageId="6" pageNumber="558">
<emphasis id="B910EAD17917936EFFD0DCC648DFFAC9" box="[125,173,1267,1288]" italics="true" pageId="6" pageNumber="558">yoshi</emphasis>
75–96 1
</th>
<td id="76855DFD79176C97FE71DCC6499AFAE2" box="[476,488,1267,1315]" gridcol="1" gridrow="3" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FD8EDCC64A5DFAE2" box="[547,559,1267,1315]" gridcol="2" gridrow="3" pageId="6" pageNumber="558">1</td>
<td id="76855DFD79176C97FDC6DCC64AD6FAE2" box="[619,676,1267,1315]" gridcol="3" gridrow="3" pageId="6" pageNumber="558">50–58</td>
<td id="76855DFD79176C97FD75DCC64A96FAE2" box="[728,740,1267,1315]" gridcol="4" gridrow="3" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FCB3DCC64B2AFAE2" box="[798,856,1267,1315]" gridcol="5" gridrow="3" pageId="6" pageNumber="558">25–30</td>
<td id="76855DFD79176C97FC26DCC64C3FFAE2" box="[907,1101,1267,1315]" gridcol="6" gridrow="3" pageId="6" pageNumber="558">0 (8–12, ♂, absent or indistinct in ♀♀)</td>
<td id="76855DFD79176C97FB2FDCC64D7BFAE2" box="[1154,1289,1267,1315]" gridcol="7" gridrow="3" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FA97DCC64D34FAE2" box="[1338,1350,1267,1315]" gridcol="8" gridrow="3" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97FA62DCC64DA9FAE2" box="[1487,1499,1267,1315]" gridcol="9" gridrow="3" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97F987DCC64E44FAE2" box="[1578,1590,1267,1315]" gridcol="10" gridrow="3" pageId="6" pageNumber="558">0</td>
<td id="76855DFD79176C97F92EDCC64F58FAE2" box="[1667,1834,1267,1315]" gridcol="11" gridrow="3" pageId="6" pageNumber="558">bl C, I</td>
<td id="76855DFD79176C97F8F2DCC64F04FAE2" box="[1887,1910,1267,1315]" gridcol="12" gridrow="3" pageId="6" pageNumber="558">-</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDB36C37917936EFF26DD414C7EFA49" blockId="6.[125,1866,1357,1569]" box="[139,1036,1395,1416]" pageId="6" pageNumber="558">
<bibRefCitation id="EFF54B327917936EFF26DD414936FA49" box="[139,324,1395,1416]" pageId="6" pageNumber="566" refString="Harvey, M. B., O'Connell, K. A., Barraza, G., Riyanto, A., Kurniawan, N. &amp; Smith, E. N. (2015) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the southern Bukit Barisan range of Sumatra and an estimation of their phylogeny. Zootaxa, 4020 (3), 495 - 516. http: // dx. doi. org / 10.11646 / zootaxa. 4020.3.5" type="journal article">
Harvey 
<emphasis id="B910EAD17917936EFF79DD46488EFA49" box="[212,252,1395,1416]" italics="true" pageId="6" pageNumber="558">et al</emphasis>
. (2015)
</bibRefCitation>
did not provide ranges for adult specimens of 
<taxonomicName id="4C644D407917936EFD5DDD464B26FA49" box="[752,852,1395,1416]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="psarops">
<emphasis id="B910EAD17917936EFD5DDD464A8DFA49" box="[752,767,1395,1416]" italics="true" pageId="6" pageNumber="558">C</emphasis>
. 
<emphasis id="B910EAD17917936EFCA6DD464B26FA49" box="[779,852,1395,1416]" italics="true" pageId="6" pageNumber="558">psarops</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C644D407917936EFC2DDD464C74FA49" box="[896,1030,1395,1416]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="semicinctus">
<emphasis id="B910EAD17917936EFC2DDD464BFDFA49" box="[896,911,1395,1416]" italics="true" pageId="6" pageNumber="558">C</emphasis>
. 
<emphasis id="B910EAD17917936EFC36DD464C74FA49" box="[923,1030,1395,1416]" italics="true" pageId="6" pageNumber="558">semicinctus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB36C37917936EFF24DDAE4B4FFA14" blockId="6.[125,1866,1357,1569]" pageId="6" pageNumber="558">
In female 
<taxonomicName id="4C644D407917936EFF4BDDAF4938FA6E" box="[230,330,1434,1455]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="psarops">
<emphasis id="B910EAD17917936EFF4BDDAF4887FA6E" box="[230,245,1434,1455]" italics="true" pageId="6" pageNumber="558">C</emphasis>
. 
<emphasis id="B910EAD17917936EFEACDDAF4938FA6E" box="[257,330,1434,1455]" italics="true" pageId="6" pageNumber="558">psarops</emphasis>
</taxonomicName>
, pores can either be arranged continuously, discontinuously, or can be absent altogether. 
<bibRefCitation id="EFF54B327917936EFBD0DDAE4D44FA6E" box="[1149,1334,1434,1455]" pageId="6" pageNumber="566" refString="Harvey, M. B., O'Connell, K. A., Barraza, G., Riyanto, A., Kurniawan, N. &amp; Smith, E. N. (2015) Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the southern Bukit Barisan range of Sumatra and an estimation of their phylogeny. Zootaxa, 4020 (3), 495 - 516. http: // dx. doi. org / 10.11646 / zootaxa. 4020.3.5" type="journal article">
Harvey 
<emphasis id="B910EAD17917936EFB6BDDAF4C9CFA6E" box="[1222,1262,1434,1455]" italics="true" pageId="6" pageNumber="558">et al</emphasis>
. (2015)
</bibRefCitation>
only provided counts for total pore numbers, but failed to indicate individual counts for the pore-bearing parts of separated pore series.
</paragraph>
<paragraph id="8BDB36C37917936EFF26DDD24E2CFA3A" blockId="6.[125,1866,1357,1569]" box="[139,1630,1510,1531]" pageId="6" pageNumber="558">
In their diagnosis and 
<tableCitation id="C6E603787917936EFEFADDD249EEFA3A" box="[343,412,1511,1531]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Metric and meristic characters with abbreviations used in this study." pageId="6" pageNumber="558">Table 1</tableCitation>
, 
<bibRefCitation id="EFF54B327917936EFE05DDD24A17FA3A" box="[424,613,1510,1531]" pageId="6" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article">
Linkem 
<emphasis id="B910EAD17917936EFE59DDD34A6FFA3A" box="[500,541,1510,1531]" italics="true" pageId="6" pageNumber="558">et al</emphasis>
. (2008)
</bibRefCitation>
listed 12–13 precloacal pores for 
<taxonomicName id="4C644D407917936EFC30DDD34C75FA3A" box="[925,1031,1510,1531]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B910EAD17917936EFC30DDD34BDEFA3A" box="[925,940,1510,1531]" italics="true" pageId="6" pageNumber="558">C</emphasis>
. 
<emphasis id="B910EAD17917936EFC15DDD34C75FA3A" box="[952,1031,1510,1531]" italics="true" pageId="6" pageNumber="558">spinosus</emphasis>
</taxonomicName>
, whereas in their 
<tableCitation id="C6E603787917936EFB06DDD24C83FA3A" box="[1195,1265,1511,1531]" captionStart="TABLE 2" captionStartId="3.[151,244,151,176]" captionTargetBox="[151,1436,239,1258]" captionTargetPageId="3" captionText="TABLE 2. Metric (in mm) and meristic data for the type series of Cyrtodactylus klakahensis sp. nov. Abbreviations are defined in Table 1." httpUri="http://table.plazi.org/id/DF1B664B7912936BFF3AD8A24A45FF15" pageId="6" pageNumber="558" tableUuid="DF1B664B7912936BFF3AD8A24A45FF15">Table 2</tableCitation>
, they listed only 8–12 precloacal pores.
</paragraph>
<paragraph id="8BDB36C37917936EFF24DE384DBAF9E0" blockId="6.[125,1866,1357,1569]" box="[137,1480,1548,1569]" pageId="6" pageNumber="558">
According to the diagnosis in 
<bibRefCitation id="EFF54B327917936EFE30DE384A2FF9E0" box="[413,605,1548,1569]" pageId="6" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article">
Linkem 
<emphasis id="B910EAD17917936EFE47DE394A60F9E0" box="[490,530,1548,1569]" italics="true" pageId="6" pageNumber="558">et al</emphasis>
. (2008)
</bibRefCitation>
, 
<taxonomicName id="4C644D407917936EFDCADE394AA2F9E0" box="[615,720,1548,1569]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="558" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B910EAD17917936EFDCADE394A04F9E0" box="[615,630,1548,1569]" italics="true" pageId="6" pageNumber="558">C</emphasis>
. 
