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<document ID-DOI="http://doi.org/10.5281/zenodo.3854812" ID-GBIF-Dataset="40effc56-2cc7-40ae-9592-c7baafeccef1" ID-GBIF-Taxon="163976989" ID-ISSN="13132970" ID-Zenodo-Dep="3854812" approvalRequired="53" approvalRequired_for_bibRefs="1" approvalRequired_for_textStreams="51" approvalRequired_for_treatments="1" checkinTime="1587997431956" checkinUser="plazi" docAuthor="Lourenco, Wilson &amp; Duhem, Bernard" docDate="2009" docId="03CD87CC650BC13BFF16FEA1FC85B912" docLanguage="en" docName="ZK_article_2004.pdf" docOrigin="ZooKeys 14" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@18c8ecf5" docStyleName="zookeys.2008.journal_article" docTitle="Saharobuthus elegans Lourenco &amp; Duhem 2009, sp. n." docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="45" masterDocId="FFF4FFB46508C13CFF9AFFF7FFFEBB75" masterDocTitle="Saharo-Sindian buthid scorpions; description of two new genera and species from Occidental Sahara and Afghanistan" masterLastPageNumber="54" masterPageNumber="37" pageId="3" pageNumber="40" updateTime="1636548053746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Saharo-Sindian buthid scorpions; description of two new genera and species from Occidental Sahara and Afghanistan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lourenco, Wilson</mods:namePart>
<mods:affiliation>Museum national d Histoire naturelle, Paris, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Duhem, Bernard</mods:namePart>
<mods:affiliation>Muséum national d'Histoire naturelle, Paris ,, France</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-07-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>14</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>37</mods:start>
<mods:end>54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.14.212</mods:identifier>
<mods:identifier type="GBIF-Dataset">40effc56-2cc7-40ae-9592-c7baafeccef1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576470</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3854812" ID-GBIF-Taxon="163976989" ID-Zenodo-Dep="3854812" LSID="urn:lsid:plazi:treatment:03CD87CC650BC13BFF16FEA1FC85B912" httpUri="http://treatment.plazi.org/id/03CD87CC650BC13BFF16FEA1FC85B912" lastPageId="7" lastPageNumber="45" pageId="3" pageNumber="40">
<subSubSection box="[140,469,342,369]" pageId="3" pageNumber="40" type="nomenclature">
<paragraph blockId="3.[140,968,342,440]" box="[140,469,342,369]" pageId="3" pageNumber="40">
<heading box="[140,469,342,369]" level="2" pageId="3" pageNumber="40">
<emphasis bold="true" box="[140,469,342,369]" pageId="3" pageNumber="40">
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[140,397,342,369]" class="Arachnida" family="Buthidae" genus="Saharobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="elegans" status="sp. n.">
<emphasis bold="true" box="[140,397,342,369]" italics="true" pageId="3" pageNumber="40">Saharobuthus elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[404,469,343,369]" pageId="3" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="40" type="description">
<paragraph blockId="3.[140,968,342,440]" box="[140,968,378,405]" pageId="3" pageNumber="40">
<uri box="[140,968,378,405]" pageId="3" pageNumber="40">
urn:lsid:zoobank.org:act:
<uuid box="[421,968,378,405]" pageId="3" pageNumber="40">3C2F50F7-DD92-4854-8584-CBE4FF62B084</uuid>
</uri>
</paragraph>
<paragraph blockId="3.[140,968,342,440]" box="[140,318,413,440]" pageId="3" pageNumber="40">
Figs
<figureCitation box="[192,207,413,439]" captionStart="Figures 1-8" captionStartId="4.[140,222,1558,1580]" captionTargetBox="[310,943,177,1536]" captionTargetId="figure@4.[336,919,851,1538]" captionTargetPageId="4" captionText="Figures 1-8. Carapace and tergites I-II. 1. Saharobuthus elegans sp. n., male holotype. 2. Compsobuthus acutecarinatus, female holotype. 3. Leiurus quinquestriatus, male. 4. Mesobuthus eupeus persicus, female. 5. Pantobuthus complicatus sp. n., female holotype. 6. Odontobuthus doriae, female. 7. Buthus draa, female. 8. Hottentotta schach, male (scales = 5 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770927" httpUri="https://zenodo.org/record/3770927/files/figure.png" pageId="3" pageNumber="40">1</figureCitation>
,
<figureCitation box="[221,277,413,440]" captionStart="Figures 9-17" captionStartId="6.[140,222,1558,1580]" captionTargetBox="[164,1092,178,1502]" captionTargetId="figure@6.[137,1110,165,1526]" captionTargetPageId="6" captionText="Figures 9-17. Saharobuthus elegans sp. n., male holotype. 9. Chelicerae, dorsal aspect. 10. Sternum, genital operculum and pectines. 11. Chela, dorso-external aspect. 12-13. Patella, dorsal and external aspects. 14. Femur, dorsal aspect. 15. Cutting edge of movable finger with rows of granules.16-17. Idem for Compsobuthus acutecarinatus, female holotype (scales = 3 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770929" httpUri="https://zenodo.org/record/3770929/files/figure.png" pageId="3" pageNumber="40">9-15</figureCitation>
,
<figureCitation box="[289,318,413,439]" captionStart="Figures 29-32. 29" captionStartId="10.[140,222,1590,1612]" captionTargetBox="[164,1038,207,1546]" captionTargetId="figure@10.[140,1108,180,1565]" captionTargetPageId="10" captionText="Figures 29-32. 29. Pedipalp, dorsal aspect, Vachoniolus globimanus, male holotype (scale = 5 mm). 30- 32. Metasoma and telson, lateral aspect. 30. Saharobuthus elegans sp. n., male holotype. 31. Pantobuthus complicatus sp. n., female holotype. 32. Odontobuthus doriae, female (scales 30-32 = 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770933" httpUri="https://zenodo.org/record/3770933/files/figure.png" pageId="3" pageNumber="40">30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[140,512,483,510]" pageId="3" pageNumber="40" type="diagnosis">
<paragraph blockId="3.[140,1108,483,1672]" box="[140,512,483,510]" pageId="3" pageNumber="40">
<emphasis bold="true" box="[140,267,483,509]" pageId="3" pageNumber="40">Diagnosis:</emphasis>
as for the new genus.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="40" type="materials_examined">
<paragraph blockId="3.[140,1108,483,1672]" pageId="3" pageNumber="40">
