184 lines
19 KiB
XML
184 lines
19 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4765172" ID-GBIF-Dataset="9417b10b-5d16-4f7a-9d9d-86d80f05c38b" ID-GBIF-Taxon="182137829" ID-ISSN="1854-0392" ID-Zenodo-Dep="4765172" ID-ZooBank="79B7368A-29DC-44C2-B9AD-5AFA006FBC7F" checkinTime="1620963140635" checkinUser="felipe" docAuthor="Stark, Bill P." docDate="2014" docId="03CD878C823DFFB70F955FAE0DACFAA1" docLanguage="en" docName="Illiesia.10.2.6-16.pdf" docOrigin="Illiesia 10 (2)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Anacroneuria guanacaste Stark 2014, sp. n." docType="treatment" docVersion="5" lastPageId="4" lastPageNumber="10" masterDocId="FFF4FFF4823FFFB30C2C592C0C25FFA1" masterDocTitle="Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species" masterLastPageNumber="16" masterPageNumber="6" pageId="2" pageNumber="8" updateTime="1632022174258" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Stark, Bill P.</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Illiesia</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2014</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>10</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="issue">
|
|
<mods:number>2</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>6</mods:start>
|
|
<mods:end>16</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760479</mods:identifier>
|
|
<mods:identifier type="GBIF-Dataset">9417b10b-5d16-4f7a-9d9d-86d80f05c38b</mods:identifier>
|
|
<mods:identifier type="ISSN">1854-0392</mods:identifier>
|
|
<mods:identifier type="Zenodo-Dep">4760479</mods:identifier>
|
|
<mods:identifier type="ZooBank">79B7368A-29DC-44C2-B9AD-5AFA006FBC7F</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4765172" ID-GBIF-Taxon="182137829" ID-Zenodo-Dep="4765172" LSID="urn:lsid:plazi:treatment:03CD878C823DFFB70F955FAE0DACFAA1" httpUri="http://treatment.plazi.org/id/03CD878C823DFFB70F955FAE0DACFAA1" lastPageId="4" lastPageNumber="10" pageId="2" pageNumber="8">
|
|
<subSubSection box="[953,1325,1666,1691]" pageId="2" pageNumber="8" type="nomenclature">
|
|
<paragraph blockId="2.[953,1325,1666,1691]" box="[953,1325,1666,1691]" pageId="2" pageNumber="8">
|
|
<heading bold="true" box="[953,1325,1666,1691]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="8" reason="2">
|
|
<emphasis bold="true" box="[953,1325,1666,1691]" pageId="2" pageNumber="8">
|
|
<taxonomicName authority="Stark, 2014" authorityName="Stark" authorityYear="2014" box="[953,1253,1666,1690]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="8" phylum="Arthropoda" rank="species" species="guanacaste" status="sp. n.">
|
|
<emphasis bold="true" box="[953,1253,1666,1690]" italics="true" pageId="2" pageNumber="8">Anacroneuria guanacaste</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[1260,1325,1667,1691]" pageId="2" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection box="[1081,1197,1702,1727]" pageId="2" pageNumber="8" type="description">
|
|
<paragraph blockId="2.[1081,1197,1702,1727]" box="[1081,1197,1702,1727]" pageId="2" pageNumber="8">
|
|
(
|
|
<figureCitation box="[1089,1189,1702,1727]" captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="2" pageNumber="8">Figs. 1-5</figureCitation>
|
|
)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="2" pageNumber="8" type="materials_examined">
|
|
<paragraph blockId="2.[835,1443,1773,1904]" pageId="2" pageNumber="8">
|
|
<emphasis bold="true" box="[835,1089,1773,1797]" pageId="2" pageNumber="8">Material examined.</emphasis>
|
|
<materialsCitation ID-GBIF-Occurrence="3111908306" collectingDate="1989-03-17" collectingMethod="light trap" collectionCode="USNM" country="Costa Rica" location="Rio Tempisquito" municipality="Estacion Biologica Maritza" pageId="2" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste Province" typeStatus="holotype">
|
|
<typeStatus box="[1111,1225,1773,1797]" pageId="2" pageNumber="8">Holotype</typeStatus>
|
|
<specimenCount box="[1245,1265,1773,1797]" pageId="2" pageNumber="8" type="male">♂</specimenCount>
|
|
,
|
|
<emphasis bold="true" box="[1292,1436,1773,1797]" pageId="2" pageNumber="8">
|
|
<collectingCountry box="[1292,1436,1773,1797]" name="Costa Rica" pageId="2" pageNumber="8">Costa Rica</collectingCountry>
|
|
</emphasis>
|
|
,
|
|
<collectingRegion box="[835,1093,1809,1833]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="8">Guanacaste Province</collectingRegion>
|
|
,
|
|
<collectingMunicipality box="[1107,1438,1809,1833]" pageId="2" pageNumber="8">Estación Biológica Maritza</collectingMunicipality>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:03CD878C823DFFB70F955FAE0DACFAA1:8EBB6041823DFFB10F6F5E19083AF8EC" box="[835,1055,1845,1869]" country="Costa Rica" municipality="Estacion Biologica Maritza" name="Rio Tempisquito" pageId="2" pageNumber="8" stateProvince="Guanacaste Province">Rio Tempisquito</location>
|
|
,
|
|
<date box="[1081,1288,1845,1869]" pageId="2" pageNumber="8" value="1989-03-17">
|
|
<collectingDate box="[1081,1288,1845,1869]" pageId="2" pageNumber="8" value="1989-03-17">17 March 1989</collectingDate>
|
|
</date>
|
|
,
|
|
<collectingMethod box="[1314,1441,1845,1869]" pageId="2" pageNumber="8">light trap</collectingMethod>
|
|
(
|
|
<collectionCode box="[843,932,1880,1904]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="8" type="Museum">USNM</collectionCode>
|
|
).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.4765160" ID-Zenodo-Dep="4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="3" pageNumber="9" startId="3.[189,241,1610,1634]" targetBox="[328,1308,225,1589]" targetPageId="3">
|
|
<paragraph blockId="3.[189,1442,1610,1670]" pageId="3" pageNumber="9">
|
|
Figs. 1-5.
|
|
<taxonomicName authority="Stark, 2014" authorityName="Stark" authorityYear="2014" box="[297,569,1611,1634]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="9" phylum="Arthropoda" rank="species" species="guanacaste" status="sp.n.">
|
|
<emphasis box="[297,569,1611,1634]" italics="true" pageId="3" pageNumber="9">Anacroneuria guanacaste</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[574,630,1611,1634]" pageId="3" pageNumber="9" rank="species">sp.n.</taxonomicNameLabel>
|
|
male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection lastPageId="4" lastPageNumber="10" pageId="3" pageNumber="9" type="description">
|
|
<paragraph blockId="3.[189,797,1752,1919]" lastBlockId="3.[835,1443,1752,1918]" pageId="3" pageNumber="9">
|
|
<emphasis bold="true" box="[189,365,1752,1776]" pageId="3" pageNumber="9">Adult habitus.</emphasis>
|
|
General color brown, patterned with pale brown. Head dark brown pigment extending from behind ocelli in a broad patch covering central frons; lappets and antennae dark brown, pale callosities lateral to ocelli distinct (
|
|
<figureCitation box="[612,680,1895,1919]" captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="3" pageNumber="9">Fig. 1</figureCitation>
|
|
). Median pronotal band yellow, most of disc brown. Wing membrane pale amber, veins brown. Hind femora with pale proximal band covering basal 2/3, apical third dark brown. Tibiae banded, pale in median field, dark brown at both ends.