<emphasis id="B910EAD17917936EFD2CDE394AA2F9E0" box="[641,720,1548,1569]" italics="true" pageId="6" pageNumber="558">spinosus</emphasis>
</taxonomicName>
lacks femoral pores; it was listed as possessing 4–7 femoral pores in their 
<tableCitation id="C6E603787917936EFAD0DE384DB7F9E0" box="[1405,1477,1549,1569]" captionStart="TABLE 2" captionStartId="3.[151,244,151,176]" captionTargetBox="[151,1436,239,1258]" captionTargetPageId="3" captionText="TABLE 2. Metric (in mm) and meristic data for the type series of Cyrtodactylus klakahensis sp. nov. Abbreviations are defined in Table 1." httpUri="http://table.plazi.org/id/DF1B664B7912936BFF3AD8A24A45FF15" pageId="6" pageNumber="558" tableUuid="DF1B664B7912936BFF3AD8A24A45FF15">Table 2</tableCitation>
.
</paragraph>
<caption id="DF1B664B7916936FFF3ADF4F4AD8F82D" httpUri="https://zenodo.org/record/267813/files/figure.png" pageId="7" pageNumber="559" targetBox="[275,1312,197,1880]" targetPageId="7">
<paragraph id="8BDB36C37916936FFF3ADF4F4AD8F82D" blockId="7.[151,1435,1914,2028]" pageId="7" pageNumber="559">
<emphasis id="B910EAD17916936FFF3ADF4F4965F851" bold="true" box="[151,279,1914,1936]" pageId="7" pageNumber="559">FIGURE 1.</emphasis>
Morphological features of the holotype of 
<taxonomicName id="4C644D407916936FFD4ADF4F4B87F851" box="[743,1013,1914,1936]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="559" phylum="Chordata" rank="species" species="klakahensis" status="sp. nov.">
<emphasis id="B910EAD17916936FFD4ADF4F4B87F851" box="[743,1013,1914,1936]" italics="true" pageId="7" pageNumber="559">Cyrtodactylus klakahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357AA7916936FFC52DF4F4C26F851" box="[1023,1108,1914,1936]" pageId="7" pageNumber="559" rank="species">
<emphasis id="B910EAD17916936FFC52DF4F4C26F851" bold="true" box="[1023,1108,1914,1936]" pageId="7" pageNumber="559">sp. nov.</emphasis>
</taxonomicNameLabel>
(SMF 22476). (A) Dorsal and ventral view of the body. (B) Precloacal region showing precloacofemoral scales (bearing precloacofemoral pores) and posterior precloacal scales. (C) Lateral view of the left side of the head. (D) Ventral view of the head, showing pattern of postmental and gular scales. Photos by Sven Mecke.
</paragraph>
</caption>
<caption id="DF1B664B79199360FF3ADF924CB7F81A" httpUri="https://zenodo.org/record/267814/files/figure.png" pageId="8" pageNumber="560" targetBox="[303,1286,221,1902]" targetPageId="8">
<paragraph id="8BDB36C379199360FF3ADF924CB7F81A" blockId="8.[151,1437,1959,2011]" pageId="8" pageNumber="560">
<emphasis id="B910EAD179199360FF3ADF924961F87C" bold="true" box="[151,275,1959,1981]" pageId="8" pageNumber="560">FIGURE 2.</emphasis>
Paratype series of 
<taxonomicName id="4C644D4079199360FE75DF924A91F87C" box="[472,739,1959,1981]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="560" phylum="Chordata" rank="species" species="klakahensis" status="sp. nov.">
<emphasis id="B910EAD179199360FE75DF924A91F87C" box="[472,739,1959,1981]" italics="true" pageId="8" pageNumber="560">Cyrtodactylus klakahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357AA79199360FD47DF924B49F87C" box="[746,827,1959,1981]" pageId="8" pageNumber="560" rank="species">
<emphasis id="B910EAD179199360FD47DF924B49F87C" bold="true" box="[746,827,1959,1981]" pageId="8" pageNumber="560">sp. nov.</emphasis>
</taxonomicNameLabel>
in dorsal and ventral view. (A) SMF 22477, an adult male. (B) SMF 22478, an adult female. (C) SMF 22479, an unsexed juvenile specimen. Photos by Sven Mecke.
</paragraph>
</caption>
<paragraph id="8BDB36C379189361FF3ADC464956F9C1" blockId="9.[151,1436,1139,1536]" pageId="9" pageNumber="561">
<emphasis id="B910EAD179189361FF3ADC464961FB48" bold="true" box="[151,275,1139,1161]" pageId="9" pageNumber="561">FIGURE 3.</emphasis>
Comparison of the shape and arrangement of dorsal tubercles at midbody, the precloacal region in males, and the postmental and gular scale pattern between 
<taxonomicName id="4C644D4079189361FDFFDCA44B2EFB66" box="[594,860,1169,1191]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="561" phylum="Chordata" rank="species" species="klakahensis" status="sp. nov.">
<emphasis id="B910EAD179189361FDFFDCA44B2EFB66" box="[594,860,1169,1191]" italics="true" pageId="9" pageNumber="561">Cyrtodactylus klakahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357AA79189361FCCFDCA44BC1FB66" box="[866,947,1169,1191]" pageId="9" pageNumber="561" rank="species">
<emphasis id="B910EAD179189361FCCFDCA44BC1FB66" bold="true" box="[866,947,1169,1191]" pageId="9" pageNumber="561">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C644D4079189361FC6DDCA74C42FB66" box="[960,1072,1169,1191]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="561" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD179189361FC6DDCA74BA3FB66" box="[960,977,1170,1191]" italics="true" pageId="9" pageNumber="561">C</emphasis>
. 
<emphasis id="B910EAD179189361FC70DCA44C42FB66" box="[989,1072,1169,1191]" italics="true" pageId="9" pageNumber="561">fumosus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C644D4079189361FBC5DCA74D73FB66" box="[1128,1281,1170,1191]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="561" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD179189361FBC5DCA74C0BFB66" box="[1128,1145,1170,1191]" italics="true" pageId="9" pageNumber="561">C</emphasis>
. 
<emphasis id="B910EAD179189361FB28DCA74D73FB66" box="[1157,1281,1170,1191]" italics="true" pageId="9" pageNumber="561">marmoratus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C644D4079189361FAA3DCA4497FFB07" class="Reptilia" family="Gekkonidae" genus="Cyrtodacytlus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="561" phylum="Chordata" rank="species" species="klakahensis" status="sp. nov.">
<emphasis id="B910EAD179189361FAA3DCA4497FFB07" italics="true" pageId="9" pageNumber="561">Cyrtodacytlus klakahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357AA79189361FEB9DC854914FB07" box="[276,358,1200,1222]" pageId="9" pageNumber="561" rank="species">
<emphasis id="B910EAD179189361FEB9DC854914FB07" bold="true" box="[276,358,1200,1222]" pageId="9" pageNumber="561">sp. nov.</emphasis>
</taxonomicNameLabel>
: A) Closely arranged, trihedral, raised, and strongly keeled dorsal tubercles at midbody; B) eidonomy of precloacofemoral scales (three series), precloacal groove, and posterior precloacal scales; C) gular region, showing the presence of enlarged 2nd postmentals (photos and drawings of SMF 22476, 
<typeStatus id="54DF886179189361FC13DCC54C6EFAC7" box="[958,1052,1264,1286]" pageId="9" pageNumber="561" type="holotype">holotype</typeStatus>
). 
<taxonomicName id="4C644D4079189361FB9DDCC54D69FAC7" box="[1072,1307,1264,1286]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="561" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD179189361FB9DDCC54D69FAC7" box="[1072,1307,1264,1286]" italics="true" pageId="9" pageNumber="561">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
: D) Widely scattered, roundish, flat, and smooth dorsal tubercles at midbody; E) eidonomy of precloacofemoral scales (only one series distinctly enlarged), precloacal groove, and posterior precloacal scales; F) gular region, indicating the absence of enlarged 2nd postmentals (D, F = NMB-REPT 2662, 
<typeStatus id="54DF886179189361FD9FDD7B4AFCFAA5" box="[562,654,1358,1380]" pageId="9" pageNumber="561" type="holotype">holotype</typeStatus>
; E = BMNH 1896.12.9.3). 
<taxonomicName id="4C644D4079189361FC0ADD7B4CCAFAA5" box="[935,1208,1358,1380]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="561" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD179189361FC0ADD7B4CCAFAA5" box="[935,1208,1358,1380]" italics="true" pageId="9" pageNumber="561">Cyrtodactylus marmoratus</emphasis>
</taxonomicName>
: G) Closely arranged, slightly raised and keeled dorsal tubercles at midbody; H) eidonomy of precloacofemoral scales (three series) and precloacal groove, with posterior precloacal scales absent; I) gular region, indicating the absence of enlarged 2nd postmentals and the presence of a single pair of enlarged gular scales bordering the single pair of enlarged postmentals posteriorly (G = MTKD 8903; H, I = RMNH.RENA 2710a.1, 
<typeStatus id="54DF886179189361FDB2DDF94AF3FA23" box="[543,641,1484,1506]" pageId="9" pageNumber="561" type="lectotype">lectotype</typeStatus>
). Photos by Sven Mecke; line drawings by Felix Mader (based on photos by Sven Mecke).