<emphasis bold="true" box="[185,358,518,545]" pageId="3" pageNumber="40">Type material:</emphasis>
<materialsCitation ID-GBIF-Occurrence="2610752302" collectingDate="1964-05-25" collectorName="P. L. Dekeyser" elevation="430" location="Adrar-Sotuf" municipality="Sahara" pageId="3" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Al Gharbiyah" typeStatus="holotype">
<collectingRegion box="[365,489,519,545]" country="Egypt" name="Al Gharbiyah" pageId="3" pageNumber="40">Occidental</collectingRegion>
<collectingMunicipality box="[498,576,519,545]" pageId="3" pageNumber="40">Sahara</collectingMunicipality>
, region of
<location LSID="urn:lsid:plazi:treatment:03CD87CC650BC13BFF16FEA1FC85B912:8EBB6001650BC13FFD23FDF0FCBCB954" box="[697,834,519,545]" municipality="Sahara" name="Adrar-Sotuf" pageId="3" pageNumber="40" stateProvince="Al Gharbiyah">Adrar-Sotuf</location>
,
<quantity box="[847,922,519,545]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="40" unit="m" value="430.0">
<elevation box="[847,922,519,545]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="40" unit="m" value="430.0">430 m</elevation>
</quantity>
alt.,
<date box="[980,1107,518,545]" pageId="3" pageNumber="40" value="1964-05-25">
<collectingDate box="[980,1107,518,545]" pageId="3" pageNumber="40" value="1964-05-25">25/V/1964</collectingDate>
</date>
(
<collectorName box="[148,313,554,581]" pageId="3" pageNumber="40">P. L. Dekeyser</collectorName>
),
<specimenCount box="[331,406,554,580]" pageId="3" pageNumber="40" type="male">1 male</specimenCount>
<typeStatus box="[413,516,554,580]" pageId="3" pageNumber="40">holotype</typeStatus>
. Deposited in the collection of the Muséum national dHistoire naturelle, Paris
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="40" type="etymology">
<paragraph blockId="3.[140,1108,483,1672]" pageId="3" pageNumber="40">
<emphasis bold="true" box="[185,323,624,650]" pageId="3" pageNumber="40">Etymology:</emphasis>
The specific name is a reference to the very slender build of the new species.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="44" pageId="3" pageNumber="40" type="description">
<paragraph blockId="3.[140,1108,483,1672]" box="[185,953,695,721]" pageId="3" pageNumber="40">
Description based on the male
<typeStatus box="[537,636,695,721]" pageId="3" pageNumber="40">holotype</typeStatus>
(Measurements in
<tableCitation box="[853,936,695,721]" captionStart="Table 1" captionStartId="5.[142,209,166,190]" captionTargetBox="[149,1077,244,1475]" captionTargetId="graphics@5.[141,1106,806,874]" captionTargetPageId="5" captionText="Table 1. Morphometric values (in mm) of the male holotype of Saharobuthus elegans sp. n., and the female holotype of Pantobuthus complicatus sp. n." httpUri="http://table.plazi.org/id/DF1B6652650DC139FF14FF51FD25BBAB" pageId="3" pageNumber="40" tableUuid="DF1B6652650DC139FF14FF51FD25BBAB">Table 1</tableCitation>
).
</paragraph>
<paragraph blockId="3.[140,1108,483,1672]" pageId="3" pageNumber="40">Coloration. Basically yellowish to pale yellow, with some light reddish zones. Prosoma: carapace yellowish with carinae and granulations reddish; median and lateral eyes surrounded by black pigment. Mesosoma: yellowish with some very light greyish zones. Metasoma: all segments yellowish, with carinae reddish. Vesicle pale yellow; base of aculeus yellowish with a reddish tip. Venter yellowish; pectines pale yellow. Chelicerae yellowish without spots; fingers yellowish with reddish teeth. Pedipalps yellowish with carinae slightly reddish; rows of granules on fingers reddish. Legs yellowish with some diffused spots.</paragraph>
<paragraph blockId="3.[140,1108,483,1672]" pageId="3" pageNumber="40">
Morphology. Prosoma: Anterior margin of carapace not emarginated, almost straight. Carapace carinae strongly developed; anterior median, central median and posterior median carinae strongly marked; central median and posterior median fused; central lateral strongly marked; posterior median carinae terminating distally in a small spinoid process that extends slightly beyond the posterior margin of the carapace (process less marked than in the genus
<taxonomicName authorityName="Vachon" authorityYear="1949" box="[519,677,1187,1213]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis box="[519,677,1187,1213]" italics="true" pageId="3" pageNumber="40">Compsobuthus</emphasis>
</taxonomicName>
). Intercarinal spaces strongly granular on anterior and posterior zones; moderate to weak on the other regions. Median ocular tubercle anterior to the centre of the carapace; median eyes separated by two ocular diameters. Three pairs of lateral eyes. Mesosoma: Tergites I-VI tricarinate. Lateral carinae on I-VI strongly marked; each carina terminating distally with a spinoid process that extends slightly beyond the posterior margin of the tergite. Median carinae on I moderate; on II-VI strong, crenulate; terminating distally on each segment with a spinoid process that also extends slightly beyond the posterior margin of the tergite. Tergite VII pentacarinate, with lateral pairs of carinae strong; median carinae present on proximal half, strong. Intercarinal spaces strongly granular. Sternites: Lateral carinae absent from sternites III to
<collectionCode box="[310,343,1540,1566]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="40">VI</collectionCode>
; moderate, finely crenulate on VII. Submedian carinae on sternites III-VI absent; on VII moderate. Pectines long; pectinal tooth count 28-29. Metasoma: Segments I-II with 10 carinae, crenulate; III-IV with 8 carinae. Segment
<collectionCode box="[1000,1020,1610,1636]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="40" type="Museum">V</collectionCode>
with 5 carinae; ventromedian carinae strongly marked, with several lobate denticles; anal arc
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770927" ID-Zenodo-Dep="3770927" httpUri="https://zenodo.org/record/3770927/files/figure.png" pageId="4" pageNumber="41" startId="4.[140,222,1558,1580]" targetBox="[310,943,177,1536]" targetPageId="4">
<paragraph blockId="4.[140,1109,1558,1676]" pageId="4" pageNumber="41">
<emphasis bold="true" box="[140,269,1558,1580]" pageId="4" pageNumber="41">Figures 1-8.</emphasis>
Carapace and tergites I-II.
