410 lines
46 KiB
XML
410 lines
46 KiB
XML
<document id="70DB93A104495BA92D656BBA624CC66A" ID-DOI="10.11646/zootaxa.3683.2.2" ID-GBIF-Dataset="edc5aac3-f1f9-4ff7-b583-616556bf1407" ID-ISSN="1175-5326" ID-Zenodo-Dep="223681" ID-ZooBank="B393CCFE-94EE-438F-958D-3712BA3D4C18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460644097812" checkinUser="plazi" docAuthor="Lobo, Fernando, Laspiur, Alejandro & Acosta, Juan Carlos" docDate="2013" docId="03C97438FFDA6A780CEBF954A7D1DE5A" docLanguage="en" docName="zt03683p132.pdf" docOrigin="Zootaxa 3683 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phymaturus williamsi Lobo, Laspiur & Acosta, 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="124" masterDocId="FFF00C40FFDF6A7F0C7CFF8CA53AD972" masterDocTitle="Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina" masterLastPageNumber="132" masterPageNumber="117" pageNumber="122" updateTime="1698336511002" updateUser="plazi">
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<mods:titleInfo id="D94BF019725659B2E2EE90CE95B02811">
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<mods:title id="21DD2D6E6A6E3677B49F8558CBF2491A">Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina</mods:title>
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<mods:roleTerm id="FD308EC62CC95C99AD21AE12E974E0BC">Author</mods:roleTerm>
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<mods:namePart id="A1E2F20A6159D3CCED0E65F4CEB82259">Lobo, Fernando</mods:namePart>
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<mods:roleTerm id="E7DC0ED012B49E5687180CC8CB5A3134">Author</mods:roleTerm>
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<mods:namePart id="85D88AC6450BA5BCB02307421610B5E9">Laspiur, Alejandro</mods:namePart>
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<mods:name id="7C6FA04359A4045B06D3AAA7C9AF1852" type="personal">
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<mods:roleTerm id="579525195F3DDEFFF5EB55E6A83A529D">Author</mods:roleTerm>
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<mods:namePart id="A11510097661CB6508B2CBD53B2BDC74">Acosta, Juan Carlos</mods:namePart>
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<mods:typeOfResource id="13B23F1689CBE630F040497DEE3A1B08">text</mods:typeOfResource>
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<mods:title id="E0E8E141382A3C4E998A89974A86C01F">Zootaxa</mods:title>
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<mods:part id="20BBA51A54AA9D8A1C97EBD7E4703586">
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<mods:date id="33FFB1150DEEB40F6EED48C24D0D460E">2013</mods:date>
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<mods:number id="42B4C9F4A3CDB0CC9B27AE01193CA907">3683</mods:number>
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<mods:classification id="A6060EE7E683BF746A51AE386483EF1D">journal article</mods:classification>
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<mods:identifier id="3CD6651CCBE6C313E698B472B1063A13" type="DOI">10.11646/zootaxa.3683.2.2</mods:identifier>
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<treatment id="03C97438FFDA6A780CEBF954A7D1DE5A" ID-DOI="http://doi.org/10.5281/zenodo.5696282" ID-GBIF-Taxon="119571812" ID-Zenodo-Dep="5696282" LSID="urn:lsid:plazi:treatment:03C97438FFDA6A780CEBF954A7D1DE5A" httpUri="http://treatment.plazi.org/id/03C97438FFDA6A780CEBF954A7D1DE5A" lastPageId="7" lastPageNumber="124" pageId="5" pageNumber="122">
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<subSubSection id="C37A96A5FFDA6A7A0CEBF954A73EDF80" box="[151,516,1752,1778]" pageId="5" pageNumber="122" type="nomenclature">
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<paragraph id="8BDFC52EFFDA6A7A0CEBF954A73EDF80" blockId="5.[151,516,1752,1778]" box="[151,516,1752,1778]" pageId="5" pageNumber="122">
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<heading id="D0977242FFDA6A7A0CEBF954A73EDF80" bold="true" box="[151,516,1752,1778]" fontSize="11" level="1" pageId="5" pageNumber="122" reason="1">
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<emphasis id="B914193CFFDA6A7A0CEBF954A73EDF80" bold="true" box="[151,516,1752,1778]" pageId="5" pageNumber="122">
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<taxonomicName id="4C60BEADFFDA6A7A0CEBF954A4A5DF80" box="[151,415,1752,1778]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="122" phylum="Chordata" rank="species" species="williamsi" status="sp. nov.">
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<emphasis id="B914193CFFDA6A7A0CEBF954A4A5DF80" bold="true" box="[151,415,1752,1778]" italics="true" pageId="5" pageNumber="122">Phymaturus williamsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A227A447FFDA6A7A0DDBF954A73EDF80" box="[423,516,1752,1778]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37A96A5FFDA6A7A0CEBF893A74DDEB5" pageId="5" pageNumber="122" type="materials_examined">
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<paragraph id="8BDFC52EFFDA6A7A0CEBF893A073DE29" blockId="5.[151,1436,1823,1992]" pageId="5" pageNumber="122">
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<typeStatus id="54DB7B8CFFDA6A7A0CEBF893A43EDE45" box="[151,260,1823,1847]" pageId="5" pageNumber="122" type="holotype">
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<emphasis id="B914193CFFDA6A7A0CEBF893A43EDE45" bold="true" box="[151,260,1823,1847]" pageId="5" pageNumber="122">Holotype</emphasis>
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</typeStatus>
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: MCN 2820. Male. Quebrada vallecito,
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<quantity id="4C9868CBFFDA6A7A0EAEF893A621DE45" box="[722,795,1823,1847]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="122" unit="km" value="40.0">40 km</quantity>
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W of Calingasta town, Calingasta department, San Juan province.
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<geoCoordinate id="EE54A3E9FFDA6A7A0D7BF8CFA4AFDE29" box="[263,405,1859,1883]" direction="south" orientation="latitude" pageId="5" pageNumber="122" precision="15" value="-31.189165">31º11’21’’ S</geoCoordinate>
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;
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<geoCoordinate id="EE54A3E9FFDA6A7A0DDDF8CFA775DE29" box="[417,591,1859,1883]" direction="west" orientation="longitude" pageId="5" pageNumber="122" precision="1" value="-69.70419">69º42’15.1’’ W</geoCoordinate>
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,
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<quantity id="4C9868CBFFDA6A7A0E26F8CFA794DE29" box="[602,686,1859,1883]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="122" unit="m" value="3000.0">3000 m</quantity>
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asl. A. Laspiur, R. Acosta & J. C. Acosta cols.
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<date id="FFDEE3EEFFDA6A7A08BFF8CFA07FDE29" box="[1219,1349,1859,1883]" pageId="5" pageNumber="122" value="2008-05-11">11/05/2008</date>
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.
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</paragraph>
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<paragraph id="8BDFC52EFFDA6A7A0CB5F8EBA74DDEB5" blockId="5.[151,1436,1823,1992]" pageId="5" pageNumber="122">
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<typeStatus id="54DB7B8CFFDA6A7A0CB5F8EBA47BDE0D" box="[201,321,1895,1919]" pageId="5" pageNumber="122" type="paratype">
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<emphasis id="B914193CFFDA6A7A0CB5F8EBA47BDE0D" bold="true" box="[201,321,1895,1919]" pageId="5" pageNumber="122">Paratypes</emphasis>
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</typeStatus>
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:.MCN
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<date id="FFDEE3EEFFDA6A7A0DE5F8EBA4C2DE0D" box="[409,504,1895,1919]" pageId="5" pageNumber="122" value="2808" valueMax="2810">2808-10</date>
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,
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<date id="FFDEE3EEFFDA6A7A0E79F8EBA75FDE0D" box="[517,613,1895,1919]" pageId="5" pageNumber="122" value="2812" valueMax="2814">2812-14</date>
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, 2815,
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<date id="FFDEE3EEFFDA6A7A0EC5F8EBA623DE0D" box="[697,793,1895,1919]" pageId="5" pageNumber="122" value="2816" valueMax="2817">2816-17</date>
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, 2821. Same data as
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<typeStatus id="54DB7B8CFFDA6A7A0875F8EBA155DE0D" box="[1033,1135,1895,1919]" pageId="5" pageNumber="122" type="holotype">holotype</typeStatus>
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. MCN 3259-65. Quebrada Vallecito,
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<quantity id="4C9868CBFFDA6A7A0D77F807A46EDED1" box="[267,340,1931,1955]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="122" unit="km" value="40.0">40 km</quantity>
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W of Calingasta town, Calingasta department, San Juan province.
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<geoCoordinate id="EE54A3E9FFDA6A7A0820F807A1D9DED1" box="[1116,1251,1931,1955]" direction="south" orientation="latitude" pageId="5" pageNumber="122" precision="15" value="-31.189165">31º11’21’’S</geoCoordinate>
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;
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||
<geoCoordinate id="EE54A3E9FFDA6A7A088EF807A0A2DED1" box="[1266,1432,1931,1955]" direction="west" orientation="longitude" pageId="5" pageNumber="122" precision="1" value="-69.70419">69º42’15.1’’W</geoCoordinate>
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,
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<quantity id="4C9868CBFFDA6A7A0CEBF823A5DEDEB5" box="[151,228,1967,1991]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="122" unit="m" value="3000.0">3000m</quantity>
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asl. A. Laspiur & J. C. Acosta cols.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF1F95A6FFD96A790CEBFB5DA1A1DD95" box="[151,1179,1233,1256]" httpUri="https://zenodo.org/record/223684/files/figure.png" pageId="6" pageNumber="123" targetBox="[151,1436,249,1211]" targetPageId="6">
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<paragraph id="8BDFC52EFFD96A790CEBFB5DA1A1DD95" blockId="6.[151,1179,1233,1256]" box="[151,1179,1233,1256]" pageId="6" pageNumber="123">
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||
<emphasis id="B914193CFFD96A790CEBFB5DA428DD95" bold="true" box="[151,274,1233,1256]" pageId="6" pageNumber="123">FIGURE 3.</emphasis>
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Distribution of
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<taxonomicName id="4C60BEADFFD96A790DC9FB5DA70ADD95" box="[437,560,1233,1255]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="genus">
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<emphasis id="B914193CFFD96A790DC9FB5DA70ADD95" box="[437,560,1233,1255]" italics="true" pageId="6" pageNumber="123">Phymaturus</emphasis>
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</taxonomicName>
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species in San Juan province and near localities (Argentina).
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</paragraph>
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||
</caption>
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<subSubSection id="C37A96A5FFD96A780CB5FA82A7CDD8DE" lastPageId="7" lastPageNumber="124" pageId="6" pageNumber="123" type="diagnosis">
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||
<paragraph id="8BDFC52EFFD96A780CB5FA82A7CDD8DE" blockId="6.[151,1436,1294,2003]" lastBlockId="7.[151,1437,151,1833]" lastPageId="7" lastPageNumber="124" pageId="6" pageNumber="123">
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<emphasis id="B914193CFFD96A790CB5FA82A479DC55" bold="true" box="[201,323,1294,1319]" pageId="6" pageNumber="123">Diagnosis.</emphasis>
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<taxonomicName id="4C60BEADFFD96A790D29FA83A76FDC55" box="[341,597,1295,1319]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
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<emphasis id="B914193CFFD96A790D29FA83A76FDC55" box="[341,597,1295,1319]" italics="true" pageId="6" pageNumber="123">Phymaturus williamsi</emphasis>
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</taxonomicName>
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belongs to the
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<taxonomicName id="4C60BEADFFD96A790F50FA83A6B2DC55" box="[812,904,1295,1319]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="palluma">
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<emphasis id="B914193CFFD96A790F50FA83A6B2DC55" box="[812,904,1295,1319]" italics="true" pageId="6" pageNumber="123">palluma</emphasis>
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</taxonomicName>
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group because it shares their well-known apomorphies with all the other members: square, non-imbricate superciliaries, rugose dorsal scales of the tail, usually a fragmented subocular, and subocular and supralabials separated by two or more scale rows. This new taxon belongs to the Puna subclade of the
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<taxonomicName id="4C60BEADFFD96A790E05FAF7A7EFDCE1" box="[633,725,1403,1427]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="palluma">
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<emphasis id="B914193CFFD96A790E05FAF7A7EFDCE1" box="[633,725,1403,1427]" italics="true" pageId="6" pageNumber="123">palluma</emphasis>
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</taxonomicName>
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group (
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<bibRefCitation id="EFF1B8DFFFD96A790F52FAF6A175DCE0" author="Lobo" box="[814,1103,1402,1426]" pageId="6" pageNumber="123" refString="Lobo, F. & Quinteros, S. (2005 b) A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia, 45, 143 - 177. http: // dx. doi. org / 10.1590 / S 0031 - 10492005001300001" type="journal article" year="2005" yearSuffix="b">Lobo & Quinteros 2005b</bibRefCitation>
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) because it shows the typical dorsal “spray” pattern (southern
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<taxonomicName id="4C60BEADFFD96A790E64FA13A74EDCC5" box="[536,628,1439,1463]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="palluma">
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<emphasis id="B914193CFFD96A790E64FA13A74EDCC5" box="[536,628,1439,1463]" italics="true" pageId="6" pageNumber="123">palluma</emphasis>
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</taxonomicName>
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group members show a dorsal reticulated pattern). Within the Puna subclade,
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<taxonomicName id="4C60BEADFFD96A790D74FA48A4B1DCA9" box="[264,395,1475,1499]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
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<emphasis id="B914193CFFD96A790D74FA48A4B1DCA9" box="[264,395,1475,1499]" italics="true" pageId="6" pageNumber="123">P. williamsi</emphasis>
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</taxonomicName>
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differs from all other species in its particular character combination;
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<taxonomicName id="4C60BEADFFD96A7908DCFA48A01EDCA9" box="[1184,1316,1475,1499]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
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<emphasis id="B914193CFFD96A7908DCFA48A01EDCA9" box="[1184,1316,1475,1499]" italics="true" pageId="6" pageNumber="123">P. williamsi</emphasis>
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</taxonomicName>
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shows the dorsum of neck homogeneous melanic, not interrupted in the midline as in
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<taxonomicName id="4C60BEADFFD96A790F87FA64A1FCDC8D" box="[1019,1222,1511,1535]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="antofagastensis">
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<emphasis id="B914193CFFD96A790F87FA64A1FCDC8D" box="[1019,1222,1511,1535]" italics="true" pageId="6" pageNumber="123">P. antofagastensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C60BEADFFD96A7908ABFA64A077DC8D" box="[1239,1357,1511,1535]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="laurenti">
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<emphasis id="B914193CFFD96A7908ABFA64A077DC8D" box="[1239,1357,1511,1535]" italics="true" pageId="6" pageNumber="123">P. laurenti</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C60BEADFFD96A7909FBFA64A5E1DF51" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="punae">
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<emphasis id="B914193CFFD96A7909FBFA64A5E1DF51" italics="true" pageId="6" pageNumber="123">P. punae</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C60BEADFFD96A790C95F987A4E4DF51" box="[233,478,1547,1571]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
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<emphasis id="B914193CFFD96A790C95F987A4E4DF51" box="[233,478,1547,1571]" italics="true" pageId="6" pageNumber="123">Phymaturus williamsi</emphasis>
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</taxonomicName>
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lacks enlarged scales posterior to the cloacal opening which are present in
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<taxonomicName id="4C60BEADFFD96A790955F980A0A6DF51" box="[1321,1436,1547,1571]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="laurenti">