</paragraph>
<paragraph id="8BDB36C379189361FF6ADE124B0DF8D6" blockId="9.[151,1436,1575,2032]" pageId="9" pageNumber="561">
<emphasis id="B910EAD179189361FF6ADE124C5BF981" bold="true" box="[199,1065,1575,1600]" pageId="9" pageNumber="561">
Description of the 
<typeStatus id="54DF886179189361FE0FDE124A79F981" box="[418,523,1575,1600]" pageId="9" pageNumber="561" type="holotype">holotype</typeStatus>
. General habitus, metrics (in mm) and ratios.
</emphasis>
Adult male; SVL = 68.1; AxialL = 31.9; TailL = 61.7 (tail regenerated); ArmL = 25.9; LegL = 36.3; HeadL = 18.1; HeadW = 10.6; HeadH = 7.0; SnoutL = 7.6; OrbEarL = 5.9; OrbD = 3.7; EarL = 1.3; head length moderate (HeadL/SVL = 0.27); head rather wide (HeadW/HeadL = 0.59), clearly depressed between eyes, distinct from neck; snout rather elongate (SnoutL/ HeadL = 0.42), much longer than OrbD (SnoutL/OrbD = 2.10), canthus rostralis distinct; fore- and hindlimbs of moderate size (ArmL/SVL = 0.38; LegL/SVL = 0.53), without webbing between digits; relative length of fingers = IV&gt; III&gt; V&gt; II&gt; I, relative length of toes = IV&gt; III&gt; V&gt; II&gt; I.
</paragraph>
<paragraph id="8BDB36C379189361FF6ADF164B34F831" blockId="9.[151,1436,1575,2032]" pageId="9" pageNumber="561">
<emphasis id="B910EAD179189361FF6ADF16494EF8FD" bold="true" box="[199,316,1827,1852]" pageId="9" pageNumber="561">Scalation.</emphasis>
Dorsal scales rounded and granulate, interspersed with distinctly enlarged, trihedral, raised, strongly keeled, and irregularly arranged dorsal tubercles (Figure 3A); 19 DTR; 26 PVT; latero-dorsal tubercles most strongly keeled, tubercles on lateral portion of the trunk and PVT smaller and feebly keeled; tubercles on base of the tail largest, strongly keeled, pointed and elongate, in eight rows (the longer part of the tail is regenerated, without tubercles present); tubercles on forelimbs small, most prominent on the antebrachium; tubercles on hindlimbs similar in size and shape to latero-dorsal tubercles.
</paragraph>
<paragraph id="8BDB36C3791B9362FF6AD8A24AA5FE49" blockId="10.[151,1437,151,2035]" pageId="10" pageNumber="562">
Ventral scales distinctly larger than dorsals, juxtaposed; 38 VS; three series of enlarged PFS, lowest series possessing 40 scales, bearing 37 pores; pore series interrupted by a single, enlarged, pore-less PFS on the left femur (PFS at level of this pore-less scale irregularly arranged and/or smaller; due to an aberration likely caused by an injury); scales immediately posterior to the precloacal groove (posterior precloacal scales) enlarged, arranged in a chevron-like shape consisting of three series (from anterior to posterior: nine scales/ seven scales/ three scales) (
<figureCitation id="135F2A46791B9362FF33D9794975FEA5" box="[158,263,332,357]" captionStart="FIGURE 1" captionStartId="7.[151,250,1914,1936]" captionTargetBox="[275,1312,197,1880]" captionTargetId="figure@7.[275,1312,195,1880]" captionTargetPageId="7" captionText="FIGURE 1. Morphological features of the holotype of Cyrtodactylus klakahensis sp. nov. (SMF 22476). (A) Dorsal and ventral view of the body. (B) Precloacal region showing precloacofemoral scales (bearing precloacofemoral pores) and posterior precloacal scales. (C) Lateral view of the left side of the head. (D) Ventral view of the head, showing pattern of postmental and gular scales. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267813/files/figure.png" pageId="10" pageNumber="562">Figures 1</figureCitation>
B &amp; 3B); two domed PCT; number of lamellae under fingers: I 
<date id="FFDA1003791B9362FC7BD9794C4AFEA5" box="[982,1080,332,356]" pageId="10" pageNumber="562" value="2011-02-12">11, II 12</date>
, III 
<date id="FFDA1003791B9362FBC5D9794CA7FEA4" box="[1128,1237,332,357]" pageId="10" pageNumber="562" value="2014-04-15">14, IV 15</date>
, V 15; number of lamellae under toes: I 
<date id="FFDA1003791B9362FE3DD9454983FE49" box="[400,497,367,392]" pageId="10" pageNumber="562" value="2014-02-16">14, II 16</date>
, III 
<date id="FFDA1003791B9362FD8DD9454AFFFE49" box="[544,653,368,392]" pageId="10" pageNumber="562" value="1918-04-20">18, IV 20</date>
, V 19.
</paragraph>
<paragraph id="8BDB36C3791B9362FF6AD9A34BA5FD77" blockId="10.[151,1437,151,2035]" pageId="10" pageNumber="562">Rostral shield rectangular, about 0.6 time high as wide, partly divided by a suture dorsally, in contact with 1st SupraLab, two rostronasals and a single InterNas; nostril surrounded by rostral, 1st SupraLab, three post-nasals, and a single rostro-nasal; R10 L10 SupraLab1, R6 L5 SupraLab2, separated from the orbit by 2–3 rows of small granular scales; R9 L9 InfraLab, bordered by two rows of scales larger than granular scales on the throat; cephalic scales small, rounded, granulate and juxtaposed; tubercles on occiput and neck raised, bearing an apex; 43 IOS; 26 SC; mental triangular, wider than long (MentalL/MentalW = 1.4); one pair of enlarged 1st postmentals, followed by a pair of enlarged 2nd postmentals (Figure 3C); pair of 1st postmentals bordered by mental, 1st InfraLab, enlarged 2nd postmentals, and four GulS (Figure 3C); scales on throat minute, rounded.</paragraph>
<paragraph id="8BDB36C3791B9362FF6ADAF74AEFFC73" blockId="10.[151,1437,151,2035]" pageId="10" pageNumber="562">
<emphasis id="B910EAD1791B9362FF6ADAF7493DFD1A" bold="true" box="[199,335,706,731]" pageId="10" pageNumber="562">Coloration.</emphasis>
In preservative, ground color of dorsal surface of head and body Drab (19); head with indistinct Walnut Brown (27) colored reticulum; a Burnt Sienna (38) stripe running from the posterior border of the orbit to a point above the ear opening; light Pale Buff (1) labial scales and postmentals strongly stippled with darker color; dorsum with irregular, faint Warm Sepia (40) blotches, most visible on neck and at level of posterior margin of the forelimbs, between hindlimbs and base of tail; ground color of dorsal surface of limbs like body; venter, throat and lower surface of limbs uniformly Pale Buff (1), heavily dotted and stippled with dark markings of different size; color of regenerated tail Pale Pinkish Buff (3).