<emphasis bold="true" box="[525,545,1558,1580]" pageId="4" pageNumber="41">1.</emphasis>
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[551,740,1558,1579]" class="Arachnida" family="Buthidae" genus="Saharobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="elegans" status="sp. n.">
<emphasis box="[551,740,1558,1579]" italics="true" pageId="4" pageNumber="41">Saharobuthus elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[746,798,1559,1580]" pageId="4" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
, male holotype.
<emphasis bold="true" box="[952,972,1558,1580]" pageId="4" pageNumber="41">2.</emphasis>
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1882" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis italics="true" pageId="4" pageNumber="41">Compsobuthus acutecarinatus</emphasis>
</taxonomicName>
, female holotype.
<emphasis bold="true" box="[437,458,1590,1612]" pageId="4" pageNumber="41">3.</emphasis>
<taxonomicName baseAuthorityName="Ehrenberg" baseAuthorityYear="1828" box="[463,672,1591,1612]" class="Arachnida" family="Buthidae" genus="Leiurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="quinquestriatus">
<emphasis box="[463,672,1591,1612]" italics="true" pageId="4" pageNumber="41">Leiurus quinquestriatus</emphasis>
</taxonomicName>
, male.
<emphasis bold="true" box="[739,759,1590,1612]" pageId="4" pageNumber="41">4.</emphasis>
<taxonomicName box="[764,1004,1590,1612]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="eupeus" subSpecies="persicus">
<emphasis box="[764,1004,1590,1612]" italics="true" pageId="4" pageNumber="41">Mesobuthus eupeus persicus</emphasis>
</taxonomicName>
, female.
<emphasis bold="true" box="[1088,1108,1590,1612]" pageId="4" pageNumber="41">5.</emphasis>
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[140,359,1622,1643]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis box="[140,359,1622,1643]" italics="true" pageId="4" pageNumber="41">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[367,419,1623,1644]" pageId="4" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
, female holotype.
<emphasis bold="true" box="[592,612,1622,1644]" pageId="4" pageNumber="41">6.</emphasis>
<taxonomicName baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[618,807,1622,1643]" class="Arachnida" family="Buthidae" genus="Odontobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="doriae">
<emphasis box="[618,807,1622,1643]" italics="true" pageId="4" pageNumber="41">Odontobuthus doriae</emphasis>
</taxonomicName>
, female.
<emphasis bold="true" box="[893,913,1622,1644]" pageId="4" pageNumber="41">7.</emphasis>
<taxonomicName box="[919,1028,1622,1643]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="draa">
<emphasis box="[919,1028,1622,1643]" italics="true" pageId="4" pageNumber="41">Buthus draa</emphasis>
</taxonomicName>
, female.
<emphasis bold="true" box="[140,160,1654,1676]" pageId="4" pageNumber="41">8.</emphasis>
<taxonomicName baseAuthorityName="Birula" baseAuthorityYear="1905" box="[166,330,1654,1676]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="schach">
<emphasis box="[166,330,1654,1676]" italics="true" pageId="4" pageNumber="41">Hottentotta schach</emphasis>
</taxonomicName>
, male (scales = 5 mm).
</paragraph>
</caption>
<caption ID-Table-UUID="DF1B6652650DC139FF14FF51FD25BBAB" httpUri="http://table.plazi.org/id/DF1B6652650DC139FF14FF51FD25BBAB" pageId="5" pageNumber="42" startId="5.[142,209,166,190]" targetBox="[149,1077,244,1475]" targetIsTable="true" targetPageId="5">
<paragraph blockId="5.[142,1105,166,222]" box="[142,1105,166,190]" pageId="5" pageNumber="42">
<emphasis bold="true" box="[142,240,166,190]" pageId="5" pageNumber="42">Table 1.</emphasis>
Morphometric values (in mm) of the male
<typeStatus box="[697,787,166,190]" pageId="5" pageNumber="42">holotype</typeStatus>
of
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[824,1033,166,190]" class="Arachnida" family="Buthidae" genus="Saharobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="elegans" status="sp. n.">
<emphasis box="[824,1033,166,190]" italics="true" pageId="5" pageNumber="42">Saharobuthus elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1041,1099,167,190]" pageId="5" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
,
</paragraph>
<paragraph blockId="5.[142,1105,166,222]" box="[142,731,198,222]" pageId="5" pageNumber="42">
and the female
<typeStatus box="[300,390,198,222]" pageId="5" pageNumber="42">holotype</typeStatus>
of
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[425,668,198,222]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis box="[425,668,198,222]" italics="true" pageId="5" pageNumber="42">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[675,731,199,222]" pageId="5" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="42">
<table box="[149,1077,244,1475]" gridcols="3" gridrows="37" pageId="5" pageNumber="42">
<tr box="[149,1077,244,266]" gridrow="0" pageId="5" pageNumber="42" rowspan-0="1">
