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<document id="49859C8969A685860B367EAE312B7296" ID-DOI="10.5281/zenodo.281568" ID-GBIF-Dataset="eb4d0071-03e7-4670-b730-43e20478afa1" ID-ISSN="1175-5326" ID-Zenodo-Dep="281568" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460492821005" checkinUser="plazi" docAuthor="Saigusa, Toyohei" docDate="2012" docId="03C68C2E8077FF8C92C998AA3E287FE1" docLanguage="en" docName="zt03353p068.pdf" docOrigin="Zootaxa 3353" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empis (Planempis) lindebergi Saigusa, sp. nov." docType="treatment" docVersion="8" lastPageNumber="63" masterDocId="FFFFF4568072FF84925E9E2B3B1A7E66" masterDocTitle="Revision of the type series of Empis (Planempis) mandarina Frey, the type species of Planempis (Diptera: Empididae)" masterLastPageNumber="68" masterPageNumber="55" pageNumber="60" updateTime="1698310423591" updateUser="plazi">
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<mods:title id="A958C6CCE1E5D4CDCE7FE44E2A63B147">Revision of the type series of Empis (Planempis) mandarina Frey, the type species of Planempis (Diptera: Empididae)</mods:title>
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<mods:namePart id="1402B7BE854F2EFE76AEEFE748600BD1">Saigusa, Toyohei</mods:namePart>
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<mods:typeOfResource id="8413F1843025CE06E58C5D1543E83A3C">text</mods:typeOfResource>
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<mods:date id="7605240F5B6E7315A3D6CD55CBF412E0">2012</mods:date>
<mods:detail id="5DCD9C05C4F0681159FCA0AEE7E2B0A5" type="volume">
<mods:number id="409607B51645698031479732A3DDEC1F">3353</mods:number>
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<mods:classification id="903D7A9DC305E2298BC9AC62701B3617">journal article</mods:classification>
<mods:identifier id="52F0DEF8A59ABFCC9BA65F5A25097FF3" type="DOI">10.5281/zenodo.281568</mods:identifier>
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<treatment id="03C68C2E8077FF8C92C998AA3E287FE1" ID-DOI="http://doi.org/10.5281/zenodo.6180352" ID-GBIF-Taxon="119519896" ID-Zenodo-Dep="6180352" LSID="urn:lsid:plazi:treatment:03C68C2E8077FF8C92C998AA3E287FE1" httpUri="http://treatment.plazi.org/id/03C68C2E8077FF8C92C998AA3E287FE1" lastPageId="8" lastPageNumber="63" pageId="5" pageNumber="60">
<subSubSection id="C3756EB38077FF8192C998AA3ABA78DA" pageId="5" pageNumber="60" type="nomenclature">
<paragraph id="8BD03D388077FF8192C998AA39C878FD" blockId="5.[151,722,1665,1724]" box="[151,722,1665,1691]" pageId="5" pageNumber="60">
<heading id="D0988A548077FF8192C998AA39C878FD" bold="true" box="[151,722,1665,1691]" fontSize="11" level="1" pageId="5" pageNumber="60" reason="1">
<emphasis id="B91BE12A8077FF8192C998AA39C878FD" bold="true" box="[151,722,1665,1691]" pageId="5" pageNumber="60">
<taxonomicName id="4C6F46BB8077FF8192C998AA397278FD" authority="Saigusa" authorityName="Saigusa" box="[151,616,1665,1691]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="lindebergi" status="sp. nov." subGenus="Planempis">
<emphasis id="B91BE12A8077FF8192C998AA3BFE78FD" bold="true" box="[151,228,1665,1691]" italics="true" pageId="5" pageNumber="60">Empis</emphasis>
(
<emphasis id="B91BE12A8077FF8192AA98AA3A6978FD" bold="true" box="[244,371,1665,1691]" italics="true" pageId="5" pageNumber="60">Planempis</emphasis>
)
<emphasis id="B91BE12A8077FF8193DA98AA3AE478FD" bold="true" box="[388,510,1665,1691]" italics="true" pageId="5" pageNumber="60">lindebergi</emphasis>
Saigusa
</taxonomicName>
,
<taxonomicNameLabel id="A2285C518077FF81902A98AA39C878FD" box="[628,722,1665,1691]" pageId="5" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD03D388077FF8192C9988F3ABA78DA" blockId="5.[151,722,1665,1724]" box="[151,416,1700,1724]" pageId="5" pageNumber="60">
(
<figureCitation id="135421BD8077FF8192C1988F3BF378DA" box="[159,233,1700,1724]" captionStart="FIGURE 1" captionStartId="2.[151,250,1769,1792]" captionTargetBox="[340,1246,591,1738]" captionTargetId="figure@2.[340,1247,591,1745]" captionTargetPageId="2" captionText="FIGURE 1. Empis (Planempis) spp., male; A – C: habitus, lateral aspect; D-F: anterior part of body, dorsal aspect. A, D: E. (P.) mandarina Frey, lectotype male; B, E: E. (P.) lindebergi sp. nov., holotype male; C, F: E. (P.) janssoni sp. nov." httpUri="https://zenodo.org/record/281569/files/figure.png" pageId="5" pageNumber="60">Figs. 1</figureCitation>