</paragraph>
<paragraph id="8BDB36C3791B9362FF6ADB884CE6FB38" blockId="10.[151,1437,151,2035]" pageId="10" pageNumber="562">
<emphasis id="B910EAD1791B9362FF6ADB884949FC17" bold="true" box="[199,315,957,982]" pageId="10" pageNumber="562">Variation.</emphasis>
<typeStatus id="54DF8861791B9362FEE8DB8A49C0FC16" box="[325,434,959,983]" pageId="10" pageNumber="562" type="paratype">Paratypes</typeStatus>
similar to 
<typeStatus id="54DF8861791B9362FD86DB884AFFFC17" box="[555,653,957,982]" pageId="10" pageNumber="562" type="holotype">holotype</typeStatus>
except as follows: 
<typeStatus id="54DF8861791B9362FCC8DB8A4BB5FC16" box="[869,967,959,983]" pageId="10" pageNumber="562" type="paratype">Paratype</typeStatus>
SMF 22477 (adult male; 
<figureCitation id="135F2A46791B9362FB4ADB884D34FC17" box="[1255,1350,957,982]" captionStart="FIGURE 2" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[303,1286,221,1902]" captionTargetId="figure@8.[285,1298,193,1927]" captionTargetPageId="8" captionText="FIGURE 2. Paratype series of Cyrtodactylus klakahensis sp. nov. in dorsal and ventral view. (A) SMF 22477, an adult male. (B) SMF 22478, an adult female. (C) SMF 22479, an unsexed juvenile specimen. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267814/files/figure.png" pageId="10" pageNumber="562">Figure 2</figureCitation>
A) with 17 DTR; 29 PVT; dorsal scales on original part of tail arranged in whorls, each ending in a row of four tubercles; 36 VS; a series of 38 PFS bearing 38 PFP in a continuous series; posterior precloacal scales arranged in a chevronlike shape consisting of four series of scales (from anterior to posterior: nine scales/ nine scales/ six scales/ three scales); three domed PCT; number of lamellae under fingers: I 
<date id="FFDA1003791B9362FCFBDC7B4BC4FBA7" box="[854,950,1102,1126]" pageId="10" pageNumber="562" value="2014-02-14">14, II 14</date>
, III 
<date id="FFDA1003791B9362FC48DC7B4C22FBA7" box="[997,1104,1101,1126]" pageId="10" pageNumber="562" value="2014-04-15">14, IV 15</date>
, V 15 (counted on left side of the body; some fingers on right side damaged); number of lamellae under toes: I 
<date id="FFDA1003791B9362FB8DDC474CF3FB4A" box="[1056,1153,1138,1163]" pageId="10" pageNumber="562" value="2012-02-15">12, II 15</date>
, III 
<date id="FFDA1003791B9362FB1DDC474D6EFB4B" box="[1200,1308,1138,1163]" pageId="10" pageNumber="562" value="1916-04-17">16, IV 17</date>
, V 15; R11 L10 SupraLab1, R5 L5 SupraLab2; R9 L10 InfraLab; 27 SC; a single InterNas; a second dark stripe running from nostril to occiput, interrupted only by the orbit; a dark V-shaped collar present, followed by nine pairs of dark paravertebral blotches, larger than lateral ones, which may be fused to form bars anteriorly.
</paragraph>
<paragraph id="8BDB36C3791B9362FF6ADD304D42FA39" blockId="10.[151,1437,151,2035]" pageId="10" pageNumber="562">
<typeStatus id="54DF8861791B9362FF6ADD30495BFADC" box="[199,297,1285,1309]" pageId="10" pageNumber="562" type="paratype">Paratype</typeStatus>
SMF 22478 (adult female; 
<figureCitation id="135F2A46791B9362FDC6DD314ABEFADC" box="[619,716,1284,1309]" captionStart="FIGURE 2" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[303,1286,221,1902]" captionTargetId="figure@8.[285,1298,193,1927]" captionTargetPageId="8" captionText="FIGURE 2. Paratype series of Cyrtodactylus klakahensis sp. nov. in dorsal and ventral view. (A) SMF 22477, an adult male. (B) SMF 22478, an adult female. (C) SMF 22479, an unsexed juvenile specimen. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267814/files/figure.png" pageId="10" pageNumber="562">Figure 2</figureCitation>
B) with 17 DTR; 33 PVT; dorsal scales on original part of tail arranged in whorls, each ending in a row of four to six tubercles; 36 VS; 38 PFS, lacking pores; no sign of a precloacal groove; posterior precloacal scales arranged in a chevron-like shape consisting of three series of scales (from anterior to posterior: nine scales/ seven scales/ five scales); two domed PCT; number of lamellae under fingers: I 
<date id="FFDA1003791B9362FEAADDA0491EFA6C" box="[263,364,1429,1453]" pageId="10" pageNumber="562" value="2013-02-14">13, II 14</date>
, III 
<date id="FFDA1003791B9362FE33DDA04A7CFA6C" box="[414,526,1428,1453]" pageId="10" pageNumber="562" value="1918-04-16">18, IV 16</date>
, V 13; number of lamellae under toes I 
<date id="FFDA1003791B9362FC76DDA04C32FA6C" box="[987,1088,1428,1453]" pageId="10" pageNumber="562" value="2013-02-15">13, II 15</date>
, III 
<date id="FFDA1003791B9362FBDFDDA04C90FA6C" box="[1138,1250,1429,1453]" pageId="10" pageNumber="562" value="1919-04-17">19, IV 17</date>
, V 19; R10 L11 SupraLab1, R5 L6 SupraLab2; R8 L10 InfraLab; 39 IOS; 33 SC; three InterNas; eight pairs of dark paravertebral blotches, which may be fused to form bars; original part of tail with six Burnt Sienna (38) colored bands.
</paragraph>
<paragraph id="8BDB36C3791B9362FF6ADE31489DF8FE" blockId="10.[151,1437,151,2035]" pageId="10" pageNumber="562">
<typeStatus id="54DF8861791B9362FF6ADE31495BF9DD" box="[199,297,1540,1564]" pageId="10" pageNumber="562" type="paratype">Paratype</typeStatus>
SMF 22479 (unsexed juvenile; 
<figureCitation id="135F2A46791B9362FD39DE364A81F9DD" box="[660,755,1539,1564]" captionStart="FIGURE 2" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[303,1286,221,1902]" captionTargetId="figure@8.[285,1298,193,1927]" captionTargetPageId="8" captionText="FIGURE 2. Paratype series of Cyrtodactylus klakahensis sp. nov. in dorsal and ventral view. (A) SMF 22477, an adult male. (B) SMF 22478, an adult female. (C) SMF 22479, an unsexed juvenile specimen. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267814/files/figure.png" pageId="10" pageNumber="562">Figure 2</figureCitation>
C) with 15 DTR; dorsal surface on entirely original tail with scales arranged in whorls, ending in a row of tubercles, tubercles present on proximal quarter of tail only; 35 VS; 38 slightly enlarged PFS, lacking pores; no sign of a precloacal groove; posterior precloacal scales arranged in a chevron-like shape consisting of three series of scales (from anterior to posterior: eight scales/ eight scales/ five scales); two domed PCT; number of lamellae under fingers: I 
<date id="FFDA1003791B9362FCF3DEA14BB6F96D" box="[862,964,1683,1708]" pageId="10" pageNumber="562" value="2014-02-15">14, II 15</date>
, III 
<date id="FFDA1003791B9362FC5BDEA14C15F96D" box="[1014,1127,1684,1708]" pageId="10" pageNumber="562" value="1917-04-18">17, IV 18</date>
, V 15; number of lamellae under toes: I 
<date id="FFDA1003791B9362FE85DE8D49FBF911" box="[296,393,1720,1744]" pageId="10" pageNumber="562" value="2012-02-14">12, II 14</date>
, III 
<date id="FFDA1003791B9362FE1ADE824A51F911" box="[439,547,1719,1744]" pageId="10" pageNumber="562" value="1916-04-17">16, IV 17</date>
, V 15; R11 L10; SupraLab1, R5 L6 SupraLab2; R9 L8 InfraLab; 36 IOS; 28 SC; a single InterNas; a second dark stripe running from nostril to occiput, interrupted only by the orbit; a dark Verona Brown (37) V-shaped collar present; tail with 15 Verona Brown (37) colored bands, becoming Burnt Sienna (38) distally.
</paragraph>
</subSubSection>
<subSubSection id="C37E6548791B9362FF6ADF7F4C64F846" pageId="10" pageNumber="562" type="etymology">
<paragraph id="8BDB36C3791B9362FF6ADF7F4C64F846" blockId="10.[151,1437,151,2035]" pageId="10" pageNumber="562">
<emphasis id="B910EAD1791B9362FF6ADF7F493CF8A2" bold="true" box="[199,334,1866,1891]" pageId="10" pageNumber="562">Etymology.</emphasis>
The specific epithet is a Latinized, toponymic adjective referring to the 
<typeStatus id="54DF8861791B9362FB1DDF7E4C92F8A2" box="[1200,1248,1867,1891]" pageId="10" pageNumber="562">type</typeStatus>
locality Klakah (Lumajang Regency, Jawa Timur Province, Java, 
<collectingCountry id="F3737653791B9362FD6EDF5B4B40F846" box="[707,818,1902,1927]" name="Indonesia" pageId="10" pageNumber="562">Indonesia</collectingCountry>
) of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C37E6548791B9363FF6ADFA74D21FF14" lastPageId="11" lastPageNumber="563" pageId="10" pageNumber="562" type="distribution">
<paragraph id="8BDB36C3791B9363FF6ADFA74D21FF14" blockId="10.[151,1437,151,2035]" lastBlockId="11.[151,1437,151,213]" lastPageId="11" lastPageNumber="563" pageId="10" pageNumber="562">
<emphasis id="B910EAD1791B9362FF6ADFA74A2EF86A" bold="true" box="[199,604,1938,1963]" pageId="10" pageNumber="562">Distribution and Natural History.</emphasis>
<taxonomicName id="4C644D40791B9362FDC9DFA74BFEF86A" box="[612,908,1938,1963]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="562" phylum="Chordata" rank="species" species="klakahensis">
<emphasis id="B910EAD1791B9362FDC9DFA74BFEF86A" box="[612,908,1938,1963]" italics="true" pageId="10" pageNumber="562">Cyrtodactylus klakahensis</emphasis>
</taxonomicName>
is currently only known from its 
<typeStatus id="54DF8861791B9362FAA6DFA64D49F86A" box="[1291,1339,1939,1963]" pageId="10" pageNumber="562">type</typeStatus>
locality, Klakah, Lumajang Regency, Jawa Timur Province, Java, 
<collectingCountry id="F3737653791B9362FC8BDF834BE1F80E" box="[806,915,1974,1999]" name="Indonesia" pageId="10" pageNumber="562">Indonesia</collectingCountry>
(
<figureCitation id="135F2A46791B9362FC0EDF834C7AF80E" box="[931,1032,1974,1999]" captionStart="FIGURE 4" captionStartId="11.[151,250,1115,1137]" captionTargetBox="[151,1436,265,1061]" captionTargetId="figure@11.[151,1436,265,1093]" captionTargetPageId="11" captionText="FIGURE 4. Map of Java illustrating the type-, potential type-, and paratype localities of Cyrtodactylus spp. The black triangle marks the type locality of C. klakahensis. The black diamond marks the type locality of C. semiadii (paratype locality identified by a white diamond). The exact type locality of C. marmoratus in Java is unknown, but the type specimens were, in all probability, collected from western Java. The potential type locality (indicated by a black star) may lie within the mountainous area around Bogor, but a revision of the C. marmoratus species-complex is needed to narrowly restrict the type locality of C. marmoratus sensu stricto. Base map modified from Wikipedia by Sven Mecke." httpUri="https://zenodo.org/record/267815/files/figure.png" pageId="10" pageNumber="562">Figure 4</figureCitation>
). Klakah is located in the lowlands (elevation ca. 
