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<document ID-DOI="10.11646/zootaxa.5258.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7773275" ID-ZooBank="D5B78BB6-E7AB-4CB8-B202-AA617E06F915" approvalRequired="9" approvalRequired_for_taxonomicNames="5" approvalRequired_for_textStreams="1" approvalRequired_for_treatments="3" checkinTime="1679898889705" checkinUser="plazi" docAuthor="Srinivasan, Pandiarajan, Sivaruban, T., Barathy, S. &amp; Isack, Rajasekaran" docDate="2023" docId="03C41A4F5D0360032A804489CB992F9A" docLanguage="en" docName="zootaxa.5258.1.2.pdf" docOrigin="Zootaxa 5258 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5258.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clypeocaenis kaveri Balasubramanian and Muthukatturaja 2021" docType="treatment" docVersion="2" lastPageNumber="71" masterDocId="FFFD62375D1C60232A174607CF3D287F" masterDocTitle="Contribution to the knowledge of family Caenidae (Insecta: Ephemeroptera) from Madurai District, Tamil Nadu" masterLastPageNumber="75" masterPageNumber="39" pageNumber="70" updateTime="1679899625077" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Contribution to the knowledge of family Caenidae (Insecta: Ephemeroptera) from Madurai District, Tamil Nadu</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Srinivasan, Pandiarajan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8118-3256</mods:nameIdentifier>
<mods:affiliation>PG &amp; Research department of Zoology, The American College (affiliated to Madurai Kamaraj University), Madurai- 625002, India. Department of Zoology, Fatima College (affiliated to Madurai Kamaraj University), Madurai- 625018, India. barathyruban @ gmail. com; https: // orcid. org / 0000 - 0002 - 9464 - 6464 &amp; srini 15.05.1996 @ gmail. com; https: // orcid. org / 0000 - 0001 - 8118 - 3256</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sivaruban, T.</mods:namePart>
<mods:affiliation>PG &amp; Research department of Zoology, The American College (affiliated to Madurai Kamaraj University), Madurai- 625002, India. Department of Zoology, Fatima College (affiliated to Madurai Kamaraj University), Madurai- 625018, India. barathyruban @ gmail. com; https: // orcid. org / 0000 - 0002 - 9464 - 6464</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Barathy, S.</mods:namePart>
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<mods:nameIdentifier type="email">barathyruban@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Isack, Rajasekaran</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-9952-4335</mods:nameIdentifier>
<mods:affiliation>PG &amp; Research department of Zoology, The American College (affiliated to Madurai Kamaraj University), Madurai- 625002, India. Department of Zoology, Fatima College (affiliated to Madurai Kamaraj University), Madurai- 625018, India. barathyruban @ gmail. com; https: // orcid. org / 0000 - 0002 - 9464 - 6464 &amp; iceisack 143 @ gmail. com; https: // orcid. org / 0000 - 0002 - 9952 - 4335</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-27</mods:number>
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<mods:number>5258</mods:number>
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<mods:number>1</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7773228" ID-Zenodo-Dep="7773228" LSID="urn:lsid:plazi:treatment:03C41A4F5D0360032A804489CB992F9A" httpUri="http://treatment.plazi.org/id/03C41A4F5D0360032A804489CB992F9A" lastPageId="32" lastPageNumber="71" pageId="31" pageNumber="70">
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<paragraph blockId="31.[151,965,654,717]" box="[151,965,654,681]" pageId="31" pageNumber="70">
<heading bold="true" box="[151,965,654,681]" fontSize="11" level="1" pageId="31" pageNumber="70" reason="1">
<taxonomicName authority="Balasubramanian and Muthukatturaja, 2021" authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[151,965,654,681]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="70" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis bold="true" box="[151,965,654,681]" pageId="31" pageNumber="70">
<emphasis bold="true" box="[151,393,654,681]" italics="true" pageId="31" pageNumber="70">Clypeocaenis kaveri</emphasis>
<bibRefCitation author="Balasubramanian, C. &amp; Muthukatturaja, M." box="[400,965,654,681]" pageId="31" pageNumber="70" pagination="377 - 388" refId="ref17247" refString="Balasubramanian, C. &amp; Muthukatturaja, M. (2021) Two additional new species of Clypeocaenis Soldan, 1978 (Ephemeroptera: Caenidae) from the Western Ghats of Peninsular India. Zootaxa, 4915 (3), 377 - 388. https: // doi. org / 10.11646 / zootaxa. 4915.3.6" type="journal article" year="2021">Balasubramanian and Muthukatturaja, 2021</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection box="[151,330,690,717]" pageId="31" pageNumber="70" type="description">