<th box="[431,700,244,266]" gridcol="1" gridrow="0" pageId="5" pageNumber="42">
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[431,642,244,266]" class="Arachnida" family="Buthidae" genus="Saharobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="elegans" status="sp. n.">
<emphasis bold="true" box="[431,642,244,266]" italics="true" pageId="5" pageNumber="42">Saharobuthus elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[647,700,244,265]" pageId="5" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</th>
<th box="[775,1077,244,266]" gridcol="2" gridrow="0" pageId="5" pageNumber="42">
<emphasis bold="true" box="[775,1077,244,266]" pageId="5" pageNumber="42">
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[775,1019,244,266]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis bold="true" box="[775,1019,244,266]" italics="true" pageId="5" pageNumber="42">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1024,1077,244,265]" pageId="5" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr box="[149,1077,278,299]" gridrow="1" pageId="5" pageNumber="42" rowspan-0="1">
<td box="[431,700,278,299]" gridcol="1" gridrow="1" pageId="5" pageNumber="42">
<emphasis bold="true" box="[524,619,278,299]" pageId="5" pageNumber="42">-holotype</emphasis>
</td>
<td box="[775,1077,278,299]" gridcol="2" gridrow="1" pageId="5" pageNumber="42">
<emphasis bold="true" box="[882,977,278,299]" pageId="5" pageNumber="42">-holotype</emphasis>
</td>
</tr>
<tr box="[149,1077,311,334]" gridrow="2" pageId="5" pageNumber="42">
<th box="[149,327,311,334]" gridcol="0" gridrow="2" pageId="5" pageNumber="42">Total length</th>
<td box="[431,700,311,334]" gridcol="1" gridrow="2" pageId="5" pageNumber="42">49.4(56.0*)</td>
<td box="[775,1077,311,334]" gridcol="2" gridrow="2" pageId="5" pageNumber="42">66.3(73.5*)</td>
</tr>
<tr box="[149,1077,345,366]" gridrow="3" pageId="5" pageNumber="42">
<th box="[149,1077,345,366]" colspan="3" colspanRight="2" gridcol="0" gridrow="3" pageId="5" pageNumber="42">Carapace</th>
</tr>
<tr box="[149,1077,379,401]" gridrow="4" pageId="5" pageNumber="42">
<th box="[149,327,379,401]" gridcol="0" gridrow="4" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,379,401]" gridcol="1" gridrow="4" pageId="5" pageNumber="42">6.7</td>
<td box="[775,1077,379,401]" gridcol="2" gridrow="4" pageId="5" pageNumber="42">7.9</td>
</tr>
<tr box="[149,1077,412,434]" gridrow="5" pageId="5" pageNumber="42">
<th box="[149,327,412,434]" gridcol="0" gridrow="5" pageId="5" pageNumber="42">- anterior width</th>
<td box="[431,700,412,434]" gridcol="1" gridrow="5" pageId="5" pageNumber="42">4.3</td>
<td box="[775,1077,412,434]" gridcol="2" gridrow="5" pageId="5" pageNumber="42">5.2</td>
</tr>
<tr box="[149,1077,446,468]" gridrow="6" pageId="5" pageNumber="42">
<th box="[149,327,446,468]" gridcol="0" gridrow="6" pageId="5" pageNumber="42">- posterior width</th>
<td box="[431,700,446,468]" gridcol="1" gridrow="6" pageId="5" pageNumber="42">7.1</td>
<td box="[775,1077,446,468]" gridcol="2" gridrow="6" pageId="5" pageNumber="42">8.8</td>
</tr>
<tr box="[149,1077,479,501]" gridrow="7" pageId="5" pageNumber="42">
<th box="[149,1077,479,501]" colspan="3" colspanRight="2" gridcol="0" gridrow="7" pageId="5" pageNumber="42">Metasomal segment I</th>
</tr>
<tr box="[149,1077,513,535]" gridrow="8" pageId="5" pageNumber="42">
<th box="[149,327,513,535]" gridcol="0" gridrow="8" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,513,535]" gridcol="1" gridrow="8" pageId="5" pageNumber="42">4.8</td>
<td box="[775,1077,513,535]" gridcol="2" gridrow="8" pageId="5" pageNumber="42">5.1</td>
</tr>
<tr box="[149,1077,547,569]" gridrow="9" pageId="5" pageNumber="42">
<th box="[149,327,547,569]" gridcol="0" gridrow="9" pageId="5" pageNumber="42">- width</th>
<td box="[431,700,547,569]" gridcol="1" gridrow="9" pageId="5" pageNumber="42">4.5</td>
<td box="[775,1077,547,569]" gridcol="2" gridrow="9" pageId="5" pageNumber="42">5.7</td>
</tr>
<tr box="[149,1077,580,602]" gridrow="10" pageId="5" pageNumber="42">
<th box="[149,1077,580,602]" colspan="3" colspanRight="2" gridcol="0" gridrow="10" pageId="5" pageNumber="42">Metasomal segment II</th>
</tr>
<tr box="[149,1077,614,636]" gridrow="11" pageId="5" pageNumber="42">
<th box="[149,327,614,636]" gridcol="0" gridrow="11" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,614,636]" gridcol="1" gridrow="11" pageId="5" pageNumber="42">5.6</td>
<td box="[775,1077,614,636]" gridcol="2" gridrow="11" pageId="5" pageNumber="42">5.6</td>
</tr>
<tr box="[149,1077,647,669]" gridrow="12" pageId="5" pageNumber="42">
<th box="[149,327,647,669]" gridcol="0" gridrow="12" pageId="5" pageNumber="42">- width</th>
<td box="[431,700,647,669]" gridcol="1" gridrow="12" pageId="5" pageNumber="42">4.3</td>
<td box="[775,1077,647,669]" gridcol="2" gridrow="12" pageId="5" pageNumber="42">5.5</td>
</tr>
<tr box="[149,1077,681,703]" gridrow="13" pageId="5" pageNumber="42">