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<emphasis id="B914193CFFD96A790955F980A0A6DF51" box="[1321,1436,1547,1571]" italics="true" pageId="6" pageNumber="123">P. laurenti</emphasis>
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</taxonomicName>
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and
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||
<taxonomicName id="4C60BEADFFD96A790CB4F9BCA464DF35" box="[200,350,1584,1607]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFD96A790CB4F9BCA464DF35" box="[200,350,1584,1607]" italics="true" pageId="6" pageNumber="123">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
. Tarsal scales are strongly keeled in
|
||
<taxonomicName id="4C60BEADFFD96A790F74F9BCA6B6DF35" box="[776,908,1583,1607]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790F74F9BCA6B6DF35" box="[776,908,1583,1607]" italics="true" pageId="6" pageNumber="123">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
but slightly keeled in
|
||
<taxonomicName id="4C60BEADFFD96A7908ECF9BCA03EDF35" box="[1168,1284,1583,1607]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFD96A7908ECF9BCA03EDF35" box="[1168,1284,1583,1607]" italics="true" pageId="6" pageNumber="123">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C60BEADFFD96A790968F9A3A5C4DF19" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790968F9A3A5C4DF19" italics="true" pageId="6" pageNumber="123">Phymaturus williamsi</emphasis>
|
||
</taxonomicName>
|
||
exhibits an “aggregate” dorsal pattern, unlike the homogeneous spray of most Puna species; a similar condition is only found in
|
||
<taxonomicName id="4C60BEADFFD96A790DB9F9F4A7AADFFD" box="[453,656,1655,1679]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFD96A790DB9F9F4A7AADFFD" box="[453,656,1655,1679]" italics="true" pageId="6" pageNumber="123">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 12D in
|
||
<bibRefCitation id="EFF1B8DFFFD96A790F51F9FBA16ADFFD" author="Lobo" box="[813,1104,1655,1679]" pageId="6" pageNumber="123" refString="Lobo, F. & Quinteros, S. (2005 b) A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia, 45, 143 - 177. http: // dx. doi. org / 10.1590 / S 0031 - 10492005001300001" type="journal article" year="2005" yearSuffix="b">Lobo & Quinteros 2005b</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C60BEADFFD96A790818F9FBA061DFFD" box="[1124,1371,1655,1679]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790818F9FBA061DFFD" box="[1124,1371,1655,1679]" italics="true" pageId="6" pageNumber="123">Phymaturus williamsi</emphasis>
|
||
</taxonomicName>
|
||
lacks enlarged scales on the anterior margin of the antehumeral fold and in the centre of chest, as in
|
||
<taxonomicName id="4C60BEADFFD96A7908ADF910A0A6DFC1" box="[1233,1436,1691,1715]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFD96A7908ADF910A0A6DFC1" box="[1233,1436,1691,1715]" italics="true" pageId="6" pageNumber="123">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B914193CFFD96A790CBAF94CA47BDFA5" box="[198,321,1727,1751]" italics="true" pageId="6" pageNumber="123">
|
||
<taxonomicName id="4C60BEADFFD96A790CBAF94CA407DFA5" box="[198,317,1727,1751]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="laurenti">P. laurenti</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Flank coloration in females is absent in
|
||
<taxonomicName id="4C60BEADFFD96A790F7BF94CA6B0DFA5" box="[775,906,1727,1751]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790F7BF94CA6B0DFA5" box="[775,906,1727,1751]" italics="true" pageId="6" pageNumber="123">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
but is present in
|
||
<taxonomicName id="4C60BEADFFD96A790836F94CA029DFA5" box="[1098,1299,1727,1751]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFD96A790836F94CA029DFA5" box="[1098,1299,1727,1751]" italics="true" pageId="6" pageNumber="123">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
(yellow),
|
||
<taxonomicName id="4C60BEADFFD96A7909FBF94CA5D5DF89" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFD96A7909FBF94CA5D5DF89" italics="true" pageId="6" pageNumber="123">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFD96A790D50F968A4F5DF89" box="[300,463,1763,1787]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFD96A790D50F968A4F5DF89" box="[300,463,1763,1787]" italics="true" pageId="6" pageNumber="123">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
(orange). A light-gray vertebral stripe, which is absent in
|
||
<taxonomicName id="4C60BEADFFD96A790801F968A070DF89" box="[1149,1354,1763,1787]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFD96A790801F968A070DF89" box="[1149,1354,1763,1787]" italics="true" pageId="6" pageNumber="123">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFD96A7909FBF968A5D5DE6D" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFD96A7909FBF968A5D5DE6D" italics="true" pageId="6" pageNumber="123">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
, is present in
|
||
<taxonomicName id="4C60BEADFFD96A790DE7F884A71BDE6D" box="[411,545,1799,1823]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790DE7F884A71BDE6D" box="[411,545,1799,1823]" italics="true" pageId="6" pageNumber="123">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
; a dark gray vertebral stripe can be present in individuals of
|
||
<emphasis id="B914193CFFD96A790880F884A0A6DE6D" box="[1276,1436,1800,1823]" italics="true" pageId="6" pageNumber="123">
|
||
<taxonomicName id="4C60BEADFFD96A790880F884A0A3DE6D" box="[1276,1433,1800,1823]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="aguanegra">P. aguanegra</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Females of
|
||
<taxonomicName id="4C60BEADFFD96A790D67F8A0A4A7DE31" box="[283,413,1835,1859]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790D67F8A0A4A7DE31" box="[283,413,1835,1859]" italics="true" pageId="6" pageNumber="123">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
lack white transversal stripes on the dorsal pattern, as in females of
|
||
<taxonomicName id="4C60BEADFFD96A7908DEF8A0A056DE31" box="[1186,1388,1835,1859]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFD96A7908DEF8A0A056DE31" box="[1186,1388,1835,1859]" italics="true" pageId="6" pageNumber="123">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFD96A790CEBF8DCA436DE15" box="[151,268,1871,1895]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFD96A790CEBF8DCA436DE15" box="[151,268,1871,1895]" italics="true" pageId="6" pageNumber="123">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
(not all individuals).