<quantity id="4C9C9B26791B9362FE95DFEE49F0F832" box="[312,386,2011,2035]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="562" unit="m" value="200.0">200 m</quantity>
) between the Tengger and Iyang-Argapura mountain massifs. Although the species range is probably not restricted to Klakah, it may exhibit a relatively limited distribution in central Jawa Timur Province and therefore should be regarded as endemic to the region, until evidence to the contrary becomes available.
</paragraph>
</subSubSection>
<caption id="DF1B664B791A9363FF3ADC6E4BC9FACB" httpUri="https://zenodo.org/record/267815/files/figure.png" pageId="11" pageNumber="563" targetBox="[151,1436,265,1061]" targetPageId="11">
<paragraph id="8BDB36C3791A9363FF3ADC6E4BC9FACB" blockId="11.[151,1436,1115,1290]" pageId="11" pageNumber="563">
<emphasis id="B910EAD1791A9363FF3ADC6E4961FBB0" bold="true" box="[151,275,1115,1137]" pageId="11" pageNumber="563">FIGURE 4.</emphasis>
Map of Java illustrating the type-, potential type-, and paratype localities of 
<taxonomicName id="4C644D40791A9363FBB2DC6E4CDFFBB0" box="[1055,1197,1115,1137]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="genus">
<emphasis id="B910EAD1791A9363FBB2DC6E4CDFFBB0" box="[1055,1197,1115,1137]" italics="true" pageId="11" pageNumber="563">Cyrtodactylus</emphasis>
</taxonomicName>
spp. The black triangle marks the type locality of 
<taxonomicName id="4C644D40791A9363FE30DC4F4A42FB4E" box="[413,560,1145,1167]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="klakahensis">
<emphasis id="B910EAD1791A9363FE30DC4F49DCFB4E" box="[413,430,1146,1167]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FE17DC4C4A42FB4E" box="[442,560,1145,1167]" italics="true" pageId="11" pageNumber="563">klakahensis</emphasis>
</taxonomicName>
. The black diamond marks the type locality of 
<taxonomicName id="4C644D40791A9363FBA6DC4F4C0CFB4E" box="[1035,1150,1145,1167]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="semiadii">
<emphasis id="B910EAD1791A9363FBA6DC4F4C69FB4E" box="[1035,1051,1146,1167]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FB85DC4C4C0CFB4E" box="[1064,1150,1145,1167]" italics="true" pageId="11" pageNumber="563">semiadii</emphasis>
</taxonomicName>
(paratype locality identified by a white diamond). The exact type locality of 
<taxonomicName id="4C644D40791A9363FD0DDCAC4B4FFB6F" box="[672,829,1177,1198]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD1791A9363FD0DDCAC4AC3FB6F" box="[672,689,1177,1198]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FD6CDCAC4B4FFB6F" box="[705,829,1177,1198]" italics="true" pageId="11" pageNumber="563">marmoratus</emphasis>
</taxonomicName>
in Java is unknown, but the type specimens were, in all probability, collected from western Java. The potential type locality (indicated by a black star) may lie within the mountainous area around Bogor, but a revision of the 
<taxonomicName id="4C644D40791A9363FD95DCE34AA0FB2A" box="[568,722,1238,1259]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD1791A9363FD95DCE34A3BFB2A" box="[568,585,1238,1259]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FDFBDCE34AA0FB2A" box="[598,722,1238,1259]" italics="true" pageId="11" pageNumber="563">marmoratus</emphasis>
</taxonomicName>
species-complex is needed to narrowly restrict the type locality of 
<taxonomicName id="4C644D40791A9363FA29DCE34923FACB" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="subSpecies" species="marmoratus" subSpecies="sensu">
<emphasis id="B910EAD1791A9363FA29DCE34DE7FB2A" box="[1412,1429,1238,1259]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FF3ADCC04923FACB" box="[151,337,1269,1290]" italics="true" pageId="11" pageNumber="563">
marmoratus 
<taxonomicNameLabel id="A22357AA791A9363FEB4DCC04923FACB" box="[281,337,1269,1290]" pageId="11" pageNumber="563" sensu="stricto">sensu</taxonomicNameLabel>
</emphasis>
</taxonomicName>
stricto. Base map modified from Wikipedia by Sven Mecke.
</paragraph>
</caption>
<subSubSection id="C37E6548791A9365FF6ADD044892FC82" lastPageId="13" lastPageNumber="565" pageId="11" pageNumber="563" type="discussion">
<paragraph id="8BDB36C3791A9365FF6ADD044951FF37" blockId="11.[151,1437,1329,2038]" lastBlockId="13.[151,1436,151,835]" lastPageId="13" lastPageNumber="565" pageId="11" pageNumber="563">
<emphasis id="B910EAD1791A9363FF6ADD044B31FA8B" bold="true" box="[199,835,1329,1354]" pageId="11" pageNumber="563">
Remarks on the taxonomy of 
<taxonomicName id="4C644D40791A9363FD9FDD044B4EFA8B" box="[562,828,1329,1354]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FD9FDD044B4EFA8B" bold="true" box="[562,828,1329,1354]" italics="true" pageId="11" pageNumber="563">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
.
</emphasis>
The bent-toed gecko species 
<taxonomicName id="4C644D40791A9363FB37DD044DEEFA8B" box="[1178,1436,1329,1354]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FB37DD044DEEFA8B" box="[1178,1436,1329,1354]" italics="true" pageId="11" pageNumber="563">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
(proposed vernacular name: Sulawesi bent-toed gecko) was described by Friedrich (‘Fritz’) Müller (
<date id="FFDA1003791A9363FABEDD604DE6FAAF" box="[1299,1428,1365,1390]" pageId="11" pageNumber="563" value="1834" valueMax="1895">1834–1895</date>
) based on a single adult female (NMB-REPT 2662; 
<figureCitation id="135F2A46791A9363FD4CDD4C4B33FA53" box="[737,833,1401,1426]" captionStart="FIGURE 5" captionStartId="12.[151,250,1964,1986]" captionTargetBox="[214,1327,235,1861]" captionTargetId="figure@12.[169,1413,193,1927]" captionTargetPageId="12" captionText="FIGURE 5. Holotype of Cyrtodactylus fumosus (NMB-REPT 2662; adult female) in (A) dorsal, (B) lateral, and (C) ventral view. Photos by Sven Mecke." httpUri="https://zenodo.org/record/267816/files/figure.png" pageId="11" pageNumber="563">Figure 5</figureCitation>
) collected by Karl Friedrich (‘Fritz’) Sarasin (
<date id="FFDA1003791A9363FAF9DD4C48A2FA77" pageId="11" pageNumber="563" value="1859" valueMax="1941">1859– 1941</date>
) and Paul Benedict Sarasin (
<date id="FFDA1003791A9363FDBBDDA84AE5FA77" box="[534,663,1437,1462]" pageId="11" pageNumber="563" value="1856" valueMax="1929">1856–1929</date>
) in the “Bulawa Mountains” (North Sulawesi, elevation ca. 
