treatments-xml/data/03/C4/87/03C487CEF67D2C6A8AD63F43C36DFDCB.xml

<document id="DC681893930FAFA8CCEBC6D5208DBE4F" ID-CLB-Dataset="30302" ID-DOI="10.11646/zootaxa.4407.1.6" ID-GBIF-Dataset="de87eb6f-f4eb-46af-8e43-95ca6ab2b672" ID-ISSN="1175-5326" ID-Zenodo-Dep="1216486" ID-ZooBank="951098C7-18EF-4C9B-AC19-1AEAE2E64ECD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1523433465060" checkinUser="plazi" docAuthor="Orrico, Victor G. D., Dias, Iuri R. &amp; Marciano-Jr, Euvaldo" docDate="2018" docId="03C487CEF67D2C6A8AD63F43C36DFDCB" docLanguage="en" docName="zootaxa.4407.1.5.pdf" docOrigin="Zootaxa 4407 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllodytes praeceptor Orrico &amp; Dias &amp; Marciano-Jr 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="107" masterDocId="FFFDFFB6F67C2C6C8A413A0CC27AFF94" masterDocTitle="Another new species of Phyllodytes (Anura: Hylidae) from the Atlantic Forest of northeastern Brazil" masterLastPageNumber="110" masterPageNumber="101" pageNumber="102" updateTime="1698535969158" updateUser="ExternalLinkService">
<mods:mods id="0C37862856F77D0F6B7AB50B78A897CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6E3F9650E9D299FC3335239AC09B053">
<mods:title id="84EF3155A1F7F14B729C9508EBD29826">Another new species of Phyllodytes (Anura: Hylidae) from the Atlantic Forest of northeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="8E70F26C9F1BE8811B3E7EAAADBA6ABF" type="personal">
<mods:role id="A7987B15F8506498082CEB83BE0311CC">
<mods:roleTerm id="9AC4099FF68393560CEBBA1C331214E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D7CBC2FF3902A4C7C536EBF1AD181A48">Orrico, Victor G. D.</mods:namePart>
</mods:name>
<mods:name id="6B6213F038525ECB5CF891CA7D55B4FE" type="personal">
<mods:role id="44D07871C777AE1DAF700BB469D75D99">
<mods:roleTerm id="7B576FA07C1ACCB5DB6762CAC3D77A38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47F3447ABB44E1BD61DA49606FF92B66">Dias, Iuri R.</mods:namePart>
</mods:name>
<mods:name id="936E3DFC65B4939883949C45511C2B47" type="personal">
<mods:role id="4BD6BBA6C3FFE08DD98D16293F906CF1">
<mods:roleTerm id="2FDABD745FFCB7341305445B57C05205">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61AC58AAEE2D33A75BC4B274F4BD3CCC">Marciano-Jr, Euvaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="DE4DA52247281B142BE6B09F1C37FD77">text</mods:typeOfResource>
<mods:relatedItem id="E977C0F8B6382B472A180E897ABC1954" type="host">
<mods:titleInfo id="998BA2AD8FAA18235841A33F506732FF">
<mods:title id="F22A943816889D46F9E9806DD741EAEF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="676A5B8F36B4DE3F82E23DD1BE0B67E4">
<mods:date id="36C947A07DA56C0E6404DFDB4B2B5712">2018</mods:date>
<mods:detail id="BE4D92C8093FAADABAFB7ABB9CF49C14" type="pubDate">
<mods:number id="DD7ACE38D973FF1AC17BD968458D1902">2018-04-09</mods:number>
</mods:detail>
<mods:detail id="770B53E80EB71D9657691E4D3219EF2D" type="volume">
<mods:number id="5D8D0EF5247C15F92AAB89C466045964">4407</mods:number>
</mods:detail>
<mods:detail id="5055448C242C33EACEC550709479C8C7" type="issue">
<mods:number id="317069C922579ED18D7CB2250B6E3D86">1</mods:number>
</mods:detail>
<mods:extent id="487F7D2F1523B4C9D9115002B4D8ED0A" unit="page">
<mods:start id="39A0CC89381B79680FD416CF2BE32777">101</mods:start>
<mods:end id="52D28A6EBEF3154FB02BF0D70547167D">110</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F58918BA8DE74FB7333FD8C973135FDC">journal article</mods:classification>
<mods:identifier id="2D4277A30BB81F6E83EDD5585D8AEE23" type="CLB-Dataset">30302</mods:identifier>
<mods:identifier id="F75EDB4BE683ECF46EC3DBABE3EE2286" type="DOI">10.11646/zootaxa.4407.1.6</mods:identifier>
<mods:identifier id="5B9C03F1D1DFE3A576FF94C9F94A7AD2" type="GBIF-Dataset">de87eb6f-f4eb-46af-8e43-95ca6ab2b672</mods:identifier>
<mods:identifier id="37DE9726DA12BC2D33F7889D7AA446D8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8D2C62A5E53EBEB2B95E3F37AD1E60DB" type="Zenodo-Dep">1216486</mods:identifier>
<mods:identifier id="2F1323AE60D55760943D7F6C8BF6FCE3" type="ZooBank">951098C7-18EF-4C9B-AC19-1AEAE2E64ECD</mods:identifier>
</mods:mods>
<treatment id="03C487CEF67D2C6A8AD63F43C36DFDCB" ID-DOI="http://doi.org/10.5281/zenodo.5986228" ID-GBIF-Taxon="143536579" ID-Zenodo-Dep="5986228" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB" httpUri="http://treatment.plazi.org/id/03C487CEF67D2C6A8AD63F43C36DFDCB" lastPageId="6" lastPageNumber="107" pageId="1" pageNumber="102">
<subSubSection id="C3776553F67D2C6D8AD63F43C319FA18" pageId="1" pageNumber="102" type="nomenclature">
<paragraph id="8BD236D8F67D2C6D8AD63F43C387FAFC" blockId="1.[151,509,1359,1421]" box="[151,509,1359,1384]" pageId="1" pageNumber="102">
<heading id="D09A81B4F67D2C6D8AD63F43C387FAFC" bold="true" box="[151,509,1359,1384]" fontSize="10" level="2" pageId="1" pageNumber="102" reason="2">
<emphasis id="B919EACAF67D2C6D8AD63F43C387FAFC" bold="true" box="[151,509,1359,1384]" pageId="1" pageNumber="102">
<taxonomicName id="4C6D4D5BF67D2C6D8AD63F43C3E7FAF3" ID-CoL="4H7DT" authority="Orrico &amp; Dias &amp; Marciano-Jr, 2018" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[151,413,1359,1384]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="1" pageNumber="102" phylum="Chordata" rank="species" species="praeceptor" status="sp. nov.">
<emphasis id="B919EACAF67D2C6D8AD63F43C3E7FAF3" bold="true" box="[151,413,1359,1384]" italics="true" pageId="1" pageNumber="102">Phyllodytes praeceptor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B1F67D2C6D8BE53F43C387FAFC" box="[420,509,1359,1384]" pageId="1" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD236D8F67D2C6D8AD63F78C319FA18" blockId="1.[151,509,1359,1421]" box="[151,355,1396,1421]" pageId="1" pageNumber="102">
(
<figureCitation id="13562A5DF67D2C6D8ADE3F78C29FFA18" box="[159,229,1396,1421]" captionStart="FIGURE 1" captionStartId="3.[151,250,1148,1170]" captionTargetBox="[151,1436,373,1127]" captionTargetId="figure@3.[151,1436,373,1127]" captionTargetPageId="3" captionText="FIGURE 1. Phyllodytes praeceptor sp. nov., holotype (MZUESC 17484; SVL 22.8 mm), dorsal and ventral views." httpUri="https://zenodo.org/record/1216488/files/figure.png" pageId="1" pageNumber="102">Figs 1</figureCitation>
, 
<figureCitation id="13562A5DF67D2C6D8AB23F78C37BFA18" box="[243,257,1396,1420]" captionStart="FIGURE2" captionStartId="4.[151,250,1300,1322]" captionTargetBox="[157,1430,313,1273]" captionTargetId="figure@4.[151,1436,307,1279]" captionTargetPageId="4" captionText="FIGURE2. Phyllodytes praeceptor sp. nov., holotype(MZUESC 17484; SVL 22.8 mm, scalebar = 2 mm).A) Dorsalview of the head; B) Lateral view of the head (left side); C) Left hindlimb (notice the tubercle near the heel); D) Plantar view of the left hand." httpUri="https://zenodo.org/record/1216490/files/figure.png" pageId="1" pageNumber="102">2</figureCitation>
and 
<figureCitation id="13562A5DF67D2C6D8B793F78C321FA18" box="[312,347,1396,1420]" captionStart="FIGURE 3" captionStartId="4.[151,250,1987,2009]" captionTargetBox="[157,1429,1443,1949]" captionTargetId="figure@4.[151,1435,1437,1955]" captionTargetPageId="4" captionText="FIGURE 3. Phyllodytes praeceptor sp. nov. in life.A) holotype (MZUESC 17484) and B) specimen not collected." httpUri="https://zenodo.org/record/1216492/files/figure.png" pageId="1" pageNumber="102">3A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776553F67D2C6D8AD63FB2C069FA61" pageId="1" pageNumber="102" type="reference_group">
<paragraph id="8BD236D8F67D2C6D8AD63FB2C069FA61" blockId="1.[151,793,1468,1525]" pageId="1" pageNumber="102">
<taxonomicName id="4C6D4D5BF67D2C6D8AD63FB2C377FA47" ID-CoL="6PN4" authorityName="Wagler" authorityYear="1830" box="[151,269,1470,1491]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="1" pageNumber="102" phylum="Chordata" rank="genus">
<emphasis id="B919EACAF67D2C6D8AD63FB2C377FA47" box="[151,269,1470,1491]" italics="true" pageId="1" pageNumber="102">Phyllodytes</emphasis>
</taxonomicName>
sp.— 
<bibRefCitation id="EFFC4B29F67D2C6D8B063FB1C043FA47" box="[327,569,1468,1492]" pageId="1" pageNumber="102" refId="ref5560" refString="Faivovich, J., Haddad, C. F. B., Garcia, P. C. A., Frost, D. R., Campbell, J. A. &amp; Wheeler, W. C. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1 - 240. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 294 [0001: SROTFF] 2.0. CO; 2" type="journal article">
Faivovich 
<emphasis id="B919EACAF67D2C6D8BF43FB1C397FA40" box="[437,493,1468,1492]" italics="true" pageId="1" pageNumber="102">et al.</emphasis>
(2005)
</bibRefCitation>
; 
<bibRefCitation id="EFFC4B29F67D2C6D88053FB2C16FFA47" box="[580,789,1468,1492]" pageId="1" pageNumber="102" refId="ref5970" refString="Jowers, M. J., Downie, J. R. &amp; Cohen, B. L. (2008) The golden tree frog of Trinidad, Phyllodytes auratus (Anura: Hylidae): systematic and conservation status. Studies on Neotropical Fauna and Environment, 43, 181 - 188. https: // doi. org / 10.1080 / 01650520801965490" type="journal article">
Jowers 
<emphasis id="B919EACAF67D2C6D88D03FB1C0B3FA40" box="[657,713,1468,1492]" italics="true" pageId="1" pageNumber="102">et al.</emphasis>
(2008)
</bibRefCitation>
. 
<taxonomicName id="4C6D4D5BF67D2C6D8AD63FD3C377FA60" ID-CoL="6PN4" authorityName="Wagler" authorityYear="1830" box="[151,269,1503,1524]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="1" pageNumber="102" phylum="Chordata" rank="genus">
<emphasis id="B919EACAF67D2C6D8AD63FD3C377FA60" box="[151,269,1503,1524]" italics="true" pageId="1" pageNumber="102">Phyllodytes</emphasis>
</taxonomicName>
sp. 4—Dias 
<emphasis id="B919EACAF67D2C6D8BD03FD2C3B3FA61" box="[401,457,1501,1525]" italics="true" pageId="1" pageNumber="102">et al.</emphasis>
(2014)
</paragraph>
</subSubSection>
<subSubSection id="C3776553F67D2C6E8AD63C2EC15CFE61" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102" type="materials_examined">
<paragraph id="8BD236D8F67D2C6D8AD63C2EC066F917" blockId="1.[151,1436,1570,2027]" pageId="1" pageNumber="102">
<materialsCitation id="3B053C85F67D2C6D8AD63C2EC066F917" ID-GBIF-Occurrence="1837804017" collectingDate="2015-03-02" collectionCode="MZUESC" collectorName="da Jiboia &amp; Iuri R. &amp; Carlos Augusto S." country="Brazil" county="Elisio Medrado" elevation="684" latitude="-12.869436" location="Serra da Jiboia" longLatPrecision="1" longitude="-39.461494" municipality="Santa Terezinha" pageId="1" pageNumber="102" specimenCode="MZUESC 17484" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="54D6887AF67D2C6D8AD63C2EC37EF9AF" box="[151,260,1570,1595]" pageId="1" pageNumber="102" type="holotype">
<emphasis id="B919EACAF67D2C6D8AD63C2EC37EF9AF" bold="true" box="[151,260,1570,1595]" pageId="1" pageNumber="102">Holotype</emphasis>
</typeStatus>
. 
