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<mods:title id="0FCCA2E2F5C6AEEAB18C81446DFCA2CF">Integrative taxonomy reveals hidden diversity in the Catharus fuscater (Passeriformes: Turdidae) complex in Central and South America</mods:title>
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<mods:namePart id="5942B04174A5B929C6949D5BD81EF2CE">Weckstein, Jason D.</mods:namePart>
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<treatment id="03A087D8836CFFC3B1AD4EDD2140FA24" ID-DOI="http://doi.org/10.5281/zenodo.8328703" ID-GBIF-Taxon="212969849" ID-Zenodo-Dep="8328703" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24" httpUri="http://treatment.plazi.org/id/03A087D8836CFFC3B1AD4EDD2140FA24" lastPageId="27" lastPageNumber="255" pageId="22" pageNumber="250">
<subSubSection id="C3136545836CFFCEB1AD4EDD226AFC4C" box="[255,620,970,997]" pageId="22" pageNumber="250" type="nomenclature">
<paragraph id="8BB636CE836CFFCEB1AD4EDD226AFC4C" blockId="22.[255,620,970,997]" box="[255,620,970,997]" pageId="22" pageNumber="250">
<heading id="D0FE81A2836CFFCEB1AD4EDD226AFC4C" bold="true" box="[255,620,970,997]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="250" reason="2">
<taxonomicName id="4C094D4D836CFFCEB1AD4EDD226AFC4C" authority="Lawrence 1888" authorityName="Lawrence" authorityYear="1888" box="[255,620,970,997]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC836CFFCEB1AD4EDD226AFC4C" bold="true" box="[255,620,970,997]" pageId="22" pageNumber="250">
<emphasis id="B97DEADC836CFFCEB1AD4EDD21C3FC4C" bold="true" box="[255,453,970,997]" italics="true" pageId="22" pageNumber="250">Catharus berlepschi</emphasis>
<bibRefCitation id="EF984B3F836CFFCEB09B4EDC226AFC4C" author="Lawrence GN" box="[457,620,971,997]" pageId="22" pageNumber="250" pagination="503" refId="ref32168" refString="Lawrence GN. Description of a new species of bird of the genus Catharus, from Ecuador. Proceedings of the United States National Museum ' 1887 ' (= 1888); [1887]: 503." type="journal article" year="1888">Lawrence 1888</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C3136545836CFFCEB0544EEE2260FBBA" box="[262,614,1017,1043]" pageId="22" pageNumber="250" type="vernacular_names">
<paragraph id="8BB636CE836CFFCEB0544EEE2260FBBA" blockId="22.[262,614,1017,1043]" box="[262,614,1017,1043]" pageId="22" pageNumber="250">
<emphasis id="B97DEADC836CFFCEB0544EEE2260FBBA" bold="true" box="[262,614,1017,1043]" pageId="22" pageNumber="250">Trans-Andean nightingale-thrush</emphasis>
</paragraph>
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<subSubSection id="C3136545836CFFCEB022493021FDFBE8" box="[368,507,1063,1090]" pageId="22" pageNumber="250" type="description">
<paragraph id="8BB636CE836CFFCEB022493021FDFBE8" blockId="22.[368,507,1063,1090]" box="[368,507,1063,1090]" pageId="22" pageNumber="250">
<emphasis id="B97DEADC836CFFCEB022493021FDFBE8" bold="true" box="[368,507,1063,1090]" pageId="22" pageNumber="250">
(
<figureCitation id="13322A4B836CFFCEB029493021F6FBEB" box="[379,496,1063,1090]" captionStart-0="Figure 15" captionStart-1="Figure 16" captionStart-2="Figure 17" captionStart-3="Figure 18" captionStartId-0="23.[129,194,1708,1732]" captionStartId-1="24.[113,178,1708,1732]" captionStartId-2="25.[129,194,909,933]" captionStartId-3="25.[129,194,1954,1978]" captionTargetBox-0="[257,1346,156,1680]" captionTargetBox-1="[285,1290,156,1680]" captionTargetBox-2="[149,757,345,881]" captionTargetBox-3="[283,1360,1042,1926]" captionTargetId-0="figure-75@23.[247,1356,144,1680]" captionTargetId-1="figure-124@24.[282,1290,144,1680]" captionTargetId-2="figure-247@25.[149,757,320,881]" captionTargetId-3="figure-265@25.