<quantity id="4C9C9B26791A9363FA90DDAB4DE6FA77" box="[1341,1428,1438,1462]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="563" unit="m" value="1200.0">1200 m</quantity>
) (Müller 1895; the 1894 volume was issued in 1895). In his second article published in the 
<emphasis id="B910EAD1791A9363FB6EDDF44A46FA3F" italics="true" pageId="11" pageNumber="563">Verhandlungen der naturforschenden Gesellschaft Basel</emphasis>
(Reptilien und Amphibien aus Celebes, part II), 
<bibRefCitation id="EFF54B32791A9363FBF3DDD04D78FA3F" author="Muller" box="[1118,1290,1509,1534]" pageId="11" pageNumber="567" refString="Muller, F. (1895 b) Reptilien und Amphibien aus Celebes, (II. Bericht). Verhandlungen der naturforschenden Gesellschaft in Basel, 10, 862 - 869." type="journal article" year="1895" yearSuffix="b">Müller (1895b)</bibRefCitation>
mentioned a second specimen of the species (NMB-REPT 2663) from “Masarang” but referred to NMB-REPT 2662 as “ 
<emphasis id="B910EAD1791A9363FF0EDE184908F987" box="[163,378,1581,1606]" italics="true" pageId="11" pageNumber="563">Original-Exemplar</emphasis>
” (i.e., the 
<typeStatus id="54DF8861791A9363FE5DDE184A25F987" box="[496,599,1581,1606]" pageId="11" pageNumber="563" type="holotype">holotype</typeStatus>
). We therefore recognize NMB-REPT 2662 as the only 
<typeStatus id="54DF8861791A9363FB6ADE1B4C85F987" box="[1223,1271,1582,1606]" pageId="11" pageNumber="563">type</typeStatus>
specimen (see 
<bibRefCitation id="EFF54B32791A9363FF3ADE67491BF9AB" author="Brongersma" box="[151,361,1618,1642]" pageId="11" pageNumber="566" refString="Brongersma, L. D. (1934) Contributions to Indo-Australian herpetology. Zoologische Mededelingen, 17, 161 - 251." type="journal article" year="1934">Brongersma 1934</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791A9363FED6DE674A6BF9AB" author="Kramer" box="[379,537,1618,1642]" pageId="11" pageNumber="567" refString="Kramer, E. (1979) Typenkatalog der Echsen im Naturhistorischen Museum Basel (BM). Stand 1978. Revue Suisse de Zoologie, 86 (1), 159 - 166. http: // dx. doi. org / 10.5962 / bhl. part. 82283" type="journal article" year="1979">Kramer 1979</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791A9363FD86DE644AC2F9AB" author="Koch" box="[555,688,1617,1642]" pageId="11" pageNumber="567" refString="Koch, A. (2012) Discovery, Diversity, and Distribution of the Amphibians and Reptiles of Sulawesi and its Offshore Islands. Chimaira, Frankfurt am Main, 374 pp." type="book" year="2012">Koch 2012</bibRefCitation>
) in disagreement with de 
<bibRefCitation id="EFF54B32791A9363FC5FDE644CDFF9AB" author="Rooij" box="[1010,1197,1617,1642]" pageId="11" pageNumber="566" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Vol. I: Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, 384 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">Rooij (1915:17)</bibRefCitation>
who referred to an unspecified number of 
<typeStatus id="54DF8861791A9363FE31DE4349BEF94F" box="[412,460,1654,1678]" pageId="11" pageNumber="563">type</typeStatus>
specimens in a footnote. 
<bibRefCitation id="EFF54B32791A9363FD42DE404BC4F94F" author="Boulenger" box="[751,950,1653,1678]" pageId="11" pageNumber="566" refString="Boulenger, G. A. (1897) A catalogue of the reptiles and batrachians of Celebes with special reference to the collections made by Drs P &amp; F Sarasin in 1893 - 1896. Proceedings of the Royal Society of London, 1897, 193 - 237." type="proceedings paper" year="1897">Boulenger (1897)</bibRefCitation>
provided a more detailed description of 
<taxonomicName id="4C644D40791A9363FA2FDE434881F973" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FA2FDE434DE7F94C" box="[1410,1429,1654,1677]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FF3ADEAF4881F973" box="[151,243,1690,1714]" italics="true" pageId="11" pageNumber="563">fumosus</emphasis>
</taxonomicName>
based on four specimens, including NMB-REPT 
<date id="FFDA1003791A9363FC87DEAC4BFEF973" box="[810,908,1689,1714]" pageId="11" pageNumber="563" value="2662" valueMax="2663">2662–63</date>
and two additional North Sulawesi specimens donated by Paul Sarasin, BMNH 1895.2.27.7 and BMNH 1896.12.9.3, and corrected the 
<typeStatus id="54DF8861791A9363FB6ADE8B4C85F917" box="[1223,1271,1726,1750]" pageId="11" pageNumber="563">type</typeStatus>
locality from Boelawa Mountains (= Huidu Matabulawa) to Bone Mountains (= Pegunungan Bone). De 
<bibRefCitation id="EFF54B32791A9363FB0CDED44D40F93B" author="Rooij" box="[1185,1330,1761,1786]" pageId="11" pageNumber="566" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Vol. I: Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, 384 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">Rooij (1915)</bibRefCitation>
recorded 
<taxonomicName id="4C644D40791A9363FF3ADF334964F8DF" box="[151,278,1798,1822]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FF3ADF3348D8F8DC" box="[151,170,1798,1821]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FF17DF334964F8DF" box="[186,278,1798,1822]" italics="true" pageId="11" pageNumber="563">fumosus</emphasis>
</taxonomicName>
for Java based on a single specimen but 
<bibRefCitation id="EFF54B32791A9363FD5EDF334BBDF8DF" author="Brongersma" box="[755,975,1798,1822]" pageId="11" pageNumber="566" refString="Brongersma, L. D. (1934) Contributions to Indo-Australian herpetology. Zoologische Mededelingen, 17, 161 - 251." type="journal article" year="1934">Brongersma (1934)</bibRefCitation>
subsequently referred to this apparently misidentified voucher as 
<taxonomicName id="4C644D40791A9363FE02DF1F4A2AF880" box="[431,600,1834,1857]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD1791A9363FE02DF1F49B0F880" box="[431,450,1834,1857]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FE62DF1F4A2AF880" box="[463,600,1834,1857]" italics="true" pageId="11" pageNumber="563">marmoratus</emphasis>
</taxonomicName>
and restricted 
<taxonomicName id="4C644D40791A9363FD51DF1F4B0AF883" box="[764,888,1834,1858]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FD51DF1F4B7DF880" box="[764,783,1834,1857]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FCB1DF1F4B0AF883" box="[796,888,1834,1858]" italics="true" pageId="11" pageNumber="563">fumosus</emphasis>
</taxonomicName>
to Sulawesi. 
<bibRefCitation id="EFF54B32791A9363FBA0DF1F4CC8F883" author="Mertens" box="[1037,1210,1834,1858]" pageId="11" pageNumber="567" refString="Mertens, R. (1934) Die Amphibien und Reptilien der Deutschen Limnologischen Sunda-Expedition. In: Thienemann- Festband, A. (Ed.), Tropische Binnengewasser. Archiv fur Hydrobiologie, 40, pp. 677 - 701." type="journal article" year="1934">Mertens (1934)</bibRefCitation>
once again listed 
<taxonomicName id="4C644D40791A9363FA2FDF1F4881F8A7" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FA2FDF1F4DE7F880" box="[1410,1429,1834,1857]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FF3ADF7B4881F8A7" box="[151,243,1870,1894]" italics="true" pageId="11" pageNumber="563">fumosus</emphasis>
</taxonomicName>
as occuring on Java (Klakah, Lumajang) and clearly indicated that this assignment would be provisional with the genus being in need of a revision. Several subsequent authors have listed 
<taxonomicName id="4C644D40791A9363FBDDDF474C83F84B" box="[1136,1265,1906,1930]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FBDDDF474CF1F848" box="[1136,1155,1906,1929]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FB38DF474C83F84B" box="[1173,1265,1906,1930]" italics="true" pageId="11" pageNumber="563">fumosus</emphasis>
</taxonomicName>
as part of the Javanese herpetofauna (e.g., 
<bibRefCitation id="EFF54B32791A9363FE4ADFA04B4EF86F" author="Manthey" box="[487,828,1941,1966]" pageId="11" pageNumber="567" refString="Manthey, U. &amp; Grossmann, W. (1997) Amphibien &amp; Reptilien Sudostasiens. Natur und Tier-Verlag, Munster, Germany, 512 pp." type="book" year="1997">Manthey &amp; Grossmann 1997</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791A9363FCE1DFA04C42F86F" author="Hayden" box="[844,1072,1941,1966]" pageId="11" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Hayden 
<emphasis id="B910EAD1791A9363FC03DFA24B91F86F" box="[942,995,1941,1966]" italics="true" pageId="11" pageNumber="563">et al</emphasis>
. 2008
</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791A9363FBEDDFA04D64F86F" author="Oliver" box="[1088,1302,1941,1966]" pageId="11" pageNumber="567" refString="Oliver, P., Edgar, P., Mumpuni, Iskandar, D. T. &amp; Lilley, R. (2009) A new species of bent-toed gecko (Cyrtodactylus: Gekkonidae) from Seram Island, Indonesia. Zootaxa, 2115, 47 - 55." type="journal article" year="2009">
Oliver 
<emphasis id="B910EAD1791A9363FB3EDFA24CBBF86F" box="[1171,1225,1941,1966]" italics="true" pageId="11" pageNumber="563">et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791A9363FA88DFA34DE5F86F" author="Das" box="[1317,1431,1942,1966]" pageId="11" pageNumber="566" refString="Das, I. (2010) A Field Guide to the Reptiles of South-east Asia. New Holland Publisher, London, 376 pp." type="book" year="2010">Das 2010</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791A9363FF3ADF8C4903F813" author="Riyanto" box="[151,369,1977,2002]" pageId="11" pageNumber="567" refString="Riyanto, A., Bauer, A. M. &amp; Yudha, D. S. (2014) A new small karst-dwelling species of Cyrtodactylus from Java, Indonesia. Zootaxa, 3785 (4), 589 - 599. http: // dx. doi. org / 10.11646 / zootaxa. 3785.4.7" type="journal article" year="2014">
Riyanto 
<emphasis id="B910EAD1791A9363FF5ADF8F495BF813" box="[247,297,1977,2002]" italics="true" pageId="11" pageNumber="563">et al</emphasis>
. 2014
</bibRefCitation>
) but these records seem to be either based on erroneous data provided in the literature (e.g., de 
<bibRefCitation id="EFF54B32791A9363FF3ADFE84965F837" author="Rooij" box="[151,279,2013,2038]" pageId="11" pageNumber="566" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. Vol. I: Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, 384 pp. http: // dx. doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">Rooij 1915</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791A9363FE8EDFEB49E2F837" author="Das" box="[291,400,2014,2038]" pageId="11" pageNumber="566" refString="Das, I. (2010) A Field Guide to the Reptiles of South-east Asia. New Holland Publisher, London, 376 pp." type="book" year="2010">Das 2010</bibRefCitation>
) and/or misidentified specimens. 