<th box="[149,1077,681,703]" colspan="3" colspanRight="2" gridcol="0" gridrow="13" pageId="5" pageNumber="42">Metasomal segment III</th>
</tr>
<tr box="[149,1077,714,737]" gridrow="14" pageId="5" pageNumber="42">
<th box="[149,327,714,737]" gridcol="0" gridrow="14" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,714,737]" gridcol="1" gridrow="14" pageId="5" pageNumber="42">5.7</td>
<td box="[775,1077,714,737]" gridcol="2" gridrow="14" pageId="5" pageNumber="42">5.8</td>
</tr>
<tr box="[149,1077,748,770]" gridrow="15" pageId="5" pageNumber="42">
<th box="[149,327,748,770]" gridcol="0" gridrow="15" pageId="5" pageNumber="42">- width</th>
<td box="[431,700,748,770]" gridcol="1" gridrow="15" pageId="5" pageNumber="42">4.3</td>
<td box="[775,1077,748,770]" gridcol="2" gridrow="15" pageId="5" pageNumber="42">5.3</td>
</tr>
<tr box="[149,1077,782,804]" gridrow="16" pageId="5" pageNumber="42">
<th box="[149,1077,782,804]" colspan="3" colspanRight="2" gridcol="0" gridrow="16" pageId="5" pageNumber="42">Metasomal segment IV</th>
</tr>
<tr box="[149,1077,815,837]" gridrow="17" pageId="5" pageNumber="42">
<th box="[149,327,815,837]" gridcol="0" gridrow="17" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,815,837]" gridcol="1" gridrow="17" pageId="5" pageNumber="42">6.6</td>
<td box="[775,1077,815,837]" gridcol="2" gridrow="17" pageId="5" pageNumber="42">6.9</td>
</tr>
<tr box="[149,1077,849,871]" gridrow="18" pageId="5" pageNumber="42">
<th box="[149,327,849,871]" gridcol="0" gridrow="18" pageId="5" pageNumber="42">- width</th>
<td box="[431,700,849,871]" gridcol="1" gridrow="18" pageId="5" pageNumber="42">4.1</td>
<td box="[775,1077,849,871]" gridcol="2" gridrow="18" pageId="5" pageNumber="42">5.2</td>
</tr>
<tr box="[149,1077,882,904]" gridrow="19" pageId="5" pageNumber="42">
<th box="[149,1077,882,904]" colspan="3" colspanRight="2" gridcol="0" gridrow="19" pageId="5" pageNumber="42">Metasomal segment V</th>
</tr>
<tr box="[149,1077,916,938]" gridrow="20" pageId="5" pageNumber="42">
<th box="[149,327,916,938]" gridcol="0" gridrow="20" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,916,938]" gridcol="1" gridrow="20" pageId="5" pageNumber="42">7.8</td>
<td box="[775,1077,916,938]" gridcol="2" gridrow="20" pageId="5" pageNumber="42">8.0</td>
</tr>
<tr box="[149,1077,950,972]" gridrow="21" pageId="5" pageNumber="42">
<th box="[149,327,950,972]" gridcol="0" gridrow="21" pageId="5" pageNumber="42">- width</th>
<td box="[431,700,950,972]" gridcol="1" gridrow="21" pageId="5" pageNumber="42">3.7</td>
<td box="[775,1077,950,972]" gridcol="2" gridrow="21" pageId="5" pageNumber="42">4.6</td>
</tr>
<tr box="[149,1077,983,1005]" gridrow="22" pageId="5" pageNumber="42">
<th box="[149,327,983,1005]" gridcol="0" gridrow="22" pageId="5" pageNumber="42">- depth</th>
<td box="[431,700,983,1005]" gridcol="1" gridrow="22" pageId="5" pageNumber="42">3.1</td>
<td box="[775,1077,983,1005]" gridcol="2" gridrow="22" pageId="5" pageNumber="42">4.0</td>
</tr>
<tr box="[149,1077,1017,1039]" gridrow="23" pageId="5" pageNumber="42">
<th box="[149,1077,1017,1039]" colspan="3" colspanRight="2" gridcol="0" gridrow="23" pageId="5" pageNumber="42">Telson</th>
</tr>
<tr box="[149,1077,1050,1072]" gridrow="24" pageId="5" pageNumber="42">
<th box="[149,327,1050,1072]" gridcol="0" gridrow="24" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,1050,1072]" gridcol="1" gridrow="24" pageId="5" pageNumber="42">6.6</td>
<td box="[775,1077,1050,1072]" gridcol="2" gridrow="24" pageId="5" pageNumber="42">7.2</td>
</tr>
<tr box="[149,1077,1084,1106]" gridrow="25" pageId="5" pageNumber="42">
<th box="[149,327,1084,1106]" gridcol="0" gridrow="25" pageId="5" pageNumber="42">- width</th>
<td box="[431,700,1084,1106]" gridcol="1" gridrow="25" pageId="5" pageNumber="42">2.5</td>
<td box="[775,1077,1084,1106]" gridcol="2" gridrow="25" pageId="5" pageNumber="42">4.9</td>
</tr>
<tr box="[149,1077,1117,1139]" gridrow="26" pageId="5" pageNumber="42">
<th box="[149,327,1117,1139]" gridcol="0" gridrow="26" pageId="5" pageNumber="42">- depth</th>
<td box="[431,700,1117,1139]" gridcol="1" gridrow="26" pageId="5" pageNumber="42">2.3</td>
<td box="[775,1077,1117,1139]" gridcol="2" gridrow="26" pageId="5" pageNumber="42">4.3</td>
</tr>
<tr box="[149,1077,1151,1173]" gridrow="27" pageId="5" pageNumber="42">
<th box="[149,1077,1151,1173]" colspan="3" colspanRight="2" gridcol="0" gridrow="27" pageId="5" pageNumber="42">Pedipalp</th>
</tr>
<tr box="[149,1077,1184,1207]" gridrow="28" pageId="5" pageNumber="42">
<th box="[149,327,1184,1207]" gridcol="0" gridrow="28" pageId="5" pageNumber="42">- Femur length</th>
<td box="[431,700,1184,1207]" gridcol="1" gridrow="28" pageId="5" pageNumber="42">5.7</td>
<td box="[775,1077,1184,1207]" gridcol="2" gridrow="28" pageId="5" pageNumber="42">5.4</td>
</tr>
<tr box="[149,1077,1218,1240]" gridrow="29" pageId="5" pageNumber="42">