<specimenCode id="DBCB9EA3F67D2C6D8B533C2FC3AEF9AF" box="[274,468,1571,1595]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 17484</specimenCode>
, adult male (
<figureCitation id="13562A5DF67D2C6D88253C2EC0DCF9AF" box="[612,678,1570,1595]" captionStart="FIGURE 1" captionStartId="3.[151,250,1148,1170]" captionTargetBox="[151,1436,373,1127]" captionTargetId="figure@3.[151,1436,373,1127]" captionTargetPageId="3" captionText="FIGURE 1. Phyllodytes praeceptor sp. nov., holotype (MZUESC 17484; SVL 22.8 mm), dorsal and ventral views." httpUri="https://zenodo.org/record/1216488/files/figure.png" pageId="1" pageNumber="102">Fig. 1</figureCitation>
), collected at 
<location id="8EB26003F67D2C6D89073C2FC197F9AF" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67D2C6D89073C2FC197F9AF" box="[838,1005,1570,1595]" country="Brazil" county="Elisio Medrado" latitude="-12.869436" longLatPrecision="1" longitude="-39.461494" municipality="Santa Terezinha" name="Serra da Jiboia" pageId="1" pageNumber="102" stateProvince="Bahia">
Serra 
<collectorName id="2698530EF67D2C6D89C83C2EC197F9AF" box="[905,1005,1570,1595]" pageId="1" pageNumber="102">da Jibóia</collectorName>
</location>
(
<geoCoordinate id="EE59501FF67D2C6D89BD3C2EC6E3F9AF" box="[1020,1177,1570,1595]" direction="south" orientation="latitude" pageId="1" pageNumber="102" precision="1" value="-12.869436">12.869436° S</geoCoordinate>
; 
<geoCoordinate id="EE59501FF67D2C6D8EE73C2EC730F9AE" box="[1190,1354,1570,1595]" direction="west" orientation="longitude" pageId="1" pageNumber="102" precision="1" value="-39.461494">39.461494° W</geoCoordinate>
, 
<elevation id="0040D1EBF67D2C6D8F143C2EC7E6F9AE" box="[1365,1436,1570,1595]" metricMagnitude="2" metricUnit="m" metricValue="6.84" pageId="1" pageNumber="102" unit="m" value="684.0">
<quantity id="4C959B3DF67D2C6D8F143C2EC7E6F9AE" box="[1365,1436,1570,1595]" metricMagnitude="2" metricUnit="m" metricValue="6.84" pageId="1" pageNumber="102" unit="m" value="684.0">684 m</quantity>
</elevation>
a.s.l), municipality of 
<collectingMunicipality id="6BB6ACA2F67D2C6D8BD53C4BC033F9CB" box="[404,585,1606,1631]" pageId="1" pageNumber="102">Santa Terezinha</collectingMunicipality>
and 
<collectingCounty id="62B34E54F67D2C6D88C23C4AC14CF9CB" box="[643,822,1606,1631]" pageId="1" pageNumber="102">Elísio Medrado</collectingCounty>
, State of 
<collectingRegion id="49A9F83AF67D2C6D89E23C4AC19DF9CB" box="[931,999,1606,1631]" country="Brazil" name="Bahia" pageId="1" pageNumber="102">Bahia</collectingRegion>
, 
<collectingCountry id="F37A7648F67D2C6D89B53C4AC641F9CB" box="[1012,1083,1606,1631]" name="Brazil" pageId="1" pageNumber="102">Brazil</collectingCountry>
, on 0 
<collectingDate id="EF97E9F0F67D2C6D8E3D3C4BC765F9CB" box="[1148,1311,1606,1631]" pageId="1" pageNumber="102" value="2015-03-02">
<date id="FFD31018F67D2C6D8E3D3C4BC765F9CB" box="[1148,1311,1606,1631]" pageId="1" pageNumber="102" value="2015-03-02">2 March 2015</date>
</collectingDate>
, by 
<collectorName id="2698530EF67D2C6D8F103C4AC7E6F9CA" box="[1361,1436,1606,1631]" pageId="1" pageNumber="102">Iuri R.</collectorName>
Dias and 
<collectorName id="2698530EF67D2C6D8B403C66C3B7F917" box="[257,461,1642,1667]" pageId="1" pageNumber="102">Carlos Augusto S.</collectorName>
Costa.
</materialsCitation>
</paragraph>
<paragraph id="8BD236D8F67D2C6E8A863C82C15CFE61" blockId="1.[151,1436,1570,2027]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102">
<emphasis id="B919EACAF67D2C6D8A863C82C345F933" bold="true" box="[199,319,1678,1703]" pageId="1" pageNumber="102">Paratypes</emphasis>
. Twenty eight specimens: MZUESC 17485, adult male, collected with the holotype; 
<materialsCitation id="3B053C85F67D2C6D8EBE3C83C7E2F95F" ID-GBIF-Occurrence="1837804009" collectingDate="2009-06-08" collectionCode="CFBH" collectorName="Adriano B. &amp; Kury" country="Brazil" location="Bahia" pageId="1" pageNumber="102" specimenCode="CFBH 23671" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67D2C6D8EBE3C83C7E2F933" box="[1279,1432,1678,1703]" collectionCode="CFBH" country="Brazil" httpUri="http://grbio.org/cool/3q7f-dnij" name="Universidade Estadual Paulista" pageId="1" pageNumber="102">CFBH 23671</specimenCode>
, adult male, collected at the Municipality of Una, State of 
<collectingRegion id="49A9F83AF67D2C6D897C3CBEC1FBF95F" box="[829,897,1714,1739]" country="Brazil" name="Bahia" pageId="1" pageNumber="102">Bahia</collectingRegion>
, 
<collectingCountry id="F37A7648F67D2C6D89D13CBEC1ADF95F" box="[912,983,1714,1739]" name="Brazil" pageId="1" pageNumber="102">Brazil</collectingCountry>
, on 
<collectingDate id="EF97E9F0F67D2C6D8E4C3CBFC6E4F95F" box="[1037,1182,1715,1739]" pageId="1" pageNumber="102" value="2009-06-08">
<date id="FFD31018F67D2C6D8E4C3CBFC6E4F95F" box="[1037,1182,1715,1739]" pageId="1" pageNumber="102" value="2009-06-08">8 June 2009</date>
</collectingDate>
, by 
<collectorName id="2698530EF67D2C6D8E923CBEC7E2F95F" box="[1235,1432,1714,1739]" pageId="1" pageNumber="102">Adriano B. Kury</collectorName>
</materialsCitation>
, 
<materialsCitation id="3B053C85F67D2C6D8AD63CDBC763F887" ID-GBIF-Occurrence="1837804016" collectingDate="2008-07" collectionCode="MZUESC" collectorName="Mirco Sole" country="Brazil" county="Orrico" location="Municipality of Ilheus" municipality="Ponta da Tulha" pageId="1" pageNumber="102" specimenCode="MZUESC 7297-7299" specimenCount="1" stateProvince="Amazonas">
<collectingRegion id="49A9F83AF67D2C6D8AD63CDBC377F97B" box="[151,269,1751,1775]" country="Brazil" name="Amazonas" pageId="1" pageNumber="102">Amazonas</collectingRegion>
Chagas Jr., Alessandro Giupponi, Denis R. Pedroso, and Victor G. D. 
<collectingCounty id="62B34E54F67D2C6D8E003CDAC6F4F97B" box="[1089,1166,1750,1775]" pageId="1" pageNumber="102">Orrico</collectingCounty>
; 
<specimenCode id="DBCB9EA3F67D2C6D8EDD3CDBC7E2F97B" box="[1180,1432,1751,1775]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 7297–7299</specimenCode>
, adult males, collected at “ 
<collectingMunicipality id="6BB6ACA2F67D2C6D8B8C3CF7C0FDF887" box="[461,647,1786,1811]" pageId="1" pageNumber="102">Ponta da Tulha</collectingMunicipality>
”, 
<location id="8EB26003F67D2C6D88E23CF6C1D1F887" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67D2C6D88E23CF6C1D1F887" box="[675,939,1786,1811]" country="Brazil" county="Orrico" municipality="Ponta da Tulha" name="Municipality of Ilheus" pageId="1" pageNumber="102" stateProvince="Amazonas">Municipality of Ilhéus</location>
, on 
<collectingDate id="EF97E9F0F67D2C6D89A53CF6C620F887" box="[996,1114,1786,1811]" pageId="1" pageNumber="102" value="2008-07">
<date id="FFD31018F67D2C6D89A53CF6C620F887" box="[996,1114,1786,1811]" pageId="1" pageNumber="102" value="2008-07">July 2008</date>
</collectingDate>
, by 
<collectorName id="2698530EF67D2C6D8ED23CF6C763F887" box="[1171,1305,1786,1811]" pageId="1" pageNumber="102">Mirco Solé</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B053C85F67D2C6D8F6B3CF7C643F8CF" ID-GBIF-Occurrence="1837804019" collectingDate="2010-11-29" collectionCode="MZUESC" collectorName="Tadeu Medeiros" country="Brazil" location="Fazenda Capitao" pageId="1" pageNumber="102" specimenCode="MZUESC 9169-9170, MZUESC 9175" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67D2C6D8F6B3CF7C36FF8A3" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 9169–9170</specimenCode>
and 
<specimenCode id="DBCB9EA3F67D2C6D8B223D13C05BF8A3" box="[355,545,1822,1847]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 9175</specimenCode>
, adult males and female, respectively, collected at “ 
<location id="8EB26003F67D2C6D8EFE3D12C7F4F8A3" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67D2C6D8EFE3D12C7F4F8A3" box="[1215,1422,1822,1847]" country="Brazil" name="Fazenda Capitao" pageId="1" pageNumber="102" stateProvince="Bahia">Fazenda Capitão</location>
”, Municipality of Itacaré, State of 
<collectingRegion id="49A9F83AF67D2C6D88453D4EC032F8CF" box="[516,584,1858,1883]" country="Brazil" name="Bahia" pageId="1" pageNumber="102">Bahia</collectingRegion>
, on 
<collectingDate id="EF97E9F0F67D2C6D88363D4FC128F8CF" box="[631,850,1858,1883]" pageId="1" pageNumber="102" value="2010-11-29">
<date id="FFD31018F67D2C6D88363D4FC128F8CF" box="[631,850,1858,1883]" pageId="1" pageNumber="102" value="2010-11-29">29 November 2010</date>
</collectingDate>
, by 
<collectorName id="2698530EF67D2C6D89C13D4EC643F8CF" box="[896,1081,1858,1883]" pageId="1" pageNumber="102">Tadeu Medeiros</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B053C85F67D2C6D8E023D4FC318F837" ID-GBIF-Occurrence="1837804010" collectingDate="2011-08-21" collectionCode="MZUESC" country="Brazil" location="Municipality of Ilheus" municipality="Ponta da Tulha" pageId="1" pageNumber="102" specimenCode="MZUESC 10498-10499" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67D2C6D8E023D4FC723F8CF" box="[1091,1369,1859,1883]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 10498–10499</specimenCode>
, adult males, collected at “ 
<collectingMunicipality id="6BB6ACA2F67D2C6D8B363D6BC05CF8EB" box="[375,550,1894,1919]" pageId="1" pageNumber="102">Ponta da Tulha</collectingMunicipality>
”, 
<location id="8EB26003F67D2C6D887D3D6AC143F8EB" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67D2C6D887D3D6AC143F8EB" box="[572,825,1894,1919]" country="Brazil" municipality="Ponta da Tulha" name="Municipality of Ilheus" pageId="1" pageNumber="102" stateProvince="Bahia">Municipality of Ilhéus</location>
, on 
<collectingDate id="EF97E9F0F67D2C6D89263D6BC662F8EB" box="[871,1048,1895,1919]" pageId="1" pageNumber="102" value="2011-08-21">
<date id="FFD31018F67D2C6D89263D6BC662F8EB" box="[871,1048,1895,1919]" pageId="1" pageNumber="102" value="2011-08-21">21 August 2011</date>
</collectingDate>
by Mirco Solé, Euvaldo Marciano Jr and Iuri Dias
</materialsCitation>
; 
<materialsCitation id="3B053C85F67D2C6D8B373D87C08FF852" ID-GBIF-Occurrence="1837804011" collectingDate="2012-06" collectionCode="MZUESC" collectorName="Euvaldo Marciano Jr" country="Brazil" location="Municipality of Maragogipe" municipality="Sao Roque do Paraguacu" pageId="1" pageNumber="102" specimenCode="MZUESC 10626" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67D2C6D8B373D87C03AF837" box="[374,576,1930,1955]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 10626</specimenCode>
, adult male, collected at “ 
<collectingMunicipality id="6BB6ACA2F67D2C6D89CD3D87C6BDF837" box="[908,1223,1930,1955]" pageId="1" pageNumber="102">São Roque do Paraguaçu</collectingMunicipality>
”, 
<location id="8EB26003F67D2C6D8EA53D86C35BF853" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67D2C6D8EA53D86C35BF853" country="Brazil" municipality="Sao Roque do Paraguacu" name="Municipality of Maragogipe" pageId="1" pageNumber="102" stateProvince="Bahia">Municipality of Maragogipe</location>
, on 
<collectingDate id="EF97E9F0F67D2C6D8B143DA3C3B7F853" box="[341,461,1967,1991]" pageId="1" pageNumber="102" value="2012-06">
<date id="FFD31018F67D2C6D8B143DA3C3B7F853" box="[341,461,1967,1991]" pageId="1" pageNumber="102" value="2012-06">June 2012</date>
</collectingDate>
, by 
<collectorName id="2698530EF67D2C6D88403DA2C08FF852" box="[513,757,1966,1991]" pageId="1" pageNumber="102">Euvaldo Marciano Jr</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B053C85F67D2C6D89453DA3C600F87F" ID-GBIF-Occurrence="1837804012" collectingDate="2011-02-21" collectionCode="MZUESC" collectorName="Iuri R. &amp; Dias &amp; Mirco Sole" country="Brazil" location="Municipality of Ilheus" municipality="Lagoa Encantada" pageId="1" pageNumber="102" specimenCode="MZUESC 11584-11585" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67D2C6D89453DA3C660F853" box="[772,1050,1966,1991]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 11584–11585</specimenCode>
, adult males, collected at “ 
<collectingMunicipality id="6BB6ACA2F67D2C6D8F153DA3C36BF87F" pageId="1" pageNumber="102">Lagoa Encantada</collectingMunicipality>
”, 
<location id="8EB26003F67D2C6D8B643DDEC05BF87F" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67D2C6D8B643DDEC05BF87F" box="[293,545,2002,2027]" country="Brazil" municipality="Lagoa Encantada" name="Municipality of Ilheus" pageId="1" pageNumber="102" stateProvince="Bahia">Municipality of Ilhéus</location>
, on 
<collectingDate id="EF97E9F0F67D2C6D880F3DDFC16FF87F" box="[590,789,2002,2027]" pageId="1" pageNumber="102" value="2011-02-21">
<date id="FFD31018F67D2C6D880F3DDFC16FF87F" box="[590,789,2002,2027]" pageId="1" pageNumber="102" value="2011-02-21">21 February 2011</date>
</collectingDate>
, by 
<collectorName id="2698530EF67D2C6D89023DDEC1BCF87F" box="[835,966,2002,2027]" pageId="1" pageNumber="102">Iuri R. Dias</collectorName>
and 
<collectorName id="2698530EF67D2C6D89BB3DDEC600F87F" box="[1018,1146,2002,2027]" pageId="1" pageNumber="102">Mirco Solé</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B053C85F67D2C6E8EC43DDFC0D2FF41" ID-GBIF-Occurrence="1837804008" collectingDate="2012-02-18" collectionCode="MZUESC" country="Brazil" lastPageId="2" lastPageNumber="103" location="Bahia" pageId="1" pageNumber="102" specimenCode="MZUESC 11610-11612" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67D2C6D8EC43DDFC7E2F87F" box="[1157,1432,2002,2027]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="1" pageNumber="102">MZUESC 11610–11612</specimenCode>