|
||
<taxonomicName id="4C60BEADFFD96A790E79F8C3A7C6DE15" box="[517,764,1871,1895]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790E79F8C3A7C6DE15" box="[517,764,1871,1895]" italics="true" pageId="6" pageNumber="123">Phymaturus williamsi</emphasis>
|
||
</taxonomicName>
|
||
exhibits a tricolor dorsal pattern, with two
|
||
<typeStatus id="54DB7B8CFFD96A79088CF8DCA011DE1A" box="[1264,1323,1872,1896]" pageId="6" pageNumber="123">types</typeStatus>
|
||
of brown and a scattered ferriferous oxide spotting; this pattern is not found in any other species of the group, with the exception of
|
||
<taxonomicName id="4C60BEADFFD96A790D51F814A4FEDEDD" box="[301,452,1944,1967]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFD96A790D51F814A4FEDEDD" box="[301,452,1944,1967]" italics="true" pageId="6" pageNumber="123">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C60BEADFFD96A790D83F814A793DEDD" box="[511,681,1943,1967]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="paihuanense">
|
||
<emphasis id="B914193CFFD96A790D83F814A793DEDD" box="[511,681,1943,1967]" italics="true" pageId="6" pageNumber="123">P. paihuanense</emphasis>
|
||
</taxonomicName>
|
||
. A scapular spot is mostly absent (only one subadult male with a vanishing spot) in
|
||
<taxonomicName id="4C60BEADFFD96A790D19F830A4DDDEA1" box="[357,487,1979,2003]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD96A790D19F830A4DDDEA1" box="[357,487,1979,2003]" italics="true" pageId="6" pageNumber="123">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
as in most of Puna species, with the exception of
|
||
<taxonomicName id="4C60BEADFFD96A79086CF830A195DEA1" box="[1040,1199,1979,2003]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="123" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFD96A79086CF830A195DEA1" box="[1040,1199,1979,2003]" italics="true" pageId="6" pageNumber="123">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
. Preocular scale is in contact with canthal scale in
|
||
<taxonomicName id="4C60BEADFFD86A780D8FFF15A740D9C2" box="[499,634,152,176]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD86A780D8FFF15A740D9C2" box="[499,634,152,176]" italics="true" pageId="7" pageNumber="124">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
, whereas it is separated by a smaller scale in
|
||
<taxonomicName id="4C60BEADFFD86A7808DAFF15A04ED9C2" box="[1190,1396,152,176]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFD86A7808DAFF15A04ED9C2" box="[1190,1396,152,176]" italics="true" pageId="7" pageNumber="124">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFD86A7809FBFF15A42AD9A6" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFD86A7809FBFF15A42AD9A6" italics="true" pageId="7" pageNumber="124">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFD86A780D5DFF31A4ADD9A6" box="[289,407,188,212]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFD86A780D5DFF31A4ADD9A6" box="[289,407,188,212]" italics="true" pageId="7" pageNumber="124">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C60BEADFFD86A780DA6FF31A707D9A6" box="[474,573,189,212]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="punae">
|
||
<emphasis id="B914193CFFD86A780DA6FF31A707D9A6" box="[474,573,189,212]" italics="true" pageId="7" pageNumber="124">P. punae</emphasis>
|
||
</taxonomicName>
|
||
. Rostral scale can be divided in
|
||
<taxonomicName id="4C60BEADFFD86A780FC1FF31A179D9A6" box="[957,1091,188,212]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD86A780FC1FF31A179D9A6" box="[957,1091,188,212]" italics="true" pageId="7" pageNumber="124">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
but is always undivided in
|
||
<taxonomicName id="4C60BEADFFD86A7809FBFF31A42AD98A" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFD86A7809FBFF31A42AD98A" italics="true" pageId="7" pageNumber="124">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C60BEADFFD86A780D58FF6DA4A7D98A" box="[292,413,224,248]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="B914193CFFD86A780D58FF6DA4A7D98A" box="[292,413,224,248]" italics="true" pageId="7" pageNumber="124">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C60BEADFFD86A780D9BFF6DA7B7D98A" box="[487,653,224,248]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="mallimaccii">
|
||
<emphasis id="B914193CFFD86A780D9BFF6DA7B7D98A" box="[487,653,224,248]" italics="true" pageId="7" pageNumber="124">P. mallimaccii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C60BEADFFD86A780EDEFF6DA610D98A" box="[674,810,224,248]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD86A780EDEFF6DA610D98A" box="[674,810,224,248]" italics="true" pageId="7" pageNumber="124">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
shows the largest number of scales counted around midbody within the Puna subclade (x= 213.4; 186–235); the remaining species show a mean below 200 scales. Neither males nor females of
|
||
<taxonomicName id="4C60BEADFFD86A780D98FEA5A7B7D832" box="[484,653,296,320]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="paihuanense">
|
||
<emphasis id="B914193CFFD86A780D98FEA5A7B7D832" box="[484,653,296,320]" italics="true" pageId="7" pageNumber="124">P. paihuanense</emphasis>
|
||
</taxonomicName>
|
||
exhibit head melanism (present in
|
||
<taxonomicName id="4C60BEADFFD86A780865FEA5A1A6D832" box="[1049,1180,296,320]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD86A780865FEA5A1A6D832" box="[1049,1180,296,320]" italics="true" pageId="7" pageNumber="124">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
); males of this species show enlarged scales on the anterior margin of the antehumeral fold (absent in
|
||
<taxonomicName id="4C60BEADFFD86A780871FEC1A1B5D816" box="[1037,1167,332,356]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD86A780871FEC1A1B5D816" box="[1037,1167,332,356]" italics="true" pageId="7" pageNumber="124">P. williamsi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C60BEADFFD86A7808DBFEC0A0A6D816" box="[1191,1436,332,356]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="williamsi">
|
||
<emphasis id="B914193CFFD86A7808DBFEC0A0A6D816" box="[1191,1436,332,356]" italics="true" pageId="7" pageNumber="124">Phymaturus williamsi</emphasis>
|
||
</taxonomicName>
|
||
differ of
|
||
<taxonomicName id="4C60BEADFFD86A780C87FEFDA4ABD8FA" box="[251,401,368,392]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="damasense">
|
||
<emphasis id="B914193CFFD86A780C87FEFDA4ABD8FA" box="[251,401,368,392]" italics="true" pageId="7" pageNumber="124">P. damasense</emphasis>
|
||
</taxonomicName>
|
||
in the presence of non-projected scales in the anterior border of auditory meatus (present in
|
||
<taxonomicName id="4C60BEADFFD86A780CEBFE19A417D8DE" box="[151,301,404,428]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="damasense">
|
||
<emphasis id="B914193CFFD86A780CEBFE19A417D8DE" box="[151,301,404,428]" italics="true" pageId="7" pageNumber="124">P. damasense</emphasis>
|
||
</taxonomicName>
|
||
) and color pattern of males and females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A96A5FFD86A780CBBFE3BA060DFCE" pageId="7" pageNumber="124" type="description">
|
||
<paragraph id="8BDFC52EFFD86A780CBBFE3BA772DC7E" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="124">
|
||
<emphasis id="B914193CFFD86A780CBBFE3BA4DCD8A2" bold="true" box="[199,486,439,464]" pageId="7" pageNumber="124">
|
||
Description of
|
||
<typeStatus id="54DB7B8CFFD86A780D04FE34A4DBD8A2" box="[376,481,440,464]" pageId="7" pageNumber="124" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male. SVL 103.0 mm. Head length
|
||
<quantity id="4C9868CBFFD86A780F01FE3BA6DED8BD" box="[893,996,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="7" pageNumber="124" unit="mm" value="18.4">18.4 mm</quantity>
|
||
. Head width
|
||
<quantity id="4C9868CBFFD86A780807FE3BA1D8D8BD" box="[1147,1250,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="7" pageNumber="124" unit="mm" value="18.2">18.2 mm</quantity>
|
||
. Head height (at parietal)
|
||
<quantity id="4C9868CBFFD86A780C87FE50A459D886" box="[251,355,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="7" pageNumber="124" unit="mm" value="10.4">10.4 mm</quantity>
|
||
. Axilla-groin
|
||
<quantity id="4C9868CBFFD86A780E7CFE50A75FD886" box="[512,613,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="5.14" pageId="7" pageNumber="124" unit="mm" value="51.4">51.4 mm</quantity>
|
||
(49.8 % of SVL). Tail length (complete, not regenerated)
|
||
<quantity id="4C9868CBFFD86A78088AFE50A060D886" box="[1270,1370,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="9.77" pageId="7" pageNumber="124" unit="mm" value="97.7">97.7 mm</quantity>
|
||
(0.95 times SVL). Body moderately wide, trunk width: 40.0 mm (38.9 % of SVL). Twenty-four smooth dorsal head scales. Four, three (three times), two (two times) organs in postrostrals. Nasal bordered by nine scales, not in contact with rostral. Canthal separated from nasal by two scales. Loreal region flat. Eleven enlarged supralabial scales, none in contact with subocular. Eleven enlarged infralabials. Auditory meatus oval with three-four small conical projecting scales on the anterior margin. Auricular scale absent. Ten convex, juxtaposed temporals. Rostral undivided. Mental subpentagonal, in contact with seven scales. Interparietal bordered by 10 scales. Frontal region without an azygous scale. Supraorbital semicircles inconspicuous. No distinctly enlarged supraoculars. Twelve imbricate flat superciliaries. Subocular fragmented in three scales, separated from supralabials by one to two rows of lorilabials. Thirteen to 15 lorilabials, 12 contacting subocular on the right side. Preocular scale separated from lorilabial row by three scales. Scales of throat round, flat, and juxtaposed. One hundred and three gular scales between auditory meata. Lateral nuchal folds well developed, with granular scales over longitudinal fold. Antehumeral pocket well developed. Seventy-five scales between auditory meatus and shoulder. In ventral view, gular fold not well developed and posterior gular folds present with their anterior margins without enlarged scales on their borders. Enlarged scales in the centre of chest absent. Dorsal scales round, smooth, juxtaposed. Forty dorsal scales along midline of the trunk in a distance equivalent to head length. Scales around midbody: 220. Middorsal scales not enlarged compared to those on flanks. Ventral scales larger than dorsals. Ventral scales between mental and precloacal pores: 199. Eight precloacal pores in an undivided row with two supernumerary pores. Brachial and antebrachial scales smooth with rounded posterior margins. Supracarpals laminar, round, smooth. Subdigital lamellae of fingers with three keels. Number of subdigital lamellae of fingers I: 11; II: 13; III: 22; IV: 23; V: 15. Claws moderately long. Supradigital lamellae convex, imbricate. Infracarpals and infratarsals with round margins and 2–3 keels. Supracarpals and supratarsals smooth, with round posterior margins. Subdigital lamellae of toes I: 11; II: 17; III: 22; IV: 27; V: 20.
|
||
</paragraph>
|
||
<paragraph id="8BDFC52EFFD86A780CB5FA95A417DCEE" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="124">
|
||
<emphasis id="B914193CFFD86A780CB5FA95A4D7DC42" box="[201,493,1304,1328]" italics="true" pageId="7" pageNumber="124">Pattern of body and limbs</emphasis>
|
||
(
|
||
<figureCitation id="135BD9ABFFD86A780D80FA94A770DC42" box="[508,586,1304,1328]" captionStart="FIGURE 4. A" captionStartId="8.[151,250,1244,1267]" captionTargetBox="[151,1436,176,1229]" captionTargetId="figure@8.[151,1436,165,1229]" captionTargetPageId="8" captionText="FIGURE 4. A. Dorsal view of the holotype specimen of Phymaturus williamsi (MCN 2820); B. Ventral view of the same specimen. SVL: 103 mm." httpUri="https://zenodo.org/record/223685/files/figure.png" pageId="7" pageNumber="124">Figs. 4</figureCitation>
|
||
and
|
||
<figureCitation id="135BD9ABFFD86A780EFDFA94A7ABDC42" box="[641,657,1304,1328]" captionStart="FIGURE 5" captionStartId="9.[151,250,1188,1211]" captionTargetBox="[159,1428,277,1160]" captionTargetId="figure@9.[151,1436,265,1167]" captionTargetPageId="9" captionText="FIGURE 5. Patterns of coloration in life of Phymaturus williamsi sp. nov. A and B: Males; C and D: Females." httpUri="https://zenodo.org/record/223686/files/figure.png" pageId="7" pageNumber="124">5</figureCitation>
|
||
): This species exhibits an aggregate pattern of brown spots similar to that of
|
||
<taxonomicName id="4C60BEADFFD86A780C99FAB1A494DC26" box="[229,430,1340,1364]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="antofagastensis">
|
||
<emphasis id="B914193CFFD86A780C99FAB1A494DC26" box="[229,430,1340,1364]" italics="true" pageId="7" pageNumber="124">P. antofagastensis</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 12D in
|
||
<bibRefCitation id="EFF1B8DFFFD86A780E38FAB0A658DC26" author="Lobo" box="[580,866,1340,1364]" pageId="7" pageNumber="124" refString="Lobo, F. & Quinteros, S. (2005 b) A morphology-based phylogeny of Phymaturus (Iguania: Liolaemidae) with the description of four new species from Argentina. Papeis Avulsos de Zoologia, 45, 143 - 177. http: // dx. doi. org / 10.1590 / S 0031 - 10492005001300001" type="journal article" year="2005" yearSuffix="b">Lobo & Quinteros 2005b</bibRefCitation>
|
||
). Dorsum of neck and head is more melanic than
|
||
<emphasis id="B914193CFFD86A7809FAFAB1A415DC0A" italics="true" pageId="7" pageNumber="124">
|
||
in
|
||
<taxonomicName id="4C60BEADFFD86A780CEBFAEDA415DC0A" box="[151,303,1377,1400]" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="aguanegra">P. aguanegra</taxonomicName>
|
||
</emphasis>
|
||
individuals. In this species a vertebral line is usually conspicuous, dividing the dark coloration of neck (
|
||
<figureCitation id="135BD9ABFFD86A780CA7FA08A425DCEE" box="[219,287,1412,1436]" captionStart="FIGURE 5" captionStartId="9.[151,250,1188,1211]" captionTargetBox="[159,1428,277,1160]" captionTargetId="figure@9.[151,1436,265,1167]" captionTargetPageId="9" captionText="FIGURE 5. Patterns of coloration in life of Phymaturus williamsi sp. nov. A and B: Males; C and D: Females." httpUri="https://zenodo.org/record/223686/files/figure.png" pageId="7" pageNumber="124">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDFC52EFFD86A780CB5FA24A13DDC96" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="124">
|
||
<emphasis id="B914193CFFD86A780CB5FA24A4AFDCB2" box="[201,405,1448,1472]" italics="true" pageId="7" pageNumber="124">Coloration in life</emphasis>
|
||
(
|
||
<figureCitation id="135BD9ABFFD86A780DD5FA24A4C9DCB2" box="[425,499,1448,1472]" captionStart="FIGURE 5" captionStartId="9.[151,250,1188,1211]" captionTargetBox="[159,1428,277,1160]" captionTargetId="figure@9.[151,1436,265,1167]" captionTargetPageId="9" captionText="FIGURE 5. Patterns of coloration in life of Phymaturus williamsi sp. nov. A and B: Males; C and D: Females." httpUri="https://zenodo.org/record/223686/files/figure.png" pageId="7" pageNumber="124">Fig. 5</figureCitation>
|
||
): The general coloration varies between brown-orange to light yellow; as in
|
||
<taxonomicName id="4C60BEADFFD86A7809FBFA25A42ADC96" class="Reptilia" family="Liolaemidae" genus="Phymaturus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="124" phylum="Chordata" rank="species" species="aguanegra">
|
||
<emphasis id="B914193CFFD86A7809FBFA25A42ADC96" italics="true" pageId="7" pageNumber="124">P. aguanegra</emphasis>
|
||
</taxonomicName>
|
||
, this dorsal coloration exhibits the characteristic ferric oxide shade.
|
||
</paragraph>
|
||
<paragraph id="8BDFC52EFFD86A780CB5FA7DA060DFCE" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="124">
|
||
<emphasis id="B914193CFFD86A780CB5FA7DA400DF7A" box="[201,314,1521,1544]" italics="true" pageId="7" pageNumber="124">Variation:</emphasis>
|
||
Based on
|
||
<specimenCount id="9D660EA7FFD86A780DD2FA7CA732DF7A" box="[430,520,1520,1544]" pageId="7" pageNumber="124" type="adult">20 adult</specimenCount>
|
||
specimens (
|
||
<specimenCount id="9D660EA7FFD86A780EEFFA7CA7C2DF7A" box="[659,760,1520,1544]" pageId="7" pageNumber="124" type="male">10 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D660EA7FFD86A780F50FA7CA691DF7A" box="[812,939,1520,1544]" pageId="7" pageNumber="124" type="female">10 females</specimenCount>
|
||
). SVL
|
||
<quantity id="4C9868CBFFD86A780F84FA7CA192DF7A" box="[1016,1192,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="9.995" metricValueMax="10.45" metricValueMin="9.54" pageId="7" pageNumber="124" unit="mm" value="99.95" valueMax="104.5" valueMin="95.4">95.4–104.5 mm</quantity>
|
||
(x = 100.2; SD = 3.1) for adult specimens only. Head length 16.3–18.4% (x = 17.4%; SD = 0.1) of SVL. Tail length 0.86–1.05 (x = 0.92; SD = 0.06) times SVL. Scales around midbody 186–235 (x = 213.4; SD = 13.0). Dorsal head scales 22–27 (x = 24.0; SD = 1.4). Ventrals 169–209 (x = 191.1; SD = 11.4). Scales surrounding interparietal 8–10 (x = 8.8; SD = 0.9). Scales of neck along longitudinal fold from posterior border of auditory meatus to shoulder 64–89 (x = 76.1; SD = 7.4). Gulars 82–110 (x = 95.2; SD = 7.7). Scales between rostral and frontal 9–14 (x = 10.6; SD = 1.6).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A96A5FFD86A780CB5F944A1B1DE77" pageId="7" pageNumber="124" type="etymology">
|
||
<paragraph id="8BDFC52EFFD86A780CB5F944A1B1DE77" blockId="7.[151,1437,151,1833]" pageId="7" pageNumber="124">
|
||
<emphasis id="B914193CFFD86A780CB5F944A46ADF92" bold="true" box="[201,336,1736,1760]" pageId="7" pageNumber="124">Etymology.</emphasis>
|
||
We name this new species in honour of our
|
||
<collectingCountry id="F37785BEFFD86A780F20F944A6F1DF92" box="[860,971,1736,1760]" name="Argentina" pageId="7" pageNumber="124">Argentine</collectingCountry>
|
||
colleague and friend Jorge Williams, in recognition of his effort and dedication for the development of herpetology in our country.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A96A5FFD86A780CB5F883A7D1DE5A" box="[201,747,1807,1833]" pageId="7" pageNumber="124" type="distribution">
|
||
<paragraph id="8BDFC52EFFD86A780CB5F883A7D1DE5A" blockId="7.[151,1437,151,1833]" box="[201,747,1807,1833]" pageId="7" pageNumber="124">
|
||
<emphasis id="B914193CFFD86A780CB5F883A458DE5A" bold="true" box="[201,354,1807,1832]" pageId="7" pageNumber="124">Distribution.</emphasis>
|
||
Only known from its
|
||
<typeStatus id="54DB7B8CFFD86A780E26F89DA7B0DE5B" box="[602,650,1809,1833]" pageId="7" pageNumber="124">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |