treatments-xml/data/03/C4/87/03C487B9332FFF83FF3BF8E4FBA3F82A.xml

<document id="65ED2A059985CF4A009E9665B05CD06E" ID-CLB-Dataset="8849" ID-DOI="10.11646/zootaxa.4908.1.7" ID-GBIF-Dataset="567b6953-7707-4078-a4d3-54bfe787094c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4435860" ID-ZooBank="694FB3A3-3940-47EF-86BF-DFB96B4BBE99" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610525669478" checkinUser="plazi" docAuthor="Hsu, Yu-Feng, Lo, Yik-Fui Philip &amp; Lin, Rung-Juen" docDate="2021" docId="03C487B9332FFF83FF3BF8E4FBA3F82A" docLanguage="en" docName="zootaxa.4908.1.7.pdf" docOrigin="Zootaxa 4908 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chrysozephyrus mushaellus subsp. paolongkoui Hsu &amp; Lo &amp; Lin 2021, subsp. nov." docType="treatment" docVersion="7" lastPageNumber="118" masterDocId="FFFDFFC1332DFF86FFACFFE9FFCAFFAC" masterDocTitle="On a New Insular Subspecies of Chrysozephyrus mushaellus (Matsumura) (Lepidoptera: Lycaenidae) Discovered from Hainan" masterLastPageNumber="122" masterPageNumber="113" pageNumber="115" updateTime="1698925216341" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C2EC84F156D63297C5FF99394CD25FEB">On a New Insular Subspecies of Chrysozephyrus mushaellus (Matsumura) (Lepidoptera: Lycaenidae) Discovered from Hainan</mods:title>
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<mods:namePart id="EC42ED680E7851090BFE9A068CC28594">Hsu, Yu-Feng</mods:namePart>
<mods:affiliation id="74FF085907B988BA02B25701EBB8C579">Department of Life Science, National Taiwan Normal University, Taipei, Taiwan 116, R. O. C.</mods:affiliation>
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<mods:namePart id="1DD416A0E7D750E2F70CCBF046C0C6DD">Lo, Yik-Fui Philip</mods:namePart>
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<mods:affiliation id="62512B23838E7DD6D6EB897E3FC86E59">Kadoorie Conservation China, Kadoorie Farm and Botanic Garden, Lam Kam Road, Tai Po, New Territories, Hong Kong. philiplo @ kfbg. org, hkbutterfly @ gmail. com; https: // orcid. org / 0000 - 0001 - 6979 - 7108</mods:affiliation>
<mods:nameIdentifier id="1881A67FF22EC017D8AE8A28092CB257" type="email">hkbutterfly@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F1ACE69153A88E8736C8B11167438F91">Lin, Rung-Juen</mods:namePart>
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<mods:affiliation id="92C3AF5DAD71D283C449FEA017DCA977">Department of Life Science, National Taiwan Normal University, Taipei, Taiwan 116, R. O. C. &amp; puji. lin @ gmail. com; https: // orcid. org / 0000 - 0001 - 6114 - 5324</mods:affiliation>
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<mods:date id="6CA518A0A14821EF4711849C6BE22DCF">2021</mods:date>
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<treatment id="03C487B9332FFF83FF3BF8E4FBA3F82A" ID-DOI="http://doi.org/10.5281/zenodo.4443099" ID-GBIF-Taxon="176596750" ID-Zenodo-Dep="4443099" LSID="urn:lsid:plazi:treatment:03C487B9332FFF83FF3BF8E4FBA3F82A" httpUri="http://treatment.plazi.org/id/03C487B9332FFF83FF3BF8E4FBA3F82A" lastPageId="5" lastPageNumber="118" pageId="2" pageNumber="115">
<subSubSection id="C3776524332FFF84FF3BF8E4FC6FF88B" box="[151,933,1805,1832]" pageId="2" pageNumber="115" type="nomenclature">
<paragraph id="8BD236AF332FFF84FF3BF8E4FC6FF88B" blockId="2.[151,933,1805,1868]" box="[151,933,1805,1832]" pageId="2" pageNumber="115">
<heading id="D09A81C3332FFF84FF3BF8E4FC6FF88B" bold="true" box="[151,933,1805,1832]" fontSize="11" level="1" pageId="2" pageNumber="115" reason="1">
<emphasis id="B919EABD332FFF84FF3BF8E4FC6FF88B" bold="true" box="[151,933,1805,1832]" pageId="2" pageNumber="115">
<taxonomicName id="4C6D4D2C332FFF84FF3BF8E4FCC5F88B" authority="Hsu &amp; Lo" authorityName="Hsu &amp; Lo &amp; Lin" authorityYear="2021" box="[151,783,1805,1832]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="subSpecies" species="mushaellus" status="subsp. nov." subSpecies="paolongkoui">
<emphasis id="B919EABD332FFF84FF3BF8E4FD40F884" bold="true" box="[151,650,1805,1832]" italics="true" pageId="2" pageNumber="115">Chrysozephyrus mushaellus paolongkoui</emphasis>
Hsu &amp; Lo
</taxonomicName>
, 
<taxonomicNameLabel id="A22A57C6332FFF84FCB0F8E4FC6FF88B" box="[796,933,1805,1831]" pageId="2" pageNumber="115" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3776524332FFF84FF3BF8D8FDD6F8E7" box="[151,540,1841,1868]" pageId="2" pageNumber="115" type="description">
<paragraph id="8BD236AF332FFF84FF3BF8D8FDD6F8E7" blockId="2.[151,933,1805,1868]" box="[151,540,1841,1868]" pageId="2" pageNumber="115">
(
<figureCitation id="13562A2A332FFF84FF33F8D8FEC1F8E7" box="[159,267,1841,1867]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1906,1930]" captionTargetBox="[163,1425,501,1873]" captionTargetId="figure-159@4.[151,1436,489,1883]" captionTargetPageId="4" captionText="FIGURES 1–6. Male adults of Chrysozephyrus mushaellus (Matsumura). 1, upperside of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., holotype, 2, underside of holotype, 3, upperside of C. m. mushaellus (Taiwan), 4, underside of C. m. mushaellus (Taiwan), 5, upperside of C. m. rileyi (Guangdong), 6, underside of C. m. rileyi (Guangdong). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4435862" httpUri="https://zenodo.org/record/4435862/files/figure.png" pageId="2" pageNumber="115">Figs. 1, 2</figureCitation>
, 
<figureCitation id="13562A2A332FFF84FEBBF8DBFE88F8E7" box="[279,322,1841,1868]" captionStart="FIGURES 7–12" captionStartId="5.[151,264,1578,1602]" captionTargetBox="[158,1422,200,1540]" captionTargetId="figure-17@5.[151,1436,181,1554]" captionTargetPageId="5" captionText="FIGURES 7–12. Female adults of Chrysozephyrus mushaellus (Matsumura). 7, upperside of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., paratype, 8, underside of paratype, 9, upperside of C. m. mushaellus (Taiwan), 10, underside of C. m. mushaellus (Taiwan), 11, upperside of C. m. rileyi (Guangdong), 12, underside of C. m. rileyi (Guangdong). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4435866" httpUri="https://zenodo.org/record/4435866/files/figure.png" pageId="2" pageNumber="115">7, 8</figureCitation>