|
|
</paragraph>
|
|
<paragraph blockId="4.[189,798,225,1281]" pageId="4" pageNumber="10">
|
|
<emphasis bold="true" box="[189,258,225,249]" pageId="4" pageNumber="10">Male.</emphasis>
|
|
Forewing length
|
|
<quantity box="[483,573,225,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="10" unit="mm" value="12.0">12 mm</quantity>
|
|
. Hammer thimble shaped, height slightly greater than apical diameter (
|
|
<figureCitation box="[197,269,296,320]" captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="4" pageNumber="10">Fig. 2</figureCitation>
|
|
). Aedeagal apex simple, notched on apical margin; ventral membranous lobes conspicuous (
|
|
<figureCitation captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="4" pageNumber="10">Fig. 3</figureCitation>
|
|
). In lateral aspect, apex is somewhat foot-shaped, but with a slightly bulging process at ankle (
|
|
<figureCitation box="[716,783,403,427]" captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="4" pageNumber="10">Fig. 4</figureCitation>
|
|
). Dorsal keel consists of an obscure series of low chevron-like lines (
|
|
<figureCitation box="[420,490,474,498]" captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="4" pageNumber="10">Fig. 5</figureCitation>
|
|
), but median line absent. Hooks slender.
|
|
</paragraph>
|
|
<paragraph blockId="4.[189,798,225,1281]" box="[189,416,545,569]" pageId="4" pageNumber="10">
|
|
<emphasis bold="true" box="[189,283,545,569]" pageId="4" pageNumber="10">Female.</emphasis>
|
|
Unknown.
|
|
</paragraph>
|
|
<paragraph blockId="4.[189,798,225,1281]" box="[189,396,581,605]" pageId="4" pageNumber="10">
|
|
<emphasis bold="true" box="[189,264,581,605]" pageId="4" pageNumber="10">Larva.</emphasis>
|
|
Unknown.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="4" pageNumber="10" type="etymology">
|
|
<paragraph blockId="4.[189,798,225,1281]" pageId="4" pageNumber="10">
|
|
<emphasis bold="true" box="[189,327,617,641]" pageId="4" pageNumber="10">Etymology.</emphasis>
|
|
The species name, based on
|
|
<collectingRegion country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="10">Guanacaste Province</collectingRegion>
|
|
, is used as a noun in apposition.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="4" pageNumber="10" type="diagnosis">
|
|
<paragraph blockId="4.[189,798,225,1281]" pageId="4" pageNumber="10">
|
|
<emphasis bold="true" box="[189,319,688,712]" pageId="4" pageNumber="10">Diagnosis.</emphasis>
|
|
In the
|
|
<bibRefCitation author="Stark, B. P." box="[395,533,688,712]" pageId="4" pageNumber="10" pagination="551 - 603" refId="ref5572" refString="Stark, B. P. 1998. The Anacroneuria of Costa Rica and Panama (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 111: 551 - 603." type="journal article" year="1998">Stark (1998)</bibRefCitation>
|
|
key this species runs to couplet 18 where an impasse occurs due to the intermediate forewing length. If the impasse is resolved by choosing the greater length (
|
|
<quantity box="[665,748,795,819]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="10" unit="mm" value="13.0">13 mm</quantity>
|
|
), no resolution is gained, although a determination of
|
|
<taxonomicName authorityName="Stark" authorityYear="1998" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="10" phylum="Arthropoda" rank="species" species="marca">
|
|
<emphasis italics="true" pageId="4" pageNumber="10">A. marca</emphasis>
|
|
</taxonomicName>
|
|
or
|
|
<taxonomicName authority="Stark, 1998" authorityName="Stark" authorityYear="1998" box="[297,554,866,890]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="10" phylum="Arthropoda" rank="species" species="ventana">
|
|
<emphasis box="[297,417,866,890]" italics="true" pageId="4" pageNumber="10">A. ventana</emphasis>
|
|
<bibRefCitation author="Stark, B. P." box="[424,554,866,890]" pageId="4" pageNumber="10" pagination="551 - 603" refId="ref5572" refString="Stark, B. P. 1998. The Anacroneuria of Costa Rica and Panama (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 111: 551 - 603." type="journal article" year="1998">Stark, 1998</bibRefCitation>
|
|
</taxonomicName>
|
|
can be forced; if the smaller length (
|
|
<quantity box="[372,454,901,925]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="10" unit="mm" value="10.0">10 mm</quantity>
|
|
) is chosen a determination of
|
|
<taxonomicName authority="Stark, 1998" authorityName="Stark" authorityYear="1998" box="[189,435,937,961]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="10" phylum="Arthropoda" rank="species" species="varilla">
|
|
<emphasis box="[189,295,937,961]" italics="true" pageId="4" pageNumber="10">A. varilla</emphasis>
|
|
<bibRefCitation author="Stark, B. P." box="[304,435,937,961]" pageId="4" pageNumber="10" pagination="551 - 603" refId="ref5572" refString="Stark, B. P. 1998. The Anacroneuria of Costa Rica and Panama (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 111: 551 - 603." type="journal article" year="1998">Stark, 1998</bibRefCitation>
|
|
</taxonomicName>
|
|
is made. In some respects,
|
|
<taxonomicName authorityName="Stark" authorityYear="2014" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="10" phylum="Arthropoda" rank="species" species="guanacaste">
|
|
<emphasis italics="true" pageId="4" pageNumber="10">A. guanacaste</emphasis>
|
|
</taxonomicName>
|
|
is intermediate between these species with a slender, thimble-shaped hammer like that of
|
|
<taxonomicName authorityName="Stark" authorityYear="1998" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="10" phylum="Arthropoda" rank="species" species="marca">
|
|
<emphasis italics="true" pageId="4" pageNumber="10">A. marca</emphasis>
|
|
</taxonomicName>
|
|
, and with an aedeagal apex more similar to that of
|
|
<taxonomicName authorityName="Stark" authorityYear="1998" box="[278,382,1079,1103]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="10" phylum="Arthropoda" rank="species" species="varilla">
|
|
<emphasis box="[278,382,1079,1103]" italics="true" pageId="4" pageNumber="10">A. varilla</emphasis>
|
|
</taxonomicName>
|
|
. The new species is distinguished from both by the apical notch in the aedeagal tip (
|
|
<figureCitation captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="4" pageNumber="10">Fig. 3</figureCitation>
|
|
), and by the chevron- like dorsal keel (
|
|
<figureCitation box="[664,731,1150,1174]" captionStart="Figs" captionStartId="3.[189,241,1610,1634]" captionTargetBox="[328,1308,225,1589]" captionTargetId="figure-153@3.[325,1308,215,1589]" captionText="Figs. 1-5. Anacroneuria guanacaste sp.n. male. 1. Head and pronotum. 2. Sternum 9 with hammer. 3. Aedeagus, ventral. 4. Aedeagus, lateral. 5. Aaedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765160" httpUri="https://zenodo.org/record/4765160/files/figure.png" pageId="4" pageNumber="10">Fig. 5</figureCitation>
|
|
). The species is probably a near relative of
|
|
<taxonomicName authority="Jewet, 1958" authorityName="Jewet" authorityYear="1958" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="10" phylum="Arthropoda" rank="species" species="flavominuta">
|
|
<emphasis box="[633,797,1185,1209]" italics="true" pageId="4" pageNumber="10">A. flavominuta</emphasis>
|
|
Jewet, 1958
|
|
</taxonomicName>
|
|
and other Mexican species (
|
|
<bibRefCitation author="Stark, B. P. & B. C. Kondratieff" pageId="4" pageNumber="10" pagination="1 - 64" refId="ref5653" refString="Stark, B. P. & B. C. Kondratieff. 2004. Anacroneuria from Mexico and upper Mesoamerica (Plecoptera: Perlidae). Monographs of the Western North American Naturalist, 2: 1 - 64." type="journal article" year="2004">Stark & Kondratieff 2004</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |