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<document id="CCB3C0782A15578C67E40D051474E53D" ID-DOI="10.11646/zootaxa.3826.1.3" ID-GBIF-Dataset="6f4b1535-6c6b-4446-aa26-b1ba83ad9126" ID-ISSN="1175-5326" ID-Zenodo-Dep="228537" ID-ZooBank="53549D58-8F38-47B5-879F-0245E900C131" IM.bibliography_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1460444968431" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent &amp; Welcomme, Jean-Loup" docDate="2014" docId="03A087E00E54FFAFFF7FFF79FCEFFAF9" docLanguage="en" docName="zt03826p138.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sindhiluta lakhraensis" docType="treatment" docVersion="11" lastPageNumber="130" masterDocId="FF99FF980E49FFB2FFE8FFEEFFDAFFAB" masterDocTitle="Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography" masterLastPageNumber="138" masterPageNumber="101" pageNumber="130" updateTime="1729174894255" updateUser="ExternalLinkService">
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<mods:title id="1BFAF09B0618EDA2BF57BAC84B128ECB">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:namePart id="57C257F85A1BEBCD7CCABAAD1913AB75">Merle, Didier</mods:namePart>
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<mods:namePart id="73813BBBC81B730ADA99EC41FBE86016">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="24E7AD930734111C8F1F8CD8CDDC224B">Métais, Grégoire</mods:namePart>
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<mods:namePart id="F173D16F82A5F2B00EA57C6DF6460AFB">Bartolini, Annachiara</mods:namePart>
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<mods:namePart id="5B5005BF9684A720645C6F797F59CCC0">Lashari, Rafiq A.</mods:namePart>
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<mods:namePart id="273AB0A70D6B9140B46715CC7CF8274F">Brohi, Imdad A.</mods:namePart>
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<mods:namePart id="3B5E323E5F2F8528E6D7B66B448557E6">Solangi, Sarfraz H.</mods:namePart>
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<mods:namePart id="97407BC6308E12FBFBB1C066F5B606B2">Marivaux, Laurent</mods:namePart>
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<mods:namePart id="E496CBE893E3FEB887EDD0B3D468313B">Welcomme, Jean-Loup</mods:namePart>
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<mods:date id="B84DD6794F09D6A031E4917899E627A8">2014</mods:date>
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<mods:number id="E1E069E9491A0CCD5040B3BEB05E03E8">2014-06-27</mods:number>
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<mods:number id="98CDB6F8478CA4F07BAC952992724849">3826</mods:number>
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<treatment id="03A087E00E54FFAFFF7FFF79FCEFFAF9" ID-DOI="http://doi.org/10.5281/zenodo.6137864" ID-GBIF-Taxon="119509273" ID-Zenodo-Dep="6137864" LSID="urn:lsid:plazi:treatment:03A087E00E54FFAFFF7FFF79FCEFFAF9" httpUri="http://treatment.plazi.org/id/03A087E00E54FFAFFF7FFF79FCEFFAF9" lastPageNumber="130" pageId="29" pageNumber="130">
<subSubSection id="C313657D0E54FFAFFF7FFF79FEF1FF78" pageId="29" pageNumber="130" type="nomenclature">
<paragraph id="8BB636F60E54FFAFFF7FFF79FDD4FF19" blockId="29.[151,526,151,211]" box="[151,526,151,178]" pageId="29" pageNumber="130">
<heading id="D0FE819A0E54FFAFFF7FFF79FDD4FF19" bold="true" box="[151,526,151,178]" fontSize="11" level="1" pageId="29" pageNumber="130" reason="1">
<emphasis id="B97DEAE40E54FFAFFF7FFF79FDD4FF19" bold="true" box="[151,526,151,178]" pageId="29" pageNumber="130">
<taxonomicName id="4C094D750E54FFAFFF7FFF79FE73FF1A" authority="Merle et al., 2014" box="[151,425,151,177]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="lakhraensis" status="sp. nov.">
<emphasis id="B97DEAE40E54FFAFFF7FFF79FE73FF1A" bold="true" box="[151,425,151,177]" italics="true" pageId="29" pageNumber="130">Sindhiluta lakhraensis</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB636F60E54FFAFFF7FFF54FEF1FF78" blockId="29.[151,526,151,211]" box="[151,299,186,211]" pageId="29" pageNumber="130">
(
<figureCitation id="13322A730E54FFAFFF77FF54FEF8FF78" box="[159,290,186,211]" captionStart="FIGURE 12. A F" captionStartId="30.[151,250,1850,1872]" captionTargetBox="[242,1337,193,1802]" captionTargetId="figure@30.[242,1337,193,1804]" captionTargetPageId="30" captionText="FIGURE 12. A F. Sindhiluta lakhraensis gen. nov., sp. nov., A B (holotype, CPAG. RAN. I. 48, ventral (A) and dorsal (B) views, stn 4), C D (paratype 1, CPAG. RAN. I. 49, ventral and dorsal view, stn 4), E F (paratype 2, CPAG. RAN. I. 50, dorsal view of the broken specimen showing the columellar folds (E) and enlarged view of the spire (F); G M. Pakiluta solangii gen. nov., sp. nov., G H (holotype, CPAG. RAN. I. 51, ventral (G) and dorsal (H) views, stn 4), I J (paratype 3, CPAG. RAN. I. 54, spire (I), and enlarged view of the spire (J), stn 5), K L (paratype 2, CPAG. RAN. I. 53, spire (K) and enlarged view of the spire (L), stn 4), M (paratype 1, CPAG. RAN. I. 52, ventral view, stn 4). Scale bars: A I, K, M = 10 mm; J, L = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.228549" httpUri="https://zenodo.org/record/228549/files/figure.png" pageId="29" pageNumber="130">Fig. 12AF</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E54FFAFFF7FFEECFD3AFEB0" box="[151,736,258,283]" pageId="29" pageNumber="130" type="etymology">
<paragraph id="8BB636F60E54FFAFFF7FFEECFD3AFEB0" blockId="29.[151,1437,258,1363]" box="[151,736,258,283]" pageId="29" pageNumber="130">
<emphasis id="B97DEAE40E54FFAFFF7FFEECFEC4FEB0" bold="true" box="[151,286,258,283]" pageId="29" pageNumber="130">Etymology.</emphasis>
From the
<typeStatus id="54B288540E54FFAFFE79FEEDFE1BFEB0" box="[401,449,259,283]" pageId="29" pageNumber="130">type</typeStatus>
locality (Lakhra village).
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E54FFAFFF2FFEC8FD5DFE00" pageId="29" pageNumber="130" type="materials_examined">
<paragraph id="8BB636F60E54FFAFFF2FFEC8FB6DFE94" blockId="29.[151,1437,258,1363]" box="[199,1207,294,319]" pageId="29" pageNumber="130">
<emphasis id="B97DEAE40E54FFAFFF2FFEC8FEBFFE94" bold="true" box="[199,357,294,319]" pageId="29" pageNumber="130">
<typeStatus id="54B288540E54FFAFFF2FFEC8FED8FE94" box="[199,258,294,319]" pageId="29" pageNumber="130">Type</typeStatus>
locality.
</emphasis>
Stn 4: Lakhra Dome, Lakhra village section, base of the Lakhra Formation.
</paragraph>
<paragraph id="8BB636F60E54FFAFFF2FFEA4FB97FE2C" blockId="29.[151,1437,258,1363]" pageId="29" pageNumber="130">
<emphasis id="B97DEAE40E54FFAFFF2FFEA4FEAEFEC8" bold="true" box="[199,372,330,355]" pageId="29" pageNumber="130">
<typeStatus id="54B288540E54FFAFFF2FFEA4FED8FEC8" box="[199,258,330,355]" pageId="29" pageNumber="130">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B288540E54FFAFFE92FEA4FE38FEC8" box="[378,482,330,355]" pageId="29" pageNumber="130" type="holotype">Holotype</typeStatus>
(stn 4:
<collectionCode id="ED18AE330E54FFAFFDDEFEA5FD5CFEC8" box="[566,646,331,355]" pageId="29" pageNumber="130">CPAG</collectionCode>
.RAN.
<date id="FFB710360E54FFAFFD23FEA5FD29FEC8" box="[715,755,331,355]" pageId="29" pageNumber="130" value="1948-01">I.48</date>
, cast
<collectionCode id="ED18AE330E54FFAFFCD8FEA4FC51FEC8" LSID="urn:lsid:biocol.org:col:34988" box="[816,907,330,355]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="130">MNHN</collectionCode>
.F.A50386),
<typeStatus id="54B288540E54FFAFFBFAFEA5FBA8FEC8" box="[1042,1138,331,355]" pageId="29" pageNumber="130" type="paratype">paratype</typeStatus>
1 (stn 4:
<collectionCode id="ED18AE330E54FFAFFB33FEA5FAF0FEC8" box="[1243,1322,331,355]" pageId="29" pageNumber="130">CPAG</collectionCode>
.RAN.
<date id="FFB710360E54FFAFFA87FEA5FA42FEC8" box="[1391,1432,331,355]" pageId="29" pageNumber="130" value="1949-01">I.49</date>
, cast
<collectionCode id="ED18AE330E54FFAFFF21FE80FEFEFE2C" LSID="urn:lsid:biocol.org:col:34988" box="[201,292,366,391]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="130">MNHN</collectionCode>
.F.
<accessionNumber id="945AAB150E54FFAFFED3FE80FE43FE2C" box="[315,409,366,391]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A50387" pageId="29" pageNumber="130" type="EnaNcbi">A50387</accessionNumber>
),
<typeStatus id="54B288540E54FFAFFE44FE81FDD6FE2C" box="[428,524,367,391]" pageId="29" pageNumber="130" type="paratype">paratype</typeStatus>
2 (stn 4:
<collectionCode id="ED18AE330E54FFAFFD9EFE80FD1FFE2C" box="[630,709,366,391]" pageId="29" pageNumber="130">CPAG</collectionCode>
.RAN.
<date id="FFB710360E54FFAFFCE2FE80FCE9FE2C" box="[778,819,366,391]" pageId="29" pageNumber="130" value="1950-01">I.50</date>
, cast
<collectionCode id="ED18AE330E54FFAFFC99FE80FC16FE2C" LSID="urn:lsid:biocol.org:col:34988" box="[881,972,366,391]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="130">MNHN</collectionCode>
.F.A50388).
</paragraph>
<paragraph id="8BB636F60E54FFAFFF2FFE7CFD5DFE00" blockId="29.[151,1437,258,1363]" box="[199,647,402,427]" pageId="29" pageNumber="130">
<emphasis id="B97DEAE40E54FFAFFF2FFE7CFE5BFE00" bold="true" box="[199,385,402,427]" pageId="29" pageNumber="130">Other material.</emphasis>
4 spm (stn 4:
<collectionCode id="ED18AE330E54FFAFFDC9FE7DFDA6FE00" LSID="urn:lsid:biocol.org:col:34988" box="[545,636,403,427]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="130">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E54FFAFFF26FE58FCEFFAF9" pageId="29" pageNumber="130" type="description">
<paragraph id="8BB636F60E54FFAFFF26FE58FE39FCB8" blockId="29.[151,1437,258,1363]" pageId="29" pageNumber="130">
<emphasis id="B97DEAE40E54FFAFFF26FE58FEBAFE64" bold="true" box="[206,352,438,463]" pageId="29" pageNumber="130">Description.</emphasis>
Shell elongate, H 5658, D
<quantity id="4CF19B130E54FFAFFD48FE59FCC3FE65" box="[672,793,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="29" pageNumber="130" unit="mm" value="22.0" valueMax="23.0" valueMin="21.0">2123 mm</quantity>
(
<typeStatus id="54B288540E54FFAFFCCFFE58FC51FE64" box="[807,907,438,463]" pageId="29" pageNumber="130" type="holotype">holotype</typeStatus>
H 53.6 not complete, D
<quantity id="4CF19B130E54FFAFFB49FE59FAD0FE64" box="[1185,1290,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="3.19" pageId="29" pageNumber="130" unit="mm" value="31.9">31.9 mm</quantity>
). Protoconch not preserved. Teleoconch of 7 whorls. Spire relatively high, occupying 31% of total shell height. Spiral whorls convex to subcarinate. Last whorl narrow, convex posteriorly, not excavated anteriorly. Suture linear with weak undulation between bases of costae. Axial sculpture of strong but rounded costae on spire. Costae orthocline, subcarinate at shoulder, not spinous. On penultimate whorl, costae decreasing in prominence, becoming obsolete, disappearing on last whorl. First whorl: not preserved; second whorl: 11 costae; third whorl: 10 costae; fourth and fifth whorls: 910 costae; sixth whorl: 89 costae, becoming obsolete; seventh whorl: no costae. No apparent spiral sculpture. Aperture narrow, lenticular, occupying 63% of total height, 26% of diameter. Outer lip thin, not thickened externally. Parietal callus thick posteriorly. Four strong, oblique columellar folds, anterior-most weaker. Siphonal canal not preserved.
</paragraph>
<paragraph id="8BB636F60E54FFAFFF2FFCF0FEC4FA84" blockId="29.[151,1437,258,1363]" pageId="29" pageNumber="130">
<emphasis id="B97DEAE40E54FFAFFF2FFCF0FEB7FC9C" bold="true" box="[199,365,798,823]" pageId="29" pageNumber="130">Comparisons.</emphasis>
From the Ypresian of the Aquitaine Basin,
<taxonomicName id="4C094D750E54FFAFFC6AFCF0FB3AFC9C" authority="Rouault, 1850" authorityName="Rouault" authorityYear="1850" box="[898,1248,798,823]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="prevosti">
<emphasis id="B97DEAE40E54FFAFFC6AFCF0FBF5FC9D" box="[898,1071,798,823]" italics="true" pageId="29" pageNumber="130">Voluta prevosti</emphasis>
Rouault, 1850
</taxonomicName>
[
<taxonomicName id="4C094D750E54FFAFFB1EFCF1FEE7FCF0" authority="Rouault, 1848" authorityName="Rouault" authorityYear="1848" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="prevosti">
<emphasis id="B97DEAE40E54FFAFFB1EFCF1FA41FC9D" box="[1270,1435,799,822]" italics="true" pageId="29" pageNumber="130">Mitra prevosti</emphasis>
Rouault, 1848
</taxonomicName>
is a
<emphasis id="B97DEAE40E54FFAFFE60FCAAFDF7FCF0" box="[392,557,834,859]" italics="true" pageId="29" pageNumber="130">nomen nudum</emphasis>
] is the single Paleogene volutid that shares rounded, subcarinate costae disappearing progressively at the end of growth with
<taxonomicName id="4C094D750E54FFAFFD1BFC88FCBCFCD4" box="[755,870,870,895]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E54FFAFFD1BFC88FCBCFCD4" box="[755,870,870,895]" italics="true" pageId="29" pageNumber="130">Sindhiluta</emphasis>
</taxonomicName>
. In addition, as in
<taxonomicName id="4C094D750E54FFAFFBD4FC89FB00FCD4" box="[1084,1242,870,895]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="lakhraensis" status="sp. nov.">
<emphasis id="B97DEAE40E54FFAFFBD4FC89FB00FCD4" box="[1084,1242,870,895]" italics="true" pageId="29" pageNumber="130">S. lakhraensis</emphasis>
</taxonomicName>
<emphasis id="B97DEAE40E54FFAFFB0AFC88FAE6FCD4" bold="true" box="[1250,1340,870,895]" pageId="29" pageNumber="130">sp. nov.</emphasis>
, the last whorls of
<taxonomicName id="4C094D750E54FFAFFEF8FC62FE52FC08" box="[272,392,908,931]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="prevosti">
<emphasis id="B97DEAE40E54FFAFFEF8FC62FE52FC08" box="[272,392,908,931]" italics="true" pageId="29" pageNumber="130">V. prevosti</emphasis>
</taxonomicName>
are rounded and not shouldered posteriorly, its base is not excavated, its outer lip is not thickened and a parietal callus is present. Regarding the columellar folds, their numbers vary between three and six, but their construction is rather similar to that of
<taxonomicName id="4C094D750E54FFAFFD50FC3DFC8FFC40" box="[696,853,978,1003]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="lakhraensis" status="sp. nov.">
<emphasis id="B97DEAE40E54FFAFFD50FC3DFC8FFC40" box="[696,853,978,1003]" italics="true" pageId="29" pageNumber="130">S. lakhraensis</emphasis>
</taxonomicName>
<emphasis id="B97DEAE40E54FFAFFCB5FC3CFC62FC40" bold="true" box="[861,952,978,1003]" pageId="29" pageNumber="130">sp. nov.</emphasis>
The most anterior fold is present in young specimens, but is weaker than the three adapical folds or tends to disappear in adult specimens. In
<taxonomicName id="4C094D750E54FFAFFB15FC19FA46FBA4" box="[1277,1436,1014,1039]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="lakhraensis" status="sp. nov.">
<emphasis id="B97DEAE40E54FFAFFB15FC19FA46FBA4" box="[1277,1436,1014,1039]" italics="true" pageId="29" pageNumber="130">S. lakhraensis</emphasis>
</taxonomicName>
<emphasis id="B97DEAE40E54FFAFFF7FFBF4FF28FB98" bold="true" box="[151,242,1050,1075]" pageId="29" pageNumber="130">sp. nov.</emphasis>
, this anterior fold is also weaker even if it persists at the end of growth. Adapically, the columella of
<taxonomicName id="4C094D750E54FFAFFA6FFBF2FF2AFBFD" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="prevosti">
<emphasis id="B97DEAE40E54FFAFFA6FFBF2FF2AFBFD" italics="true" pageId="29" pageNumber="130">V. prevosti</emphasis>
</taxonomicName>
displays three strong folds as in
<taxonomicName id="4C094D750E54FFAFFD88FBD1FD27FBFC" box="[608,765,1086,1111]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="lakhraensis">
<emphasis id="B97DEAE40E54FFAFFD88FBD1FD27FBFC" box="[608,765,1086,1111]" italics="true" pageId="29" pageNumber="130">S. lakhraensis</emphasis>
</taxonomicName>
. In its posterior part, the columella displays two weak folds in a few specimens, but they are missing in
<taxonomicName id="4C094D750E54FFAFFD6AFB8DFCFAFBD0" box="[642,800,1122,1147]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="lakhraensis">
<emphasis id="B97DEAE40E54FFAFFD6AFB8DFCFAFBD0" box="[642,800,1122,1147]" italics="true" pageId="29" pageNumber="130">S. lakhraensis</emphasis>
</taxonomicName>
. Considering the large number of shared characters with the
<collectingCountry id="F31E76660E54FFAFFF2DFB68FEF6FB34" box="[197,300,1158,1183]" name="Pakistan" pageId="29" pageNumber="130">Pakistani</collectingCountry>
species, we do not hesitate to assign
<taxonomicName id="4C094D750E54FFAFFD1BFB68FC45FB35" box="[755,927,1158,1183]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="130" phylum="Mollusca" rank="species" species="prevosti">
<emphasis id="B97DEAE40E54FFAFFD1BFB68FC45FB35" box="[755,927,1158,1183]" italics="true" pageId="29" pageNumber="130">Voluta prevosti</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="4C094D750E54FFAFFBA0FB68FB61FB34" box="[1096,1211,1158,1183]" class="Gastropoda" family="Volutidae" genus="Sindhiluta" kingdom="Animalia" order="Neogastropoda" pageId="29" pageNumber="128" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E54FFAFFBA0FB68FB61FB34" box="[1096,1211,1158,1183]" italics="true" pageId="29" pageNumber="130">Sindhiluta</emphasis>
</taxonomicName>
. On biogeographic grounds, it seems unlikely that these species, located at two extremities of Tethyan Ocean, belong to a same genus. However, the both extremities of the Tethyan Ocean share numerous common genera and even some common species (
<bibRefCitation id="EF984B070E54FFAFFF1EFB1CFE56FAA0" author="Pacaud" box="[246,396,1266,1291]" firstAuthor="Pacaud" pageId="29" pageNumber="130" refString="Pacaud, J. - M. (1997) Nouvelles donnees sur le genre Popenoeum (Mollusca, Pseudolividae). Remarques taxonomiques sur une espece ubiquiste du Paleocene inferieur, Popenoeum ambiguum (Binkhorst, 1861: Pyrula) nov. comb. Cossmanniana, 5 (1 - 2), 1 - 28." type="journal article" year="1997">Pacaud 1997</bibRefCitation>
). In addition, examples of range disjunction in tropical marine area are also known (
<bibRefCitation id="EF984B070E54FFAFFAD6FB1DFEC8FA84" author="Lozouet" etAl="et al." firstAuthor="Lozouet" pageId="29" pageNumber="130" refString="Lozouet, P., Ledon, D. &amp; Lesport, J. F. (1994) Le genre Lindapterys (Neogastropoda, Muricidae): un exemple de disjonction de distribution en domaine tropical marin. Geobios, 27 (1), 39 - 50. http: // dx. doi. org / 10.1016 / s 0016 - 6995 (06) 80210 - x" type="journal article" year="1994">
Lozouet
<emphasis id="B97DEAE40E54FFAFFF7FFAF9FF15FA84" box="[151,207,1302,1327]" italics="true" pageId="29" pageNumber="130">et al.</emphasis>
1994
</bibRefCitation>
).
</paragraph>
<paragraph id="8BB636F60E54FFAFFF2FFAD4FCEFFAF9" blockId="29.[151,1437,258,1363]" box="[199,821,1338,1363]" pageId="29" pageNumber="130">
<emphasis id="B97DEAE40E54FFAFFF2FFAD4FE62FAF8" bold="true" box="[199,440,1338,1363]" pageId="29" pageNumber="130">Stratigraphic range.</emphasis>
Lakhra formation: Lakhra Dome.
</paragraph>
</subSubSection>
</treatment>
</document>