, 
<figureCitation id="13562A2A332FFF84FEE3F8D8FE5DF8E7" box="[335,407,1841,1867]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1187,1211]" captionTargetBox="[151,1436,181,1164]" captionTargetId="figure-19@6.[151,1436,181,1164]" captionTargetPageId="6" captionText="FIGURES 13–16. Immatures of Chrysozephyrus mushaellus paolongkoui Hsu &amp; Lo, ssp. nov., 13, egg on near base of dormant buds of Lithocarpus litseifolius, 14, final (4th) instar larva (green form), 15, final (4th) instar larva (brown form), 16, pupa (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4435868" httpUri="https://zenodo.org/record/4435868/files/figure.png" pageId="2" pageNumber="115">13–16</figureCitation>
, 
<figureCitation id="13562A2A332FFF84FE0FF8D8FDDEF8E7" box="[419,532,1841,1867]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="2" pageNumber="115">17, 20, 23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776524332FFF85FF3BF890FF24FEED" lastPageId="3" lastPageNumber="116" pageId="2" pageNumber="115" type="materials_examined">
<paragraph id="8BD236AF332FFF84FF3BF890FB6BF81B" blockId="2.[151,1436,1913,2014]" pageId="2" pageNumber="115">
<emphasis id="B919EABD332FFF84FF3BF890FE85F83F" bold="true" box="[151,335,1913,1939]" pageId="2" pageNumber="115">Type materials.</emphasis>
<materialsCitation id="3B053CF2332FFF84FEF9F890FB6BF81B" ID-GBIF-Occurrence="3017385301" collectingDate="2016-01-31" collectingDateMax="2016-04-10" collectingDateMin="2016-01-31" collectionCode="IOZ" collectorName="Coll. Y. F. Hsu &amp; Y. F. Lo" country="China" county="Baisha Co." elevation="1500" location="Yinggeling" pageId="2" pageNumber="115" specimenCode="HSU 16" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<emphasis id="B919EABD332FFF84FEF9F890FE03F83F" bold="true" box="[341,457,1913,1939]" pageId="2" pageNumber="115">
<typeStatus id="54D6880D332FFF84FEF9F890FE0EF83F" box="[341,452,1913,1939]" pageId="2" pageNumber="115" type="holotype">Holotype</typeStatus>
.
</emphasis>
³: 
<collectingCountry id="F37A763F332FFF84FE5FF890FD9AF83F" box="[499,592,1913,1939]" name="China" pageId="2" pageNumber="115">CHINA</collectingCountry>
: 
<collectingRegion id="49A9F84D332FFF84FDF5F890FCEAF83F" box="[601,800,1913,1939]" country="China" name="Hainan" pageId="2" pageNumber="115">HAINAN PROV.</collectingRegion>
, 
<collectingCounty id="62B34E23332FFF84FC85F890FC6CF83F" box="[809,934,1913,1939]" pageId="2" pageNumber="115">Baisha Co.</collectingCounty>
, 
<location id="8EB26074332FFF84FC1CF890FBE5F83F" LSID="urn:lsid:plazi:treatment:03C487B9332FFF83FF3BF8E4FBA3F82A:8EB26074332FFF84FC1CF890FBE5F83F" box="[944,1071,1913,1939]" country="China" county="Baisha Co." name="Yinggeling" pageId="2" pageNumber="115" stateProvince="Hainan">Yinggeling</location>
, 
<quantity id="4C959B4A332FFF84FB95F890FB40F838" box="[1081,1162,1913,1940]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="115" unit="m" value="1500.0">
<elevation id="0040D19C332FFF84FB95F890FB40F838" box="[1081,1162,1913,1940]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="115" unit="m" value="1500.0">1500m</elevation>
</quantity>
, 
<date id="FFD3106F332FFF84FB38F890FAC5F83F" box="[1172,1295,1913,1940]" pageId="2" pageNumber="115" value="2016-01-31">
<collectingDate id="EF97E987332FFF84FB38F890FAC5F83F" box="[1172,1295,1913,1940]" pageId="2" pageNumber="115" value="2016-01-31">31. I. 2016</collectingDate>
</date>
, reared from 
<taxonomicName id="4C6D4D2C332FFF84FF3BF874FE51F81B" authorityName="Chun" authorityYear="1928" baseAuthorityName="Hance" box="[151,411,1949,1975]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="115" phylum="Tracheophyta" rank="species" species="litseifolius">
<emphasis id="B919EABD332FFF84FF3BF874FE51F81B" box="[151,411,1949,1975]" italics="true" pageId="2" pageNumber="115">Lithocarpus litseifolius</emphasis>
</taxonomicName>
(
<specimenCode id="DBCB9ED4332FFF84FE05F874FDCCF814" box="[425,518,1949,1976]" collectionCode="HSU" country="0" httpUri="http://grbio.org/cool/h9vj-e322" name="Humboldt State University" pageId="2" pageNumber="115">HSU 16</specimenCode>
A47), emgd. 
<date id="FFD3106F332FFF84FD35F874FCECF81B" box="[665,806,1949,1976]" pageId="2" pageNumber="115" value="2016-04-10">
<collectingDate id="EF97E987332FFF84FD35F874FCECF81B" box="[665,806,1949,1976]" pageId="2" pageNumber="115" value="2016-04-10">10. IV. 2016</collectingDate>
</date>
, 
<collectorName id="26985379332FFF84FC9DF874FC1FF814" box="[817,981,1949,1976]" pageId="2" pageNumber="115">Coll. Y. F. Hsu</collectorName>
&amp; 
<collectorName id="26985379332FFF84FC54F877FB9BF814" box="[1016,1105,1950,1976]" pageId="2" pageNumber="115">Y. F. Lo</collectorName>
(
<collectionCode id="ED7CAE6A332FFF84FBCCF874FB5EF81B" box="[1120,1172,1949,1975]" pageId="2" pageNumber="115">IOZ</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD236AF332FFF84FF6BF828FA56F870" blockId="2.[151,1436,1913,2014]" box="[199,1436,1985,2014]" pageId="2" pageNumber="115">
<materialsCitation id="3B053CF2332FFF84FF6BF828FA56F870" ID-GBIF-Occurrence="4433881301" box="[199,1436,1985,2014]" collectingDate="2016-04-11" collectingDateMax="2016-04-22" collectingDateMin="2016-04-11" collectionCode="IOZ" collectorName="Coll. Y. F. Hsu &amp; Y. F. Lo" country="China" county="Baisha Co." elevation="1500" location="Yinggeling" municipality="Jianfengling" pageId="2" pageNumber="115" specimenCode="HSU 15" specimenCount="3" specimenCount-female="3" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="B919EABD332FFF84FF6BF828FE8CF877" bold="true" box="[199,326,1985,2011]" pageId="2" pageNumber="115">
<typeStatus id="54D6880D332FFF84FF6BF828FE8BF877" box="[199,321,1985,2011]" pageId="2" pageNumber="115" type="paratype">Paratypes</typeStatus>
.
</emphasis>
1³ 
<specimenCount id="9D6BFD26332FFF84FEC5F82AFE59F872" box="[361,403,1987,2014]" pageId="2" pageNumber="115" type="female">3♀</specimenCount>
, same data as for holotype, emgd. 
<date id="FFD3106F332FFF84FCB6F828FC02F877" box="[794,968,1985,2012]" pageId="2" pageNumber="115" value="2016-04-11" valueMax="2016-04-22" valueMin="2016-04-11">
<collectingDate id="EF97E987332FFF84FCB6F828FC02F877" box="[794,968,1985,2012]" pageId="2" pageNumber="115" value="2016-04-11" valueMax="2016-04-22" valueMin="2016-04-11">11-22. IV. 2016</collectingDate>
</date>
(1³ dissected, genitalia preparation YFH
</materialsCitation>
</paragraph>
<paragraph id="8BD236AF332EFF85FF3BFF7FFF24FEED" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="116">
1557, DNA vouvher 160073); 2³ 
<specimenCount id="9D6BFD26332EFF85FDB9FF71FDF1FF1F" box="[533,571,152,179]" pageId="3" pageNumber="116" type="female">7♀</specimenCount>
, same locality as for holotype, 
<date id="FFD3106F332EFF85FC08FF7EFBEEFF1D" box="[932,1060,150,177]" pageId="3" pageNumber="116" value="2015-01-30">
<collectingDate id="EF97E987332EFF85FC08FF7EFBEEFF1D" box="[932,1060,150,177]" pageId="3" pageNumber="116" value="2015-01-30">30. I. 2015</collectingDate>
</date>
, reared from 
<taxonomicName id="4C6D4D2C332EFF85FB12FF7EFA99FF1D" authorityName="Chun" authorityYear="1928" baseAuthorityName="Hance" box="[1214,1363,151,177]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="116" phylum="Tracheophyta" rank="species" species="litseifolius">
<emphasis id="B919EABD332EFF85FB12FF7EFA99FF1D" box="[1214,1363,151,177]" italics="true" pageId="3" pageNumber="116">L. litseifolius</emphasis>
</taxonomicName>
(
<specimenCode id="DBCB9ED4332EFF85FACFFF7EFF78FF79" collectionCode="HSU" country="0" httpUri="http://grbio.org/cool/h9vj-e322" name="Humboldt State University" pageId="3" pageNumber="116">HSU 15</specimenCode>
A35), emgd. 
<date id="FFD3106F332EFF85FEE9FF52FE20FF79" box="[325,490,186,213]" pageId="3" pageNumber="116" value="2015-03-04" valueMax="2015-03-10" valueMin="2015-03-04">
<collectingDate id="EF97E987332EFF85FEE9FF52FE20FF79" box="[325,490,186,213]" pageId="3" pageNumber="116" value="2015-03-04" valueMax="2015-03-10" valueMin="2015-03-04">4-10. III. 2015</collectingDate>
</date>
, 
<collectorName id="26985379332EFF85FE58FF52FD5CFF79" box="[500,662,187,213]" pageId="3" pageNumber="116">Coll. Y. F. Hsu</collectorName>
&amp; 
<collectorName id="26985379332EFF85FD1BFF52FCC5FF79" box="[695,783,187,213]" pageId="3" pageNumber="116">Y. F. Lo</collectorName>
(
<collectionCode id="ED7CAE6A332EFF85FCB1FF52FC9AFF79" box="[797,848,187,213]" pageId="3" pageNumber="116">IOZ</collectionCode>
, 
<collectionCode id="ED7CAE6A332EFF85FCF6FF52FC62FF79" box="[858,936,187,213]" country="China" name="Kadoorie Farm and Botanic Garden" pageId="3" pageNumber="116" type="Herbarium">KFBG</collectionCode>
, 
<collectionCode id="ED7CAE6A332EFF85FC1EFF52FBC8FF79" box="[946,1026,187,213]" country="China" httpUri="http://grbio.org/cool/6zr8-87q6" name="National Taiwan Normal University" pageId="3" pageNumber="116">NTNU</collectionCode>
) (
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dissected, genitalia preparation YFH 1579; 
<specimenCount id="9D6BFD26332EFF85FEB2FF36FE8BFF55" box="[286,321,223,249]" pageId="3" pageNumber="116" type="female">1♀</specimenCount>
DNA vouvher 170163); 
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, 
<collectingCountry id="F37A763F332EFF85FD21FF36FD20FF55" box="[653,746,223,249]" name="China" pageId="3" pageNumber="116">CHINA</collectingCountry>
: 
<collectingRegion id="49A9F84D332EFF85FD59FF36FC77FF55" box="[757,957,223,249]" country="China" name="Hainan" pageId="3" pageNumber="116">HAINAN PROV.</collectingRegion>
, 
<collectingCounty id="62B34E23332EFF85FC64FF36FB85FF55" box="[968,1103,223,249]" pageId="3" pageNumber="116">Ledong Co.</collectingCounty>
, 
<collectingMunicipality id="6BB6ACD5332EFF85FBF7FF36FB22FF55" box="[1115,1256,223,249]" pageId="3" pageNumber="116">Jianfengling</collectingMunicipality>
, 
<quantity id="4C959B4A332EFF85FB58FF36FA8EFF55" box="[1268,1348,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="116" unit="m" value="1000.0">
<elevation id="0040D19C332EFF85FB58FF36FA8EFF55" box="[1268,1348,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="116" unit="m" value="1000.0">1000m</elevation>
</quantity>
, 
<date id="FFD3106F332EFF85FAFDFF36FF18FEB1" pageId="3" pageNumber="116" value="2006-04-18">
<collectingDate id="EF97E987332EFF85FAFDFF36FF18FEB1" pageId="3" pageNumber="116" value="2006-04-18">18. IV. 2006</collectingDate>
</date>
, 
<collectorName id="26985379332EFF85FF71FEEAFEB8FEB1" box="[221,370,259,285]" pageId="3" pageNumber="116">Coll. Y. F. Lo</collectorName>
(
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); 
<specimenCount id="9D6BFD26332EFF85FE48FEEAFDCDFEB1" box="[484,519,259,285]" pageId="3" pageNumber="116" type="female">1♀</specimenCount>
, 
<collectingCountry id="F37A763F332EFF85FDB9FEEAFDB8FEB1" box="[533,626,259,285]" name="China" pageId="3" pageNumber="116">CHINA</collectingCountry>
: 
<collectingRegion id="49A9F84D332EFF85FDD0FEEAFC89FEB1" box="[636,835,259,285]" country="China" name="Hainan" pageId="3" pageNumber="116">HAINAN PROV.</collectingRegion>
, 
<location id="8EB26074332EFF85FCE1FEEAFC19FEB1" LSID="urn:lsid:plazi:treatment:03C487B9332FFF83FF3BF8E4FBA3F82A:8EB26074332EFF85FCE1FEEAFC19FEB1" box="[845,979,259,285]" country="China" county="Ledong Co." municipality="Jianfengling" name="Ledong Co." pageId="3" pageNumber="116" stateProvince="Hainan">Ledong Co.</location>
, 
<location id="8EB26074332EFF85FC72FEEAFBA1FEB1" LSID="urn:lsid:plazi:treatment:03C487B9332FFF83FF3BF8E4FBA3F82A:8EB26074332EFF85FC72FEEAFBA1FEB1" box="[990,1131,259,285]" country="China" county="Ledong Co." municipality="Jianfengling" name="Jianfengling" pageId="3" pageNumber="116" stateProvince="Hainan">Jianfengling</location>
, 
<date id="FFD3106F332EFF85FBDAFEEAFB6FFEB1" box="[1142,1189,259,285]" pageId="3" pageNumber="116" value="1985-05-04">
4. 
<collectionCode id="ED7CAE6A332EFF85FB3DFEEAFB6FFEB1" box="[1169,1189,259,285]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="116" type="Museum">V</collectionCode>
</date>
<materialsCitation id="3B053CF2332EFF85FB0AFEEAFF20FEED" ID-GBIF-Occurrence="3017385302" collectionCode="RITF" collectorName="Coll. M. B. Gu" country="China" pageId="3" pageNumber="116" specimenCount="1" stateProvince="Hainan">
. 1985, 
<collectorName id="26985379332EFF85FB59FEEAFA56FEB1" box="[1269,1436,259,285]" pageId="3" pageNumber="116">Coll. M. B. Gu</collectorName>
(
<collectionCode id="ED7CAE6A332EFF85FF33FECEFF28FEED" box="[159,226,295,321]" pageId="3" pageNumber="116">RITF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3776524332EFF85FF6BFEA3FAFDF839" pageId="3" pageNumber="116" type="description">
<paragraph id="8BD236AF332EFF85FF6BFEA3FAD3FB45" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="116">
<emphasis id="B919EABD332EFF85FF6BFEA3FE9BFEC8" bold="true" box="[199,337,330,356]" pageId="3" pageNumber="116">Description</emphasis>
. Male (
<figureCitation id="13562A2A332EFF85FE05FEA2FDDCFEC9" box="[425,534,331,357]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1906,1930]" captionTargetBox="[163,1425,501,1873]" captionTargetId="figure-159@4.[151,1436,489,1883]" captionTargetPageId="4" captionText="FIGURES 1–6. Male adults of Chrysozephyrus mushaellus (Matsumura). 1, upperside of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., holotype, 2, underside of holotype, 3, upperside of C. m. mushaellus (Taiwan), 4, underside of C. m. mushaellus (Taiwan), 5, upperside of C. m. rileyi (Guangdong), 6, underside of C. m. rileyi (Guangdong). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4435862" httpUri="https://zenodo.org/record/4435862/files/figure.png" pageId="3" pageNumber="116">Figs. 1–2</figureCitation>
): FL 
<quantity id="4C959B4A332EFF85FDFEFEA2FD3FFEC9" box="[594,757,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" metricValueMax="1.92" metricValueMin="1.8199999999999998" pageId="3" pageNumber="116" unit="mm" value="18.7" valueMax="19.2" valueMin="18.2">18.2–19.2 mm</quantity>
(mean, 18.7 ± 
<quantity id="4C959B4A332EFF85FC33FEA2FC3DFEC9" box="[927,1015,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="116" unit="mm" value="0.4">0.4 mm</quantity>
, n = 4); AL 9.0– 
<quantity id="4C959B4A332EFF85FB1EFEA2FACDFEC9" box="[1202,1287,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="3" pageNumber="116" unit="mm" value="9.3">9.3 mm</quantity>
(mean, 9.1 ± 
<quantity id="4C959B4A332EFF85FF3BFE86FF25FE25" box="[151,239,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="116" unit="mm" value="0.2">0.2 mm</quantity>
, n = 4). Head: Hairy, vertex, frons dark brown; a narrow white rim surrounding eye; eye semi-oval, densely covered with long, buff setae; labial palpus porrect, with 3rd segment pointed downwards, covered with dark brown scalings and hairs; maxillary palpus reduced, inconspicuous; proboscis unscaled; antenna smoothly scaled, naked at terminal end of nudum, dark brown with lateral white dots on each flagellomere. Thorax: Dark brown dorsad, white ventrad; legs with tarsus of fore leg stubby; white, mottled with brown. Forewing: Termen, costa slightly convex dorsum straight. Ground color of upperside dark brown, overlaid with large area of metallic green tinged with yellow scaling proximally, covering most of forewing, leaving a dark brown area apicaly and margin along termen. Fringe white, prominent. Ground color of underside buff. Discal spot forming brown bar edged with brown. Distal band of central symmetry system represented as tilted white line edged with brown proximally, strongly dislocated inwards in cell Cu 
<subScript id="17E934EA332EFF85FEC8FD2CFEA6FD7F" attach="left" box="[356,364,709,723]" fontSize="6" pageId="3" pageNumber="116">2</subScript>
. Submarginal band and &quot;g&quot;-element (per 
<bibRefCitation id="EFFC4B5E332EFF85FCEDFD5AFC1CFD61" author="Nijhout, H. F." box="[833,982,691,717]" pageId="3" pageNumber="116" refId="ref4899" refString="Nijhout, H. F. (1991) The development and evolution of butterfly wing patterns. Smithsonian Institution Press, Washington and London, xvi + 297 pp." type="book" year="1991">Nijhout 1991</bibRefCitation>
) fused into prominent, dark brown band edged with white. Fringe white, prominent. Hindwing: Contour of wing slightly produced at distal end of M 
<subScript id="17E934EA332EFF85FAC8FD00FAA6FD5B" attach="left" box="[1380,1388,745,759]" fontSize="6" pageId="3" pageNumber="116">1</subScript>
and Cu 
<subScript id="17E934EA332EFF85FF14FCE4FF0AFCB7" attach="left" box="[184,192,781,795]" fontSize="6" pageId="3" pageNumber="116">1</subScript>
; Cu 
<subScript id="17E934EA332EFF85FF5CFCE4FF32FCB7" attach="left" box="[240,248,781,795]" fontSize="6" pageId="3" pageNumber="116">2</subScript>
bearing long, tail-like projection. Metallic scaling on upperside less extensive than that of forewing, with dark brown areas including a patch along costa, narrow bands along termen and dorsum. Narrow, metallic blue line at distal ends of cells along termen. Ground color of underside buff. Discal spot forming brown bar edged with white. Distal band of central symmetry system forming prominent white line edged with dark brown proximally, forming a roughly straight line from dorsum to vein Cu 
<subScript id="17E934EA332EFF85FD5BFC74FD35FC07" attach="left" box="[759,767,925,939]" fontSize="6" pageId="3" pageNumber="116">1</subScript>
, running outwards in cell Cu 
<subScript id="17E934EA332EFF85FB93FC74FB8DFC07" attach="left" box="[1087,1095,925,939]" fontSize="6" pageId="3" pageNumber="116">1</subScript>
, dislocated proximally as tickshaped or a straight line in cell Cu 
<subScript id="17E934EA332EFF85FDBEFC28FDD0FC63" attach="left" box="[530,538,961,975]" fontSize="6" pageId="3" pageNumber="116">2</subScript>
, retracing into straight line in cell 1A+2A. Submarginal band consisting of faint, broad, white band, a black, rounded spot enclosed within orange circle in cell Cu 
<subScript id="17E934EA332EFF85FB8DFC0CFBE3FC5F" attach="left" box="[1057,1065,997,1011]" fontSize="6" pageId="3" pageNumber="116">1</subScript>
, and a tornal orange patch edged by black and metallic blue scaling anteriorly and posteriorly; &quot;g&quot;-element consisting of a string of arch-like, faint white markings. Fringe white, prominent. Abdomen: Dark brown dorsally, gray laterally, white ventrally. 
<emphasis id="B919EABD332EFF85FA82FBF3FA56FB98" bold="true" box="[1326,1436,1050,1076]" pageId="3" pageNumber="116">Genitalia</emphasis>
(
<figureCitation id="13562A2A332EFF85FF33FBD6FE9AFBF5" box="[159,336,1087,1113]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="3" pageNumber="116">Figs. 17, 20, 23</figureCitation>
): Ring-shaped sclerites of 9+10 segments with posterior end nearly straight; saccus large, flap-like; brachium (falx) flatten, dagger-axe like, bearing a prominent distal spine pointing posteriad, serrate along anterior margin; valva with harpe as broad lobe; ampulla stout basally, with distal end forming down-curved, digitate process. Juxta broad, V-shaped. Phallus elongate, with aedeagus longer than phallobase. Aedeagus up-rasing, tubular, slightly enlarged at both ends; distal end angled with dorsal opening, signa as a patch of minute spines.
</paragraph>
<paragraph id="8BD236AF332EFF85FF6BFB1AFB46F981" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="116">
<emphasis id="B919EABD332EFF85FF6BFB1AFED4FAA1" bold="true" box="[199,286,1267,1293]" pageId="3" pageNumber="116">Female</emphasis>
(
<figureCitation id="13562A2A332EFF85FE82FB1AFE51FAA1" box="[302,411,1267,1293]" captionStart="FIGURES 7–12" captionStartId="5.[151,264,1578,1602]" captionTargetBox="[158,1422,200,1540]" captionTargetId="figure-17@5.[151,1436,181,1554]" captionTargetPageId="5" captionText="FIGURES 7–12. Female adults of Chrysozephyrus mushaellus (Matsumura). 7, upperside of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., paratype, 8, underside of paratype, 9, upperside of C. m. mushaellus (Taiwan), 10, underside of C. m. mushaellus (Taiwan), 11, upperside of C. m. rileyi (Guangdong), 12, underside of C. m. rileyi (Guangdong). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4435866" httpUri="https://zenodo.org/record/4435866/files/figure.png" pageId="3" pageNumber="116">Figs. 7–8</figureCitation>
). FL 
<quantity id="4C959B4A332EFF85FE74FB1AFDB6FAA1" box="[472,636,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="1.875" metricValueMax="1.93" metricValueMin="1.8199999999999998" pageId="3" pageNumber="116" unit="mm" value="18.75" valueMax="19.3" valueMin="18.2">18.2–19.3 mm</quantity>
(mean, 18.9 ± 
<quantity id="4C959B4A332EFF85FC8BFB1AFC4AFAA1" box="[807,896,1267,1293]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="116" unit="mm" value="0.4">0.4 mm</quantity>
, n = 10); AL 
<quantity id="4C959B4A332EFF85FBB5FB1AFB6BFAA1" box="[1049,1185,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="8.899999999999999" metricValueMax="9.2" metricValueMin="8.6" pageId="3" pageNumber="116" unit="mm" value="8.899999999999999" valueMax="9.2" valueMin="8.6">8.6–9.2 mm</quantity>
(mean, 9.0 ± 
<quantity id="4C959B4A332EFF85FA93FB1AFA52FAA1" box="[1343,1432,1267,1293]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="116" unit="mm" value="0.2">0.2 mm</quantity>
, n = 10). Body, wing patterns of underside as described for male except ground color darker. Fore leg with tarsus segmented. Wing upperside ground color brown with prominent, proximal, metallic patch blue tinged with purple on forewing. Metallic blue line at distal ends of cells along termen more prominent than that of male. 
<emphasis id="B919EABD332EFF85FA82FAB7FA56FAD4" bold="true" box="[1326,1436,1374,1400]" pageId="3" pageNumber="116">Genitalia</emphasis>
(Fig. 26): Corpus bursae ovate, bearing a pair of invaginated, horn-like signa on opposite walls. Ductus bursae largely membraneous, with posterior end containing internal sclerotized band. Sterigma forming a weakly sclerotized, dome-like structure; a funnel-shaped sclerite present around ostium bursae with a pair of ventral lobes with rounded, posterior extentions. Apophyses posteriores elongate, with posterior end abruptly flattened to sclerotized band. Papillae anales lobate, setose, weakly sclerotized near base of apophyses posteriores
</paragraph>
<paragraph id="8BD236AF332EFF85FF6BF9DEFAFDF839" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="116">
<emphasis id="B919EABD332EFF85FF6BF9DEFE81F9FD" bold="true" box="[199,331,1591,1617]" pageId="3" pageNumber="116">Immatures</emphasis>
. Egg (
<figureCitation id="13562A2A332EFF85FE3FF9DEFE29F9FD" box="[403,483,1591,1617]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1187,1211]" captionTargetBox="[151,1436,181,1164]" captionTargetId="figure-19@6.[151,1436,181,1164]" captionTargetPageId="6" captionText="FIGURES 13–16. Immatures of Chrysozephyrus mushaellus paolongkoui Hsu &amp; Lo, ssp. nov., 13, egg on near base of dormant buds of Lithocarpus litseifolius, 14, final (4th) instar larva (green form), 15, final (4th) instar larva (brown form), 16, pupa (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4435868" httpUri="https://zenodo.org/record/4435868/files/figure.png" pageId="3" pageNumber="116">Fig. 13</figureCitation>
) approximately 0.67 ± 
<quantity id="4C959B4A332EFF85FD4AF9DEFC83F9FD" box="[742,841,1591,1617]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="116" unit="mm" value="0.04">0.04 mm</quantity>
in diameter, 0.38 ± 
<quantity id="4C959B4A332EFF85FB89F9DEFB42F9FD" box="[1061,1160,1591,1617]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="116" unit="mm" value="0.02">0.02 mm</quantity>
in height (n = 28), hemispherical but compressed, white, surface with sculpture forming framework with regularly arranged, short, stub-like processes. Larvae (
<figureCitation id="13562A2A332EFF85FEDCF996FE32F935" box="[368,504,1662,1689]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1187,1211]" captionTargetBox="[151,1436,181,1164]" captionTargetId="figure-19@6.[151,1436,181,1164]" captionTargetPageId="6" captionText="FIGURES 13–16. Immatures of Chrysozephyrus mushaellus paolongkoui Hsu &amp; Lo, ssp. nov., 13, egg on near base of dormant buds of Lithocarpus litseifolius, 14, final (4th) instar larva (green form), 15, final (4th) instar larva (brown form), 16, pupa (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4435868" httpUri="https://zenodo.org/record/4435868/files/figure.png" pageId="3" pageNumber="116">Figs. 14–15</figureCitation>
) with four instars. Head dark brown, glossy. Body slug-like, of typically lycaenid form, with surface bearing transparent setae; T1 with anterior margin slightly concave medially, anal lobe with posterior margin rounded. Coloration of body green or brown. White or yellow chevrons present subdorsally, dots of same color present throughout body. Double, medial, longitudinal white or yellow lines present by dorsal vessel. T1 shield trapezoid. Spiracles dark brown. Full-grown larva 17.08 ± 
<quantity id="4C959B4A332EFF85FCC5F8E6FC06F885" box="[873,972,1807,1833]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="116" unit="mm" value="0.6">0.60 mm</quantity>
in length (n = 6). Pupae (
<figureCitation id="13562A2A332EFF85FB4FF8E6FAFCF886" box="[1251,1334,1807,1834]" captionStart="FIGURES 13–16" captionStartId="6.[151,264,1187,1211]" captionTargetBox="[151,1436,181,1164]" captionTargetId="figure-19@6.[151,1436,181,1164]" captionTargetPageId="6" captionText="FIGURES 13–16. Immatures of Chrysozephyrus mushaellus paolongkoui Hsu &amp; Lo, ssp. nov., 13, egg on near base of dormant buds of Lithocarpus litseifolius, 14, final (4th) instar larva (green form), 15, final (4th) instar larva (brown form), 16, pupa (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4435868" httpUri="https://zenodo.org/record/4435868/files/figure.png" pageId="3" pageNumber="116">Fig. 16</figureCitation>
): 11.47 ± 
<quantity id="4C959B4A332EFF85FF3BF8DAFF31F8E1" box="[151,251,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="116" unit="mm" value="0.34">0.34 mm</quantity>
in length, 4.57 ± 
<quantity id="4C959B4A332EFF85FE13F8DAFDE9F8E1" box="[447,547,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="116" unit="mm" value="0.14">0.14 mm</quantity>
in width of T2, 5.48 ± 
<quantity id="4C959B4A332EFF85FC88F8DAFC4DF8E1" box="[804,903,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="116" unit="mm" value="0.1">0.10 mm</quantity>
in width of A3, 4.62 ± 
<quantity id="4C959B4A332EFF85FB26F8DAFB24F8E1" box="[1162,1262,1843,1869]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="116" unit="mm" value="0.17">0.17 mm</quantity>
in height of A3 (n = 6); of typical lycaenid form, brown tinged with yellow, mottled with dark brown dorsally, a dark brown medial line present dorsad on abdomen; wings with more extensive dark brown markings; spiracles cream white.
</paragraph>
</subSubSection>
<subSubSection id="C3776524332EFF82FF6BF877FD12FE7D" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116" type="diagnosis">
<paragraph id="8BD236AF332EFF82FF6BF877FD75FE01" blockId="3.[151,1437,150,2013]" lastBlockId="4.[151,1437,150,465]" lastPageId="4" lastPageNumber="117" pageId="3" pageNumber="116">
<emphasis id="B919EABD332EFF85FF6BF877FE88F814" bold="true" box="[199,322,1950,1976]" pageId="3" pageNumber="116">Diagnosis.</emphasis>
The taxon 
<emphasis id="B919EABD332EFF85FE10F876FD82F815" box="[444,584,1951,1977]" italics="true" pageId="3" pageNumber="116">paolongkoui</emphasis>
<emphasis id="B919EABD332EFF85FDE2F876FD7BF815" bold="true" box="[590,689,1951,1977]" pageId="3" pageNumber="116">ssp. nov.</emphasis>
can be distinguished from other subspecies of 
<taxonomicName id="4C6D4D2C332EFF85FB19F876FA98F815" baseAuthorityName="Matsumura" baseAuthorityYear="1938" box="[1205,1362,1951,1977]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="mushaellus">
<emphasis id="B919EABD332EFF85FB19F876FA98F815" box="[1205,1362,1951,1977]" italics="true" pageId="3" pageNumber="116">C. mushaellus</emphasis>
</taxonomicName>
by the following features: 1) prominent dark brown apical area is present in male 
<emphasis id="B919EABD332EFF85FC77F82AFBADF871" box="[987,1127,1987,2013]" italics="true" pageId="3" pageNumber="116">paolongkoui</emphasis>
(
<figureCitation id="13562A2A332EFF85FBDAF82AFB7CF871" box="[1142,1206,1987,2013]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1906,1930]" captionTargetBox="[163,1425,501,1873]" captionTargetId="figure-159@4.[151,1436,489,1883]" captionTargetPageId="4" captionText="FIGURES 1–6. Male adults of Chrysozephyrus mushaellus (Matsumura). 1, upperside of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., holotype, 2, underside of holotype, 3, upperside of C. m. mushaellus (Taiwan), 4, underside of C. m. mushaellus (Taiwan), 5, upperside of C. m. rileyi (Guangdong), 6, underside of C. m. rileyi (Guangdong). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4435862" httpUri="https://zenodo.org/record/4435862/files/figure.png" pageId="3" pageNumber="116">Fig. 1</figureCitation>
) but not in the other subspecies (
<figureCitation id="13562A2A3329FF82FEB2FF7EFEB6FF1D" box="[286,380,150,177]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1906,1930]" captionTargetBox="[163,1425,501,1873]" captionTargetId="figure-159@4.[151,1436,489,1883]" captionTargetPageId="4" captionText="FIGURES 1–6. Male adults of Chrysozephyrus mushaellus (Matsumura). 1, upperside of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., holotype, 2, underside of holotype, 3, upperside of C. m. mushaellus (Taiwan), 4, underside of C. m. mushaellus (Taiwan), 5, upperside of C. m. rileyi (Guangdong), 6, underside of C. m. rileyi (Guangdong). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4435862" httpUri="https://zenodo.org/record/4435862/files/figure.png" pageId="4" pageNumber="117">Fig. 3, 5</figureCitation>
); 2) brachium (falx) of 
<emphasis id="B919EABD3329FF82FD2DFF7EFCC7FF1D" box="[641,781,151,177]" italics="true" pageId="4" pageNumber="117">paolongkoui</emphasis>
with length of distal spine is slender and shorter than basal portion of brachum (
<figureCitation id="13562A2A3329FF82FED1FF52FE05FF79" box="[381,463,187,213]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="4" pageNumber="117">Fig. 17</figureCitation>
), whereas that of the nominotypical subspecies is elongate, longer than basal portion of brachim (
<figureCitation id="13562A2A3329FF82FEAEFF36FE9FFF55" box="[258,341,223,249]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="4" pageNumber="117">Fig. 18</figureCitation>
); that of continental subspecies is stout and short (
<figureCitation id="13562A2A3329FF82FC28FF36FC1DFF55" box="[900,983,223,249]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="4" pageNumber="117">Fig. 19</figureCitation>
); 3) ampulla of valva is bent downwards in 
<emphasis id="B919EABD3329FF82FF18FEEAFE8AFEB1" box="[180,320,259,285]" italics="true" pageId="4" pageNumber="117">paolongkoui</emphasis>
(
<figureCitation id="13562A2A3329FF82FEFCFEEAFE6EFEB1" box="[336,420,259,285]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="4" pageNumber="117">Fig. 20</figureCitation>
), in contrast to curved downwars in the nominotypical subspecies (
<figureCitation id="13562A2A3329FF82FB35FEEAFB21FEB1" box="[1177,1259,259,285]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="4" pageNumber="117">Fig. 21</figureCitation>
) and straight in continental subspecies (
<figureCitation id="13562A2A3329FF82FE0FFECEFE3DFEED" box="[419,503,295,321]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1001,1025]" captionTargetBox="[231,1355,181,973]" captionTargetId="figure-17@7.[227,1360,181,977]" captionTargetPageId="7" captionText="FIGURES 17–25. Male genitalia of Chrysozephyrus mushaellus (Matsumura). 17, tegumen of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 18, teguman of C. m. mushaellus, 19, tegumen of C. m. rileyi (Guangdong), 20, left valvae of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 21, left valvae of C. m. mushaellus, 22, leftvalvae of C. m. rileyi (Guangdong), 23, phallus of C. m. paolongkoui Hsu &amp; Lo, ssp. nov., 24, phallus of C. m. mushaellus, 25, phallus of C. m. rileyi (Guangdong)." figureDoi="http://doi.org/10.5281/zenodo.4435870" httpUri="https://zenodo.org/record/4435870/files/figure.png" pageId="4" pageNumber="117">Fig. 22</figureCitation>
); 4) ventral lobes of sclerite surrounding ostium bursae forming rounded extension in 
<emphasis id="B919EABD3329FF82FF18FEA2FE8AFEC9" box="[180,320,331,357]" italics="true" pageId="4" pageNumber="117">paolongkoui</emphasis>
(Fig. 26), whereas there is no extention in other subspecies (Figs. 27–28); 5) sclerotized portion of ductus bursae is shorter than width of sclerite around ostium bursae in 
<emphasis id="B919EABD3329FF82FC15FE86FB8FFE25" box="[953,1093,367,393]" italics="true" pageId="4" pageNumber="117">paolongkoui</emphasis>
(Fig. 26), whereas it is longer than the sclerite in other subspecies (Fig. 27–28).
</paragraph>
<paragraph id="8BD236AF3329FF82FF6BFE5EFD12FE7D" blockId="4.[151,1437,150,465]" box="[199,728,439,465]" pageId="4" pageNumber="117">
<emphasis id="B919EABD3329FF82FF6BFE5EFE8FFE7D" bold="true" box="[199,325,439,465]" pageId="4" pageNumber="117">Hostplant.</emphasis>
<taxonomicName id="4C6D4D2C3329FF82FEE7FE5EFD1EFE7D" authorityName="Chun" authorityYear="1928" baseAuthorityName="Hance" box="[331,724,439,465]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="117" phylum="Tracheophyta" rank="species" species="litseifolius">
<emphasis id="B919EABD3329FF82FEE7FE5EFD85FE7D" box="[331,591,439,465]" italics="true" pageId="4" pageNumber="117">Lithocarpus litseifolius</emphasis>
(Fagaceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF1266273329FF82FF3BF89BFB56F87E" ID-DOI="http://doi.org/10.5281/zenodo.4435862" ID-Zenodo-Dep="4435862" httpUri="https://zenodo.org/record/4435862/files/figure.png" pageId="4" pageNumber="117" startId="4.[151,264,1906,1930]" targetBox="[163,1425,501,1873]" targetPageId="4">
<paragraph id="8BD236AF3329FF82FF3BF89BFB56F87E" blockId="4.[151,1437,1906,2002]" pageId="4" pageNumber="117">
<emphasis id="B919EABD3329FF82FF3BF89BFEF6F826" bold="true" box="[151,316,1906,1930]" pageId="4" pageNumber="117">FIGURES 1–6.</emphasis>
Male adults of 
<taxonomicName id="4C6D4D2C3329FF82FE4CF89BFC46F826" authority="(Matsumura)" baseAuthorityName="Matsumura" baseAuthorityYear="1938" box="[480,908,1906,1930]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="mushaellus">
<emphasis id="B919EABD3329FF82FE4CF89BFD37F826" box="[480,765,1906,1930]" italics="true" pageId="4" pageNumber="117">Chrysozephyrus mushaellus</emphasis>
(Matsumura)
</taxonomicName>
. 1, upperside of 
<emphasis id="B919EABD3329FF82FB95F89BFB3DF826" box="[1081,1271,1906,1930]" italics="true" pageId="4" pageNumber="117">C. m. paolongkoui</emphasis>
Hsu &amp; Lo, 
<emphasis id="B919EABD3329FF82FAD8F89BFF09F802" bold="true" pageId="4" pageNumber="117">ssp. nov.</emphasis>
, holotype, 2, underside of holotype, 3, upperside of 
<emphasis id="B919EABD3329FF82FD7DF87FFCCDF802" box="[721,775,1942,1966]" italics="true" pageId="4" pageNumber="117">C. m.</emphasis>
<taxonomicName id="4C6D4D2C3329FF82FCA1F87FFC28F802" authority="(Taiwan)" baseAuthorityName="Taiwan" baseAuthorityYear="1938" box="[781,994,1942,1966]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="mushaellus">
<emphasis id="B919EABD3329FF82FCA1F87FFCB5F802" box="[781,895,1942,1966]" italics="true" pageId="4" pageNumber="117">mushaellus</emphasis>
(Taiwan)
</taxonomicName>
, 4, underside of 
<emphasis id="B919EABD3329FF82FB24F87FFB74F802" box="[1160,1214,1942,1966]" italics="true" pageId="4" pageNumber="117">C. m.</emphasis>
<taxonomicName id="4C6D4D2C3329FF82FB6FF87FFA52F802" authority="(Taiwan)" baseAuthorityName="Taiwan" baseAuthorityYear="1938" box="[1219,1432,1942,1966]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="mushaellus">
<emphasis id="B919EABD3329FF82FB6FF87FFAFFF802" box="[1219,1333,1942,1966]" italics="true" pageId="4" pageNumber="117">mushaellus</emphasis>
(Taiwan)
</taxonomicName>
, 5, upperside of 
<emphasis id="B919EABD3329FF82FE99F853FEAEF87E" box="[309,356,1978,2002]" italics="true" pageId="4" pageNumber="117">C. m</emphasis>
. 
<emphasis id="B919EABD3329FF82FEDDF853FE6CF87E" box="[369,422,1978,2002]" italics="true" pageId="4" pageNumber="117">rileyi</emphasis>
(Guangdong), 6, underside of 
<emphasis id="B919EABD3329FF82FD72F853FC9AF87E" box="[734,848,1978,2002]" italics="true" pageId="4" pageNumber="117">C. m. rileyi</emphasis>
(Guangdong). Scale bar = 1 cm.
</paragraph>
</caption>
<caption id="DF1266273328FF83FF3BF9C3FB7FF926" ID-DOI="http://doi.org/10.5281/zenodo.4435866" ID-Zenodo-Dep="4435866" httpUri="https://zenodo.org/record/4435866/files/figure.png" pageId="5" pageNumber="118" startId="5.[151,264,1578,1602]" targetBox="[158,1422,200,1540]" targetPageId="5">
<paragraph id="8BD236AF3328FF83FF3BF9C3FB7FF926" blockId="5.[151,1437,1578,1674]" pageId="5" pageNumber="118">
<emphasis id="B919EABD3328FF83FF3BF9C3FE8DF9EE" bold="true" box="[151,327,1578,1602]" pageId="5" pageNumber="118">FIGURES 7–12.</emphasis>
Female adults of 
<taxonomicName id="4C6D4D2C3328FF83FE55F9C3FC6BF9EE" authority="(Matsumura)" baseAuthorityName="Matsumura" baseAuthorityYear="1938" box="[505,929,1578,1602]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="mushaellus">
<emphasis id="B919EABD3328FF83FE55F9C3FCDEF9EE" box="[505,788,1578,1602]" italics="true" pageId="5" pageNumber="118">Chrysozephyrus mushaellus</emphasis>
(Matsumura)
</taxonomicName>
. 7, upperside of 
<emphasis id="B919EABD3328FF83FBE9F9C2FB35F9EE" box="[1093,1279,1578,1602]" italics="true" pageId="5" pageNumber="118">C. m. paolongkoui</emphasis>
Hsu &amp; Lo, 
<emphasis id="B919EABD3328FF83FAD8F9C3FA57F9EE" bold="true" box="[1396,1437,1578,1602]" pageId="5" pageNumber="118">ssp.</emphasis>
nov., paratype, 8, underside of paratype, 9, upperside of 
<emphasis id="B919EABD3328FF83FD65F9A6FD35F9CA" box="[713,767,1615,1638]" italics="true" pageId="5" pageNumber="118">C. m.</emphasis>
<taxonomicName id="4C6D4D2C3328FF83FCAFF9A7FC12F9CA" authority="(Taiwan)" baseAuthorityName="Taiwan" baseAuthorityYear="1938" box="[771,984,1614,1638]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="mushaellus">
<emphasis id="B919EABD3328FF83FCAFF9A7FCBFF9CA" box="[771,885,1614,1638]" italics="true" pageId="5" pageNumber="118">mushaellus</emphasis>
(Taiwan)
</taxonomicName>
, 10, underside of 
<emphasis id="B919EABD3328FF83FB25F9A6FB75F9CA" box="[1161,1215,1615,1638]" italics="true" pageId="5" pageNumber="118">C. m.</emphasis>
<taxonomicName id="4C6D4D2C3328FF83FB6FF9A7FA52F9CA" authority="(Taiwan)" baseAuthorityName="Taiwan" baseAuthorityYear="1938" box="[1219,1432,1614,1638]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="mushaellus">
<emphasis id="B919EABD3328FF83FB6FF9A7FAFFF9CA" box="[1219,1333,1614,1638]" italics="true" pageId="5" pageNumber="118">mushaellus</emphasis>
(Taiwan)
</taxonomicName>
, 11, upperside of C. 
<emphasis id="B919EABD3328FF83FEF2F99AFEBAF926" box="[350,368,1651,1674]" italics="true" pageId="5" pageNumber="118">m</emphasis>
. 
<emphasis id="B919EABD3328FF83FED1F99BFE78F926" box="[381,434,1650,1674]" italics="true" pageId="5" pageNumber="118">rileyi</emphasis>
(Guangdong), 12, underside of 
<emphasis id="B919EABD3328FF83FD5BF99AFCC2F926" box="[759,776,1651,1674]" italics="true" pageId="5" pageNumber="118">C</emphasis>
. 
<emphasis id="B919EABD3328FF83FCB8F99AFCECF926" box="[788,806,1651,1674]" italics="true" pageId="5" pageNumber="118">m</emphasis>
. 
<emphasis id="B919EABD3328FF83FC9FF99BFCA2F926" box="[819,872,1650,1674]" italics="true" pageId="5" pageNumber="118">rileyi</emphasis>
(Guangdong). Scale bar = 1 cm.
</paragraph>
</caption>
<subSubSection id="C37765243328FF83FF6BF951FE44F892" pageId="5" pageNumber="118" type="distribution">
<paragraph id="8BD236AF3328FF83FF6BF951FBDAF95A" blockId="5.[151,1436,1720,1927]" pageId="5" pageNumber="118">
<emphasis id="B919EABD3328FF83FF6BF951FEABF97E" bold="true" box="[199,353,1720,1746]" pageId="5" pageNumber="118">Distribution.</emphasis>
Known from montane zone of 
<collectingRegion id="49A9F84D3328FF83FD67F951FCD6F97E" box="[715,796,1720,1746]" country="China" name="Hainan" pageId="5" pageNumber="118">Hainan</collectingRegion>
approximately 
<quantity id="4C959B4A3328FF83FC63F951FBA6F97F" box="[975,1132,1720,1747]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="5" pageNumber="118" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">1000– 1500m</quantity>
in elevation. The distributional range of 
<taxonomicName id="4C6D4D2C3328FF83FEEDF934FE28F95A" box="[321,482,1756,1783]" class="Insecta" family="Lycaenidae" genus="Chrysozephyrus" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="muchaellus">
<emphasis id="B919EABD3328FF83FEEDF934FE28F95A" box="[321,482,1756,1783]" italics="true" pageId="5" pageNumber="118">C. muchaellus</emphasis>
</taxonomicName>
including this new subspecies is given in 
<figureCitation id="13562A2A3328FF83FC15F935FBC1F95A" box="[953,1035,1756,1782]" captionStart="FIGURE 29" captionStartId="8.[151,250,1086,1110]" captionTargetBox="[161,1425,187,1051]" captionTargetId="figure-19@8.[151,1436,181,1062]" captionTargetPageId="8" captionText="FIGURE 29. Known distribution of Chrysozephyrus mushaellus (Matsumura), with no subspecies specified (red dots). Data sources: Uchida (1999), Koiwaya (2007) and NTNU specimens." figureDoi="http://doi.org/10.5281/zenodo.4435874" httpUri="https://zenodo.org/record/4435874/files/figure.png" pageId="5" pageNumber="118">Fig. 29</figureCitation>
.
</paragraph>
<paragraph id="8BD236AF3328FF83FF6BF8E9FE44F892" blockId="5.[151,1436,1720,1927]" pageId="5" pageNumber="118">
<emphasis id="B919EABD3328FF83FF6BF8E9FE83F8B6" bold="true" box="[199,329,1792,1818]" pageId="5" pageNumber="118">Phenology.</emphasis>
Univoltine. Adults are on wing from April to May. Overwintering as egg near bases of dormant buds on the hostplant.
</paragraph>
</subSubSection>
<subSubSection id="C37765243328FF83FF6BF8A1FBA3F82A" pageId="5" pageNumber="118" type="etymology">
<paragraph id="8BD236AF3328FF83FF6BF8A1FBA3F82A" blockId="5.[151,1436,1720,1927]" pageId="5" pageNumber="118">
<emphasis id="B919EABD3328FF83FF6BF8A1FE84F8CE" bold="true" box="[199,334,1864,1890]" pageId="5" pageNumber="118">Etymology.</emphasis>
The subspecific name is derived from 
<emphasis id="B919EABD3328FF83FCAAF8A1FC5EF8CE" box="[774,916,1864,1890]" italics="true" pageId="5" pageNumber="118">Paolongkou,</emphasis>
a mythological figure who is believed to give birth of the Li people indigenous to 
<collectingRegion id="49A9F84D3328FF83FD87F885FDB6F82A" box="[555,636,1900,1926]" country="China" name="Hainan" pageId="5" pageNumber="118">Hainan</collectingRegion>
Island according to some legendary stories.
</paragraph>
</subSubSection>
</treatment>
</document>