<th box="[149,327,1218,1240]" gridcol="0" gridrow="29" pageId="5" pageNumber="42">- Femur width</th>
<td box="[431,700,1218,1240]" gridcol="1" gridrow="29" pageId="5" pageNumber="42">1.7</td>
<td box="[775,1077,1218,1240]" gridcol="2" gridrow="29" pageId="5" pageNumber="42">2.3</td>
</tr>
<tr box="[149,1077,1252,1274]" gridrow="30" pageId="5" pageNumber="42">
<th box="[149,327,1252,1274]" gridcol="0" gridrow="30" pageId="5" pageNumber="42">- Patella length</th>
<td box="[431,700,1252,1274]" gridcol="1" gridrow="30" pageId="5" pageNumber="42">6.8</td>
<td box="[775,1077,1252,1274]" gridcol="2" gridrow="30" pageId="5" pageNumber="42">6.2</td>
</tr>
<tr box="[149,1077,1285,1307]" gridrow="31" pageId="5" pageNumber="42">
<th box="[149,327,1285,1307]" gridcol="0" gridrow="31" pageId="5" pageNumber="42">- Patella width</th>
<td box="[431,700,1285,1307]" gridcol="1" gridrow="31" pageId="5" pageNumber="42">2.5</td>
<td box="[775,1077,1285,1307]" gridcol="2" gridrow="31" pageId="5" pageNumber="42">3.1</td>
</tr>
<tr box="[149,1077,1319,1341]" gridrow="32" pageId="5" pageNumber="42">
<th box="[149,327,1319,1341]" gridcol="0" gridrow="32" pageId="5" pageNumber="42">- Chela length</th>
<td box="[431,700,1319,1341]" gridcol="1" gridrow="32" pageId="5" pageNumber="42">11.8</td>
<td box="[775,1077,1319,1341]" gridcol="2" gridrow="32" pageId="5" pageNumber="42">11.3</td>
</tr>
<tr box="[149,1077,1352,1374]" gridrow="33" pageId="5" pageNumber="42">
<th box="[149,327,1352,1374]" gridcol="0" gridrow="33" pageId="5" pageNumber="42">- Chela width</th>
<td box="[431,700,1352,1374]" gridcol="1" gridrow="33" pageId="5" pageNumber="42">3.1</td>
<td box="[775,1077,1352,1374]" gridcol="2" gridrow="33" pageId="5" pageNumber="42">4.5</td>
</tr>
<tr box="[149,1077,1386,1408]" gridrow="34" pageId="5" pageNumber="42">
<th box="[149,327,1386,1408]" gridcol="0" gridrow="34" pageId="5" pageNumber="42">- Chela depth</th>
<td box="[431,700,1386,1408]" gridcol="1" gridrow="34" pageId="5" pageNumber="42">3.4</td>
<td box="[775,1077,1386,1408]" gridcol="2" gridrow="34" pageId="5" pageNumber="42">5.2</td>
</tr>
<tr box="[149,1077,1420,1442]" gridrow="35" pageId="5" pageNumber="42">
<th box="[149,1077,1420,1442]" colspan="3" colspanRight="2" gridcol="0" gridrow="35" pageId="5" pageNumber="42">Movable finger</th>
</tr>
<tr box="[149,1077,1453,1475]" gridrow="36" pageId="5" pageNumber="42">
<th box="[149,327,1453,1475]" gridcol="0" gridrow="36" pageId="5" pageNumber="42">- length</th>
<td box="[431,700,1453,1475]" gridcol="1" gridrow="36" pageId="5" pageNumber="42">8.0</td>
<td box="[775,1077,1453,1475]" gridcol="2" gridrow="36" pageId="5" pageNumber="42">6.8</td>
</tr>
</table>
</paragraph>
<paragraph blockId="5.[142,467,1496,1518]" box="[142,467,1496,1518]" pageId="5" pageNumber="42">
<tableNote box="[142,467,1496,1518]" pageId="5" pageNumber="42" targetBox="[149,948,916,1475]" targetPageId="5">* value including the telson length.</tableNote>
</paragraph>
<subSection pageId="5" pageNumber="42" type="multiple">
<paragraph blockId="5.[140,1108,1575,1672]" pageId="5" pageNumber="42">composed of 10 ventral teeth and four rather uniform lateral lobes. Dorsal furrows of all segments moderately developed, smooth; intercarinal spaces weakly to moderately granular. Telson weakly granular to smooth; vesicle with a pear-like shape, and a long</paragraph>
</subSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770929" ID-Zenodo-Dep="3770929" httpUri="https://zenodo.org/record/3770929/files/figure.png" pageId="6" pageNumber="43" startId="6.[140,222,1558,1580]" targetBox="[164,1092,178,1502]" targetPageId="6">
<paragraph blockId="6.[140,1108,1558,1676]" pageId="6" pageNumber="43">
<emphasis bold="true" box="[140,285,1558,1580]" pageId="6" pageNumber="43">Figures 9-1 7.</emphasis>
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[293,484,1558,1579]" class="Arachnida" family="Buthidae" genus="Saharobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="elegans" status="sp. n.">
<emphasis box="[293,484,1558,1579]" italics="true" pageId="6" pageNumber="43">Saharobuthus elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[492,546,1559,1580]" pageId="6" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
, male holotype.
<emphasis bold="true" box="[707,727,1558,1580]" pageId="6" pageNumber="43">9.</emphasis>
Chelicerae, dorsal aspect.
<emphasis bold="true" box="[980,1013,1558,1580]" pageId="6" pageNumber="43">1 0.</emphasis>
Sternum, genital operculum and pectines.
<emphasis bold="true" box="[453,486,1590,1612]" pageId="6" pageNumber="43">11.</emphasis>
Chela, dorso-external aspect.
<emphasis bold="true" box="[775,843,1590,1612]" pageId="6" pageNumber="43">1 2-1 3.</emphasis>
Patella, dorsal and external aspects.
<emphasis bold="true" box="[215,248,1622,1644]" pageId="6" pageNumber="43">1 4.</emphasis>
Femur, dorsal aspect.
<emphasis bold="true" box="[451,484,1622,1644]" pageId="6" pageNumber="43">1 5.</emphasis>
Cutting edge of movable finger with rows of granules.