<paragraph blockId="31.[151,965,654,717]" box="[151,330,690,717]" pageId="31" pageNumber="70">
(
<figureCitation box="[159,322,690,717]" captionStart-0="FIGURES 157159" captionStart-1="FIGURES 160163" captionStart-2="FIGURES 164167" captionStart-3="FIGURES 168171" captionStartId-0="31.[152,270,1961,1985]" captionStartId-1="32.[152,270,1673,1697]" captionStartId-2="33.[152,270,1035,1059]" captionStartId-3="33.[152,270,1976,2000]" captionTargetBox-0="[151,1436,1029,1937]" captionTargetBox-1="[151,1436,741,1649]" captionTargetBox-2="[209,1381,181,1011]" captionTargetBox-3="[204,1383,1119,1952]" captionTargetId-0="figure-260@31.[151,1436,1029,1937]" captionTargetId-1="figure-244@32.[151,1436,741,1649]" captionTargetId-2="figure-19@33.[207,1381,181,1011]" captionTargetId-3="figure-62@33.[204,1383,1119,1952]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="33" captionText-0="FIGURES 157159. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (157) female imago; (158) abdominal terga of female imago; (159) larva." captionText-1="FIGURES 160163. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (160) prosternal triangle; (161) foreleg of female subimaginal exuvia; (162) microtichia in foretibia of female subimaginal exuvia; (163) microtichia in foretarsus of female subimaginal exuvia." captionText-2="FIGURES 164167. Larva of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (164) dorsal view of tergalius II; (165) SEM view of microtrichia in tergalius II; (166) shape of sternum IX; (167) detail of shagreen field in sternum IX." captionText-3="FIGURES 168171. SEM view of eggs of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (168) entire view; (169, 170) epithema; (171) micropyle." figureDoi-0="http://doi.org/10.5281/zenodo.7773357" figureDoi-1="http://doi.org/10.5281/zenodo.7773359" figureDoi-2="http://doi.org/10.5281/zenodo.7773361" figureDoi-3="http://doi.org/10.5281/zenodo.7773365" httpUri-0="https://zenodo.org/record/7773357/files/figure.png" httpUri-1="https://zenodo.org/record/7773359/files/figure.png" httpUri-2="https://zenodo.org/record/7773361/files/figure.png" httpUri-3="https://zenodo.org/record/7773365/files/figure.png" pageId="31" pageNumber="70">Figs. 157171</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="32" lastPageNumber="71" pageId="31" pageNumber="70" type="materials_examined">
<paragraph blockId="31.[151,1436,762,1005]" pageId="31" pageNumber="70">
<emphasis bold="true" box="[151,400,762,788]" pageId="31" pageNumber="70">Materials examined.</emphasis>
<materialsCitation collectingDate="2022-07-08" collectingDateMax="2022-08-01" collectingDateMin="2022-07-08" collectionCode="R, AMC" collectorName="P. Srinivasan &amp; Isack" country="India" county="South" elevation="144" latitude="10.142222" location="Vaigai river" longLatPrecision="21" longitude="78.11639" municipality="Madurai District" pageId="31" pageNumber="70" specimenCount="16" specimenCount-female="1" specimenCount-larva="15" stateProvince="Tamil Nadu">
1 L-S-IJ, 3 L-S-I
<specimenCount box="[633,647,763,789]" pageId="31" pageNumber="70" type="female"></specimenCount>
,
<specimenCount box="[664,761,762,789]" pageId="31" pageNumber="70" type="larva">5 larvae</specimenCount>
,
<collectingCounty box="[777,842,763,789]" pageId="31" pageNumber="70">South</collectingCounty>
<collectingCountry box="[854,914,763,789]" name="India" pageId="31" pageNumber="70">India</collectingCountry>
,
<collectingRegion box="[929,1069,763,789]" country="India" name="Tamil Nadu" pageId="31" pageNumber="70">Tamil Nadu</collectingRegion>
,
<collectingMunicipality box="[1085,1279,763,789]" pageId="31" pageNumber="70">Madurai District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03C41A4F5D0360032A804489CB992F9A:8EB2FD825D03603C2F1944FCCAA52B6A" box="[1294,1432,763,789]" country="India" county="South" latitude="10.142222" longLatPrecision="21" longitude="78.11639" municipality="Madurai District" name="Vaigai river" pageId="31" pageNumber="70" stateProvince="Tamil Nadu">Vaigai river</location>
, 9°9552N,
<geoCoordinate box="[285,418,800,827]" degrees="78" direction="east" minutes="06" orientation="longitude" pageId="31" pageNumber="70" precision="15" seconds="59" value="78.11639">78°0659E</geoCoordinate>
,
<quantity box="[429,501,799,825]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="31" pageNumber="70" unit="m" value="144.0">
<elevation box="[429,501,799,825]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="31" pageNumber="70" unit="m" value="144.0">144 m</elevation>
</quantity>
,
<date box="[513,638,799,825]" pageId="31" pageNumber="70" value="2022-07-08">
<collectingDate box="[513,638,799,825]" pageId="31" pageNumber="70" value="2022-07-08">8.VII.2022</collectingDate>
</date>
, colls.