[241,1361,1022,1926]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionTargetPageId-3="25" captionText-0="Figure 15. Ventral and dorsal views of the adult male plumages in C. b. berlepschi, C. b. caniceps, C. o.tenebris ssp. nov., C. b. nebulus ssp.nov., and C. mentalis. Photograph by M.R.H." captionText-1="Figure 16. Ventral and dorsal views of the adult female plumages in C. b. berlepschi, C. b. caniceps, C. o.tenebris ssp. nov., C. b. nebulus ssp.nov., and C. mentalis. Photograph by M.R.H." captionText-2="Figure 17. Side view of adult plumages in C. b. nebulus ssp. nov. Photograph by M.R.H." captionText-3="Figure 18. Ventral and dorsal views of juvenile plumages in C. hellmayri, C. b. nebulus ssp.nov., and C. b. berlepschi. Photograph by M.R.H." figureDoi-0="http://doi.org/10.5281/zenodo.8326332" figureDoi-1="http://doi.org/10.5281/zenodo.8326334" figureDoi-2="http://doi.org/10.5281/zenodo.8326336" figureDoi-3="http://doi.org/10.5281/zenodo.8326338" httpUri-0="https://zenodo.org/record/8326332/files/figure.png" httpUri-1="https://zenodo.org/record/8326334/files/figure.png" httpUri-2="https://zenodo.org/record/8326336/files/figure.png" httpUri-3="https://zenodo.org/record/8326338/files/figure.png" pageId="22" pageNumber="250">Figs 1518</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3136545836CFFCEB1D2497922FCF979" pageId="22" pageNumber="250" type="reference_group">
<paragraph id="8BB636CE836CFFCEB1D2497921FDFAC8" blockId="22.[127,763,1133,1659]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB1D24979228AFB2F" ID-CoL="69G37" authority="Sclater 1859 a: 136" authorityName="Sclater" authorityPageNumber="136" authorityYear="1859" box="[128,652,1133,1158]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="species" species="fuscater">
<emphasis id="B97DEADC836CFFCEB1D249792137FB2F" box="[128,305,1134,1158]" italics="true" pageId="22" pageNumber="250">Catharus fuscater</emphasis>
Sclater 1859a: 136, 1859b: 324
</taxonomicName>
(in part);
<bibRefCitation id="EF984B3F836CFFCEB1CA499A2251FB0C" author="von Berlepsch HG &amp; Taczanowski L" box="[152,599,1165,1189]" pageId="22" pageNumber="250" pagination="281 - 313" refId="ref29489" refString="von Berlepsch HG, Taczanowski L. Deuxieme liste des Oiseaux recueillis dans l'Ecuadeur occidental par MM. Stolzmann et Siemiradski. Proceedings of the Zoological Society of London 1884; 1884: 281 - 313." type="journal article" year="1884">Berlepsch and Taczanowski 1884: 283</bibRefCitation>
; Salvin 1895,
<bibRefCitation id="EF984B3F836CFFCEB1CA49BB2184FB6D" author="Goodfellow W" box="[152,386,1196,1220]" pageId="22" pageNumber="250" pagination="300 - 19" refId="ref30955" refString="Goodfellow W. Results of an ornithological journey through Colombia and Ecuador. Ibis 1901; 43: 300 - 19." type="journal article" year="1901">Goodfellow 1901: 309</bibRefCitation>
; Meyer de Schauensee 1966: 412 (in part); Meyer de Schauensee 1970: 342 (in part); Ridgley and Tudor 1989: 110 (in part);
<bibRefCitation id="EF984B3F836CFFCEB0B149FC20C4FA8B" author="Clements JF &amp; Shany N" pageId="22" pageNumber="250" refId="ref30124" refString="Clements JF, Shany N. A Field Guide to the Birds of Peru. Temecula, CA: Ibis Publishing Company, 2001." type="book" year="2001">Clements and Shany 2001: 193</bibRefCitation>
; Solano-Ugalde
<emphasis id="B97DEADC836CFFCEB039481C219AFA8B" box="[363,412,1290,1314]" italics="true" pageId="22" pageNumber="250">et al.</emphasis>
2007; Schulenberg
<emphasis id="B97DEADC836CFFCEB338481C229DFA8B" box="[618,667,1290,1314]" italics="true" pageId="22" pageNumber="250">et al.</emphasis>
2007 (in part);
<bibRefCitation id="EF984B3F836CFFCEB18C483E216BFAEB" author="Kikuchi DW" box="[222,365,1321,1346]" pageId="22" pageNumber="250" pagination="523 - 9" refId="ref31839" refString="Kikuchi DW. Terrestrial and understorey insectivorous birds of a Peruvian cloud forest: species richness, abundance, density, territory size and biomass. Journal of Tropical Ecology 2009; 25: 523 - 9." type="journal article" year="2009">Kikuchi 2009</bibRefCitation>
; Schmitt
<emphasis id="B97DEADC836CFFCEB088483D2208FAE8" box="[474,526,1321,1345]" italics="true" pageId="22" pageNumber="250">et al.</emphasis>
2013;
<bibRefCitation id="EF984B3F836CFFCEB30C483E20CAFAC8" author="Astudillo PX &amp; Tinoco BA &amp; Siddons DC" pageId="22" pageNumber="250" pagination="2 - 12" refId="ref29240" refString="Astudillo PX, Tinoco BA, Siddons DC. The avifauna of Cajas National Park and Mazan Reserve, southern Ecuador, with notes on new records. Cotinga 2015; 37: 2 - 12." type="journal article" year="2015">
Astudillo
<emphasis id="B97DEADC836CFFCEB395483D22FDFAE8" box="[711,763,1321,1345]" italics="true" pageId="22" pageNumber="250">et al.</emphasis>
2015
</bibRefCitation>
; Remsen
<emphasis id="B97DEADC836CFFCEB063485D2166FAC8" box="[305,352,1353,1377]" italics="true" pageId="22" pageNumber="250">et al.</emphasis>
2023 (in part).