<emphasis bold="true" box="[986,1054,1622,1644]" pageId="6" pageNumber="43">1 6-1 7.</emphasis>
Idem for
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1882" box="[172,434,1654,1676]" class="Arachnida" family="Buthidae" genus="Compsobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis box="[172,434,1654,1676]" italics="true" pageId="6" pageNumber="43">Compsobuthus acutecarinatus</emphasis>
</taxonomicName>
, female holotype (scales = 3 mm).
</paragraph>
</caption>
<subSection pageId="7" pageNumber="44" type="multiple">
<paragraph blockId="7.[140,1108,167,616]" pageId="7" pageNumber="44">
aculeus longer than vesicle; subaculear tooth absent. Chelicerae with two reduced but not fused denticles at the base of the movable finger (
<bibRefCitation author="Vachon" box="[755,922,202,229]" firstAuthor="Vachon" pageId="7" pageNumber="44" pagination="161 - 166" refId="ref7658" refString="Vachon M (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris 2 e ser. 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
). Pedipalps: Trichobothrial pattern orthobothriotaxic,
<typeStatus box="[580,628,238,264]" pageId="7" pageNumber="44">type</typeStatus>
A (
<bibRefCitation author="Vachon" box="[669,833,237,263]" firstAuthor="Vachon" pageId="7" pageNumber="44" pagination="857 - 958" refId="ref7700" refString="Vachon M (1974) Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 3 e ser. n ° 140, Zool. 104: 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
); dorsal trichobothria of femur in β (beta) configuration (
<bibRefCitation author="Vachon" box="[500,664,272,299]" firstAuthor="Vachon" pageId="7" pageNumber="44" pagination="1597 - 1599" refId="ref7764" refString="Vachon M (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus de l'Academie des Sciences, Paris, ser. D 281: 1597 - 1599." type="journal article" year="1975">Vachon, 1975</bibRefCitation>
). Femur pentacarinate; carinae strongly
</paragraph>
<paragraph blockId="7.[140,1108,167,616]" pageId="7" pageNumber="44">crenulate. Patella with eight carinae, moderate; dorsointernal carinae with 2-3 spinoid granules. Chela short with moderately elongated fingers; all carinae vestigial. Dentate margins on movable and fixed fingers composed of 11-12 oblique rows of granules; outer and inner accessory granules present on both fingers; four granules, two small and two large, located proximally to the terminal granule on the movable finger; basal region of movable finger with a strong lobe. Legs: Ventral aspect of tarsi with two series of spine-like setae. Tibial spurs present on legs III and IV, strongly marked. Pedal spurs present on all legs, moderate to strongly marked.</paragraph>
<subSubSection box="[185,891,589,616]" pageId="7" pageNumber="44" type="distribution">
<paragraph blockId="7.[140,1108,167,616]" box="[185,891,589,616]" pageId="7" pageNumber="44">
<emphasis bold="true" box="[185,483,589,615]" pageId="7" pageNumber="44">Geographic distribution.</emphasis>
Only known from the
<typeStatus box="[748,796,590,616]" pageId="7" pageNumber="44">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3854816" ID-GBIF-Taxon="163976987" ID-Zenodo-Dep="3854816" LSID="urn:lsid:plazi:treatment:03CD87CC650BC13BFF16FEA1FC85B912" httpUri="http://treatment.plazi.org/id/03CD87CC650BC13BFF16FEA1FC85B912" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
<subSubSection box="[140,462,694,721]" pageId="7" pageNumber="44" type="nomenclature">
<paragraph blockId="7.[140,968,694,757]" box="[140,462,694,721]" pageId="7" pageNumber="44">
<heading bold="true" box="[140,462,694,721]" fontSize="11" level="2" pageId="7" pageNumber="44" reason="0">
<emphasis bold="true" box="[140,462,694,721]" pageId="7" pageNumber="44">
Genus
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[223,374,694,721]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[223,374,694,721]" italics="true" pageId="7" pageNumber="44">Pantobuthus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[381,462,695,721]" pageId="7" pageNumber="44" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,968,730,757]" pageId="7" pageNumber="44" type="description">
<paragraph blockId="7.[140,968,694,757]" box="[140,968,730,757]" pageId="7" pageNumber="44">
<uri box="[140,968,730,757]" pageId="7" pageNumber="44">
urn:lsid:zoobank.org:act:
<uuid box="[421,968,730,757]" pageId="7" pageNumber="44">3C2F50F7-DD92-4854-8584-CBE4FF62B084</uuid>
</uri>
</paragraph>
</subSubSection>
<subSection pageId="7" pageNumber="44" type="multiple">
<subSubSection pageId="7" pageNumber="44" type="diagnosis">
<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">
<emphasis bold="true" box="[140,258,800,826]" pageId="7" pageNumber="44">Diagnosis</emphasis>
. Large scorpions, 66.3 mm in total length (excluding telson). Coloration generally pale yellow with regions of very light grey. Carapace carinae moderately marked; lyra configuration rather incomplete; median ocular tubercle at the centre of carapace; three pairs of lateral eyes situated in a posterior position in relation to the ante-
</paragraph>
<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">
rior edge; third pair totally vestigial. Sternum triangular, wider than long. Pedipalps very short and bulky; chela strongly globular; dentate margins on fixed and movable fingers of pedipalp chela composed of 9-10 oblique rows of granules. Outer and inner accessory granules present on both fingers, strongly marked. Pectinal tooth count 24-23; fulcra present; basal middle lamellae not dilated. Chelicerae with one basal denticle on the fixed finger; basal denticles of the movable finger strongly reduced. Metasomal segments II to IV with strong spinoid granules on ventral carinae; telson with a strongly globular vesicle; aculeus much shorter than vesicle and strongly curved; subaculear tooth absent. Legs: tarsi with two series of short spine-like setae; internal series with 5-7 setae, external reduced to 1-3 setae or totally absent; tibial spur strong on legs III and IV; pedal spurs moderate to strong on legs I to IV. Trichobothrial pattern A-β (beta), orthobothriotaxy. Trichobothrium
<emphasis bold="true" box="[327,358,1328,1354]" pageId="7" pageNumber="44">db</emphasis>
of chela totally displaced over the internal face of fixed finger.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="44" type="etymology">
<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">Derivatio nominis: generic name refers to the multiple characteristics of buthid genera present in the new genus.</paragraph>
</subSubSection>
<subSubSection box="[185,676,1433,1460]" pageId="7" pageNumber="44" type="type_taxon">
<paragraph blockId="7.[140,1108,800,1672]" box="[185,676,1433,1460]" pageId="7" pageNumber="44">
<emphasis bold="true" box="[185,332,1433,1460]" pageId="7" pageNumber="44">
<typeStatus box="[185,243,1434,1460]" pageId="7" pageNumber="44">Type</typeStatus>
species
</emphasis>
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[340,607,1434,1460]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis box="[340,607,1434,1460]" italics="true" pageId="7" pageNumber="44">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[615,676,1434,1460]" pageId="7" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44" type="discussion">
<paragraph blockId="7.[140,1108,800,1672]" pageId="7" pageNumber="44">
<emphasis bold="true" box="[185,507,1469,1495]" pageId="7" pageNumber="44">Affinities of the new genus.</emphasis>
The new genus
<taxonomicName authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[689,822,1469,1495]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[689,822,1469,1495]" italics="true" pageId="7" pageNumber="44">Pantobuthus</emphasis>
</taxonomicName>
shows affinities with several genera, such as
<taxonomicName authorityName="Vachon" authorityYear="1950" box="[376,529,1504,1530]" class="Arachnida" family="Buthidae" genus="Odontobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[376,529,1504,1530]" italics="true" pageId="7" pageNumber="44">Odontobuthus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Leach" authorityYear="1815" box="[548,623,1504,1530]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[548,623,1504,1530]" italics="true" pageId="7" pageNumber="44">Buthus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Vachon" authorityYear="1950" box="[641,767,1504,1530]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[641,767,1504,1530]" italics="true" pageId="7" pageNumber="44">Mesobuthus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Birula" authorityYear="1908" box="[785,908,1505,1530]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[785,908,1505,1530]" italics="true" pageId="7" pageNumber="44">Hottentotta</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" box="[973,1100,1504,1530]" class="Arachnida" family="Buthidae" genus="Vachoniolus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[973,1100,1504,1530]" italics="true" pageId="7" pageNumber="44">Vachoniolus</emphasis>
</taxonomicName>
, which are also elements of the North African and Palaearctic faunas. The moderately marked carapace carinae which form only an incomplete lyra configuration associates the new genus to
<taxonomicName authorityName="Birula" authorityYear="1908" box="[345,468,1611,1636]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[345,468,1611,1636]" italics="true" pageId="7" pageNumber="44">Hottentotta</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Vachon" authorityYear="1950" box="[528,654,1610,1636]" class="Arachnida" family="Buthidae" genus="Mesobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[528,654,1610,1636]" italics="true" pageId="7" pageNumber="44">Mesobuthus</emphasis>
</taxonomicName>
, but metasomal segments II to IV with strong spinoid granules on the ventral carinae, are more similar to those of the genus
</paragraph>
<paragraph blockId="8.[140,1108,167,510]" pageId="8" pageNumber="45">
<taxonomicName authorityName="Vachon" authorityYear="1950" box="[140,293,167,193]" class="Arachnida" family="Buthidae" genus="Odontobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis box="[140,293,167,193]" italics="true" pageId="8" pageNumber="45">Odontobuthus</emphasis>
</taxonomicName>
. Th e very central position of the median ocular tubercle on the carapace and the shape of the telson with a strongly globular vesicle and a very short aculeus associates
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[254,387,237,263]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[254,387,237,263]" italics="true" pageId="8" pageNumber="45">Pantobuthus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[396,474,238,264]" pageId="8" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
with
<taxonomicName authorityName="Leach" authorityYear="1815" box="[544,619,237,263]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis box="[544,619,237,263]" italics="true" pageId="8" pageNumber="45">Buthus</emphasis>
</taxonomicName>
. In this last genus, however, both carapace and mesosomal carinae are strongly developed and the lyra configuration is totally
</paragraph>
<paragraph blockId="8.[140,1108,167,510]" pageId="8" pageNumber="45">
complete. Th e very short and bulky pedipalps again associate
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[831,964,307,333]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[831,964,307,333]" italics="true" pageId="8" pageNumber="45">Pantobuthus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[972,1049,308,334]" pageId="8" pageNumber="45" rank="genus">gen. n.</taxonomicNameLabel>
with
<taxonomicName authorityName="Leach" authorityYear="1815" box="[140,215,343,369]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis box="[140,215,343,369]" italics="true" pageId="8" pageNumber="45">Buthus</emphasis>
</taxonomicName>
. However, the very strongly globular chela recall those of the male
<taxonomicName authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" box="[973,1100,343,369]" class="Arachnida" family="Buthidae" genus="Vachoniolus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis box="[973,1100,343,369]" italics="true" pageId="8" pageNumber="45">Vachoniolus</emphasis>
</taxonomicName>
. The male of
<taxonomicName authority="Lourenco &amp; Duhem, 2009" authorityName="Lourenco &amp; Duhem" authorityYear="2009" box="[277,539,378,404]" class="Arachnida" family="Buthidae" genus="Pantobuthus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="complicatus" status="sp. n.">
<emphasis box="[277,539,378,404]" italics="true" pageId="8" pageNumber="45">Pantobuthus complicatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[546,605,378,404]" pageId="8" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
remains unknown. Finally, the total displacement of chela trichobothrium
<emphasis bold="true" box="[490,521,413,439]" pageId="8" pageNumber="45">db</emphasis>
to the internal surface of fixed finger, the retreated position of lateral eyes and the tarsi with two series of short spine-like setae, but with external series reduced or totally absent appear to be particular characteristics of this genus.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</subSection>
</subSubSection>
</treatment>
</document>