B, E, 3B, 4, 5C)
</paragraph>
</subSubSection>
<subSubSection id="C3756EB38077FF8192C998C639C97938" pageId="5" pageNumber="60" type="reference_group">
<paragraph id="8BD03D388077FF8192C998C639CE7964" blockId="5.[151,1049,1772,1886]" box="[151,724,1772,1794]" pageId="5" pageNumber="60">
<treatmentCitationGroup id="AB7F1A168077FF8192C998C639CE7964" box="[151,724,1772,1794]" pageId="5" pageNumber="60">
<taxonomicName id="4C6F46BB8077FF8192C998C639777967" ID-CoL="8BZLZ" authority="Frey, 1953: 40" authorityName="Frey" authorityPageNumber="40" authorityYear="1953" box="[151,621,1772,1794]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
<emphasis id="B91BE12A8077FF8192C998C63BCD7964" box="[151,215,1773,1794]" italics="true" pageId="5" pageNumber="60">Empis</emphasis>
(
<emphasis id="B91BE12A8077FF8192BB98C73A4A7964" box="[229,336,1772,1794]" italics="true" pageId="5" pageNumber="60">Planempis</emphasis>
)
<emphasis id="B91BE12A8077FF81930098C73AD77964" box="[350,461,1772,1794]" italics="true" pageId="5" pageNumber="60">mandarina</emphasis>
:
<treatmentCitation id="0ACE1B298077FF81938598C639777967" author="Frey" box="[475,621,1772,1794]" page="40" pageId="5" pageNumber="60" year="1953">
<bibRefCitation id="EFFE40C98077FF81938598C639517964" author="Frey" box="[475,587,1772,1794]" pageId="5" pageNumber="60" refString="Frey, R. (1953) Studien uber ostasiatiche Dipteren. I. Die Gattung Empis L. Notulae Entomologicae, 33, 29 - 57." type="journal article" year="1953">Frey, 1953</bibRefCitation>
: 40
</treatmentCitation>
</taxonomicName>
(partim.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD03D388077FF8192C9992739457946" blockId="5.[151,1049,1772,1886]" box="[151,607,1802,1825]" pageId="5" pageNumber="60">
<treatmentCitationGroup id="AB7F1A168077FF8192C9992739457946" box="[151,607,1802,1825]" pageId="5" pageNumber="60">
<taxonomicName id="4C6F46BB8077FF8192C999273AE27946" ID-CoL="8BZLZ" authority="Frey, 1954: 417" authorityName="Frey" authorityPageNumber="417" authorityYear="1954" box="[151,504,1802,1825]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina">
<emphasis id="B91BE12A8077FF8192C999273A517947" box="[151,331,1803,1825]" italics="true" pageId="5" pageNumber="60">Empis mandarina</emphasis>
:
<treatmentCitation id="0ACE1B298077FF81930799273AE27946" author="Frey" box="[345,504,1802,1825]" page="417" pageId="5" pageNumber="60" year="1954">
<bibRefCitation id="EFFE40C98077FF81930799273AD27946" author="Frey" box="[345,456,1802,1825]" pageId="5" pageNumber="60" refString="Frey, R. (1954) 28. Empididae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region IV, 4. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 400 - 432, pl. 35 - 36." type="journal article" year="1954">Frey, 1954</bibRefCitation>
: 417
</treatmentCitation>
</taxonomicName>
(partim.).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD03D388077FF8192C9990139C97938" blockId="5.[151,1049,1772,1886]" pageId="5" pageNumber="60">
<treatmentCitationGroup id="AB7F1A168077FF8192C9990139C97938" pageId="5" pageNumber="60">
<taxonomicName id="4C6F46BB8077FF8192C9990139567959" ID-CoL="8BZLZ" authority="Frey, 1954" authorityName="Frey" authorityYear="1954" box="[151,588,1833,1855]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
<emphasis id="B91BE12A8077FF8192C999013BCD7959" box="[151,215,1834,1855]" italics="true" pageId="5" pageNumber="60">Empis</emphasis>
(
<emphasis id="B91BE12A8077FF8192BB99023A4A7959" box="[229,336,1833,1855]" italics="true" pageId="5" pageNumber="60">Planempis</emphasis>
)
<emphasis id="B91BE12A8077FF81930099023AD77959" box="[350,461,1833,1855]" italics="true" pageId="5" pageNumber="60">mandarina</emphasis>
:
<bibRefCitation id="EFFE40C98077FF819385990139567959" author="Frey" box="[475,588,1833,1855]" pageId="5" pageNumber="60" refString="Frey, R. (1954) 28. Empididae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region IV, 4. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 400 - 432, pl. 35 - 36." type="journal article" year="1954">Frey, 1954</bibRefCitation>
</taxonomicName>
., Taf. XXXV, Figs. 287, 288 (male genitalia).
<taxonomicName id="4C6F46BB8077FF8192C999623977793B" ID-CoL="8BZLZ" authority="Frey, 1955: 11" authorityName="Frey" authorityPageNumber="11" authorityYear="1955" box="[151,621,1864,1886]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
<emphasis id="B91BE12A8077FF8192C999623BCD7938" box="[151,215,1865,1886]" italics="true" pageId="5" pageNumber="60">Empis</emphasis>
(
<emphasis id="B91BE12A8077FF8192BB99633A4A7938" box="[229,336,1864,1886]" italics="true" pageId="5" pageNumber="60">Planempis</emphasis>
)
<emphasis id="B91BE12A8077FF81930099633AD77938" box="[350,461,1864,1886]" italics="true" pageId="5" pageNumber="60">mandarina</emphasis>
:
<treatmentCitation id="0ACE1B298077FF81938599623977793B" author="Frey" box="[475,621,1864,1886]" page="11" pageId="5" pageNumber="60" year="1955">
<bibRefCitation id="EFFE40C98077FF819385996239517938" author="Frey" box="[475,587,1864,1886]" pageId="5" pageNumber="60" refString="Frey, R. (1955) Studien uber ostasiatische Dipteren IV. Hilara Meig. (Suppl.), Empis L. (Suppl.), Platypalpus Macq. Notulae Entomologicae, 35, 1 - 14." type="journal article" year="1955">Frey, 1955</bibRefCitation>
: 11
</treatmentCitation>
</taxonomicName>
(partim.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3756EB38077FF8192C999A13FE079A1" pageId="5" pageNumber="60" type="diagnosis">
<paragraph id="8BD03D388077FF8192C999A13FE079A1" blockId="5.[151,1435,1930,1991]" pageId="5" pageNumber="60">
<emphasis id="B91BE12A8077FF8192C999A13A1079C5" bold="true" box="[151,266,1930,1955]" pageId="5" pageNumber="60">Diagnosis</emphasis>
. More or less large-sized blackish species of
<emphasis id="B91BE12A8077FF81915B99A7380479C5" box="[773,798,1932,1955]" italics="true" pageId="5" pageNumber="60">E.</emphasis>
(
<taxonomicName id="4C6F46BB8077FF81917099A038BE79C5" box="[814,932,1931,1955]" class="Insecta" family="Empididae" genus="Planempis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B91BE12A8077FF81917099A038BE79C5" box="[814,932,1931,1955]" italics="true" pageId="5" pageNumber="60">Planempis</emphasis>
</taxonomicName>
) with holoptic male eyes, black slender legs, entirely infuscated wings, dull blackish mesonotum without acrostichals, and flat-topped male cerci.
</paragraph>
</subSubSection>
<caption id="DF106DB08074FF8292C9989A3F507880" httpUri="https://zenodo.org/record/281572/files/figure.png" pageId="6" pageNumber="61" targetBox="[159,1403,188,1662]" targetPageId="6">
<paragraph id="8BD03D388074FF8292C9989A3F507880" blockId="6.[151,1436,1713,1766]" pageId="6" pageNumber="61">
<emphasis id="B91BE12A8074FF8292C9989A3A0878A1" bold="true" box="[151,274,1713,1736]" pageId="6" pageNumber="61">FIGURE 4.</emphasis>
<taxonomicName id="4C6F46BB8074FF8293479898395D78AE" box="[281,583,1714,1736]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="lindebergi" status="sp. nov." subGenus="Planempis">
<emphasis id="B91BE12A8074FF82934798983A4278AE" box="[281,344,1715,1736]" italics="true" pageId="6" pageNumber="61">Empis</emphasis>
(
<emphasis id="B91BE12A8074FF82933898993ACB78AE" box="[358,465,1714,1736]" italics="true" pageId="6" pageNumber="61">Planempis</emphasis>
)
<emphasis id="B91BE12A8074FF8293BE9899395D78AE" box="[480,583,1714,1736]" italics="true" pageId="6" pageNumber="61">lindebergi</emphasis>
</taxonomicName>
<emphasis id="B91BE12A8074FF8290139899398778A1" bold="true" box="[589,669,1714,1735]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="A2285C518074FF8290139899398778A1" box="[589,669,1714,1735]" pageId="6" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect.
</paragraph>
</caption>
<subSubSection id="C3756EB38074FF8392979926396378A2" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61" type="description">
<paragraph id="8BD03D388074FF839297992638A57E91" blockId="6.[151,1436,1805,2011]" lastBlockId="7.[151,1437,151,1985]" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
<emphasis id="B91BE12A8074FF82929799263A4E7940" bold="true" box="[201,340,1805,1830]" pageId="6" pageNumber="61">Description</emphasis>
.
<emphasis id="B91BE12A8074FF82933B992539097940" bold="true" box="[357,531,1806,1830]" pageId="6" pageNumber="61">
Male
<typeStatus id="54D4839A8074FF8293F2992539097940" box="[428,531,1806,1830]" pageId="6" pageNumber="61" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="135421BD8074FF829078992539737940" box="[550,617,1806,1830]" captionStart="FIGURE 1" captionStartId="2.[151,250,1769,1792]" captionTargetBox="[340,1246,591,1738]" captionTargetId="figure@2.[340,1247,591,1745]" captionTargetPageId="2" captionText="FIGURE 1. Empis (Planempis) spp., male; A – C: habitus, lateral aspect; D-F: anterior part of body, dorsal aspect. A, D: E. (P.) mandarina Frey, lectotype male; B, E: E. (P.) lindebergi sp. nov., holotype male; C, F: E. (P.) janssoni sp. nov." httpUri="https://zenodo.org/record/281569/files/figure.png" pageId="6" pageNumber="61">Fig. 1</figureCitation>
B). Head black, shrivelled, compound eyes touching for long distance nearly half as long as frons leaving small triangular upper and lower parts, upper facets enlarged, ca. 1.5X as large as lower facets in diameter; frons and face dull black, clypeus blackish brown, polished; occiput densely blackish grey pollinose, with postocular ciliation of rather short black bristles, row of 7 moderately strong occipital bristles and on precervical swelling many long yellow hairs, a few similar hairs on lower postocular area. Antenna slightly longer than shrivelled head length; basal two segments blackish brown, short black setose; basiflagellomere and stylus black; former 3.5X as long as basal thickness; relative lengths of 3 antennal segments and stylus ca. 2:1:3:2. Maxillary palpus more or less concealed in buccal cavity, slender and black, bearing a few black setae beneath; labrum 1.4X as long as shrivelled head height, polished blackish brown.
</paragraph>
<paragraph id="8BD03D388075FF8392979F2F3E867C06" blockId="7.[151,1437,151,1985]" pageId="7" pageNumber="62">
Thorax (
<figureCitation id="135421BD8075FF83936F9F2F3A6D7F7A" box="[305,375,260,284]" captionStart="FIGURE 1" captionStartId="2.[151,250,1769,1792]" captionTargetBox="[340,1246,591,1738]" captionTargetId="figure@2.[340,1247,591,1745]" captionTargetPageId="2" captionText="FIGURE 1. Empis (Planempis) spp., male; A – C: habitus, lateral aspect; D-F: anterior part of body, dorsal aspect. A, D: E. (P.) mandarina Frey, lectotype male; B, E: E. (P.) lindebergi sp. nov., holotype male; C, F: E. (P.) janssoni sp. nov." httpUri="https://zenodo.org/record/281569/files/figure.png" pageId="7" pageNumber="62">Fig. 1</figureCitation>
E) blackish brown; mesonotum densely pollinose, dull black, subdorsal stripes between acrostichal and dorsocentral stripes slightly brownish and 0.66X as wide as acrostichal stripe; humeral callus and postalar callus brown, scutellum and pleura blackish brown, latter dark grey pollinose, anepimeron brownish. Chaetotaxy: Most thoracic setae black; acrostichals completely absent; dorsocentrals irregularly biserial, consisting of sparsely set short black setae 0.75X as long as longer diameter of prothoracic spiracle, ending in strong presutellar bristle; humerus with weak black posterior bristle and many yellow fine hairs mixing 2 black ones on anterior area, notopleural depression clothed with fine yellow hairs and with 3—4 strong black bristles; hair-like black posthumeral seta; 1 supra-alar and 1 fairly strong postalar bristles; scutellum with 3 pairs of scutellars, inner two pairs equally strong; anterior pronotum with back setae, proepisternal hairs yellow; prosternum bare; laterotergal hairs dense and yellow, arising from oblique oval area, and mixing black seta on upper part of the area.
</paragraph>
<paragraph id="8BD03D388075FF8392979C463E877D3A" blockId="7.[151,1437,151,1985]" pageId="7" pageNumber="62">
Legs slender, simple, black and thinly dark grey pollinose. Coxae clothed with black bristles and hairs. Femora short setose above, almost bare beneath; f1 and f2 with av and pv row of short fine setae 0.33X as long as femora thickness, mixing a few stronger pv bristles 0.5X as long as femora thickness towards apex. Tibiae short setose; t1 with fine short weak ad bristle; t2 and t3 (
<figureCitation id="135421BD8075FF8390339CFC39B47C89" box="[621,686,727,751]" captionStart="FIGURE 5" captionStartId="8.[151,250,1180,1203]" captionTargetBox="[340,1247,447,1151]" captionTargetId="figure@8.[340,1247,447,1155]" captionTargetPageId="8" captionText="FIGURE 5. Hind legs of Empis (Planempis) spp., male; A-D: left hind leg excluding coxae; B: middle part of hind tibia showing ventral erect pile; E: hind femur showing pale posteroventral hairs; A, B: E. (P.) mandarina Frey, lectotype; C: E. (P.) lindebergi sp. nov., holotype; D, E: E. (P.) janssoni sp. nov., holotype." httpUri="https://zenodo.org/record/281573/files/figure.png" pageId="7" pageNumber="62">Fig. 5</figureCitation>
C) with ad and av rows of several bristles 0.5X as long as t2 thickness, t3 weakly dilated on apical 0.25, clothed with suberect black pile-like hairs on ventral surface. Tarsi short setose; fore and mid tarsi slender, metatarsi bearing a few short av bristles; hind metatarsus somewhat swollen, 0.5X as long as t3, almost as thick as apical part of t3, densely short setose beneath, bearing 2—3 ad and pd short bristles.
</paragraph>
<paragraph id="8BD03D388075FF8392979D4C3F287D97" blockId="7.[151,1437,151,1985]" pageId="7" pageNumber="62">
Wing (
<figureCitation id="135421BD8075FF8393479D4C3A447D19" box="[281,350,871,895]" captionStart="FIGURE 3" captionStartId="5.[151,250,786,809]" captionTargetBox="[340,1242,207,757]" captionTargetId="figure@5.[340,1247,193,765]" captionTargetPageId="5" captionText="FIGURE 3. Wings of Empis (Planempis) spp. A: E. (P.) mandarina Frey, lectotype male; B: E. (P.) lindebergi sp. nov., holotype male; C: E. (P.) janssoni sp. nov., holotype male; D: ditto, paratype female 1; E: ditto, paratype female 2; F: E. (P.) sp. female syntype of E. (P.) mandarina sensu Frey from Kuatun." httpUri="https://zenodo.org/record/281571/files/figure.png" pageId="7" pageNumber="62">Fig. 3</figureCitation>
B) as illustrated, 2.9X as long as wide, weakly tapered towards tip, and distinctly infuscated brownish; pterostigma and veins dark brown; M1, M2, M4 and CuA+CuP weakened near wing margin, M2 abbreviated distally incomplete; R4 forming ca. 60˚ to R5 basally; outer margin of cell r2+3 about 0.66X as long as that of cell r4; discal cell rather short; cell cua sharply pointed distally. Halter dark brown.
</paragraph>
<paragraph id="8BD03D388075FF8392979A2B3A947AAA" blockId="7.[151,1437,151,1985]" pageId="7" pageNumber="62">Abdomen black to blackish brown, thinly dark brown pollinose, and clothed with yellow hairs longer on lateral parts of terga 1–4; setae on dorsal parts of terga very short and black. Abdominal tergum 8 0.25X as long as abdominal tergum 7, fused with abruptly narrowed lateral areas of sternum, weakly expanded posteriorly around dorsomedian part, bearing row of 9 short setae; abdominal sternum 8 0.66X as long as sternum 7, weakly produced sublaterally forwards, and bearing 2—3 irregular rows of posterior setae, of which longest ones slightly shorter than 0.5 length of sternum.</paragraph>
<paragraph id="8BD03D388075FF8392979AF3391378C6" blockId="7.[151,1437,151,1985]" pageId="7" pageNumber="62">
Male genitalia (
<figureCitation id="135421BD8075FF8393249AF33AA57A96" box="[378,447,1240,1264]" captionStart="FIGURE 4" captionStartId="6.[151,250,1713,1736]" captionTargetBox="[159,1403,188,1662]" captionTargetId="figure@6.[151,1436,169,1689]" captionTargetPageId="6" captionText="FIGURE 4. Empis (Planempis) lindebergi sp. nov., holotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect." httpUri="https://zenodo.org/record/281572/files/figure.png" pageId="7" pageNumber="62">Fig. 4</figureCitation>
A): Flat-topped
<typeStatus id="54D4839A8075FF83902C9AF239BC7A97" box="[626,678,1241,1265]" pageId="7" pageNumber="62">type</typeStatus>
. Cercus dark reddish brown, darker on ventral process, and clothed with brown hairs; epandrial lobe blackish brown, bearing black setae. Cerci (
<figureCitation id="135421BD8075FF8391A29AD73F567B72" box="[1020,1100,1276,1300]" captionStart="FIGURE 4" captionStartId="6.[151,250,1713,1736]" captionTargetBox="[159,1403,188,1662]" captionTargetId="figure@6.[151,1436,169,1689]" captionTargetPageId="6" captionText="FIGURE 4. Empis (Planempis) lindebergi sp. nov., holotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect." httpUri="https://zenodo.org/record/281572/files/figure.png" pageId="7" pageNumber="62">Figs. 4</figureCitation>
B, C) with dorsal surface flattened, combined cerci viewed from above sub-pentagonal with lateral margins parallel on basal 0.66, then gently converging distally, and surrounding inverted triangular fenestrula, of which posterior open space almost 0.5X as long as wide; dorsal process of cercus weakly curved inwardly in dorsal aspect, in lateral aspect simple, weakly curved ventrally, but without ventral drooping and only with single concavity at tip; middle swelling of cercus moderately large; ventral process of cercus moderately long, gently widened distally and truncate at tip; supra-anal area with large sclerotization warped along posterior margin, produced into median projection, with irregular transverse wrinkles; subanal area with pair of weak oval sclerotizations; epandrial lobe with roundly produced oblique distal margin, and bearing dense bristles; hypandrium narrow and separated ventrally; phallus outside from coelom (
<figureCitation id="135421BD8075FF8392C1986B3BF4783E" box="[159,238,1600,1624]" captionStart="FIGURE 4" captionStartId="6.[151,250,1713,1736]" captionTargetBox="[159,1403,188,1662]" captionTargetId="figure@6.[151,1436,169,1689]" captionTargetPageId="6" captionText="FIGURE 4. Empis (Planempis) lindebergi sp. nov., holotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect." httpUri="https://zenodo.org/record/281572/files/figure.png" pageId="7" pageNumber="62">Figs. 4</figureCitation>
D, E) moderately long, middle 0.33 swollen, with pair of triangular lamellate lateral expansions and dorsal keel triangularly produced at middle, its distal 0.33 gradually narrowed towards middle, then much tapered and strongly recurved anterodorsally.
</paragraph>
<paragraph id="8BD03D388075FF8392979887396378A2" blockId="7.[151,1437,151,1985]" box="[201,633,1708,1732]" pageId="7" pageNumber="62">
Length: Body
<quantity id="4C9790DD8075FF83933498873AD978A2" box="[362,451,1708,1732]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="7" pageNumber="62" unit="mm" value="9.5">9.5 mm</quantity>
; wing
<quantity id="4C9790DD8075FF8390509887396F78A2" box="[526,629,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="7" pageNumber="62" unit="mm" value="10.6">10.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756EB38075FF83929798FA39C279FA" pageId="7" pageNumber="62" type="materials_examined">
<paragraph id="8BD03D388075FF83929798FA3E75791E" blockId="7.[151,1437,151,1985]" pageId="7" pageNumber="62">
<emphasis id="B91BE12A8075FF83929798FA3904788F" bold="true" box="[201,542,1744,1769]" pageId="7" pageNumber="62">
<typeStatus id="54D4839A8075FF83929798FA3A1E788F" box="[201,260,1745,1769]" pageId="7" pageNumber="62">Type</typeStatus>
material.
<typeStatus id="54D4839A8075FF8393DE98FB3904788F" box="[384,542,1744,1769]" pageId="7" pageNumber="62" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
ɗ, labelled: N.E.
<collectingCountry id="F3787DA88075FF83908098FB3853788E" box="[734,841,1744,1768]" name="Myanmar" pageId="7" pageNumber="62">BURMA</collectingCountry>
/Kambaiti,
<quantity id="4C9790DD8075FF83919598FB3F02788E" box="[971,1048,1744,1768]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="62" unit="m" value="2000.0">2000m</quantity>
/
<date id="FFD11BF88075FF83967798FB3FB8788E" box="[1065,1186,1744,1768]" pageId="7" pageNumber="62" value="1934-05-25">25/5. 1934</date>
<emphasis id="B91BE12A8075FF8396F498FB3E1E788E" box="[1194,1284,1744,1768]" italics="true" pageId="7" pageNumber="62">Malaise</emphasis>
// Spec.typ.// pinx [handwritten]//
<typeStatus id="54D4839A8075FF8393D298DF3AE9796A" box="[396,499,1780,1804]" pageId="7" pageNumber="62" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C6F46BB8075FF83905498DE39FF796A" authority="(Planempis)" baseAuthorityName="Planempis" box="[522,741,1780,1804]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B91BE12A8075FF83905498DE394A796A" box="[522,592,1781,1804]" italics="true" pageId="7" pageNumber="62">Empis</emphasis>
(
<emphasis id="B91BE12A8075FF83903998DF39C7796A" box="[615,733,1780,1804]" italics="true" pageId="7" pageNumber="62">Planempis</emphasis>
)
</taxonomicName>
/
<taxonomicName id="4C6F46BB8075FF8390A298DF3874796A" box="[764,878,1780,1804]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="lindebergi">
<emphasis id="B91BE12A8075FF8390A298DF3874796A" box="[764,878,1780,1804]" italics="true" pageId="7" pageNumber="62">lindebergi</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFFE40C98075FF8391DA98DF3F32796A" author="Saigusa" box="[900,1064,1780,1804]" pageId="7" pageNumber="62" refString="Liu, X., Saigusa, T. &amp; Yang, D. (2012) Two new species of Empis (Planempis) from Oriental China, with an updated key to species of China (Diptera: Empidoidea). Zootaxa, 3239, 51 - 57." type="journal article" year="2012">Saigusa, 2012</bibRefCitation>
// [
<typeStatus id="54D4839A8075FF83961298DF3E20796A" box="[1100,1338,1780,1804]" pageId="7" pageNumber="62" type="paralectotype">PARALECTOTYPE</typeStatus>
]/
<taxonomicName id="4C6F46BB8075FF83970898DE3A047956" authority="(Planempis)" baseAuthorityName="Planempis" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B91BE12A8075FF83970898DE3E86796A" box="[1366,1436,1781,1804]" italics="true" pageId="7" pageNumber="62">Empis</emphasis>
(
<emphasis id="B91BE12A8075FF8392FE99333A0C7956" box="[160,278,1816,1840]" italics="true" pageId="7" pageNumber="62">Planempis</emphasis>
)
</taxonomicName>
/
<taxonomicName id="4C6F46BB8075FF8393709933393C7956" authority="Frey, 1953" authorityName="Frey" authorityYear="1953" box="[302,550,1816,1841]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="mandarina">
<emphasis id="B91BE12A8075FF83937099333AB37956" box="[302,425,1816,1840]" italics="true" pageId="7" pageNumber="62">mandarina</emphasis>
<bibRefCitation id="EFFE40C98075FF8393EE9932393C7956" author="Frey" box="[432,550,1816,1841]" pageId="7" pageNumber="62" refString="Frey, R. (1953) Studien uber ostasiatiche Dipteren. I. Die Gattung Empis L. Notulae Entomologicae, 33, 29 - 57." type="journal article" year="1953">Frey, 1953</bibRefCitation>
</taxonomicName>
/ designated by/
<bibRefCitation id="EFFE40C98075FF839080993238827956" author="Saigusa" box="[734,920,1816,1840]" pageId="7" pageNumber="62" refString="Liu, X., Saigusa, T. &amp; Yang, D. (2012) Two new species of Empis (Planempis) from Oriental China, with an updated key to species of China (Diptera: Empidoidea). Zootaxa, 3239, 51 - 57." type="journal article" year="2012">T. Saigusa, 2012</bibRefCitation>
(male genitalia and posterior part of abdomen macerated and preserved in vial with glycerol, and left wing detached from body and mounted between cover slips with
<collectingCountry id="F3787DA88075FF83928E994B3A3E791E" box="[208,292,1888,1912]" name="Canada" pageId="7" pageNumber="62">Canada</collectingCountry>
balsam; left hind leg detached from body and mounted on minuten, attached to main pin) (
<collectionCode id="ED7EA5FD8075FF839741994B3E79791E" box="[1311,1379,1888,1912]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="7" pageNumber="62">ZMH</collectionCode>
).
</paragraph>
<paragraph id="8BD03D388075FF83929799AF39C279FA" blockId="7.[151,1437,151,1985]" box="[201,728,1924,1949]" pageId="7" pageNumber="62">
<emphasis id="B91BE12A8075FF83929799AF3A7879FB" bold="true" box="[201,354,1924,1949]" pageId="7" pageNumber="62">Type-locality</emphasis>
. N. E.
<collectingCountry id="F3787DA88075FF8393EF99AE391879FA" box="[433,514,1925,1948]" name="Myanmar" pageId="7" pageNumber="62">Burma</collectingCountry>
: Kambaiti
<quantity id="4C9790DD8075FF83902299AF39CE79FA" box="[636,724,1924,1948]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="62" unit="m" value="2000.0">2000 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756EB38075FF83929799833AC679A7" box="[201,476,1960,1985]" pageId="7" pageNumber="62" type="distribution">
<paragraph id="8BD03D388075FF83929799833AC679A7" blockId="7.[151,1437,151,1985]" box="[201,476,1960,1985]" pageId="7" pageNumber="62">
<emphasis id="B91BE12A8075FF83929799833A4179A7" bold="true" box="[201,347,1960,1985]" pageId="7" pageNumber="62">Distribution</emphasis>
.
<collectingCountry id="F3787DA88075FF83933799823AC279A7" box="[361,472,1961,1985]" name="Myanmar" pageId="7" pageNumber="62">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756EB3807AFF8C92979EB338D97E91" pageId="8" pageNumber="63" type="etymology">
<paragraph id="8BD03D38807AFF8C92979EB338D97E91" blockId="8.[151,1436,151,391]" pageId="8" pageNumber="63">
<emphasis id="B91BE12A807AFF8C92979EB33A527ED6" bold="true" box="[201,328,152,176]" pageId="8" pageNumber="63">Etymology</emphasis>
. The new species is named after the late Dr. Bernhard Lindeberg, a taxonomist of
<taxonomicName id="4C6F46BB807AFF8C96A29EBC3E867EC9" box="[1276,1436,151,175]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
and staff member of the Zoological Museum of University of Helsinki, who assisted the author during his stay at the museum in 1985, while working on Dr. Frey’s species of
<taxonomicName id="4C6F46BB807AFF8C911C9EF438A57E91" box="[834,959,223,247]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="family">Empididae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756EB3807AFF8C92979F2F3E287FE1" pageId="8" pageNumber="63" type="discussion">
<paragraph id="8BD03D38807AFF8C92979F2F3E287FE1" blockId="8.[151,1436,151,391]" pageId="8" pageNumber="63">
<emphasis id="B91BE12A807AFF8C92979F2F3A2C7F7A" bold="true" box="[201,310,260,284]" pageId="8" pageNumber="63">Remarks</emphasis>
. The
<typeStatus id="54D4839A807AFF8C93279F2F3AC17F7A" box="[377,475,260,284]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
is one of two male
<typeStatus id="54D4839A807AFF8C909C9F2E383E7F7B" box="[706,804,261,285]" pageId="8" pageNumber="63" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C6F46BB807AFF8C91139F2E38F37F7A" box="[845,1001,260,284]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="mandarina">
<emphasis id="B91BE12A807AFF8C91139F2E38F37F7A" box="[845,1001,260,284]" italics="true" pageId="8" pageNumber="63">E. mandarina</emphasis>
</taxonomicName>
from Kambaiti. The male genitalia of this specimen was illustrated in
<bibRefCitation id="EFFE40C9807AFF8C905A9F0338CA7F59" author="Frey" box="[516,976,295,320]" pageId="8" pageNumber="63" refString="Frey, R. (1954) 28. Empididae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region IV, 4. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 400 - 432, pl. 35 - 36." type="journal article" year="1954">Frey (1954, Taf. XXXV, Figs. 287, 288)</bibRefCitation>
. This species is somewhat similar to the
<typeStatus id="54D4839A807AFF8C92C99F673BE47F02" box="[151,254,332,356]" pageId="8" pageNumber="63" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C6F46BB807AFF8C937D9F663AA47F02" box="[291,446,332,356]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="mandarina">
<emphasis id="B91BE12A807AFF8C937D9F663AA47F02" box="[291,446,332,356]" italics="true" pageId="8" pageNumber="63">E. mandarina</emphasis>
</taxonomicName>
, but the new species is slightly larger than
<taxonomicName id="4C6F46BB807AFF8C91FA9F663F247F02" box="[932,1086,332,356]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="mandarina">
<emphasis id="B91BE12A807AFF8C91FA9F663F247F02" box="[932,1086,332,356]" italics="true" pageId="8" pageNumber="63">E. mandarina</emphasis>
</taxonomicName>
, and distinguished by the slen- der black legs and different shape of the male cerci, of which the dorsal processes are produced ventrally.
</paragraph>
</subSubSection>
</treatment>
</document>
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