</paragraph>
<paragraph id="8BB636CE836CFFCEB1D2487F22D0FA09" blockId="22.[127,763,1133,1659]" pageId="22" pageNumber="250">
<treatmentCitationGroup id="AB1911E0836CFFCEB1D2487F22D0FA09" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB1D2487F21BFFA29" authority="Lawrence" authorityName="Lawrence" authorityYear="1888" box="[128,441,1384,1408]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC836CFFCEB1D2487F214EFA29" box="[128,328,1384,1408]" italics="true" pageId="22" pageNumber="250">Catharus berlepschi</emphasis>
Lawrence
</taxonomicName>
1887 (=1888; see:
<treatmentCitation id="0AA810DF836CFFCEB3FE487F20FFFA09" author="LeCroy M" page="39" pageId="22" pageNumber="250" year="2005">
<bibRefCitation id="EF984B3F836CFFCEB3FE487F20FFFA09" author="LeCroy M" pageId="22" pageNumber="250" pagination="1 - 132" refId="ref32207" refString="LeCroy M. Type specimens of birds in the American Museum of Natural History. Part 6, Passeriformes: Prunellidae, Turdidae, Orthonychidae, Timaliidae, Paradoxornithidae, Picathartidae, and Polioptilidae. Bulletin of the American Museum of Natural History 2005; 292: 1 - 132." type="journal article" year="2005">LeCroy 2005: 39</bibRefCitation>
</treatmentCitation>
): 503; Ridgway 1887: 504; Sharpe 1902: 182.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB636CE836CFFCEB1D248B02277FA54" blockId="22.[127,763,1133,1659]" pageId="22" pageNumber="250">
<treatmentCitationGroup id="AB1911E0836CFFCEB1D248B02277FA54" pageId="22" pageNumber="250">
<treatmentCitation id="0AA810DF836CFFCEB1D248B02249FA16" author="Chapman FM" box="[128,591,1447,1471]" page="14" pageId="22" pageNumber="250" year="1924">
<taxonomicName id="4C094D4D836CFFCEB1D248B02249FA16" ID-CoL="5G9M7" authority="Chapman 1924: 14" authorityName="Chapman" authorityPageNumber="14" authorityYear="1924" box="[128,591,1447,1471]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="subSpecies" species="fuscater" subSpecies="caniceps">
<emphasis id="B97DEADC836CFFCEB1D248B02186FA16" box="[128,384,1447,1471]" italics="true" pageId="22" pageNumber="250">Catharus fuscater caniceps</emphasis>
<bibRefCitation id="EF984B3F836CFFCEB0D448B02249FA16" author="Chapman FM" box="[390,591,1447,1471]" pageId="22" pageNumber="250" pagination="1 - 16" refId="ref30009" refString="Chapman FM. Descriptions of new birds from Ecuador, Colombia, Peru, and Bolivia. American Museum Novitates 1924; 138: 1 - 16." type="journal article" year="1924">Chapman 1924: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AA810DF836CFFCEB30B48B020C4FA77" author="Hellmayr CE" page="466" pageId="22" pageNumber="250" year="1934">
<bibRefCitation id="EF984B3F836CFFCEB30B48B020C4FA77" author="Hellmayr CE" pageId="22" pageNumber="250" pagination="1 - 531" refId="ref31543" refString="Hellmayr CE. Catalogue of birds of the Americas and the adjacent islands in Field Museum of Natural History, including all species and subspecies known to occur in North America, Mexico, Central America, the West Indies, and islands of the Caribbean Sea, the Galapagos Archipelago and other islands which may be included on account of their faunal affinities. Field Museum of Natural History, Zoological Series 13 1934; 7: 1 - 531." type="journal article" year="1934">Hellmayr 1934: 466</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AA810DF836CFFCEB18248D12187FA77" author="Bond J" box="[208,385,1478,1502]" page="241" pageId="22" pageNumber="250" year="1956">
<bibRefCitation id="EF984B3F836CFFCEB18248D12187FA77" author="Bond J" box="[208,385,1478,1502]" pageId="22" pageNumber="250" pagination="227 - 47" refId="ref29674" refString="Bond J. Additional notes on Peruvian Birds II. Proceedings of the Academy of Natural Sciences of Philadelphia 1956; 108: 227 - 47." type="journal article" year="1956">Bond 1956: 241</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AA810DF836CFFCEB0DD48D122D4FA77" author="Fjeldsa J &amp; Krabbe N" box="[399,722,1478,1502]" page="554" pageId="22" pageNumber="250" year="1990">
<bibRefCitation id="EF984B3F836CFFCEB0DD48D122D4FA77" author="Fjeldsa J &amp; Krabbe N" box="[399,722,1478,1502]" pageId="22" pageNumber="250" refId="ref30694" refString="Fjeldsa J, Krabbe N. Birds of the High Andes. Svendborg, Denmark: Zoological Museum, University of Copenhagen, and Apollo Books, 1990." type="book" year="1990">Fjeldså and Krabbe 1990: 554</bibRefCitation>
</treatmentCitation>
(in part);
<treatmentCitation id="0AA810DF836CFFCEB18B48F22183FA57" author="Collar NJ" box="[217,389,1509,1534]" page="700" pageId="22" pageNumber="250" year="2005">
<bibRefCitation id="EF984B3F836CFFCEB18B48F22183FA57" author="Collar NJ" box="[217,389,1509,1534]" pageId="22" pageNumber="250" refId="ref30149" refString="Collar NJ. Family Turdidae (Thrushes). In del Hoyo J, Elliou A, Christie DA, (eds). Handbook of the Birds of the World. Vol. 10. Cuckoo-shrikes to Thrushes, pp. 514 - 807. Barcelona: Lynx Edicions 2005." type="book" year="2005">Collar 2005: 700</bibRefCitation>
</treatmentCitation>
(in part);