, adult males, collected at the Municipality of Una, State of 
<collectingRegion id="49A9F83AF67E2C6E89743A9BC103FF24" box="[821,889,151,176]" country="Brazil" name="Bahia" pageId="2" pageNumber="103">Bahia</collectingRegion>
, on 
<date id="FFD31018F67E2C6E89EB3A94C60DFF24" box="[938,1143,151,176]" pageId="2" pageNumber="103" value="2012-02-18">
<collectingDate id="EF97E9F0F67E2C6E89EB3A94C60DFF24" box="[938,1143,151,176]" pageId="2" pageNumber="103" value="2012-02-18">18 February 2012</collectingDate>
</date>
, by Euvaldo Marciano Jr, Paulo Henrique Ribeiro and Leandro Oliveira
</materialsCitation>
; 
<materialsCitation id="3B053C85F67E2C6E88F73AB0C6BCFF63" ID-GBIF-Occurrence="1837804015" collectingDate="2012-08-11" collectionCode="MZUESC" collectorName="Euvaldo Marciano Jr" country="Brazil" location="Municipality of Igrapiuna" municipality="Reserva Michelin" pageId="2" pageNumber="103" specimenCode="MZUESC 13016, MZUESC 13026" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67E2C6E88F73AB0C10DFF41" box="[694,887,188,213]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 13016</specimenCode>
and 
<specimenCode id="DBCB9EA3F67E2C6E89F23AB0C602FF41" box="[947,1144,188,213]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 13026</specimenCode>
, adult males, collected at “ 
<collectingMunicipality id="6BB6ACA2F67E2C6E8AE03AECC30BFF6C" box="[161,369,223,248]" pageId="2" pageNumber="103">Reserva Michelin</collectingMunicipality>
”, 
<location id="8EB26003F67E2C6E8BC83AD3C0C8FF6C" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67E2C6E8BC83AD3C0C8FF6C" box="[393,690,223,248]" country="Brazil" municipality="Reserva Michelin" name="Municipality of Igrapiuna" pageId="2" pageNumber="103" stateProvince="Bahia">Municipality of Igrapiúna</location>
, on 
<collectingDate id="EF97E9F0F67E2C6E88A73AECC1DAFF6C" box="[742,928,224,248]" pageId="2" pageNumber="103" value="2012-08-11">
<date id="FFD31018F67E2C6E88A73AECC1DAFF6C" box="[742,928,224,248]" pageId="2" pageNumber="103" value="2012-08-11">11 August 2012</date>
</collectingDate>
, by 
<collectorName id="2698530EF67E2C6E89923AD3C6BCFF63" box="[979,1222,223,248]" pageId="2" pageNumber="103">Euvaldo Marciano Jr</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B053C85F67E2C6E8E953AECC384FEAB" ID-GBIF-Occurrence="1837804013" collectingDate="2012-08-13" collectionCode="MZUESC" collectorName="Euvaldo Marciano Jr" country="Brazil" location="Sao Roque do Paraguacu" pageId="2" pageNumber="103" specimenCode="MZUESC 13027" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67E2C6E8E953AECC7EDFF6C" box="[1236,1431,224,248]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 13027</specimenCode>
, adult male, collected at “ 
<location id="8EB26003F67E2C6E8BF63B08C09AFE88" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67E2C6E8BF63B08C09AFE88" box="[439,736,260,285]" country="Brazil" name="Sao Roque do Paraguacu" pageId="2" pageNumber="103" stateProvince="Bahia">São Roque do Paraguaçu</location>
”, Municipality of Maragogipe, State of 
<collectingRegion id="49A9F83AF67E2C6E8EEF3B08C688FE89" box="[1198,1266,260,285]" country="Brazil" name="Bahia" pageId="2" pageNumber="103">Bahia</collectingRegion>
, on 
<collectingDate id="EF97E9F0F67E2C6E8F643B08C2A8FED4" pageId="2" pageNumber="103" value="2012-08-13">
<date id="FFD31018F67E2C6E8F643B08C2A8FED4" pageId="2" pageNumber="103" value="2012-08-13">13 August 2012</date>
</collectingDate>
, by 
<collectorName id="2698530EF67E2C6E8B463B2BC384FEAB" box="[263,510,295,320]" pageId="2" pageNumber="103">Euvaldo Marciano Jr</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B053C85F67E2C6E884C3B24C7E2FEF1" ID-GBIF-Occurrence="1837804018" collectingDate="2015-02-26" collectionCode="MZUESC" country="Brazil" location="Municipality of Ilheus" municipality="Acuipe" pageId="2" pageNumber="103" specimenCode="MZUESC 14199-14200, MZUESC 14207, MZUESC 14212" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67E2C6E884C3B24C15CFED4" box="[525,806,296,320]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 14199–14200</specimenCode>
, 
<specimenCode id="DBCB9EA3F67E2C6E89743B24C180FED4" box="[821,1018,296,320]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 14207</specimenCode>
, and 
<specimenCode id="DBCB9EA3F67E2C6E8E7C3B24C778FED4" box="[1085,1282,296,320]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 14212</specimenCode>
, adult males, collected at “ 
<collectingMunicipality id="6BB6ACA2F67E2C6E8B713B40C3F2FEF0" box="[304,392,332,356]" pageId="2" pageNumber="103">Acuípe</collectingMunicipality>
”, 
<location id="8EB26003F67E2C6E8BE03B40C0DCFEF1" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67E2C6E8BE03B40C0DCFEF1" box="[417,678,332,357]" country="Brazil" municipality="Acuipe" name="Municipality of Ilheus" pageId="2" pageNumber="103" stateProvince="Bahia">Municipality of Ilhéus</location>
, on 
<collectingDate id="EF97E9F0F67E2C6E889D3B40C1D7FEF1" box="[732,941,332,357]" pageId="2" pageNumber="103" value="2015-02-26">
<date id="FFD31018F67E2C6E889D3B40C1D7FEF1" box="[732,941,332,357]" pageId="2" pageNumber="103" value="2015-02-26">26 February 2015</date>
</collectingDate>
, by Euvaldo Marciano Jr and Mirco Solé
</materialsCitation>
; 
<materialsCitation id="3B053C85F67E2C6E8AD63B7CC035FE39" ID-GBIF-Occurrence="1837804007" collectingDate="2013-06-06" collectionCode="MZUESC" country="Brazil" location="Bahia" pageId="2" pageNumber="103" specimenCode="MZUESC 17481-17483" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67E2C6E8AD63B7CC3D4FE1C" box="[151,430,368,392]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 17481–17483</specimenCode>
, adult males, collected at the Municipality of Una, State of 
<collectingRegion id="49A9F83AF67E2C6E8E213B63C6DEFE1C" box="[1120,1188,367,392]" country="Brazil" name="Bahia" pageId="2" pageNumber="103">Bahia</collectingRegion>
, on 0 
<collectingDate id="EF97E9F0F67E2C6E8EA43B63C709FE1C" box="[1253,1395,367,392]" pageId="2" pageNumber="103" value="2013-06-06">
<date id="FFD31018F67E2C6E8EA43B63C709FE1C" box="[1253,1395,367,392]" pageId="2" pageNumber="103" value="2013-06-06">6 June 2013</date>
</collectingDate>
, by Euvaldo Marciano Jr and Mirco Solé
</materialsCitation>
; 
<materialsCitation id="3B053C85F67E2C6E88213B98C15CFE61" ID-GBIF-Occurrence="1837804014" collectingDate="2015-03-23" collectingDateMax="2015-03-31" collectingDateMin="2015-03-23" collectionCode="MZUESC" collectorName="Iuri R." country="Brazil" location="Municipality of Igrapiuna" municipality="Reserva Michelin" pageId="2" pageNumber="103" specimenCode="MZUESC 17486-17487" specimenCount="1" stateProvince="Bahia">
<specimenCode id="DBCB9EA3F67E2C6E88213B98C100FE39" box="[608,890,404,429]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="103">MZUESC 17486–17487</specimenCode>
, adult males, collected at “ 
<collectingMunicipality id="6BB6ACA2F67E2C6E8EFA3B98C7F7FE39" box="[1211,1421,404,429]" pageId="2" pageNumber="103">Reserva Michelin</collectingMunicipality>
”, 
<location id="8EB26003F67E2C6E8AD63BBBC3BEFE44" LSID="urn:lsid:plazi:treatment:03C487CEF67D2C6A8AD63F43C36DFDCB:8EB26003F67E2C6E8AD63BBBC3BEFE44" box="[151,452,439,464]" country="Brazil" municipality="Reserva Michelin" name="Municipality of Igrapiuna" pageId="2" pageNumber="103" stateProvince="Bahia">Municipality of Igrapiúna</location>
, between 
<collectingDate id="EF97E9F0F67E2C6E887C3BB4C164FE44" box="[573,798,439,464]" pageId="2" pageNumber="103" value="2015-03-23" valueMax="2015-03-31" valueMin="2015-03-23">
<date id="FFD31018F67E2C6E887C3BB4C164FE44" box="[573,798,439,464]" pageId="2" pageNumber="103" value="2015-03-23" valueMax="2015-03-31" valueMin="2015-03-23">23–31 March 2015</date>
</collectingDate>
, by 
<collectorName id="2698530EF67E2C6E89143BBBC1D8FE5B" box="[853,930,439,464]" pageId="2" pageNumber="103">Iuri R.</collectorName>
Dias, Euvaldo Marciano Jr, Caio V. Mira- Mendes, Francisco F.R. Oliveira and Leandro Drummond.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3776553F67E2C6E8A863BF3C707FDF4" pageId="2" pageNumber="103" type="description">
<paragraph id="8BD236D8F67E2C6E8A863BF3C6DFFD8C" blockId="2.[151,1437,151,2013]" box="[199,1189,511,536]" pageId="2" pageNumber="103">
<emphasis id="B919EACAF67E2C6E8A863BF3C3DDFD8C" bold="true" box="[199,423,511,536]" pageId="2" pageNumber="103">Referred specimen</emphasis>
. MRT 6144, not examined, collected at “Brazil: Bahia, Uruci-Una”.
</paragraph>
<paragraph id="8BD236D8F67E2C6E8A863828C707FDF4" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="103">
<emphasis id="B919EACAF67E2C6E8A863828C3CCFDA9" bold="true" box="[199,438,548,573]" pageId="2" pageNumber="103">Generic placement.</emphasis>
A member of 
<taxonomicName id="4C6D4D5BF67E2C6E88343828C082FDA8" authorityName="Wagler" authorityYear="1830" box="[629,760,548,572]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="genus">
<emphasis id="B919EACAF67E2C6E88343828C082FDA8" box="[629,760,548,572]" italics="true" pageId="2" pageNumber="103">Phyllodytes</emphasis>
</taxonomicName>
, by possessing fang-like odontoids; molecular data also supports the placement of 
<taxonomicName id="4C6D4D5BF67E2C6E8BFF3845C02EFDF4" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[446,596,585,608]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF67E2C6E8BFF3845C02EFDF4" box="[446,596,585,608]" italics="true" pageId="2" pageNumber="103">P. praeceptor</emphasis>
</taxonomicName>
as sister taxon of 
<taxonomicName id="4C6D4D5BF67E2C6E895E3845C1EEFDF4" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[799,916,584,608]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="luteolus">
<emphasis id="B919EACAF67E2C6E895E3845C1EEFDF4" box="[799,916,584,608]" italics="true" pageId="2" pageNumber="103">P. luteolus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFC4B29F67E2C6E89E3384BC6ECFDF4" box="[930,1174,583,608]" pageId="2" pageNumber="103" refId="ref5560" refString="Faivovich, J., Haddad, C. F. B., Garcia, P. C. A., Frost, D. R., Campbell, J. A. &amp; Wheeler, W. C. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1 - 240. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 294 [0001: SROTFF] 2.0. CO; 2" type="journal article">
Faivovich 
<emphasis id="B919EACAF67E2C6E8E5A3845C629FDF4" box="[1051,1107,584,608]" italics="true" pageId="2" pageNumber="103">et al.</emphasis>
2005
</bibRefCitation>
; 
<bibRefCitation id="EFFC4B29F67E2C6E8EE03844C70BFDF4" box="[1185,1393,584,608]" pageId="2" pageNumber="103" refId="ref5970" refString="Jowers, M. J., Downie, J. R. &amp; Cohen, B. L. (2008) The golden tree frog of Trinidad, Phyllodytes auratus (Anura: Hylidae): systematic and conservation status. Studies on Neotropical Fauna and Environment, 43, 181 - 188. https: // doi. org / 10.1080 / 01650520801965490" type="journal article">
Jowers 
<emphasis id="B919EACAF67E2C6E8EB43845C754FDF4" box="[1269,1326,584,608]" italics="true" pageId="2" pageNumber="103">et al.</emphasis>
2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3776553F67E2C6E8A863860C769F9E1" pageId="2" pageNumber="103" type="diagnosis">
<paragraph id="8BD236D8F67E2C6E8A863860C74DFC30" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="103">
<emphasis id="B919EACAF67E2C6E8A863860C340FD11" bold="true" box="[199,314,620,645]" pageId="2" pageNumber="103">Diagnosis</emphasis>
. A medium-sized species (SVL 
<quantity id="4C959B3DF67E2C6E88FB3860C11BFD10" box="[698,865,620,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" metricValueMax="2.58" metricValueMin="2.07" pageId="2" pageNumber="103" unit="mm" value="23.25" valueMax="25.8" valueMin="20.7">20.7–25.8 mm</quantity>
in males) characterized by: (1) dorsum of body, arms, and legs cream with scattered dark brown spots; (2) a dark brown stripe extending from the eyes; (3) snout short, semicircular (to truncated) in dorsal view, vertical (to truncated) in profile; (4) vomerine teeth in two dense patches, disposed in a transverse plane, behind the choanae; (5) one anterior large, and additional smaller subequal odontoids on each side of the mandible; (6) dorsal skin smooth; (7) vocal sac externally visible; (8) venter areolate with two paramedial rows of developed tubercles; (9) small tubercles in the ventral side of the thighs, the most medial more developed; (10) a tubercle at the tibio-tarsal articulation; (11) unpigmented nuptial pad visible, rounded, moderately expanded, (12) advertisement call consisting of a series of 6–12 pulsed notes, average duration of 
<geoCoordinate id="EE59501FF67E2C6E8B583980C326FC30" box="[281,348,908,933]" direction="south" orientation="latitude" pageId="2" pageNumber="103" precision="555" value="-5.34">5.34 s</geoCoordinate>
with notes lasting 
<geoCoordinate id="EE59501FF67E2C6E88713980C0FBFC30" box="[560,641,908,933]" direction="south" orientation="latitude" pageId="2" pageNumber="103" precision="55" value="-0.295">0.295 s</geoCoordinate>
and dominant frequency ranging between 2928.5–3273.0 Hz.
</paragraph>
<paragraph id="8BD236D8F67E2C6E8A8639A3C769F9E1" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="103">
<emphasis id="B919EACAF67E2C6E8A8639A3C032FC5C" bold="true" box="[199,584,943,968]" pageId="2" pageNumber="103">Comparison with other species.</emphasis>
Character states of 
<taxonomicName id="4C6D4D5BF67E2C6E897439BCC641FC5C" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[821,1083,944,968]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF67E2C6E897439BCC641FC5C" box="[821,1083,944,968]" italics="true" pageId="2" pageNumber="103">Phyllodytes praeceptor</emphasis>
</taxonomicName>
in parentheses. All species of 
<taxonomicName id="4C6D4D5BF67E2C6E8AD639D8C362FC78" authorityName="Wagler" authorityYear="1830" box="[151,280,980,1004]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="genus">
<emphasis id="B919EACAF67E2C6E8AD639D8C362FC78" box="[151,280,980,1004]" italics="true" pageId="2" pageNumber="103">Phyllodytes</emphasis>
</taxonomicName>
have multiple tubercles on the tarsus in various arrangements (a single large tubercle on posteroventral internal margin of tarsus, near to the junction with shank) and, with the exception of 
<emphasis id="B919EACAF67E2C6E8E0539F5C689FB84" box="[1092,1267,1016,1040]" italics="true" pageId="2" pageNumber="103">
<taxonomicName id="4C6D4D5BF67E2C6E8E0539F5C695FB84" authorityName="Bokermann" authorityYear="1966" box="[1092,1263,1016,1040]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="tuberculosus">P. tuberculosus</taxonomicName>
,
</emphasis>
different snout shapes in dorsal view (semicircular to truncated). Below we list other diagnostic morphological characters. Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E8B763E4DC3DDFBCC" box="[311,423,1088,1112]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B919EACAF67E2C6E8B763E4DC3DDFBCC" box="[311,423,1088,1112]" italics="true" pageId="2" pageNumber="103">P. amadoi</emphasis>
</taxonomicName>
have snout with a small apical tubercle (absent) and two large anterior odontoids on both sides of the mandible (one larger odontoid). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E890E3E69C195FBE8" authorityName="Bokermann" authorityYear="1966" box="[847,1007,1125,1148]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B919EACAF67E2C6E890E3E69C195FBE8" box="[847,1007,1125,1148]" italics="true" pageId="2" pageNumber="103">P. acuminatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6D4D5BF67E2C6E8E713E69C685FBE8" authorityName="" authorityYear="1830" box="[1072,1279,1125,1148]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B919EACAF67E2C6E8E713E69C685FBE8" box="[1072,1279,1125,1148]" italics="true" pageId="2" pageNumber="103">P. megatympanum</emphasis>
</taxonomicName>
have a large tympanum, TD&gt; 7 % of SVL, (small, TD ~ 6 % of SVL). Specimens of 
<emphasis id="B919EACAF67E2C6E89953E85C295FB50" italics="true" pageId="2" pageNumber="103">
<taxonomicName id="4C6D4D5BF67E2C6E89953E85C608FB34" box="[980,1138,1160,1184]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="brevirostris">P. brevirostris</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF67E2C6E8E3F3E85C68FFB34" box="[1150,1269,1160,1184]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="edelmoi">P. edelmoi</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF67E2C6E8F433E85C701FB34" box="[1282,1403,1160,1184]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="kautskyi">P. kautskyi</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF67E2C6E8FC93E85C295FB50" baseAuthorityName="Wied" baseAuthorityYear="1824" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="luteolus">P. luteolus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C6D4D5BF67E2C6E8B6E3EA1C384FB50" authorityName="" authorityYear="1830" box="[303,510,1197,1220]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B919EACAF67E2C6E8B6E3EA1C384FB50" box="[303,510,1197,1220]" italics="true" pageId="2" pageNumber="103">P. megatympanum</emphasis>
</taxonomicName>
have an immaculate dorsum (spots and blotches on dorsum). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E8FC93EA1C362FB7C" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B919EACAF67E2C6E8FC93EA1C362FB7C" italics="true" pageId="2" pageNumber="103">P. gyrinaethes</emphasis>
</taxonomicName>
have a hidden tympanum (distinct) and a mottled dorsal pattern (spots and blotches on dorsum). Male specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E8B773EF9C3D4FA98" box="[310,430,1268,1292]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B919EACAF67E2C6E8B773EF9C3D4FA98" box="[310,430,1268,1292]" italics="true" pageId="2" pageNumber="103">P. kautskyi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6D4D5BF67E2C6E8BAD3EF9C0FAFA98" box="[492,640,1268,1292]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B919EACAF67E2C6E8BAD3EF9C380FA98" box="[492,506,1269,1292]" italics="true" pageId="2" pageNumber="103">P</emphasis>
. 
<emphasis id="B919EACAF67E2C6E884A3EF8C0FAFA98" box="[523,640,1268,1292]" italics="true" pageId="2" pageNumber="103">maculosus</emphasis>
</taxonomicName>
are larger, combined SVL = 38.0– 
<quantity id="4C959B3DF67E2C6E8E543EF8C601FA98" box="[1045,1147,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" pageId="2" pageNumber="103" unit="mm" value="43.5">43.5 mm</quantity>
(SVL = 
<quantity id="4C959B3DF67E2C6E8EA63EF8C7EBFA98" box="[1255,1425,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" metricValueMax="2.58" metricValueMin="2.07" pageId="2" pageNumber="103" unit="mm" value="23.25" valueMax="25.8" valueMin="20.7">20.7–25.8 mm</quantity>
). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E8B723F15C3DFFAA4" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[307,421,1304,1328]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="luteolus">
<emphasis id="B919EACAF67E2C6E8B723F15C3DFFAA4" box="[307,421,1304,1328]" italics="true" pageId="2" pageNumber="103">P. luteolus</emphasis>
</taxonomicName>
have a snout mucronated in dorsal view (semicircular to truncated) and acute in lateral view (vertical to truncated), rows of tubercles in the forearm (inconspicuous), venter with various rows of developed tubercles (only the two paramedial are well-developed). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E89F83F6DC636FAEC" box="[953,1100,1376,1400]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B919EACAF67E2C6E89F83F6DC636FAEC" box="[953,1100,1376,1400]" italics="true" pageId="2" pageNumber="103">P. maculosus</emphasis>
</taxonomicName>
have a deeply mottled dorsal pattern (spots and blotches on dorsum). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E89653F89C18FFA08" authorityName="" authorityYear="1830" box="[804,1013,1413,1436]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B919EACAF67E2C6E89653F89C18FFA08" box="[804,1013,1413,1436]" italics="true" pageId="2" pageNumber="103">P. megatympanum</emphasis>
</taxonomicName>
have a bright yellow groin in live specimens (no conspicuous coloration). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E88B43FA5C1CAFA54" box="[757,944,1448,1472]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B919EACAF67E2C6E88B43FA5C1CAFA54" box="[757,944,1448,1472]" italics="true" pageId="2" pageNumber="103">P. melanomystax</emphasis>
</taxonomicName>
have a conspicuous large dark brown stripe on the snout and 
<emphasis id="B919EACAF67E2C6E8B253FC0C05BFA70" box="[356,545,1484,1508]" italics="true" pageId="2" pageNumber="103">canthus rostralis</emphasis>
(absent or thin faded stripe in 
<taxonomicName id="4C6D4D5BF67E2C6E89D03FC1C653FA70" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[913,1065,1485,1508]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF67E2C6E89D03FC1C653FA70" box="[913,1065,1485,1508]" italics="true" pageId="2" pageNumber="103">P. praeceptor</emphasis>
</taxonomicName>
) and lack organized paramedial ventral rows of enlarged tubercles (present). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E89713FFDC1C3F99C" box="[816,953,1521,1544]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B919EACAF67E2C6E89713FFDC1C3F99C" box="[816,953,1521,1544]" italics="true" pageId="2" pageNumber="103">P. punctatus</emphasis>
</taxonomicName>
lack organized paramedial ventral rows of enlarged tubercles (present). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E88CC3C19C140F9B8" authorityName="Bokermann" authorityYear="1966" box="[653,826,1556,1580]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B919EACAF67E2C6E88CC3C19C140F9B8" box="[653,826,1556,1580]" italics="true" pageId="2" pageNumber="103">P. tuberculosus</emphasis>
</taxonomicName>
lack organized paramedial ventral rows of enlarged tubercles (present) and have supranumerary tubercles in the plantar surface (absent). Specimens of 
<taxonomicName id="4C6D4D5BF67E2C6E8E993C35C71BF9C4" baseAuthorityName="Peters" baseAuthorityYear="1873" box="[1240,1377,1592,1616]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="103" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B919EACAF67E2C6E8E993C35C71BF9C4" box="[1240,1377,1592,1616]" italics="true" pageId="2" pageNumber="103">P. wuchereri</emphasis>
</taxonomicName>
have a dorsal pattern consisting of large black and white stripes and blotches (spots and blotches on dorsum).
</paragraph>
</subSubSection>
<subSubSection id="C3776553F67E2C688A863C73C612FF63" lastPageId="4" lastPageNumber="105" pageId="2" pageNumber="103" type="description">
<paragraph id="8BD236D8F67E2C6F8A863C73C1EAFED4" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103">
<emphasis id="B919EACAF67E2C6E8A863C73C39BF90C" bold="true" box="[199,481,1663,1688]" pageId="2" pageNumber="103">
Description of 
<typeStatus id="54D6887AF67E2C6E8B353C73C3A6F90C" box="[372,476,1663,1688]" pageId="2" pageNumber="103" type="holotype">holotype</typeStatus>
.
</emphasis>
An adult male in good state of preservation, with a piece of muscle removed from the right thigh for molecular analyses. Raw measurements are detailed in 
<tableCitation id="C6EF0363F67E2C6E89E03CA8C18EF928" box="[929,1012,1700,1725]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Range, mean, and standard deviation (SD) of the measurements (mm) of the type specimens of Phyllodytes praeceptor sp. nov. (n = 28adult males and one female)." pageId="2" pageNumber="103">Table 1</tableCitation>
. Head wider than long (HL 89.5 % of HW; HW 37.7% of SVL; HL 33.8 % of SVL); snout truncated in dorsal and lateral views (
<figureCitation id="13562A5DF67E2C6E8EE23CCBC690F974" box="[1187,1258,1735,1760]" captionStart="FIGURE 1" captionStartId="3.[151,250,1148,1170]" captionTargetBox="[151,1436,373,1127]" captionTargetId="figure@3.[151,1436,373,1127]" captionTargetPageId="3" captionText="FIGURE 1. Phyllodytes praeceptor sp. nov., holotype (MZUESC 17484; SVL 22.8 mm), dorsal and ventral views." httpUri="https://zenodo.org/record/1216488/files/figure.png" pageId="2" pageNumber="103">Figs 1</figureCitation>
and 
<figureCitation id="13562A5DF67E2C6E8F623CC4C749F974" box="[1315,1331,1736,1760]" captionStart="FIGURE2" captionStartId="4.[151,250,1300,1322]" captionTargetBox="[157,1430,313,1273]" captionTargetId="figure@4.[151,1436,307,1279]" captionTargetPageId="4" captionText="FIGURE2. Phyllodytes praeceptor sp. nov., holotype(MZUESC 17484; SVL 22.8 mm, scalebar = 2 mm).A) Dorsalview of the head; B) Lateral view of the head (left side); C) Left hindlimb (notice the tubercle near the heel); D) Plantar view of the left hand." httpUri="https://zenodo.org/record/1216490/files/figure.png" pageId="2" pageNumber="103">2</figureCitation>
); nostrils small, rounded, slightly protuberant, situated dorsally and directed anteriorly, nearer to the tip of snout than to eye; internarial distance smaller than eye–nostril distance (IND 65.2 % of END), eyes large (ED 33.8% of HL; 30.2 % of HW; 113 % of END), prominent, situated laterally, directed anterolaterally; interorbital space and dorsum of snout flat; 
<emphasis id="B919EACAF67E2C6E8A8A3D54C3FFF8E4" box="[203,389,1880,1904]" italics="true" pageId="2" pageNumber="103">canthus rostralis</emphasis>
distinct, rounded; loreal region oblique, slightly concave; tympanum evident, relatively large (TD 18.2 % of HL), nearly circular, separated from posterior border of eye by approximately the diameter of tympanum; tympanum diameter smaller than internarial distance (TD 93.3 % of IND, eye to nostril distance (TD 60.9 % of END), eye diameter (TD 53.8 % of ED), and interorbital distance (TD 31.8 % of IOD); its diameter slightly larger than width of discs on third finger (TD/DF3 = 116.7 %) and of fourth toe (TD/4TD = 127.3 %); tympanic annulus evident; supratympanic fold developed, covering dorsal edge of tympanum, nearly straight, slightly curved downwards and extending until the insertion of arm; vocal sac externally developed, subgular, single; vomerine teeth forming a straight line behind small, rounded, widely separated choanae; each side of mandible with one anterior, large, and smaller subequal odontoids; pupil horizontal.
</paragraph>
<caption id="DF126650F67F2C6F8AD63E70C756FB06" box="[151,1324,1148,1170]" httpUri="https://zenodo.org/record/1216488/files/figure.png" pageId="3" pageNumber="104" startId="3.[151,250,1148,1170]" targetBox="[151,1436,373,1127]" targetPageId="3">
<paragraph id="8BD236D8F67F2C6F8AD63E70C756FB06" blockId="3.[151,1324,1148,1170]" box="[151,1324,1148,1170]" pageId="3" pageNumber="104">
<emphasis id="B919EACAF67F2C6F8AD63E70C369FB05" bold="true" box="[151,275,1148,1170]" pageId="3" pageNumber="104">FIGURE 1.</emphasis>
<taxonomicName id="4C6D4D5BF67F2C6F8B583E70C07EFB06" authority="Orrico &amp; Dias &amp; Marciano-Jr, 2018" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[281,516,1148,1170]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="104" phylum="Chordata" rank="species" species="praeceptor" status="sp. nov.">
<emphasis id="B919EACAF67F2C6F8B583E70C07EFB06" box="[281,516,1148,1170]" italics="true" pageId="3" pageNumber="104">Phyllodytes praeceptor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B1F67F2C6F884A3E71C021FB06" box="[523,603,1149,1170]" pageId="3" pageNumber="104" rank="species">
<emphasis id="B919EACAF67F2C6F884A3E71C021FB06" bold="true" box="[523,603,1149,1170]" pageId="3" pageNumber="104">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype (MZUESC 17484; SVL 22.8 mm), dorsal and ventral views.
</paragraph>
</caption>
<paragraph id="8BD236D8F67F2C6F8A863EB6C138FA3F" blockId="3.[151,1436,1210,2029]" pageId="3" pageNumber="104">Forelimbs not hypertrophied, forearms slightly larger (in cross-section) than arms, with four outer tubercles in the forearm near to the hand being the closest to the hand the larger. Hands large (HAL 84.4 % of HL), outer and inner carpal tubercles developed, elliptical; fingers not webbed, small fringe present between fingers III and IV, and IV and V; fingers in crescent order of size, II &lt;V &lt;III &lt;IV; adhesive discs developed (adhesive discs diameter approximately two times finger width), transversally elliptical; disc on finger II, smaller than of other fingers; prepollex indistinct; nuptial pad on the internal base of the finger II, no dark nuptial excrescence; subarticular tubercles large, rounded, single; no supernumerary tubercles.</paragraph>
<paragraph id="8BD236D8F67F2C6F8A863FB9C67DF93C" blockId="3.[151,1436,1210,2029]" pageId="3" pageNumber="104">
Hind limbs long (THL+TBL+TAL+FL equals 155.3 % SVL, not hypertrophied; thigh length smaller than tibia length (THL 94.4 % of TBL; THL 44.3 % of SVL; TBL 46.9 % of SVL); sum of thigh and tibia lengths smaller than SVL (THL+TBL 91.2 % of SVL); a single large tubercle on posteroventral internal margin of tarsus, near to the junction with shank. Tarsal length smaller than foot length (TAL 73.8 % of FL). Foot length smaller than thigh and tibia lengths (FL 83.2 % of THL; FL 78.5 % of TBL). Plantar surface without supernumerary tubercles; inner metatarsal tubercle large, rounded; outer metatarsal tubercle small and rounded; toes fringed; discs on toe I, smaller than of other toes; plantar formula, I 0+ – 2½ 
<date id="FFD31018F67F2C6F88D03C9DC16DF93D" box="[657,791,1680,1705]" pageId="3" pageNumber="104">II 1 – 2 III 1</date>
½ – 2½ 
<date id="FFD31018F67F2C6F89373C9DC1DDF93D" box="[886,935,1680,1705]" pageId="3" pageNumber="104">IV 3</date>
½ – 
<date id="FFD31018F67F2C6F89983C9DC67DF93C" box="[985,1031,1681,1705]" pageId="3" pageNumber="104">2 V.</date>
</paragraph>
<paragraph id="8BD236D8F67F2C6F8A863CB8C6DCF965" blockId="3.[151,1436,1210,2029]" pageId="3" pageNumber="104">Dorsal surfaces smooth; calcar appendage and supra-cloacal crest absent. Venter and ventral surfaces of thighs warty, each thigh with a large mesal pericloacal tubercle; remaining ventral surfaces smooth.</paragraph>
<paragraph id="8BD236D8F67F2C6F8A863CF1C7FBF8C9" blockId="3.[151,1436,1210,2029]" pageId="3" pageNumber="104">In preservative, dorsum uniformly creamy with small, irregular dark patches; wide brown stripe extending from the posterior corner of the eye to the insertion of arm; all ventral surfaces white, with the exception of a few minute dark flecks in the mental region and the undersides of the jaws, tarsus, tibia and palmar, plantar surfaces.</paragraph>
<paragraph id="8BD236D8F67F2C688A863D64C612FF63" blockId="3.[151,1436,1210,2029]" lastBlockId="4.[151,1436,151,248]" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
<emphasis id="B919EACAF67F2C6F8A863D64C345F815" bold="true" box="[199,319,1896,1921]" pageId="3" pageNumber="104">Variation.</emphasis>
The type specimens are alike in morphological traits, although the female (n = 1) is larger than the males. Variation in measurements is presented in 
<tableCitation id="C6EF0363F67F2C6F88FE3D80C169F831" box="[703,787,1932,1957]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Range, mean, and standard deviation (SD) of the measurements (mm) of the type specimens of Phyllodytes praeceptor sp. nov. (n = 28adult males and one female)." pageId="3" pageNumber="104">Table 1</tableCitation>
. Lateral dark brown stripe may extend from the eyes until or after the insertion of the arm. The amount of spots on the dorsum is variable. Only one specimen has no spots on the dorsum (MZUESC 17845). Some specimens (
<emphasis id="B919EACAF67F2C6F88893DD9C08AF878" box="[712,752,2005,2028]" italics="true" pageId="3" pageNumber="104">e.g.</emphasis>
, MZUESC 7298; 10498; 11584; 13026) have black spots on limbs, especially more evident on the hind limbs. A thin faded brown stripe on snout and 
<emphasis id="B919EACAF6782C688E9B3A94C7E6FF24" box="[1242,1436,152,176]" italics="true" pageId="4" pageNumber="105">canthus rostralis</emphasis>
extending to the eyes can be found in some individuals (
<emphasis id="B919EACAF6782C6889533AB1C149FF40" box="[786,819,189,212]" italics="true" pageId="4" pageNumber="105">e.g</emphasis>
., MZUESC 9175; MZUESC 11610; MZUESC 14207; MZUESC 14212). Snout in dorsal and lateral views vary from semicircular to truncate.
</paragraph>
</subSubSection>
<caption id="DF126650F6782C688AD63F18C2B5FAF3" httpUri="https://zenodo.org/record/1216490/files/figure.png" pageId="4" pageNumber="105" startId="4.[151,250,1300,1322]" targetBox="[157,1430,313,1273]" targetPageId="4">
<paragraph id="8BD236D8F6782C688AD63F18C2B5FAF3" blockId="4.[151,1436,1300,1383]" pageId="4" pageNumber="105">
<emphasis id="B919EACAF6782C688AD63F18C368FABD" bold="true" box="[151,274,1300,1322]" pageId="4" pageNumber="105">FIGURE 2.</emphasis>
<taxonomicName id="4C6D4D5BF6782C688B593F18C079FABD" authority="Orrico &amp; Dias &amp; Marciano-Jr, 2018" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[280,515,1300,1321]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="105" phylum="Chordata" rank="species" species="praeceptor" status="sp. nov.">
<emphasis id="B919EACAF6782C688B593F18C079FABD" box="[280,515,1300,1321]" italics="true" pageId="4" pageNumber="105">Phyllodytes praeceptor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B1F6782C68884B3F18C020FABD" box="[522,602,1300,1321]" pageId="4" pageNumber="105" rank="species">
<emphasis id="B919EACAF6782C68884B3F18C020FABD" bold="true" box="[522,602,1300,1321]" pageId="4" pageNumber="105">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype (MZUESC 17484; SVL 22.8 mm, scale bar = 2 mm). A) Dorsal view of the head; B) Lateral view of the head (left side); C) Left hindlimb (notice the tubercle near the heel); D) Plantar view of the left hand.
</paragraph>
</caption>
<caption id="DF126650F6782C688AD63DCFC752F84D" box="[151,1320,1987,2010]" httpUri="https://zenodo.org/record/1216492/files/figure.png" pageId="4" pageNumber="105" startId="4.[151,250,1987,2009]" targetBox="[157,1429,1443,1949]" targetPageId="4">
<paragraph id="8BD236D8F6782C688AD63DCFC752F84D" blockId="4.[151,1320,1987,2010]" box="[151,1320,1987,2010]" pageId="4" pageNumber="105">
<emphasis id="B919EACAF6782C688AD63DCFC377F84C" bold="true" box="[151,269,1987,2009]" pageId="4" pageNumber="105">FIGURE 3</emphasis>
. 
<taxonomicName id="4C6D4D5BF6782C688B583DCFC07EF84D" authority="Orrico &amp; Dias &amp; Marciano-Jr, 2018" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[281,516,1987,2009]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="105" phylum="Chordata" rank="species" species="praeceptor" status="sp. nov.">
<emphasis id="B919EACAF6782C688B583DCFC07EF84D" box="[281,516,1987,2009]" italics="true" pageId="4" pageNumber="105">Phyllodytes praeceptor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A57B1F6782C68884A3DCFC021F84C" box="[523,603,1987,2008]" pageId="4" pageNumber="105" rank="species">
<emphasis id="B919EACAF6782C68884A3DCFC021F84C" bold="true" box="[523,603,1987,2008]" pageId="4" pageNumber="105">sp. nov.</emphasis>
</taxonomicNameLabel>
in life. A) holotype (MZUESC 17484) and B) specimen not collected.
</paragraph>
</caption>
<caption id="DF126650F6792C698AD63A94C08CFF5B" pageId="5" pageNumber="106">
<paragraph id="8BD236D8F6792C698AD63A94C08CFF5B" blockId="5.[151,1435,151,207]" pageId="5" pageNumber="106">
<emphasis id="B919EACAF6792C698AD63A94C371FF3B" bold="true" box="[151,267,152,175]" pageId="5" pageNumber="106">TABLE 1.</emphasis>
Range, mean, and standard deviation (SD) of the measurements (mm) of the type specimens of 
<taxonomicName id="4C6D4D5BF6792C698F5E3A9BC376FF5B" authority="Orrico &amp; Dias &amp; Marciano-Jr, 2018" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="praeceptor" status="sp. nov.">
<emphasis id="B919EACAF6792C698F5E3A9BC376FF5B" italics="true" pageId="5" pageNumber="106">Phyllodytes praeceptor</emphasis>
</taxonomicName>
<emphasis id="B919EACAF6792C698B533AB4C312FF5B" bold="true" box="[274,360,184,207]" pageId="5" pageNumber="106">
<taxonomicNameLabel id="A22A57B1F6792C698B533AB4C312FF5B" box="[274,360,184,207]" pageId="5" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(n = 28 adult males and one female).
</paragraph>
</caption>
<paragraph id="8BD236D8F6792C698BFB3AE6C77DFC02" pageId="5" pageNumber="106">
<table id="F96DC478F679D3938ADC3AE6C748FC02" box="[157,1330,234,918]" gridcols="6" gridrows="17" pageId="5" pageNumber="106">
<tr id="355D349AF679D3938ADC3AE6C748FE94" box="[157,1330,234,256]" gridrow="0" pageId="5" pageNumber="106" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="768C5DE6F679D3938BFB3AE6C062FE94" box="[442,536,234,256]" gridcol="1" gridrow="0" pageId="5" pageNumber="106">Males</th>
<th id="768C5DE6F679D3938EA63AE6C748FE94" box="[1255,1330,234,256]" gridcol="5" gridrow="0" pageId="5" pageNumber="106">Female</th>
</tr>
<tr id="355D349AF679D3938ADC3B18C748FEBE" box="[157,1330,276,298]" gridrow="1" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3B18C373FEBE" box="[157,265,276,298]" gridcol="0" gridrow="1" pageId="5" pageNumber="106">Characters</th>
<td id="768C5DE6F679D3938BFB3B18C062FEBE" box="[442,536,276,298]" gridcol="1" gridrow="1" pageId="5" pageNumber="106">Holotype</td>
<td id="768C5DE6F679D39388E23B18C172FEBE" box="[675,776,276,298]" gridcol="2" gridrow="1" pageId="5" pageNumber="106">Range</td>
<td id="768C5DE6F679D39389E23B18C1A7FEBE" box="[931,989,276,298]" gridcol="3" gridrow="1" pageId="5" pageNumber="106">Mean</td>
<td id="768C5DE6F679D3938E0D3B18C616FEBE" box="[1100,1132,276,298]" gridcol="4" gridrow="1" pageId="5" pageNumber="106">SD</td>
<td id="768C5DE6F679D3938EA63B18C748FEBE" box="[1255,1330,276,298]" gridcol="5" gridrow="1" pageId="5" pageNumber="106">-</td>
</tr>
<tr id="355D349AF679D3938ADC3B31C748FEC7" box="[157,1330,317,339]" gridrow="2" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3B31C373FEC7" box="[157,265,317,339]" gridcol="0" gridrow="2" pageId="5" pageNumber="106">SVL</th>
<td id="768C5DE6F679D3938BFB3B31C062FEC7" box="[442,536,317,339]" gridcol="1" gridrow="2" pageId="5" pageNumber="106">22.8</td>
<td id="768C5DE6F679D39388E23B31C172FEC7" box="[675,776,317,339]" gridcol="2" gridrow="2" pageId="5" pageNumber="106">20.7–25.8</td>
<td id="768C5DE6F679D39389E23B31C1A7FEC7" box="[931,989,317,339]" gridcol="3" gridrow="2" pageId="5" pageNumber="106">22.8</td>
<td id="768C5DE6F679D3938E0D3B31C616FEC7" box="[1100,1132,317,339]" gridcol="4" gridrow="2" pageId="5" pageNumber="106">1.2</td>
<td id="768C5DE6F679D3938EA63B31C748FEC7" box="[1255,1330,317,339]" gridcol="5" gridrow="2" pageId="5" pageNumber="106">25.9</td>
</tr>
<tr id="355D349AF679D3938ADC3B6AC748FEE8" box="[157,1330,358,380]" gridrow="3" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3B6AC373FEE8" box="[157,265,358,380]" gridcol="0" gridrow="3" pageId="5" pageNumber="106">HW</th>
<td id="768C5DE6F679D3938BFB3B6AC062FEE8" box="[442,536,358,380]" gridcol="1" gridrow="3" pageId="5" pageNumber="106">8.6</td>
<td id="768C5DE6F679D39388E23B6AC172FEE8" box="[675,776,358,380]" gridcol="2" gridrow="3" pageId="5" pageNumber="106">8.0–9.3</td>
<td id="768C5DE6F679D39389E23B6AC1A7FEE8" box="[931,989,358,380]" gridcol="3" gridrow="3" pageId="5" pageNumber="106">8.6</td>
<td id="768C5DE6F679D3938E0D3B6AC616FEE8" box="[1100,1132,358,380]" gridcol="4" gridrow="3" pageId="5" pageNumber="106">0.4</td>
<td id="768C5DE6F679D3938EA63B6AC748FEE8" box="[1255,1330,358,380]" gridcol="5" gridrow="3" pageId="5" pageNumber="106">9.5</td>
</tr>
<tr id="355D349AF679D3938ADC3B9CC748FE32" box="[157,1330,400,422]" gridrow="4" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3B9CC373FE32" box="[157,265,400,422]" gridcol="0" gridrow="4" pageId="5" pageNumber="106">HL</th>
<td id="768C5DE6F679D3938BFB3B9CC062FE32" box="[442,536,400,422]" gridcol="1" gridrow="4" pageId="5" pageNumber="106">7.7</td>
<td id="768C5DE6F679D39388E23B9CC172FE32" box="[675,776,400,422]" gridcol="2" gridrow="4" pageId="5" pageNumber="106">6.6–8.1</td>
<td id="768C5DE6F679D39389E23B9CC1A7FE32" box="[931,989,400,422]" gridcol="3" gridrow="4" pageId="5" pageNumber="106">7.5</td>
<td id="768C5DE6F679D3938E0D3B9CC616FE32" box="[1100,1132,400,422]" gridcol="4" gridrow="4" pageId="5" pageNumber="106">0.4</td>
<td id="768C5DE6F679D3938EA63B9CC748FE32" box="[1255,1330,400,422]" gridcol="5" gridrow="4" pageId="5" pageNumber="106">8.7</td>
</tr>
<tr id="355D349AF679D3938ADC3BB5C748FE5B" box="[157,1330,441,463]" gridrow="5" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3BB5C373FE5B" box="[157,265,441,463]" gridcol="0" gridrow="5" pageId="5" pageNumber="106">IND</th>
<td id="768C5DE6F679D3938BFB3BB5C062FE5B" box="[442,536,441,463]" gridcol="1" gridrow="5" pageId="5" pageNumber="106">1.5</td>
<td id="768C5DE6F679D39388E23BB5C172FE5B" box="[675,776,441,463]" gridcol="2" gridrow="5" pageId="5" pageNumber="106">1.2–1.7</td>
<td id="768C5DE6F679D39389E23BB5C1A7FE5B" box="[931,989,441,463]" gridcol="3" gridrow="5" pageId="5" pageNumber="106">1.4</td>
<td id="768C5DE6F679D3938E0D3BB5C616FE5B" box="[1100,1132,441,463]" gridcol="4" gridrow="5" pageId="5" pageNumber="106">0.1</td>
<td id="768C5DE6F679D3938EA63BB5C748FE5B" box="[1255,1330,441,463]" gridcol="5" gridrow="5" pageId="5" pageNumber="106">1.7</td>
</tr>
<tr id="355D349AF679D3938ADC3BEEC748FE6C" box="[157,1330,482,504]" gridrow="6" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3BEEC373FE6C" box="[157,265,482,504]" gridcol="0" gridrow="6" pageId="5" pageNumber="106">END</th>
<td id="768C5DE6F679D3938BFB3BEEC062FE6C" box="[442,536,482,504]" gridcol="1" gridrow="6" pageId="5" pageNumber="106">2.3</td>
<td id="768C5DE6F679D39388E23BEEC172FE6C" box="[675,776,482,504]" gridcol="2" gridrow="6" pageId="5" pageNumber="106">1.8–2.6</td>
<td id="768C5DE6F679D39389E23BEEC1A7FE6C" box="[931,989,482,504]" gridcol="3" gridrow="6" pageId="5" pageNumber="106">2.2</td>
<td id="768C5DE6F679D3938E0D3BEEC616FE6C" box="[1100,1132,482,504]" gridcol="4" gridrow="6" pageId="5" pageNumber="106">0.2</td>
<td id="768C5DE6F679D3938EA63BEEC748FE6C" box="[1255,1330,482,504]" gridcol="5" gridrow="6" pageId="5" pageNumber="106">2.6</td>
</tr>
<tr id="355D349AF679D3938ADC3800C748FDB6" box="[157,1330,524,546]" gridrow="7" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3800C373FDB6" box="[157,265,524,546]" gridcol="0" gridrow="7" pageId="5" pageNumber="106">IOD</th>
<td id="768C5DE6F679D3938BFB3800C062FDB6" box="[442,536,524,546]" gridcol="1" gridrow="7" pageId="5" pageNumber="106">4.4</td>
<td id="768C5DE6F679D39388E23800C172FDB6" box="[675,776,524,546]" gridcol="2" gridrow="7" pageId="5" pageNumber="106">4.1–5.3</td>
<td id="768C5DE6F679D39389E23800C1A7FDB6" box="[931,989,524,546]" gridcol="3" gridrow="7" pageId="5" pageNumber="106">4.5</td>
<td id="768C5DE6F679D3938E0D3800C616FDB6" box="[1100,1132,524,546]" gridcol="4" gridrow="7" pageId="5" pageNumber="106">0.3</td>
<td id="768C5DE6F679D3938EA63800C748FDB6" box="[1255,1330,524,546]" gridcol="5" gridrow="7" pageId="5" pageNumber="106">5.1</td>
</tr>
<tr id="355D349AF679D3938ADC3839C748FDDF" box="[157,1330,565,587]" gridrow="8" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3839C373FDDF" box="[157,265,565,587]" gridcol="0" gridrow="8" pageId="5" pageNumber="106">ED</th>
<td id="768C5DE6F679D3938BFB3839C062FDDF" box="[442,536,565,587]" gridcol="1" gridrow="8" pageId="5" pageNumber="106">2.6</td>
<td id="768C5DE6F679D39388E23839C172FDDF" box="[675,776,565,587]" gridcol="2" gridrow="8" pageId="5" pageNumber="106">2.6–3.1</td>
<td id="768C5DE6F679D39389E23839C1A7FDDF" box="[931,989,565,587]" gridcol="3" gridrow="8" pageId="5" pageNumber="106">2.8</td>
<td id="768C5DE6F679D3938E0D3839C616FDDF" box="[1100,1132,565,587]" gridcol="4" gridrow="8" pageId="5" pageNumber="106">0.1</td>
<td id="768C5DE6F679D3938EA63839C748FDDF" box="[1255,1330,565,587]" gridcol="5" gridrow="8" pageId="5" pageNumber="106">3.0</td>
</tr>
<tr id="355D349AF679D3938ADC3852C748FDE0" box="[157,1330,606,628]" gridrow="9" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3852C373FDE0" box="[157,265,606,628]" gridcol="0" gridrow="9" pageId="5" pageNumber="106">TD</th>
<td id="768C5DE6F679D3938BFB3852C062FDE0" box="[442,536,606,628]" gridcol="1" gridrow="9" pageId="5" pageNumber="106">1.4</td>
<td id="768C5DE6F679D39388E23852C172FDE0" box="[675,776,606,628]" gridcol="2" gridrow="9" pageId="5" pageNumber="106">1.1–1.7</td>
<td id="768C5DE6F679D39389E23852C1A7FDE0" box="[931,989,606,628]" gridcol="3" gridrow="9" pageId="5" pageNumber="106">1.4</td>
<td id="768C5DE6F679D3938E0D3852C616FDE0" box="[1100,1132,606,628]" gridcol="4" gridrow="9" pageId="5" pageNumber="106">0.1</td>
<td id="768C5DE6F679D3938EA63852C748FDE0" box="[1255,1330,606,628]" gridcol="5" gridrow="9" pageId="5" pageNumber="106">1.5</td>
</tr>
<tr id="355D349AF679D3938ADC3884C748FD0A" box="[157,1330,648,670]" gridrow="10" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3884C373FD0A" box="[157,265,648,670]" gridcol="0" gridrow="10" pageId="5" pageNumber="106">THL</th>
<td id="768C5DE6F679D3938BFB3884C062FD0A" box="[442,536,648,670]" gridcol="1" gridrow="10" pageId="5" pageNumber="106">10.1</td>
<td id="768C5DE6F679D39388E23884C172FD0A" box="[675,776,648,670]" gridcol="2" gridrow="10" pageId="5" pageNumber="106">9.5–12.0</td>
<td id="768C5DE6F679D39389E23884C1A7FD0A" box="[931,989,648,670]" gridcol="3" gridrow="10" pageId="5" pageNumber="106">10.8</td>
<td id="768C5DE6F679D3938E0D3884C616FD0A" box="[1100,1132,648,670]" gridcol="4" gridrow="10" pageId="5" pageNumber="106">0.6</td>
<td id="768C5DE6F679D3938EA63884C748FD0A" box="[1255,1330,648,670]" gridcol="5" gridrow="10" pageId="5" pageNumber="106">12.0</td>
</tr>
<tr id="355D349AF679D3938ADC38BDC748FD53" box="[157,1330,689,711]" gridrow="11" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC38BDC373FD53" box="[157,265,689,711]" gridcol="0" gridrow="11" pageId="5" pageNumber="106">TBL</th>
<td id="768C5DE6F679D3938BFB38BDC062FD53" box="[442,536,689,711]" gridcol="1" gridrow="11" pageId="5" pageNumber="106">10.7</td>
<td id="768C5DE6F679D39388E238BDC172FD53" box="[675,776,689,711]" gridcol="2" gridrow="11" pageId="5" pageNumber="106">9.8–13.2</td>
<td id="768C5DE6F679D39389E238BDC1A7FD53" box="[931,989,689,711]" gridcol="3" gridrow="11" pageId="5" pageNumber="106">11.6</td>
<td id="768C5DE6F679D3938E0D38BDC616FD53" box="[1100,1132,689,711]" gridcol="4" gridrow="11" pageId="5" pageNumber="106">0.7</td>
<td id="768C5DE6F679D3938EA638BDC748FD53" box="[1255,1330,689,711]" gridcol="5" gridrow="11" pageId="5" pageNumber="106">12.8</td>
</tr>
<tr id="355D349AF679D3938ADC38D6C748FD64" box="[157,1330,730,752]" gridrow="12" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC38D6C373FD64" box="[157,265,730,752]" gridcol="0" gridrow="12" pageId="5" pageNumber="106">TAL</th>
<td id="768C5DE6F679D3938BFB38D6C062FD64" box="[442,536,730,752]" gridcol="1" gridrow="12" pageId="5" pageNumber="106">6.2</td>
<td id="768C5DE6F679D39388E238D6C172FD64" box="[675,776,730,752]" gridcol="2" gridrow="12" pageId="5" pageNumber="106">5.6–8.0</td>
<td id="768C5DE6F679D39389E238D6C1A7FD64" box="[931,989,730,752]" gridcol="3" gridrow="12" pageId="5" pageNumber="106">6.4</td>
<td id="768C5DE6F679D3938E0D38D6C616FD64" box="[1100,1132,730,752]" gridcol="4" gridrow="12" pageId="5" pageNumber="106">0.5</td>
<td id="768C5DE6F679D3938EA638D6C748FD64" box="[1255,1330,730,752]" gridcol="5" gridrow="12" pageId="5" pageNumber="106">7.1</td>
</tr>
<tr id="355D349AF679D3938ADC3908C748FC8E" box="[157,1330,772,794]" gridrow="13" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3908C373FC8E" box="[157,265,772,794]" gridcol="0" gridrow="13" pageId="5" pageNumber="106">FL</th>
<td id="768C5DE6F679D3938BFB3908C062FC8E" box="[442,536,772,794]" gridcol="1" gridrow="13" pageId="5" pageNumber="106">8.4</td>
<td id="768C5DE6F679D39388E23908C172FC8E" box="[675,776,772,794]" gridcol="2" gridrow="13" pageId="5" pageNumber="106">7.1–9.5</td>
<td id="768C5DE6F679D39389E23908C1A7FC8E" box="[931,989,772,794]" gridcol="3" gridrow="13" pageId="5" pageNumber="106">8.6</td>
<td id="768C5DE6F679D3938E0D3908C616FC8E" box="[1100,1132,772,794]" gridcol="4" gridrow="13" pageId="5" pageNumber="106">0.6</td>
<td id="768C5DE6F679D3938EA63908C748FC8E" box="[1255,1330,772,794]" gridcol="5" gridrow="13" pageId="5" pageNumber="106">10.2</td>
</tr>
<tr id="355D349AF679D3938ADC3921C748FCD7" box="[157,1330,813,835]" gridrow="14" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC3921C373FCD7" box="[157,265,813,835]" gridcol="0" gridrow="14" pageId="5" pageNumber="106">HAL</th>
<td id="768C5DE6F679D3938BFB3921C062FCD7" box="[442,536,813,835]" gridcol="1" gridrow="14" pageId="5" pageNumber="106">6.5</td>
<td id="768C5DE6F679D39388E23921C172FCD7" box="[675,776,813,835]" gridcol="2" gridrow="14" pageId="5" pageNumber="106">5.6–7.2</td>
<td id="768C5DE6F679D39389E23921C1A7FCD7" box="[931,989,813,835]" gridcol="3" gridrow="14" pageId="5" pageNumber="106">6.4</td>
<td id="768C5DE6F679D3938E0D3921C616FCD7" box="[1100,1132,813,835]" gridcol="4" gridrow="14" pageId="5" pageNumber="106">0.4</td>
<td id="768C5DE6F679D3938EA63921C748FCD7" box="[1255,1330,813,835]" gridcol="5" gridrow="14" pageId="5" pageNumber="106">7.2</td>
</tr>
<tr id="355D349AF679D3938ADC395AC748FCF8" box="[157,1330,854,876]" gridrow="15" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC395AC373FCF8" box="[157,265,854,876]" gridcol="0" gridrow="15" pageId="5" pageNumber="106">DF3</th>
<td id="768C5DE6F679D3938BFB395AC062FCF8" box="[442,536,854,876]" gridcol="1" gridrow="15" pageId="5" pageNumber="106">1.2</td>
<td id="768C5DE6F679D39388E2395AC172FCF8" box="[675,776,854,876]" gridcol="2" gridrow="15" pageId="5" pageNumber="106">0.9–1.2</td>
<td id="768C5DE6F679D39389E2395AC1A7FCF8" box="[931,989,854,876]" gridcol="3" gridrow="15" pageId="5" pageNumber="106">1.1</td>
<td id="768C5DE6F679D3938E0D395AC616FCF8" box="[1100,1132,854,876]" gridcol="4" gridrow="15" pageId="5" pageNumber="106">0.1</td>
<td id="768C5DE6F679D3938EA6395AC748FCF8" box="[1255,1330,854,876]" gridcol="5" gridrow="15" pageId="5" pageNumber="106">1.4</td>
</tr>
<tr id="355D349AF679D3938ADC398CC748FC02" box="[157,1330,896,918]" gridrow="16" pageId="5" pageNumber="106">
<th id="768C5DE6F679D3938ADC398CC373FC02" box="[157,265,896,918]" gridcol="0" gridrow="16" pageId="5" pageNumber="106">4TD</th>
<td id="768C5DE6F679D3938BFB398CC062FC02" box="[442,536,896,918]" gridcol="1" gridrow="16" pageId="5" pageNumber="106">1.1</td>
<td id="768C5DE6F679D39388E2398CC172FC02" box="[675,776,896,918]" gridcol="2" gridrow="16" pageId="5" pageNumber="106">0.8–1.3</td>
<td id="768C5DE6F679D39389E2398CC1A7FC02" box="[931,989,896,918]" gridcol="3" gridrow="16" pageId="5" pageNumber="106">1.0</td>
<td id="768C5DE6F679D3938E0D398CC616FC02" box="[1100,1132,896,918]" gridcol="4" gridrow="16" pageId="5" pageNumber="106">0.1</td>
<td id="768C5DE6F679D3938EA6398CC748FC02" box="[1255,1330,896,918]" gridcol="5" gridrow="16" pageId="5" pageNumber="106">1.5</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3776553F6792C698A8639D9C15CFBEA" pageId="5" pageNumber="106" type="etymology">
<paragraph id="8BD236D8F6792C698A8639D9C60BFB86" blockId="5.[151,1437,981,2014]" pageId="5" pageNumber="106">
<emphasis id="B919EACAF6792C698A8639D9C334FC7A" bold="true" box="[199,334,981,1006]" pageId="5" pageNumber="106">Etymology.</emphasis>
The specific epithet is a Latin word meaning “teacher” or “instructor”. The name is given as homage to Dr. Miguel Trefaut Rodrigues for his continuous formation of herpetologists.
</paragraph>
<paragraph id="8BD236D8F6792C698A863E11C15CFBEA" blockId="5.[151,1437,981,2014]" pageId="5" pageNumber="106">
<emphasis id="B919EACAF6792C698A863E11C394FBA2" bold="true" box="[199,494,1053,1078]" pageId="5" pageNumber="106">Geographic distribution.</emphasis>
Known from the coastal lowlands of 
<collectingRegion id="49A9F83AF6792C6989D03E11C1AFFBA2" box="[913,981,1053,1078]" country="Brazil" name="Bahia" pageId="5" pageNumber="106">Bahia</collectingRegion>
, from municipality of Maragogipe to the south of the state in the municipality of Una. Specimens are also known from higher areas such as in Serra da Jiboia (
<quantity id="4C959B3DF6792C698AA93E69C34BFBE9" box="[232,305,1125,1150]" metricMagnitude="2" metricUnit="m" metricValue="6.84" pageId="5" pageNumber="106" unit="m" value="684.0">684 m</quantity>
a.s.l), Santa Terezinha municipality (
<figureCitation id="13562A5DF6792C6988923E69C163FBEA" box="[723,793,1125,1150]" captionStart="FIGURE 4" captionStartId="6.[151,250,1885,1907]" captionTargetBox="[162,1417,657,1862]" captionTargetId="figure@6.[151,1436,657,1863]" captionTargetPageId="6" captionText="FIGURE 4. Geographic distribution of Phyllodytes praeceptor sp. nov. Geographic coordinates in decimal degrees, Datum WGS 84. (Star) -12.869436 S, -39.461494 W; (1) -12.836035 S, -38.866291 W; (2) -13.828639 S, -39.218833 W; (3) - 14.32797222 S, -39.07811111 W; (4) -14.53912 S, -39.15461; (5) -14.600967 S, -39.062332 W; (6) -15.07488889 S, -39.0525 W; (7) -15.18401S, -39.06638 W." httpUri="https://zenodo.org/record/1216494/files/figure.png" pageId="5" pageNumber="106">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3776553F6792C6A8A863E85C36DFDCB" lastPageId="6" lastPageNumber="107" pageId="5" pageNumber="106" type="description">
<paragraph id="8BD236D8F6792C698A863E85C0EFFAEE" blockId="5.[151,1437,981,2014]" pageId="5" pageNumber="106">
<emphasis id="B919EACAF6792C698A863E85C3D5FB36" bold="true" box="[199,431,1161,1186]" pageId="5" pageNumber="106">Advertisement call</emphasis>
. The advertisement call (
<figureCitation id="13562A5DF6792C6988A83E85C149FB36" box="[745,819,1161,1186]" captionStart="FIGURE 5" captionStartId="7.[151,250,596,618]" captionTargetBox="[411,1176,193,571]" captionTargetId="figure@7.[410,1176,193,574]" captionTargetPageId="7" captionText="FIGURE 5. Advertisement call of Phyllodytes praeceptor (MZUESC 17484, holotype). A) Waveform and B) audiospectrogram.Recorded on 02 March 2015 at 22 h 17 min. Air temperature during recording = 22°C." httpUri="https://zenodo.org/record/1216496/files/figure.png" pageId="5" pageNumber="106">Fig. 5</figureCitation>
) of 
<taxonomicName id="4C6D4D5BF6792C6989313E85C606FB35" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[880,1148,1161,1185]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF6792C6989313E85C606FB35" box="[880,1148,1161,1185]" italics="true" pageId="5" pageNumber="106">Phyllodytes praeceptor</emphasis>
</taxonomicName>
has a total duration of 3.02– 
<geoCoordinate id="EE59501FF6792C698A963EA2C362FB51" box="[215,280,1198,1222]" direction="south" orientation="latitude" pageId="5" pageNumber="106" precision="555" value="-9.41">9.41 s</geoCoordinate>
(5.34 ± 1.53, n = 28). It is emitted at intervals of 29.5– 
<geoCoordinate id="EE59501FF6792C69893E3EA1C1B9FB51" box="[895,963,1197,1222]" direction="south" orientation="latitude" pageId="5" pageNumber="106" precision="5555" value="-99.2">99.2 s</geoCoordinate>
(48.5 ± 21.2, n = 27) and with a dominant frequency of 2928.5–3273.0 Hz (3045.4 ± 115.3, n = 28). Each call is composed of 6–12 notes (8.39 ± 1.55, n = 28) with structured pulses lasting 0.141– 
<geoCoordinate id="EE59501FF6792C6988753EF9C0FCFA99" box="[564,646,1269,1294]" direction="south" orientation="latitude" pageId="5" pageNumber="106" precision="55" value="-0.538">0.538 s</geoCoordinate>
(0.295 ± 0.075, n = 235), emitted at intervals of 0.236– 
<geoCoordinate id="EE59501FF6792C698EBC3EF9C737FA99" box="[1277,1357,1269,1294]" direction="south" orientation="latitude" pageId="5" pageNumber="106" precision="55" value="-1.057">1.057 s</geoCoordinate>
(0.389 ± 0.102, n = 207), resulting in a note rate of 1.28–2.02 (1.61 ± 0.2, n = 28) notes/s. The interval between the final two notes of the call is generally much larger than the previous intervals. The notes dominant frequency range is 1378.1–3617.6 Hz (2998.1 ± 275.1, n = 235).
</paragraph>
<paragraph id="8BD236D8F6792C698A863F8AC73EF99D" blockId="5.[151,1437,981,2014]" pageId="5" pageNumber="106">Notes are composed by isolated pulses and/or by 1–11 pulse groups with 2–8 pulses in each group. Usually, the last pulse group is the largest among them and always composed of 6–15 more aggregated pulses. The total number of pulses per note is 17–58 (28.7 ± 9.4, n = 89) and pulse emission rate is 67.2–141.8 pulses/s (98.9 ± 15.8, n = 89). The notes exhibit amplitude modulation, usually starting with low intensity and increasing during the note.</paragraph>
<paragraph id="8BD236D8F6792C698A863C19C3CBF90E" blockId="5.[151,1437,981,2014]" pageId="5" pageNumber="106">
<emphasis id="B919EACAF6792C698A863C19C0AEF9BA" bold="true" box="[199,724,1557,1582]" pageId="5" pageNumber="106">Bioacoustic comparisons with other species.</emphasis>
The advertisement call of 
<taxonomicName id="4C6D4D5BF6792C698E453C19C773F9B9" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[1028,1289,1557,1581]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF6792C698E453C19C773F9B9" box="[1028,1289,1557,1581]" italics="true" pageId="5" pageNumber="106">Phyllodytes praeceptor</emphasis>
</taxonomicName>
has a pulsed structure being readily distinguishable from the call of 
<taxonomicName id="4C6D4D5BF6792C69896D3C36C1B4F9C5" authorityName="Bokermann" authorityYear="1966" box="[812,974,1594,1617]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B919EACAF6792C69896D3C36C1B4F9C5" box="[812,974,1594,1617]" italics="true" pageId="5" pageNumber="106">P. acuminatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF6792C6989A33C36C61AF9C5" box="[994,1120,1593,1617]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B919EACAF6792C6989A33C36C61AF9C5" box="[994,1120,1593,1617]" italics="true" pageId="5" pageNumber="106">P. kautskyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF6792C698E323C36C73FF9C5" authorityName="" authorityYear="1830" box="[1139,1349,1594,1617]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B919EACAF6792C698E323C36C73FF9C5" box="[1139,1349,1594,1617]" italics="true" pageId="5" pageNumber="106">P. megatympanum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6D4D5BF6792C698FC63C36C34DF9E1" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B919EACAF6792C698FC63C36C34DF9E1" italics="true" pageId="5" pageNumber="106">P. melanomystax</emphasis>
</taxonomicName>
, which show unpulsed structure (Campos 
<emphasis id="B919EACAF6792C6989503C52C132F9E1" box="[785,840,1629,1653]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
, 2014; Nunes 
<emphasis id="B919EACAF6792C6989AA3C52C659F9E1" box="[1003,1059,1629,1653]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
, 2007; Simon &amp; Gasparini, 2003, Marciano Jr. 
<emphasis id="B919EACAF6792C698B6B3C8EC318F90D" box="[298,354,1665,1689]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
2017).
</paragraph>
<paragraph id="8BD236D8F6792C698A863CA9C6BDF8DA" blockId="5.[151,1437,981,2014]" pageId="5" pageNumber="106">
The calls of 
<emphasis id="B919EACAF6792C698B213CA9C0B4F929" box="[352,718,1701,1725]" italics="true" pageId="5" pageNumber="106">
<taxonomicName id="4C6D4D5BF6792C698B213CA9C03FF929" box="[352,581,1701,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="amadoi">Phyllodytes amadoi</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF6792C6988143CAAC0B4F929" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[597,718,1701,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="luteolus">P. luteolus</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C6D4D5BF6792C6988A13CAAC12FF929" box="[736,853,1701,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B919EACAF6792C6988A13CAAC12FF929" box="[736,853,1701,1725]" italics="true" pageId="5" pageNumber="106">P. edelmoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF6792C69892D3CAAC660F929" authorityName="Bokermann" authorityYear="1966" box="[876,1050,1701,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B919EACAF6792C69892D3CAAC660F929" box="[876,1050,1701,1725]" italics="true" pageId="5" pageNumber="106">P. tuberculosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6D4D5BF6792C698E6D3CAAC6B4F929" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[1068,1230,1701,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B919EACAF6792C698E6D3CAAC6B4F929" box="[1068,1230,1701,1725]" italics="true" pageId="5" pageNumber="106">P. gyrinaethes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6D4D5BF6792C698F4E3CAAC7E6F929" baseAuthorityName="Peters" baseAuthorityYear="1873" box="[1295,1436,1701,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B919EACAF6792C698F4E3CAAC7E6F929" box="[1295,1436,1701,1725]" italics="true" pageId="5" pageNumber="106">P wuchereri</emphasis>
</taxonomicName>
(Weygoldt 1981; Lima 
<emphasis id="B919EACAF6792C698BE23CC6C3A4F975" box="[419,478,1737,1761]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
2008; Juncá 
<emphasis id="B919EACAF6792C6988383CC6C0CEF975" box="[633,692,1737,1761]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
2012; Roberto &amp; Ávila 2013 
<bibRefCitation id="EFFC4B29F6792C698E533CC6C6ABF976" box="[1042,1233,1737,1762]" pageId="5" pageNumber="106" refId="ref5347" refString="Cruz, D., Marciano-Jr., E. &amp; Napoli, M. F. (2014) Advertisement and courtship calls of Phyllodytes wuchereri (Peters, 1873) (Anura: Hylidae) Zootaxa, 3774, 97 - 100. https: // doi. org / 10.11646 / zootaxa. 3774.1.8" type="journal article">
Cruz 
<emphasis id="B919EACAF6792C698E133CC6C6F6F975" box="[1106,1164,1737,1761]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
2014
</bibRefCitation>
; 
<bibRefCitation id="EFFC4B29F6792C698E9E3CC5C2A9F892" pageId="5" pageNumber="106" refId="ref6106" refString="Magalhaes, F. De M., Junca, F. A. &amp; Garda, A. A. (2015) Tadpole and vocalisations of Phyllodytes wuchereri (Anura: Hylidae) from Bahia, Brazil. Salamandra, 51, 83 - 90." type="journal article">
Magalhães 
<emphasis id="B919EACAF6792C698F233CC6C7E6F975" box="[1378,1436,1737,1761]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
2015
</bibRefCitation>
; Vörös 
<emphasis id="B919EACAF6792C698B7A3CE2C307F891" box="[315,381,1773,1797]" italics="true" pageId="5" pageNumber="106">et al.</emphasis>
2017) are more similar to 
<taxonomicName id="4C6D4D5BF6792C69889F3CE2C104F891" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[734,894,1774,1797]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF6792C69889F3CE2C104F891" box="[734,894,1774,1797]" italics="true" pageId="5" pageNumber="106">P. praeceptor</emphasis>
</taxonomicName>
because all of them consist of a series of multipulsionated notes and present overlap in some acoustic parameters, mainly in dominant frequency and call duration (except 
<taxonomicName id="4C6D4D5BF6792C698B153D3AC389F8D9" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[340,499,1845,1869]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B919EACAF6792C698B153D3AC389F8D9" box="[340,499,1845,1869]" italics="true" pageId="5" pageNumber="106">P. gyrinaethes</emphasis>
</taxonomicName>
, with shorter call and 
<taxonomicName id="4C6D4D5BF6792C6988AC3D3AC126F8D9" box="[749,860,1845,1869]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B919EACAF6792C6988AC3D3AC126F8D9" box="[749,860,1845,1869]" italics="true" pageId="5" pageNumber="106">P. amadoi</emphasis>
</taxonomicName>
with high dominant frequency).
</paragraph>
<paragraph id="8BD236D8F6792C698A863D55C388F84A" blockId="5.[151,1437,981,2014]" pageId="5" pageNumber="106">
The number of notes per call is higher in 
<taxonomicName id="4C6D4D5BF6792C6988DE3D56C16FF8E5" box="[671,789,1881,1905]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B919EACAF6792C6988DE3D56C16FF8E5" box="[671,789,1881,1905]" italics="true" pageId="5" pageNumber="106">P. edelmoi</emphasis>
</taxonomicName>
(22–29), 
<taxonomicName id="4C6D4D5BF6792C6989C53D56C18FF8E5" box="[900,1013,1881,1905]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B919EACAF6792C6989C53D56C18FF8E5" box="[900,1013,1881,1905]" italics="true" pageId="5" pageNumber="106">P. amadoi</emphasis>
</taxonomicName>
(13-17) and 
<taxonomicName id="4C6D4D5BF6792C698ECB3D56C74EF8E5" authorityName="Bokermann" authorityYear="1966" box="[1162,1332,1881,1905]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B919EACAF6792C698ECB3D56C74EF8E5" box="[1162,1332,1881,1905]" italics="true" pageId="5" pageNumber="106">P. tuberculosus</emphasis>
</taxonomicName>
(14–23), and their ranges do not overlap with the variation of 
<taxonomicName id="4C6D4D5BF6792C6989553D72C1CAF801" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[788,944,1918,1941]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF6792C6989553D72C1CAF801" box="[788,944,1918,1941]" italics="true" pageId="5" pageNumber="106">P. praeceptor</emphasis>
</taxonomicName>
(6–12). The mean number of notes of 
<taxonomicName id="4C6D4D5BF6792C698FC63D72C368F82D" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF6792C698FC63D72C368F82D" italics="true" pageId="5" pageNumber="106">P. praeceptor</emphasis>
</taxonomicName>
(8.39 ± 1.55, 6–12) is smaller than the mean of 
<taxonomicName id="4C6D4D5BF6792C6989743DAEC1BBF82D" baseAuthorityName="Peters" baseAuthorityYear="1873" box="[821,961,1953,1977]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B919EACAF6792C6989743DAEC1BBF82D" box="[821,961,1953,1977]" italics="true" pageId="5" pageNumber="106">P. wuchereri</emphasis>
</taxonomicName>
(16.18±3.25, 10–21) and larger than of 
<taxonomicName id="4C6D4D5BF6792C698FC63DAEC360F849" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="106" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B919EACAF6792C698FC63DAEC360F849" italics="true" pageId="5" pageNumber="106">P. gyrinaethes</emphasis>
</taxonomicName>
(4.90 ± 0.60, 4–6).
</paragraph>
<paragraph id="8BD236D8F67A2C6A8A863A9BC349FEF0" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="107">
The note duration in 
<taxonomicName id="4C6D4D5BF67A2C6A8BFA3A95C02EFF24" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[443,596,153,176]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF67A2C6A8BFA3A95C02EFF24" box="[443,596,153,176]" italics="true" pageId="6" pageNumber="107">P. praeceptor</emphasis>
</taxonomicName>
(0.295 ± 0.075) is longer than in all species of the genus with described calls: 
<taxonomicName id="4C6D4D5BF67A2C6A8A9B3AB1C32AFF40" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[218,336,188,212]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="luteolus">
<emphasis id="B919EACAF67A2C6A8A9B3AB1C32AFF40" box="[218,336,188,212]" italics="true" pageId="6" pageNumber="107">P. luteolus</emphasis>
</taxonomicName>
(0.125, no range available), 
<taxonomicName id="4C6D4D5BF67A2C6A88D83AB1C174FF40" box="[665,782,188,212]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B919EACAF67A2C6A88D83AB1C174FF40" box="[665,782,188,212]" italics="true" pageId="6" pageNumber="107">P. edelmoi</emphasis>
</taxonomicName>
(0.10 ± 0.003, 0.04–0.16), 
<taxonomicName id="4C6D4D5BF67A2C6A8E093AB1C689FF40" authorityName="Bokermann" authorityYear="1966" box="[1096,1267,188,212]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B919EACAF67A2C6A8E093AB1C689FF40" box="[1096,1267,188,212]" italics="true" pageId="6" pageNumber="107">P. tuberculosus</emphasis>
</taxonomicName>
(0.17 ± 0.047, 0.07–0.25), 
<taxonomicName id="4C6D4D5BF67A2C6A8B653AEDC3BDFF6C" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[292,455,224,248]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B919EACAF67A2C6A8B653AEDC3BDFF6C" box="[292,455,224,248]" italics="true" pageId="6" pageNumber="107">P. gyrinaethes</emphasis>
</taxonomicName>
(0.04 ± 0.01, 0.02–0.07), 
<taxonomicName id="4C6D4D5BF67A2C6A89463AEDC1E2FF6C" baseAuthorityName="Peters" baseAuthorityYear="1873" box="[775,920,224,248]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B919EACAF67A2C6A89463AEDC1E2FF6C" box="[775,920,224,248]" italics="true" pageId="6" pageNumber="107">P. wuchereri</emphasis>
</taxonomicName>
(0.19 ± 0.04, 0.11–0.32 and 0.12 ± 0.013, 0.04–0.14), 
<taxonomicName id="4C6D4D5BF67A2C6A8B613B09C3E3FE88" box="[288,409,260,284]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B919EACAF67A2C6A8B613B09C3E3FE88" box="[288,409,260,284]" italics="true" pageId="6" pageNumber="107">P. kautskyi</emphasis>
</taxonomicName>
(0.08 ± 0.012, no range available), 
<taxonomicName id="4C6D4D5BF67A2C6A89783B09C18CFE88" box="[825,1014,260,284]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B919EACAF67A2C6A89783B09C18CFE88" box="[825,1014,260,284]" italics="true" pageId="6" pageNumber="107">P. melanomystax</emphasis>
</taxonomicName>
(0.07 ± 0.04, no range available), 
<taxonomicName id="4C6D4D5BF67A2C6A8FC63B09C362FEAB" authorityName="Bokermann" authorityYear="1966" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B919EACAF67A2C6A8FC63B09C362FEAB" italics="true" pageId="6" pageNumber="107">P. acuminatus</emphasis>
</taxonomicName>
(0.10 ± 0.03, 0.03–0.17), 
<taxonomicName id="4C6D4D5BF67A2C6A88023B25C16BFED4" authorityName="" authorityYear="1830" box="[579,785,297,320]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B919EACAF67A2C6A88023B25C16BFED4" box="[579,785,297,320]" italics="true" pageId="6" pageNumber="107">P. megatympanum</emphasis>
</taxonomicName>
(0.092 ± 0.08, 0.009–0.24) and 
<taxonomicName id="4C6D4D5BF67A2C6A8EC23B25C689FED4" box="[1155,1267,296,320]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B919EACAF67A2C6A8EC23B25C689FED4" box="[1155,1267,296,320]" italics="true" pageId="6" pageNumber="107">P. amadoi</emphasis>
</taxonomicName>
(0.043 ± 0.02, 0.008–0.119).
</paragraph>
<paragraph id="8BD236D8F67A2C6A8A863B63C29EFE44" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="107">
Finally, the call of 
<taxonomicName id="4C6D4D5BF67A2C6A8BEB3B7CC0C9FE1C" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" box="[426,691,368,392]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF67A2C6A8BEB3B7CC0C9FE1C" box="[426,691,368,392]" italics="true" pageId="6" pageNumber="107">Phyllodytes praeceptor</emphasis>
</taxonomicName>
differs from all species of the genus by having an accentuated increase in amplitude from the first to the last pulse of each note (
<figureCitation id="13562A5DF67A2C6A89373B98C1C3FE39" box="[886,953,404,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,596,618]" captionTargetBox="[411,1176,193,571]" captionTargetId="figure@7.[410,1176,193,574]" captionTargetPageId="7" captionText="FIGURE 5. Advertisement call of Phyllodytes praeceptor (MZUESC 17484, holotype). A) Waveform and B) audiospectrogram.Recorded on 02 March 2015 at 22 h 17 min. Air temperature during recording = 22°C." httpUri="https://zenodo.org/record/1216496/files/figure.png" pageId="6" pageNumber="107">Fig. 5</figureCitation>
), concentrating energy at the last cluster of pulses.
</paragraph>
<paragraph id="8BD236D8F67A2C6A8A863BD0C36DFDCB" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="107">
<emphasis id="B919EACAF67A2C6A8A863BD0C3FCFE61" bold="true" box="[199,390,476,501]" pageId="6" pageNumber="107">Molecular data.</emphasis>
Uncorrected pairwise genetic distances of 803 pb of 16S generated in this study are 1 % in relation to 
<taxonomicName id="4C6D4D5BF67A2C6A8B59380CC3E1FD8C" authorityName="Wagler" authorityYear="1830" box="[280,411,512,536]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="genus">
<emphasis id="B919EACAF67A2C6A8B59380CC3E1FD8C" box="[280,411,512,536]" italics="true" pageId="6" pageNumber="107">Phyllodytes</emphasis>
</taxonomicName>
sp. (
<accessionNumber id="943EAB3BF67A2C6A8B99380CC02DFD8C" box="[472,599,512,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY843722" pageId="6" pageNumber="107">AY843722</accessionNumber>
; voucher MRT 6144) and 9.6 % to 
<taxonomicName id="4C6D4D5BF67A2C6A8E43380DC601FD8C" baseAuthorityName="Wied" baseAuthorityYear="1824" box="[1026,1147,512,536]" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="luteolus">
<emphasis id="B919EACAF67A2C6A8E43380DC601FD8C" box="[1026,1147,512,536]" italics="true" pageId="6" pageNumber="107">P. luteolus</emphasis>
</taxonomicName>
(
<accessionNumber id="943EAB3BF67A2C6A8ECC380CC774FD8C" box="[1165,1294,512,536]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY843721" pageId="6" pageNumber="107">AY843721</accessionNumber>
). Given this results, we conclude that the specimen MRT 6144, sequenced by 
<bibRefCitation id="EFFC4B29F67A2C6A89353828C60DFDA9" box="[884,1143,548,573]" pageId="6" pageNumber="107" refId="ref5560" refString="Faivovich, J., Haddad, C. F. B., Garcia, P. C. A., Frost, D. R., Campbell, J. A. &amp; Wheeler, W. C. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1 - 240. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 294 [0001: SROTFF] 2.0. CO; 2" type="journal article">
Faivovich 
<emphasis id="B919EACAF67A2C6A89AA3829C65EFDA8" box="[1003,1060,548,572]" italics="true" pageId="6" pageNumber="107">et al.</emphasis>
(2005)
</bibRefCitation>
, should be identified as 
<taxonomicName id="4C6D4D5BF67A2C6A8FC63829C36AFDF4" authorityName="Orrico &amp; Dias &amp; Marciano-Jr" authorityYear="2018" class="Amphibia" family="Hylidae" genus="Phyllodytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="107" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B919EACAF67A2C6A8FC63829C36AFDF4" italics="true" pageId="6" pageNumber="107">P. praeceptor</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>