<bibRefCitation id="EFF54B32791A9363FCB5DFE84B8DF837" box="[792,1023,2013,2038]" pageId="11" pageNumber="567" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Setiadi, I. M., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toed gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica, 64 (2), 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article">
Linkem 
<emphasis id="B910EAD1791A9363FCD8DFEA4BD4F837" box="[885,934,2013,2038]" italics="true" pageId="11" pageNumber="563">et al</emphasis>
. (2008)
</bibRefCitation>
did not consider 
<taxonomicName id="4C644D40791A9363FB6FDFEB4D4CF837" box="[1218,1342,2014,2038]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="563" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791A9363FB6FDFEB4CA7F834" box="[1218,1237,2014,2037]" italics="true" pageId="11" pageNumber="563">C</emphasis>
. 
<emphasis id="B910EAD1791A9363FB4FDFEB4D4CF837" box="[1250,1342,2014,2038]" italics="true" pageId="11" pageNumber="563">fumosus</emphasis>
</taxonomicName>
a valid species, stating that it had been synonymized with 
<taxonomicName id="4C644D40791C9365FCBDD8AD4BB0FF71" box="[784,962,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD1791C9365FCBDD8AD4B51FF6E" box="[784,803,152,175]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FC94D8AC4BB0FF71" box="[825,962,153,176]" italics="true" pageId="13" pageNumber="565">marmoratus</emphasis>
</taxonomicName>
. However, 
<bibRefCitation id="EFF54B32791C9365FBFED8AD4D44FF71" author="Brongersma" box="[1107,1334,152,176]" pageId="13" pageNumber="566" refString="Brongersma, L. D. (1934) Contributions to Indo-Australian herpetology. Zoologische Mededelingen, 17, 161 - 251." type="journal article" year="1934">Brongersma (1934)</bibRefCitation>
did not synonymize 
<taxonomicName id="4C644D40791C9365FE84D88E49DBFF12" box="[297,425,187,211]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FE84D88E494EFF13" box="[297,316,187,210]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FEE0D88E49DBFF12" box="[333,425,187,211]" italics="true" pageId="13" pageNumber="565">fumosus</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C644D40791C9365FE42D88E4AEFFF12" box="[495,669,187,211]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B910EAD1791C9365FE42D88E4A70FF13" box="[495,514,187,210]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FDB9D8894AEFFF12" box="[532,669,188,211]" italics="true" pageId="13" pageNumber="565">marmoratus</emphasis>
</taxonomicName>
but indicated both taxa were distinct (see also 
<bibRefCitation id="EFF54B32791C9365FB65D88E4DE5FF12" author="Brongersma" box="[1224,1431,186,211]" pageId="13" pageNumber="566" refString="Brongersma, L. D. (1953) Gymnodactylus marmoratus. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, 62, 172 - 175." type="journal article" year="1953">Brongersma 1953</bibRefCitation>
; 
<bibRefCitation id="EFF54B32791C9365FF3AD8E8496AFF37" author="Koch" box="[151,280,221,246]" pageId="13" pageNumber="567" refString="Koch, A. (2012) Discovery, Diversity, and Distribution of the Amphibians and Reptiles of Sulawesi and its Offshore Islands. Chimaira, Frankfurt am Main, 374 pp." type="book" year="2012">Koch 2012</bibRefCitation>
).
</paragraph>
<caption id="DF1B664B791D9364FF3ADF9949B1F821" httpUri="https://zenodo.org/record/267816/files/figure.png" pageId="12" pageNumber="564" targetBox="[214,1327,235,1861]" targetPageId="12">
<paragraph id="8BDB36C3791D9364FF3ADF9949B1F821" blockId="12.[151,1435,1964,2016]" pageId="12" pageNumber="564">
<emphasis id="B910EAD1791D9364FF3ADF994966F803" bold="true" box="[151,276,1964,1986]" pageId="12" pageNumber="564">FIGURE 5.</emphasis>
Holotype of 
<taxonomicName id="4C644D40791D9364FE0DDF994AFBF803" box="[416,649,1964,1986]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="564" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791D9364FE0DDF994AFBF803" box="[416,649,1964,1986]" italics="true" pageId="12" pageNumber="564">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
(NMB-REPT 2662; adult female) in (A) dorsal, (B) lateral, and (C) ventral view. Photos by Sven Mecke.
</paragraph>
</caption>
<paragraph id="8BDB36C3791C9365FF6AD8CA4CA1FDCA" blockId="13.[151,1436,151,835]" pageId="13" pageNumber="565">
There is considerable confusion in the literature as to whether 
<taxonomicName id="4C644D40791C9365FC39D8CA4CE5FED9" box="[916,1175,255,280]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FC39D8CA4CE5FED9" box="[916,1175,255,280]" italics="true" pageId="13" pageNumber="565">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
possesses a precloacal groove in males. 
<taxonomicName id="4C644D40791C9365FEF1D9174A2CFEFA" box="[348,606,290,315]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FEF1D9174A2CFEFA" box="[348,606,290,315]" italics="true" pageId="13" pageNumber="565">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
was referred to as a species lacking a precloacal groove by 
<bibRefCitation id="EFF54B32791C9365FABDD9174896FE9F" pageId="13" pageNumber="567" refString="Rosler, H., Richards, S. J. &amp; Gunther, R. (2007) Remarks on morphology and taxonomy of geckos of the genus Cyrtodactylus Gray, 1827, occurring east of Wallacea, with descriptions of two new species (Reptilia: Sauria: Gekkonidae). Salamandra, 43 (4), 193 - 230." type="journal article">
Rösler 
<emphasis id="B910EAD1791C9365FACFD9114DE7FEFA" box="[1378,1429,290,315]" italics="true" pageId="13" pageNumber="565">et al</emphasis>
. (2007)
</bibRefCitation>
, 
<bibRefCitation id="EFF54B32791C9365FF5CD9704A48FE9F" author="Grismer" box="[241,570,325,350]" pageId="13" pageNumber="566" refString="Grismer, L. L. &amp; Norhayati, A. (2008) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa, 1924, 53 - 68." type="journal article" year="2008">Grismer &amp; Norhayati (2008)</bibRefCitation>
, 
<bibRefCitation id="EFF54B32791C9365FDEAD9704B5DFE9F" box="[583,815,325,350]" pageId="13" pageNumber="568" refString="Welton, L. J., Siler, C. D., Diesmos, A. &amp; Brown, R. M. (2009) A new bent-toed gecko (genus: Cyrtodactylus) from southern Palawan Island, Philippines and clarification of the taxonomic status of C. annulatus. Herpetologica, 65 (3), 328 - 343. http: // dx. doi. org / 10.1655 / 08 - 057 R 1.1" type="journal article">
Welton 
<emphasis id="B910EAD1791C9365FD0DD9734AA0FE9F" box="[672,722,325,350]" italics="true" pageId="13" pageNumber="565">et al</emphasis>
. (2009)
</bibRefCitation>
, 
<bibRefCitation id="EFF54B32791C9365FC91D9704C17FE9F" author="Chan" box="[828,1125,325,350]" pageId="13" pageNumber="566" refString="Chan, K. O. &amp; Norhayati, A. (2010) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia. Zootaxa, 2389, 47 - 56." type="journal article" year="2010">Chan &amp; Norhayati (2010)</bibRefCitation>
, and 
<bibRefCitation id="EFF54B32791C9365FB0ED9704DEAFE9F" box="[1187,1432,325,350]" pageId="13" pageNumber="566" refString="Grismer, L. L., Wood, P. L. Jr. &amp; Lim, K. K. P. (2012) Cyrtodactylus majulah, a new species of bent-toed gecko Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology, 60 (2), 487 - 499." type="journal article">
Grismer 
<emphasis id="B910EAD1791C9365FAA5D9734D49FE9F" box="[1288,1339,325,350]" italics="true" pageId="13" pageNumber="565">et al</emphasis>
. (2012)
</bibRefCitation>
. 
<bibRefCitation id="EFF54B32791C9365FF3AD9524914FE41" author="Boulenger" box="[151,358,359,384]" pageId="13" pageNumber="566" refString="Boulenger, G. A. (1897) A catalogue of the reptiles and batrachians of Celebes with special reference to the collections made by Drs P &amp; F Sarasin in 1893 - 1896. Proceedings of the Royal Society of London, 1897, 193 - 237." type="proceedings paper" year="1897">Boulenger (1897)</bibRefCitation>
, who examined 
<taxonomicName id="4C644D40791C9365FD9DD95D4AC1FE41" box="[560,691,360,384]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FD9DD95D4A31FEBE" box="[560,579,360,383]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FDFAD95D4AC1FE41" box="[599,691,360,384]" italics="true" pageId="13" pageNumber="565">fumosus</emphasis>
</taxonomicName>
specimens from North Sulawesi (Bone Mountains, Masarang, Rurukan; all collected by Fritz and Paul Sarasin), reported that male individuals of this species did possess a precloacal groove. We examined the adult male specimen (BMNH 1896.12.9.3) used by Boulenger for his species account of 
<taxonomicName id="4C644D40791C9365FEB1D9E549ECFE29" box="[284,414,464,488]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FEB1D9E5495DFE26" box="[284,303,464,487]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FEEFD9E549ECFE29" box="[322,414,464,488]" italics="true" pageId="13" pageNumber="565">fumosus</emphasis>
</taxonomicName>
and confirm the presence of a precloacal groove in this species. A precloacal groove, however, may be only weakly defined in subadult male specimens, as seen in NMB-REPT 2663.
</paragraph>
<paragraph id="8BDB36C3791C9365FF6ADA234892FC82" blockId="13.[151,1436,151,835]" pageId="13" pageNumber="565">
It appears that several species may be masquerading under the name 
<taxonomicName id="4C644D40791C9365FBAFDA204D75FDEF" box="[1026,1287,533,558]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FBAFDA204D75FDEF" box="[1026,1287,533,558]" italics="true" pageId="13" pageNumber="565">Cyrtodactylus fumosus</emphasis>
</taxonomicName>
, both in the Greater and Lesser Sunda Islands (
<bibRefCitation id="EFF54B32791C9365FDEEDA024B45FD91" author="Riyanto" box="[579,823,567,592]" pageId="13" pageNumber="567" refString="Riyanto, A., Bauer, A. M. &amp; Yudha, D. S. (2014) A new small karst-dwelling species of Cyrtodactylus from Java, Indonesia. Zootaxa, 3785 (4), 589 - 599. http: // dx. doi. org / 10.11646 / zootaxa. 3785.4.7" type="journal article" year="2014">
Riyanto 
<emphasis id="B910EAD1791C9365FD01DA0C4A96FD91" box="[684,740,567,592]" italics="true" pageId="13" pageNumber="565">et al</emphasis>
. 2014
</bibRefCitation>
; pers. obs.). Above, we were able to demonstrate unequivocally that the specimens from Klakah, Lumajang Regency, Java, originally referred to as 
<taxonomicName id="4C644D40791C9365FB54DA6E4D0AFDB2" box="[1273,1400,603,627]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FB54DA6E4D7EFDB3" box="[1273,1292,603,626]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FAB1DA6E4D0AFDB2" box="[1308,1400,603,627]" italics="true" pageId="13" pageNumber="565">fumosus</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFF54B32791C9365FF3ADA4B4934FD57" author="Mertens" box="[151,326,638,662]" pageId="13" pageNumber="567" refString="Mertens, R. (1934) Die Amphibien und Reptilien der Deutschen Limnologischen Sunda-Expedition. In: Thienemann- Festband, A. (Ed.), Tropische Binnengewasser. Archiv fur Hydrobiologie, 40, pp. 677 - 701." type="journal article" year="1934">Mertens (1934)</bibRefCitation>
, represent a new species. The populations on Bali referred to as 
<taxonomicName id="4C644D40791C9365FBB4DA4B4CE4FD57" box="[1049,1174,638,662]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FBB4DA4B4C5EFD54" box="[1049,1068,638,661]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FB97DA4B4CE4FD57" box="[1082,1174,638,662]" italics="true" pageId="13" pageNumber="565">fumosus</emphasis>
</taxonomicName>
by Mc 
<bibRefCitation id="EFF54B32791C9365FB41DA4B4D1EFD57" author="Kay" box="[1260,1388,637,662]" pageId="13" pageNumber="567" refString="McKay, J. L. (2006) A Field Guide to the Amphibians and Reptiles of Bali. Krieger Publishing Company, Malabar, Florida, 138 pp." type="book" year="2006">Kay (2006)</bibRefCitation>
and additional forms from Java are likely also new species (Mecke 
<emphasis id="B910EAD1791C9365FCF2DA944BE3FD79" box="[863,913,671,696]" italics="true" pageId="13" pageNumber="565">et al</emphasis>
., in prep., Riyanto 
<emphasis id="B910EAD1791C9365FBC6DA944CEFFD79" box="[1131,1181,671,696]" italics="true" pageId="13" pageNumber="565">et al</emphasis>
., in prep.). Based on a photograph of a strongly tuberculated individual of 
<taxonomicName id="4C644D40791C9365FD4EDAF64BF4FD1A" box="[739,902,706,731]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FD4EDAF64A84FD1B" box="[739,758,707,730]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. cf. 
<emphasis id="B910EAD1791C9365FC87DAF64BF4FD1A" box="[810,902,707,731]" italics="true" pageId="13" pageNumber="565">fumosus</emphasis>
</taxonomicName>
in 
<bibRefCitation id="EFF54B32791C9365FC00DAF74C05FD1A" author="Koch" box="[941,1143,706,731]" pageId="13" pageNumber="567" refString="Koch, A. (2012) Discovery, Diversity, and Distribution of the Amphibians and Reptiles of Sulawesi and its Offshore Islands. Chimaira, Frankfurt am Main, 374 pp." type="book" year="2012">Koch (2012: 151)</bibRefCitation>
from North Sulawesi, we assume that Sulawesi 
<taxonomicName id="4C644D40791C9365FE21DAD34A7AFD3F" box="[396,520,742,766]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="species" species="fumosus">
<emphasis id="B910EAD1791C9365FE21DAD349EDFD3C" box="[396,415,742,765]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FE01DAD34A7AFD3F" box="[428,520,742,766]" italics="true" pageId="13" pageNumber="565">fumosus</emphasis>
</taxonomicName>
populations may represent a species complex as well. The taxon is in dire need of a revision, with a thorough re-description of 
<taxonomicName id="4C644D40791C9365FD08DB3D4B1CFCE1" box="[677,878,776,800]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="565" phylum="Chordata" rank="subSpecies" species="fumosus" subSpecies="sensu">
<emphasis id="B910EAD1791C9365FD08DB3D4ACAFCDE" box="[677,696,776,799]" italics="true" pageId="13" pageNumber="565">C</emphasis>
. 
<emphasis id="B910EAD1791C9365FD67DB3D4B1CFCE1" box="[714,878,776,800]" italics="true" pageId="13" pageNumber="565">
fumosus 
<taxonomicNameLabel id="A22357AA791C9365FC9DDB3C4B1CFCE1" box="[816,878,777,800]" pageId="13" pageNumber="565" sensu="stricto">sensu</taxonomicNameLabel>
</emphasis>
</taxonomicName>
stricto currently underway (Hartmann 
<emphasis id="B910EAD1791C9365FA94DB3C4D1FFCE1" box="[1337,1389,775,800]" italics="true" pageId="13" pageNumber="565">et al</emphasis>
., in prep.).
</paragraph>
</subSubSection>
</treatment>
</document>