<bibRefCitation id="EF984B3F836CFFCEB0BD48F2226DFA57" author="Halley MR" box="[495,619,1509,1534]" pageId="22" pageNumber="250" refId="ref31093" refString="Halley MR. Slaty-backed Nightingale-Thrush (Catharus fuscater), version 1.0. In: Schulenberg TS, ed. Birds of the World. Ithaca: Cornell Lab of Ornithology, 2020. hups: // doi. org / 10.2173 / bow. sbnthr 1.01" type="book" year="2020">Halley 2020</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB636CE836CFFCEB1D24B12227FF9F5" blockId="22.[127,763,1133,1659]" pageId="22" pageNumber="250">
<treatmentCitationGroup id="AB1911E0836CFFCEB1D24B12227FF9F5" pageId="22" pageNumber="250">
<treatmentCitation id="0AA810DF836CFFCEB1D24B1222D4F9B4" author="Fjeldsa J &amp; Krabbe N" box="[128,722,1540,1565]" page="554" pageId="22" pageNumber="250" year="1990">
<taxonomicName id="4C094D4D836CFFCEB1D24B1222D4F9B4" ID-CoL="5G9M8" authority="Fjeldsa and Krabbe 1990: 554" authorityName="Fjeldsa and Krabbe" authorityPageNumber="554" authorityYear="1990" box="[128,722,1540,1565]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="subSpecies" species="fuscater" subSpecies="fuscater">
<emphasis id="B97DEADC836CFFCEB1D24B122183F9B4" box="[128,389,1541,1565]" italics="true" pageId="22" pageNumber="250">Catharus fuscater fuscater</emphasis>
<bibRefCitation id="EF984B3F836CFFCEB0DC4B1322D4F9B4" author="Fjeldsa J &amp; Krabbe N" box="[398,722,1540,1565]" pageId="22" pageNumber="250" refId="ref30694" refString="Fjeldsa J, Krabbe N. Birds of the High Andes. Svendborg, Denmark: Zoological Museum, University of Copenhagen, and Apollo Books, 1990." type="book" year="1990">Fjeldså and Krabbe 1990: 554</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part);
<treatmentCitation id="0AA810DF836CFFCEB18B4B3321A7F995" author="Clement P" box="[217,417,1572,1596]" page="299" pageId="22" pageNumber="250" year="2000">
<bibRefCitation id="EF984B3F836CFFCEB18B4B3321A7F995" author="Clement P" box="[217,417,1572,1596]" pageId="22" pageNumber="250" refId="ref30112" refString="Clement P. Thrushes. Princeton: Princeton University Press 2000." type="book" year="2000">Clement 2000: 299</bibRefCitation>
</treatmentCitation>
(in part); Ridgley and Greenfield 2001: 660 (in part);
<treatmentCitation id="0AA810DF836CFFCEB03B4B542210F9F5" author="Collar NJ" box="[361,534,1603,1628]" page="700" pageId="22" pageNumber="250" year="2005">
<bibRefCitation id="EF984B3F836CFFCEB03B4B542210F9F5" author="Collar NJ" box="[361,534,1603,1628]" pageId="22" pageNumber="250" refId="ref30149" refString="Collar NJ. Family Turdidae (Thrushes). In del Hoyo J, Elliou A, Christie DA, (eds). Handbook of the Birds of the World. Vol. 10. Cuckoo-shrikes to Thrushes, pp. 514 - 807. Barcelona: Lynx Edicions 2005." type="book" year="2005">Collar 2005: 700</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB636CE836CFFCEB12D4B74224FF9D2" blockId="22.[127,763,1133,1659]" box="[127,585,1635,1659]" pageId="22" pageNumber="250">
<treatmentCitationGroup id="AB1911E0836CFFCEB12D4B74224FF9D2" box="[127,585,1635,1659]" pageId="22" pageNumber="250">
<treatmentCitation id="0AA810DF836CFFCEB12D4B742243F9D2" author="Halley MR" box="[127,581,1635,1659]" page="132" pageId="22" pageNumber="250" year="2021">
<taxonomicName id="4C094D4D836CFFCEB12D4B742243F9D2" authority="Halley 2021: 132" authorityName="Halley" authorityPageNumber="132" authorityYear="2021" box="[127,581,1635,1659]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="subSpecies" species="fuscater" subSpecies="berlepschi">
<emphasis id="B97DEADC836CFFCEB12D4B742188F9D2" box="[127,398,1635,1659]" italics="true" pageId="22" pageNumber="250">Catharus fuscater berlepschi</emphasis>
<bibRefCitation id="EF984B3F836CFFCEB0C14B742243F9D2" author="Halley MR" box="[403,581,1635,1659]" pageId="22" pageNumber="250" refId="ref31140" refString="Halley MR. Integrative systematics and evolution of seasonal migration in the nightingale-thrushes (Aves: Turdidae: Catharus). Ph. D. Thesis, Drexel University, Proºuest Dissertations Publishing. 2021." type="book" year="2021">Halley 2021: 132</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB636CE836CFFCEB1DF4BA022FCF979" blockId="22.[141,762,1719,1744]" box="[141,762,1719,1744]" pageId="22" pageNumber="250">
<emphasis id="B97DEADC836CFFCEB1DF4BA021FFF979" bold="true" box="[141,505,1719,1744]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB1DF4BA021A6F966" authority="Halley &amp; Catanach &amp; Klicka &amp; Weckstein, 2023" authorityName="Halley &amp; Catanach &amp; Klicka &amp; Weckstein" authorityYear="2023" box="[141,416,1719,1743]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="subSpecies" species="berlepschi" status="subsp. nov." subSpecies="nebulus">
<emphasis id="B97DEADC836CFFCEB1DF4BA021A6F966" bold="true" box="[141,416,1719,1743]" italics="true" pageId="22" pageNumber="250">Catharus berlepschi nebulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E57A7836CFFCEB0F64BAF21FFF979" box="[420,505,1720,1744]" pageId="22" pageNumber="250" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
(formerly Undescribed 3).
</paragraph>
</subSubSection>
<subSubSection id="C3136545836CFFCEB1DF4BF3211BF8B2" pageId="22" pageNumber="250" type="description">
<paragraph id="8BB636CE836CFFCEB1DF4BF3211BF8B2" blockId="22.[113,762,1764,1976]" pageId="22" pageNumber="250">
<uri id="FF983ACC836CFFCEB1DF4BF3211FF8B2" pageId="22" pageNumber="250">
urn:lsid:zoobank.org:act:
<uuid id="FFAF0C1B836CFFCEB0CA4BF3211FF8B2" pageId="22" pageNumber="250">EAE392CC-A763-4E5E-8F6F-68921515C9A2</uuid>
</uri>
.
</paragraph>
</subSubSection>
<subSubSection id="C3136545836CFFCEB1D24A3421C6F811" pageId="22" pageNumber="250" type="reference_group">
<paragraph id="8BB636CE836CFFCEB1D24A342100F8F3" blockId="22.[113,762,1764,1976]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB1D24A342223F892" ID-CoL="69G37" authority="Taczanowski 1874: 504" authorityName="Taczanowski" authorityPageNumber="504" authorityYear="1874" box="[128,549,1827,1851]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="species" species="fuscater">
<emphasis id="B97DEADC836CFFCEB1D24A34212AF892" box="[128,300,1827,1851]" italics="true" pageId="22" pageNumber="250">Catharus fuscater</emphasis>
Taczanowski 1874: 504
</taxonomicName>
; 1879: 222; 1882: 4; 1884: 483.
</paragraph>
<paragraph id="8BB636CE836CFFCEB1D24A7622C7F830" blockId="22.[113,762,1764,1976]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB1D24A76229DF8D0" ID-CoL="5G9M7" authority="Fjeldsa and Krabbe 1990: 554" authorityName="Fjeldsa and Krabbe" authorityPageNumber="554" authorityYear="1990" box="[128,667,1889,1914]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="subSpecies" species="fuscater" subSpecies="caniceps">
<emphasis id="B97DEADC836CFFCEB1D24A762173F8D0" box="[128,373,1889,1913]" italics="true" pageId="22" pageNumber="250">Catharus fuscater caniceps</emphasis>
<bibRefCitation id="EF984B3F836CFFCEB02A4A76229DF8D0" author="Fjeldsa J &amp; Krabbe N" box="[376,667,1889,1914]" pageId="22" pageNumber="250" refId="ref30694" refString="Fjeldsa J, Krabbe N. Birds of the High Andes. Svendborg, Denmark: Zoological Museum, University of Copenhagen, and Apollo Books, 1990." type="book" year="1990">Fjeldså and Krabbe 1990: 554</bibRefCitation>
</taxonomicName>
(in part);
<bibRefCitation id="EF984B3F836CFFCEB1CA4A962151F830" author="Clement P" box="[152,343,1921,1945]" pageId="22" pageNumber="250" refId="ref30112" refString="Clement P. Thrushes. Princeton: Princeton University Press 2000." type="book" year="2000">Clement 2000: 299</bibRefCitation>
(in part);
<bibRefCitation id="EF984B3F836CFFCEB0EE4A962264F830" author="Collar NJ" box="[444,610,1921,1945]" pageId="22" pageNumber="250" refId="ref30149" refString="Collar NJ. Family Turdidae (Thrushes). In del Hoyo J, Elliou A, Christie DA, (eds). Handbook of the Birds of the World. Vol. 10. Cuckoo-shrikes to Thrushes, pp. 514 - 807. Barcelona: Lynx Edicions 2005." type="book" year="2005">Collar 2005: 700</bibRefCitation>
(in part).
</paragraph>
<paragraph id="8BB636CE836CFFCEB12D4AB721C6F811" blockId="22.[113,762,1764,1976]" box="[127,448,1952,1976]" pageId="22" pageNumber="250">
Unnamed-3
<bibRefCitation id="EF984B3F836CFFCEB0584AB721BAF811" author="Halley MR" box="[266,444,1952,1976]" pageId="22" pageNumber="250" refId="ref31140" refString="Halley MR. Integrative systematics and evolution of seasonal migration in the nightingale-thrushes (Aves: Turdidae: Catharus). Ph. D. Thesis, Drexel University, Proºuest Dissertations Publishing. 2021." type="book" year="2021">Halley 2021: 133</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3136545836CFFCEB57C4D862509FC54" pageId="22" pageNumber="250" type="materials_examined">
<paragraph id="8BB636CE836CFFCEB57C4D8624A9FF03" blockId="22.[810,1460,144,1021]" box="[1070,1199,144,170]" pageId="22" pageNumber="250">
<emphasis id="B97DEADC836CFFCEB57C4D8624A9FF03" box="[1070,1199,144,170]" italics="true" pageId="22" pageNumber="250">
<typeStatus id="54B2886C836CFFCEB57C4D86245AFF03" box="[1070,1116,145,170]" pageId="22" pageNumber="250">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BB636CE836CFFCEB2784DA1251AFE60" blockId="22.[810,1460,144,1021]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB2784DA1240FFF67" authority=": AMNH" authorityName="Bonaparte" authorityYear="1850" box="[810,1033,182,206]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB2784DA1240FFF67" box="[810,1033,182,206]" italics="true" pageId="22" pageNumber="250">Catharus b. berlepschi</emphasis>
</taxonomicName>
:
<materialsCitation id="3B613C93836CFFCEB5484DA0246DFF66" ID-GBIF-Occurrence="4406001439" box="[1050,1131,183,207]" collectingDate="1883-01-23" country="Ecuador" location="Joseph Siemiradzki in Cayandeled" pageId="22" pageNumber="250" specimenCount="1" stateProvince="Chimborazo" typeStatus="holotype">AMNH</materialsCitation>
39096 (
<typeStatus id="54B2886C836CFFCEB5954DA1252DFF67" box="[1223,1323,182,206]" pageId="22" pageNumber="250" type="holotype">holotype</typeStatus>
), study skin, adult male, collected by
<location id="8ED66015836CFFCEB5124DC125B5FF47" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB5124DC125B5FF47" box="[1088,1459,214,238]" country="Ecuador" name="Joseph Siemiradzki in Cayandeled" pageId="22" pageNumber="250" stateProvince="Chimborazo">Joseph Siemiradzki in Cayandeled</location>
(approximately 2.117°, 78.98°; see: Paynter 1993: 34),
<collectingRegion id="49CDF82C836CFFCEB2784C0323AAFE85" box="[810,940,276,300]" country="Ecuador" name="Chimborazo" pageId="22" pageNumber="250">Chimborazo</collectingRegion>
,
<collectingCountry id="F31E765E836CFFCEB2924C032411FE85" box="[960,1047,276,300]" name="Ecuador" pageId="22" pageNumber="250">Ecuador</collectingCountry>
, on
<date id="FFB7100E836CFFCEB5044C032514FE85" box="[1110,1298,276,301]" pageId="22" pageNumber="250" value="1883-01-23">
<collectingDate id="EFF3E9E6836CFFCEB5044C032514FE85" box="[1110,1298,276,301]" pageId="22" pageNumber="250" value="1883-01-23">23 January 1883</collectingDate>
</date>
(see:
<bibRefCitation id="EF984B3F836CFFCEB4364C032397FEE5" author="LeCroy M" pageId="22" pageNumber="250" pagination="1 - 132" refId="ref32207" refString="LeCroy M. Type specimens of birds in the American Museum of Natural History. Part 6, Passeriformes: Prunellidae, Turdidae, Orthonychidae, Timaliidae, Paradoxornithidae, Picathartidae, and Polioptilidae. Bulletin of the American Museum of Natural History 2005; 292: 1 - 132." type="journal article" year="2005">LeCroy 2005: 38</bibRefCitation>
).
<materialsCitation id="3B613C93836CFFCEB2FE4C23251EFE60" ID-GBIF-Occurrence="4406001434" collectingDate="2013-06-20" collectionCode="R" country="Ecuador" pageId="22" pageNumber="250" specimenCode="AMNH 39096" specimenCount="1" specimenCount-male="1" stateProvince="Chimborazo" typeStatus="holotype">
Siemiradzki collected
<specimenCount id="9D0FFD47836CFFCEB5CE4C232516FEE5" box="[1180,1296,308,332]" pageId="22" pageNumber="250" type="male">four males</specimenCount>
and a female, which
<bibRefCitation id="EF984B3F836CFFCEB2224C442513FEC2" author="von Berlepsch HG &amp; Taczanowski L" box="[880,1301,339,363]" pageId="22" pageNumber="250" pagination="281 - 313" refId="ref29489" refString="von Berlepsch HG, Taczanowski L. Deuxieme liste des Oiseaux recueillis dans l'Ecuadeur occidental par MM. Stolzmann et Siemiradski. Proceedings of the Zoological Society of London 1884; 1884: 281 - 313." type="journal article" year="1884">Berlepsch and Taczanowski (1884: 283)</bibRefCitation>
classified as
<taxonomicName id="4C094D4D836CFFCEB4CF4C442373FE23" authorityName="Taczanowski" authorityYear="1874" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="species" species="fuscater">
<emphasis id="B97DEADC836CFFCEB4CF4C442373FE23" italics="true" pageId="22" pageNumber="250">C. fuscater</emphasis>
</taxonomicName>
. Berlepsch then sent one of the males (
<specimenCode id="DBAF9EB5836CFFCEB45E4C6525AAFE23" box="[1292,1452,370,394]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 39096</specimenCode>
) to Lawrence, who described it as a new species. For this study, M.
<collectionCode id="ED18AE0B836CFFCEB21A4CA6235AFE61" box="[840,860,433,456]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="250">R</collectionCode>
.H. examined the
<typeStatus id="54B2886C836CFFCEB55F4CA6246FFE60" box="[1037,1129,433,457]" pageId="22" pageNumber="250" type="holotype">holotype</typeStatus>
on
<date id="FFB7100E836CFFCEB5DD4CA6251EFE60" box="[1167,1304,433,457]" pageId="22" pageNumber="250" value="2013-06-20">
<collectingDate id="EFF3E9E6836CFFCEB5DD4CA6251EFE60" box="[1167,1304,433,457]" pageId="22" pageNumber="250" value="2013-06-20">20 June 2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636CE836CFFCEB2174CC72363FD2C" blockId="22.[810,1460,144,1021]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB2174CC7240BFE41" authorityName="Bonaparte" authorityYear="1850" box="[837,1037,464,488]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB2174CC7240BFE41" box="[837,1037,464,488]" italics="true" pageId="22" pageNumber="250">Catharus b. caniceps</emphasis>
</taxonomicName>
:
<materialsCitation id="3B613C93836CFFCEB5494CC72417FDCC" ID-GBIF-Occurrence="4406001499" collectingDate="1922-09-16" collectionCode="AMNH" collectorName="H. Watkins" country="Peru" location="Piura" pageId="22" pageNumber="250" specimenCode="AMNH 175530" specimenCount="1" stateProvince="Piura" typeStatus="holotype">
<specimenCode id="DBAF9EB5836CFFCEB5494CC724C6FE41" box="[1051,1216,464,488]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 175530</specimenCode>
(
<typeStatus id="54B2886C836CFFCEB59C4CC72534FE41" box="[1230,1330,464,488]" pageId="22" pageNumber="250" type="holotype">holotype</typeStatus>
), study skin, adult male, collected by
<collectorName id="26FC5318836CFFCEB5734CE72495FDA1" box="[1057,1171,496,520]" pageId="22" pageNumber="250">H. Watkins</collectorName>
(no. 6011) at Palambla (approximately 5.23°, 79.37°; see: Stephens and Traylor 1983: 153),
<collectingRegion id="49CDF82C836CFFCEB2384F3923A4FDEF" box="[874,930,558,582]" country="Peru" name="Piura" pageId="22" pageNumber="250">Piura</collectingRegion>
,
<collectingCountry id="F31E765E836CFFCEB2FD4F3823E7FDEF" box="[943,993,559,582]" name="Peru" pageId="22" pageNumber="250">Peru</collectingCountry>
, on
<date id="FFB7100E836CFFCEB5424F3924D8FDEF" box="[1040,1246,558,582]" pageId="22" pageNumber="250" value="1922-09-16">
<collectingDate id="EFF3E9E6836CFFCEB5424F3924D8FDEF" box="[1040,1246,558,582]" pageId="22" pageNumber="250" value="1922-09-16">16 September 1922</collectingDate>
</date>
(see:
<bibRefCitation id="EF984B3F836CFFCEB4734F39234EFDCF" author="LeCroy M" pageId="22" pageNumber="250" pagination="1 - 132" refId="ref32207" refString="LeCroy M. Type specimens of birds in the American Museum of Natural History. Part 6, Passeriformes: Prunellidae, Turdidae, Orthonychidae, Timaliidae, Paradoxornithidae, Picathartidae, and Polioptilidae. Bulletin of the American Museum of Natural History 2005; 292: 1 - 132." type="journal article" year="2005">LeCroy 2005: 39</bibRefCitation>
). For this study, M
</materialsCitation>
.
<materialsCitation id="3B613C93836CFFCEB5464F592366FD2C" ID-GBIF-Occurrence="4406001400" collectingDate="2013-06-20" collectionCode="R" country="Peru" pageId="22" pageNumber="250" specimenCount="1" stateProvince="Piura" typeStatus="holotype">
<collectionCode id="ED18AE0B836CFFCEB5464F59242EFDCC" box="[1044,1064,590,613]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="22" pageNumber="250">R</collectionCode>
.H. examined the
<typeStatus id="54B2886C836CFFCEB58C4F5A253CFDCC" box="[1246,1338,589,613]" pageId="22" pageNumber="250" type="holotype">holotype</typeStatus>
on
<date id="FFB7100E836CFFCEB4314F592366FD2C" pageId="22" pageNumber="250" value="2013-06-20">
<collectingDate id="EFF3E9E6836CFFCEB4314F592366FD2C" pageId="22" pageNumber="250" value="2013-06-20">20 June 2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636CE836CFFCEB2174F9B2509FC54" blockId="22.[810,1460,144,1021]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB2174F9B2417FD0D" authorityName="Bonaparte" authorityYear="1850" box="[837,1041,652,676]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB2174F9B2417FD0D" box="[837,1041,652,676]" italics="true" pageId="22" pageNumber="250">Catharus b. nebulus</emphasis>
</taxonomicName>
<materialsCitation id="3B613C93836CFFCEB54E4F9A24FFFCAB" ID-GBIF-Occurrence="4406001494" collectingDate="2010-06-23" collectionCode="FMNH" collectorName="David E. Willard" country="Peru" elevation="2400" location="Willard on Cerro Monte Alegre" municipality="Amazonas" pageId="22" pageNumber="250" specimenCode="FMNH 474413" specimenCount="2" typeStatus="holotype">
ssp. nov.:
<specimenCode id="DBAF9EB5836CFFCEB5DB4F9A2536FD0D" box="[1161,1328,652,676]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="22" pageNumber="250" type="Museum">FMNH 474413</specimenCode>
(
<typeStatus id="54B2886C836CFFCEB4114F9B25A1FD0D" box="[1347,1447,652,676]" pageId="22" pageNumber="250" type="holotype">holotype</typeStatus>
), study skin, adult male, collected and prepared by
<collectorName id="26FC5318836CFFCEB4064FBC2371FD4A" pageId="22" pageNumber="250">
David E.