<collectorName box="[715,860,799,825]" pageId="31" pageNumber="70">P. Srinivasan</collectorName>
&amp;
<collectionCode box="[895,917,799,825]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="70">R</collectionCode>
. Isack (
<collectionCode box="[1000,1067,798,825]" country="USA" httpUri="http://grbio.org/cool/2xyg-zxc2" name="Department of Biologics Research" pageId="31" pageNumber="70">AMC</collectionCode>
/ZN/257).
<specimenCount box="[1183,1289,799,825]" pageId="31" pageNumber="70" type="larva">10 larvae</specimenCount>
, South
<collectingCountry box="[1372,1432,799,825]" name="India" pageId="31" pageNumber="70">India</collectingCountry>
,
<collectingRegion box="[151,286,835,861]" country="India" name="Tamil Nadu" pageId="31" pageNumber="70">Tamil Nadu</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03C41A4F5D0360032A804489CB992F9A:8EB2FD825D03603C2B3D4544CEDB2B22" box="[298,486,835,861]" country="India" county="South" latitude="10.142222" longLatPrecision="21" longitude="78.11639" municipality="Madurai District" name="Madurai District" pageId="31" pageNumber="70" stateProvince="Tamil Nadu">Madurai District</location>
,
<location LSID="urn:lsid:plazi:treatment:03C41A4F5D0360032A804489CB992F9A:8EB2FD825D03603C2BE64544CD4B2B22" box="[497,630,835,861]" country="India" county="South" latitude="10.142222" longLatPrecision="21" longitude="78.11639" municipality="Madurai District" name="Vaigai river" pageId="31" pageNumber="70" stateProvince="Tamil Nadu">Vaigai river</location>
,
<geoCoordinate box="[641,777,836,863]" degrees="10" direction="north" minutes="08" orientation="latitude" pageId="31" pageNumber="70" precision="15" seconds="32" value="10.142222">10°0832N</geoCoordinate>
, 77°9322E,
<quantity box="[933,1006,835,861]" metricMagnitude="2" metricUnit="m" metricValue="1.92" pageId="31" pageNumber="70" unit="m" value="192.0">
<elevation box="[933,1006,835,861]" metricMagnitude="2" metricUnit="m" metricValue="1.92" pageId="31" pageNumber="70" unit="m" value="192.0">192 m</elevation>
</quantity>
,
<date box="[1018,1152,835,861]" pageId="31" pageNumber="70" value="2022-08-01">
<collectingDate box="[1018,1152,835,861]" pageId="31" pageNumber="70" value="2022-08-01">1.VIII.2022</collectingDate>
</date>
, colls.
<collectorName box="[1230,1375,835,861]" pageId="31" pageNumber="70">P. Srinivasan</collectorName>
&amp;
<collectionCode box="[1411,1433,835,861]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="70">R</collectionCode>
.
<collectorName box="[151,210,871,897]" pageId="31" pageNumber="70">Isack</collectorName>
(
<collectionCode box="[225,292,870,897]" country="USA" httpUri="http://grbio.org/cool/2xyg-zxc2" name="Department of Biologics Research" pageId="31" pageNumber="70">AMC</collectionCode>
/ZN/257)
</materialsCitation>
.
</paragraph>
<paragraph blockId="31.[151,1436,762,1005]" pageId="31" pageNumber="70">
<emphasis bold="true" box="[199,536,906,933]" italics="true" pageId="31" pageNumber="70">Additions to the description.</emphasis>
Detailed imago and larval descriptions of
<taxonomicName authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[1045,1277,907,933]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="70" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis box="[1045,1277,907,933]" italics="true" pageId="31" pageNumber="70">Clypeocaenis kaveri</emphasis>
</taxonomicName>
are given by
<bibRefCitation author="Balasubramanian, C. &amp; Muthukatturaja, M." box="[151,665,943,969]" pageId="31" pageNumber="70" pagination="377 - 388" refId="ref17247" refString="Balasubramanian, C. &amp; Muthukatturaja, M. (2021) Two additional new species of Clypeocaenis Soldan, 1978 (Ephemeroptera: Caenidae) from the Western Ghats of Peninsular India. Zootaxa, 4915 (3), 377 - 388. https: // doi. org / 10.11646 / zootaxa. 4915.3.6" type="journal article" year="2021">Balasubramanian and Muthukatturaja, 2021</bibRefCitation>
. However, several important diagnostic characters of the imago, subimago and larva were not included in the original description.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7773357" ID-Zenodo-Dep="7773357" httpUri="https://zenodo.org/record/7773357/files/figure.png" pageId="31" pageNumber="70" startId="31.[152,270,1961,1985]" targetBox="[151,1436,1029,1937]" targetPageId="31">
<paragraph blockId="31.[151,1436,1961,2021]" pageId="31" pageNumber="70">
<emphasis bold="true" box="[152,372,1961,1985]" pageId="31" pageNumber="70">FIGURES 157159.</emphasis>
<taxonomicName authority="Balasubramanian and Muthukatturaja, 2021" authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[379,1042,1961,1985]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="31" pageNumber="70" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis box="[379,585,1961,1985]" italics="true" pageId="31" pageNumber="70">Clypeocaenis kaveri</emphasis>
<bibRefCitation author="Balasubramanian, C. &amp; Muthukatturaja, M." box="[592,1042,1961,1985]" pageId="31" pageNumber="70" pagination="377 - 388" refId="ref17247" refString="Balasubramanian, C. &amp; Muthukatturaja, M. (2021) Two additional new species of Clypeocaenis Soldan, 1978 (Ephemeroptera: Caenidae) from the Western Ghats of Peninsular India. Zootaxa, 4915 (3), 377 - 388. https: // doi. org / 10.11646 / zootaxa. 4915.3.6" type="journal article" year="2021">Balasubramanian and Muthukatturaja, 2021</bibRefCitation>
</taxonomicName>
: (157) female imago; (158) abdominal terga of female imago; (159) larva.
</paragraph>
</caption>
<paragraph blockId="32.[151,1437,150,717]" box="[199,492,150,177]" pageId="32" pageNumber="71">
<emphasis bold="true" box="[199,366,150,177]" pageId="32" pageNumber="71">Female imago</emphasis>
(
<figureCitation box="[382,477,150,177]" captionStart="FIGURES 157159" captionStartId="31.[152,270,1961,1985]" captionTargetBox="[151,1436,1029,1937]" captionTargetId="figure-260@31.[151,1436,1029,1937]" captionTargetPageId="31" captionText="FIGURES 157159. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (157) female imago; (158) abdominal terga of female imago; (159) larva." figureDoi="http://doi.org/10.5281/zenodo.7773357" httpUri="https://zenodo.org/record/7773357/files/figure.png" pageId="32" pageNumber="71">Fig. 157</figureCitation>
).
</paragraph>
<paragraph blockId="32.[151,1437,150,717]" pageId="32" pageNumber="71">
Coloration: Terga whitish (eggs under the body give the appearance of yellowish color to the tergum). Posterior half of terga I
<collectionCode box="[316,346,223,249]" pageId="32" pageNumber="71">IX</collectionCode>
with dark brownish transverse band (
<figureCitation box="[766,864,222,249]" captionStart="FIGURES 157159" captionStartId="31.[152,270,1961,1985]" captionTargetBox="[151,1436,1029,1937]" captionTargetId="figure-260@31.[151,1436,1029,1937]" captionTargetPageId="31" captionText="FIGURES 157159. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (157) female imago; (158) abdominal terga of female imago; (159) larva." figureDoi="http://doi.org/10.5281/zenodo.7773357" httpUri="https://zenodo.org/record/7773357/files/figure.png" pageId="32" pageNumber="71">Fig. 158</figureCitation>
). Prosternal sclerites trapezoid with concave lateral sides with a transverse blackish strip in the fore margin and with well-developed basolateral dark pigmentation (
<figureCitation captionStart="FIGURES 160163" captionStartId="32.[152,270,1673,1697]" captionTargetBox="[151,1436,741,1649]" captionTargetId="figure-244@32.[151,1436,741,1649]" captionTargetPageId="32" captionText="FIGURES 160163. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (160) prosternal triangle; (161) foreleg of female subimaginal exuvia; (162) microtichia in foretibia of female subimaginal exuvia; (163) microtichia in foretarsus of female subimaginal exuvia." figureDoi="http://doi.org/10.5281/zenodo.7773359" httpUri="https://zenodo.org/record/7773359/files/figure.png" pageId="32" pageNumber="71">Fig. 160</figureCitation>
).
</paragraph>
<paragraph blockId="32.[151,1437,150,717]" pageId="32" pageNumber="71">
<emphasis bold="true" box="[199,326,330,356]" pageId="32" pageNumber="71">Subimago.</emphasis>
Legs. Cuticular coloration: Femora of all legs light brownish-ochre, with diffusive light and darker areas; tibiae and tarsi light brownish-ochre (
<figureCitation box="[666,767,367,393]" captionStart="FIGURES 160163" captionStartId="32.[152,270,1673,1697]" captionTargetBox="[151,1436,741,1649]" captionTargetId="figure-244@32.[151,1436,741,1649]" captionTargetPageId="32" captionText="FIGURES 160163. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (160) prosternal triangle; (161) foreleg of female subimaginal exuvia; (162) microtichia in foretibia of female subimaginal exuvia; (163) microtichia in foretarsus of female subimaginal exuvia." figureDoi="http://doi.org/10.5281/zenodo.7773359" httpUri="https://zenodo.org/record/7773359/files/figure.png" pageId="32" pageNumber="71">Fig. 161</figureCitation>
). In both sexes of all leg pairs, tibiae and tarsomeres are covered with microtrichia (
<figureCitation box="[456,621,403,429]" captionStart="FIGURES 160163" captionStartId="32.[152,270,1673,1697]" captionTargetBox="[151,1436,741,1649]" captionTargetId="figure-244@32.[151,1436,741,1649]" captionTargetPageId="32" captionText="FIGURES 160163. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (160) prosternal triangle; (161) foreleg of female subimaginal exuvia; (162) microtichia in foretibia of female subimaginal exuvia; (163) microtichia in foretarsus of female subimaginal exuvia." figureDoi="http://doi.org/10.5281/zenodo.7773359" httpUri="https://zenodo.org/record/7773359/files/figure.png" pageId="32" pageNumber="71">Figs. 162, 163</figureCitation>
).
</paragraph>
<paragraph blockId="32.[151,1437,150,717]" pageId="32" pageNumber="71">
<emphasis bold="true" box="[199,272,439,465]" pageId="32" pageNumber="71">Larva</emphasis>
(
<figureCitation box="[289,386,438,465]" captionStart="FIGURES 157159" captionStartId="31.[152,270,1961,1985]" captionTargetBox="[151,1436,1029,1937]" captionTargetId="figure-260@31.[151,1436,1029,1937]" captionTargetPageId="31" captionText="FIGURES 157159. Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (157) female imago; (158) abdominal terga of female imago; (159) larva." figureDoi="http://doi.org/10.5281/zenodo.7773357" httpUri="https://zenodo.org/record/7773357/files/figure.png" pageId="32" pageNumber="71">Fig. 159</figureCitation>
). Abdomen. Tergalius II with a well developed medial ridge, forming an elevated triangular keel (
<figureCitation box="[212,309,475,501]" captionStart="FIGURES 164167" captionStartId="33.[152,270,1035,1059]" captionTargetBox="[209,1381,181,1011]" captionTargetId="figure-19@33.[207,1381,181,1011]" captionTargetPageId="33" captionText="FIGURES 164167. Larva of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (164) dorsal view of tergalius II; (165) SEM view of microtrichia in tergalius II; (166) shape of sternum IX; (167) detail of shagreen field in sternum IX." figureDoi="http://doi.org/10.5281/zenodo.7773361" httpUri="https://zenodo.org/record/7773361/files/figure.png" pageId="32" pageNumber="71">Fig. 164</figureCitation>
). Ventral row of microtrichia ending far from the hind margin, consists of 23 irregular rows of cluster of 26 spines medially (
<figureCitation box="[426,526,510,537]" captionStart="FIGURES 164167" captionStartId="33.[152,270,1035,1059]" captionTargetBox="[209,1381,181,1011]" captionTargetId="figure-19@33.[207,1381,181,1011]" captionTargetPageId="33" captionText="FIGURES 164167. Larva of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (164) dorsal view of tergalius II; (165) SEM view of microtrichia in tergalius II; (166) shape of sternum IX; (167) detail of shagreen field in sternum IX." figureDoi="http://doi.org/10.5281/zenodo.7773361" httpUri="https://zenodo.org/record/7773361/files/figure.png" pageId="32" pageNumber="71">Fig. 165</figureCitation>
). Hind margin of sternum
<collectionCode box="[829,859,511,537]" pageId="32" pageNumber="71">IX</collectionCode>
broadly rounded with numerous indentations (
<figureCitation captionStart="FIGURES 164167" captionStartId="33.[152,270,1035,1059]" captionTargetBox="[209,1381,181,1011]" captionTargetId="figure-19@33.[207,1381,181,1011]" captionTargetPageId="33" captionText="FIGURES 164167. Larva of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (164) dorsal view of tergalius II; (165) SEM view of microtrichia in tergalius II; (166) shape of sternum IX; (167) detail of shagreen field in sternum IX." figureDoi="http://doi.org/10.5281/zenodo.7773361" httpUri="https://zenodo.org/record/7773361/files/figure.png" pageId="32" pageNumber="71">Fig. 166</figureCitation>
); shagreen on dorsal side of sternum
<collectionCode box="[630,660,547,573]" pageId="32" pageNumber="71">IX</collectionCode>
with 56 irregular rows of very small denticles parallel to the hind margin (
<figureCitation box="[245,344,583,609]" captionStart="FIGURES 164167" captionStartId="33.[152,270,1035,1059]" captionTargetBox="[209,1381,181,1011]" captionTargetId="figure-19@33.[207,1381,181,1011]" captionTargetPageId="33" captionText="FIGURES 164167. Larva of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (164) dorsal view of tergalius II; (165) SEM view of microtrichia in tergalius II; (166) shape of sternum IX; (167) detail of shagreen field in sternum IX." figureDoi="http://doi.org/10.5281/zenodo.7773361" httpUri="https://zenodo.org/record/7773361/files/figure.png" pageId="32" pageNumber="71">Fig. 167</figureCitation>
).
</paragraph>
<paragraph blockId="32.[151,1437,150,717]" pageId="32" pageNumber="71">
<emphasis bold="true" box="[199,253,619,645]" pageId="32" pageNumber="71">Egg.</emphasis>
Length: 115120 μm; width: 6570 μm. Oval-shaped (
<figureCitation box="[888,988,619,645]" captionStart="FIGURES 168171" captionStartId="33.[152,270,1976,2000]" captionTargetBox="[204,1383,1119,1952]" captionTargetId="figure-62@33.[204,1383,1119,1952]" captionTargetPageId="33" captionText="FIGURES 168171. SEM view of eggs of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (168) entire view; (169, 170) epithema; (171) micropyle." figureDoi="http://doi.org/10.5281/zenodo.7773365" httpUri="https://zenodo.org/record/7773365/files/figure.png" pageId="32" pageNumber="71">Fig. 168</figureCitation>
); chorion smooth (
<figureCitation box="[1204,1302,619,645]" captionStart="FIGURES 168171" captionStartId="33.[152,270,1976,2000]" captionTargetBox="[204,1383,1119,1952]" captionTargetId="figure-62@33.[204,1383,1119,1952]" captionTargetPageId="33" captionText="FIGURES 168171. SEM view of eggs of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (168) entire view; (169, 170) epithema; (171) micropyle." figureDoi="http://doi.org/10.5281/zenodo.7773365" httpUri="https://zenodo.org/record/7773365/files/figure.png" pageId="32" pageNumber="71">Fig. 169</figureCitation>
) with a flat epithema forming a coil of fine threads with numerous small terminal knobs on one pole (
<figureCitation box="[1197,1298,655,681]" captionStart="FIGURES 168171" captionStartId="33.[152,270,1976,2000]" captionTargetBox="[204,1383,1119,1952]" captionTargetId="figure-62@33.[204,1383,1119,1952]" captionTargetPageId="33" captionText="FIGURES 168171. SEM view of eggs of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (168) entire view; (169, 170) epithema; (171) micropyle." figureDoi="http://doi.org/10.5281/zenodo.7773365" httpUri="https://zenodo.org/record/7773365/files/figure.png" pageId="32" pageNumber="71">Fig. 170</figureCitation>
). Micropyle short, situated at the subequatorial plane with mouth slightly broadened (
<figureCitation box="[968,1065,691,717]" captionStart="FIGURES 168171" captionStartId="33.[152,270,1976,2000]" captionTargetBox="[204,1383,1119,1952]" captionTargetId="figure-62@33.[204,1383,1119,1952]" captionTargetPageId="33" captionText="FIGURES 168171. SEM view of eggs of Clypeocaenis kaveri Balasubramanian and Muthukatturaja, 2021: (168) entire view; (169, 170) epithema; (171) micropyle." figureDoi="http://doi.org/10.5281/zenodo.7773365" httpUri="https://zenodo.org/record/7773365/files/figure.png" pageId="32" pageNumber="71">Fig. 171</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7773359" ID-Zenodo-Dep="7773359" httpUri="https://zenodo.org/record/7773359/files/figure.png" pageId="32" pageNumber="71" startId="32.[152,270,1673,1697]" targetBox="[151,1436,741,1649]" targetPageId="32">
<paragraph blockId="32.[151,1436,1673,1769]" pageId="32" pageNumber="71">
<emphasis bold="true" box="[152,371,1673,1697]" pageId="32" pageNumber="71">FIGURES 160163.</emphasis>
<taxonomicName authority="Balasubramanian and Muthukatturaja, 2021" authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[377,1035,1673,1697]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis box="[377,582,1673,1697]" italics="true" pageId="32" pageNumber="71">Clypeocaenis kaveri</emphasis>
<bibRefCitation author="Balasubramanian, C. &amp; Muthukatturaja, M." box="[589,1035,1673,1697]" pageId="32" pageNumber="71" pagination="377 - 388" refId="ref17247" refString="Balasubramanian, C. &amp; Muthukatturaja, M. (2021) Two additional new species of Clypeocaenis Soldan, 1978 (Ephemeroptera: Caenidae) from the Western Ghats of Peninsular India. Zootaxa, 4915 (3), 377 - 388. https: // doi. org / 10.11646 / zootaxa. 4915.3.6" type="journal article" year="2021">Balasubramanian and Muthukatturaja, 2021</bibRefCitation>
</taxonomicName>
: (160) prosternal triangle; (161) foreleg of female subimaginal exuvia; (162) microtichia in foretibia of female subimaginal exuvia; (163) microtichia in foretarsus of female subimaginal exuvia.
</paragraph>
</caption>
<subSubSection box="[199,1055,1815,1841]" pageId="32" pageNumber="71" type="distribution">
<paragraph blockId="32.[151,1436,1815,2022]" box="[199,1055,1815,1841]" pageId="32" pageNumber="71">
<emphasis bold="true" box="[199,352,1815,1841]" pageId="32" pageNumber="71">Distribution.</emphasis>
Kaveri river (
<collectingRegion box="[510,631,1815,1841]" country="India" name="Karnataka" pageId="32" pageNumber="71">Karnataka</collectingRegion>
), Vaigai river (Madurai,
<collectingRegion box="[906,1044,1815,1841]" country="India" name="Tamil Nadu" pageId="32" pageNumber="71">Tamil Nadu</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="71" type="biology_ecology">
<paragraph blockId="32.[151,1436,1815,2022]" pageId="32" pageNumber="71">
<emphasis bold="true" box="[199,300,1851,1877]" pageId="32" pageNumber="71">Ecology.</emphasis>
Larvae of
<taxonomicName authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[415,641,1851,1877]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis box="[415,641,1851,1877]" italics="true" pageId="32" pageNumber="71">Clypeocaenis kaveri</emphasis>
</taxonomicName>
were collected in the water temperature ranges between 24°C26°C; pH 7.27.3. Substratum is mainly made of cobbles and pebbles. The larvae of
<taxonomicName authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[987,1215,1887,1913]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis box="[987,1215,1887,1913]" italics="true" pageId="32" pageNumber="71">Clypeocaenis kaveri</emphasis>
</taxonomicName>
shared their habitat with
<taxonomicName authorityName="Kluge, Srinivasan, Vasanth, Sivaruban, Barathy &amp; Isack" authorityYear="2022" box="[206,553,1923,1949]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="latus" subGenus="Euthraulus">
<emphasis box="[206,345,1923,1949]" italics="true" pageId="32" pageNumber="71">Choroterpes</emphasis>
(
<emphasis box="[360,484,1923,1949]" italics="true" pageId="32" pageNumber="71">Euthraulus</emphasis>
)
<emphasis box="[499,553,1923,1949]" italics="true" pageId="32" pageNumber="71">latus</emphasis>
</taxonomicName>
(
<taxonomicName box="[566,755,1923,1949]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Leptophlebiidae</taxonomicName>
),
<taxonomicName box="[771,936,1923,1950]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[771,899,1923,1949]" italics="true" pageId="32" pageNumber="71">Tenuibaetis</emphasis>
sp.
</taxonomicName>
(
<taxonomicName authorityName="Leach" authorityYear="1815" box="[949,1053,1923,1949]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
),
<taxonomicName authorityName="Muller-Liebenau" authorityYear="1984" box="[1070,1307,1923,1949]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="operosus">
<emphasis box="[1070,1307,1923,1949]" italics="true" pageId="32" pageNumber="71">Labiobaetis operosus</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Leach" authorityYear="1815" box="[1320,1424,1923,1949]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
),
<taxonomicName authorityName="Sivaruban, Srinivasan, Barathy and Isack" authorityYear="2022" box="[151,357,1959,1985]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="klugei">
<emphasis box="[151,357,1959,1985]" italics="true" pageId="32" pageNumber="71">Nigrobaetis klugei</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Leach" authorityYear="1815" box="[371,475,1959,1985]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
),
<taxonomicName authorityName="Sivaruban, Srinivasan, Barathy, Isack and Kluge" authorityYear="2022" box="[493,779,1959,1985]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="vaigaiensis">
<emphasis box="[493,779,1959,1985]" italics="true" pageId="32" pageNumber="71">Cheleocloeon vaigaiensis</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Leach" authorityYear="1815" box="[793,897,1959,1985]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
),
<taxonomicName authorityName="Srinivasan, Sivaruban, Barathy &amp; Isack" authorityYear="2022" box="[915,1169,1959,1985]" class="Insecta" family="Tricorythidae" genus="Tricorythus" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="meenakshi">
<emphasis box="[915,1169,1959,1985]" italics="true" pageId="32" pageNumber="71">Tricorythus meenakshi</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Lestage" authorityYear="1942" box="[1183,1341,1959,1985]" class="Insecta" family="Tricorythidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Tricorythidae</taxonomicName>
),
<taxonomicName authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="maduraiensis">
<emphasis italics="true" pageId="32" pageNumber="71">Caenis maduraiensis</emphasis>
</taxonomicName>
(
<taxonomicName box="[317,427,1995,2021]" class="Insecta" family="Caenidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Caenidae</taxonomicName>
),
<taxonomicName authority="Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack, 2023" authorityName="Srinivasan &amp; Sivaruban &amp; Barathy &amp; Isack" authorityYear="2023" box="[446,699,1995,2022]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="venkataramani" status="sp. n.">
<emphasis box="[446,699,1995,2022]" italics="true" pageId="32" pageNumber="71">Caenis venkataramani</emphasis>
</taxonomicName>
<emphasis bold="true" box="[706,769,1995,2021]" pageId="32" pageNumber="71">
<taxonomicNameLabel box="[706,769,1995,2021]" pageId="32" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[823,1051,1995,2021]" class="Insecta" family="Caenidae" genus="Clypeocaenis" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="species" species="kaveri">
<emphasis box="[823,1051,1995,2021]" italics="true" pageId="32" pageNumber="71">Clypeocaenis kaveri</emphasis>
</taxonomicName>
(
<taxonomicName box="[1066,1177,1995,2021]" class="Insecta" family="Caenidae" kingdom="Animalia" order="Ephemeroptera" pageId="32" pageNumber="71" phylum="Arthropoda" rank="family">Caenidae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>