<location id="8ED66015836CFFCEB2784FDC2371FD4A" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB2784FDC2371FD4A" box="[810,887,715,739]" country="Peru" municipality="Amazonas" name="Willard on Cerro Monte Alegre" pageId="22" pageNumber="250">Willard</location>
</collectorName>
on Cerro Monte Alegre (6.5850°, 77.5617°, elev.
<quantity id="4CF19B2B836CFFCEB2784FFD237BFCAB" box="[810,893,746,770]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="250" unit="m" value="2400.0">
<elevation id="0024D1FD836CFFCEB2784FFD237BFCAB" box="[810,893,746,770]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="250" unit="m" value="2400.0">2400 m</elevation>
</quantity>
),
<collectingMunicipality id="6BD2ACB4836CFFCEB2C34FFD23FBFCAB" box="[913,1021,746,770]" pageId="22" pageNumber="250">Amazonas</collectingMunicipality>
,
<collectingCountry id="F31E765E836CFFCEB55A4FFC243CFCAB" box="[1032,1082,747,770]" name="Peru" pageId="22" pageNumber="250">Peru</collectingCountry>
, on
<date id="FFB7100E836CFFCEB5354FFD24F3FCAB" box="[1127,1269,746,770]" pageId="22" pageNumber="250" value="2010-06-23">
<collectingDate id="EFF3E9E6836CFFCEB5354FFD24F3FCAB" box="[1127,1269,746,770]" pageId="22" pageNumber="250" value="2010-06-23">23 June 2010</collectingDate>
</date>
.
</materialsCitation>
When it was prepared, the
<typeStatus id="54B2886C836CFFCEB2C44E1E23F4FC88" box="[918,1010,777,801]" pageId="22" pageNumber="250" type="holotype">holotype</typeStatus>
weighed
<quantity id="4CF19B2B836CFFCEB5064E1E2492FC8B" box="[1108,1172,777,802]" metricMagnitude="-2" metricUnit="kg" metricValue="3.75" pageId="22" pageNumber="250" unit="g" value="37.5">37.5 g</quantity>
. Its testes were not enlarged (1 ×
<quantity id="4CF19B2B836CFFCEB20D4E3E23B2FCE8" box="[863,948,809,833]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="250" unit="mm" value="0.5">0.5 mm</quantity>
) and the skull was 50% pneumatized. In 2022, the entire dorsal surface was darker than Sepia (119), with the crown slightly darker than the back and rump. The undertail coverts were Dark Drab (119B) and the flanks were darker and cooler than Vandyke Brown (121). The breast was darker than Glaucous (79) with faint mottling. The throat was cooler than Drab-Grey (119D), contrasting with the breast.
</paragraph>
</subSubSection>
<subSubSection id="C3136545836CFFCEB54F493F24FBFA29" pageId="22" pageNumber="250" type="distribution">
<paragraph id="8BB636CE836CFFCEB54F493F24C6FBEB" blockId="22.[810,1460,1064,1408]" box="[1053,1216,1064,1090]" pageId="22" pageNumber="250">
<emphasis id="B97DEADC836CFFCEB54F493F24C6FBEB" box="[1053,1216,1064,1090]" italics="true" pageId="22" pageNumber="250">Geographic range</emphasis>
</paragraph>
<paragraph id="8BB636CE836CFFCEB2784959239FFB2F" blockId="22.[810,1460,1064,1408]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB278495923F9FBCF" authorityName="Bonaparte" authorityYear="1850" box="[810,1023,1102,1126]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB278495923F9FBCF" box="[810,1023,1102,1126]" italics="true" pageId="22" pageNumber="250">Catharus b. berlepschi</emphasis>
</taxonomicName>
:
<location id="8ED66015836CFFCEB559495824A0FBCF" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB559495824A0FBCF" box="[1035,1190,1102,1126]" country="Ecuador" name="Western Andes" pageId="22" pageNumber="250" stateProvince="Carchi">Western Andes</location>
in
<collectingCountry id="F31E765E836CFFCEB5964959251CFBCF" box="[1220,1306,1102,1126]" name="Ecuador" pageId="22" pageNumber="250">Ecuador</collectingCountry>
(
<collectingRegion id="49CDF82C836CFFCEB47549592576FBCF" box="[1319,1392,1102,1126]" country="Ecuador" name="Carchi" pageId="22" pageNumber="250">Carchi</collectingRegion>
, south to
<collectingRegion id="49CDF82C836CFFCEB2164979238BFB2F" box="[836,909,1134,1158]" country="Ecuador" name="El Oro" pageId="22" pageNumber="250">El Oro</collectingRegion>
).
</paragraph>
<paragraph id="8BB636CE836CFFCEB217499A25B2FB4D" blockId="22.[810,1460,1064,1408]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB217499A2413FB0C" authorityName="Bonaparte" authorityYear="1850" box="[837,1045,1165,1189]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB217499A2413FB0C" box="[837,1045,1165,1189]" italics="true" pageId="22" pageNumber="250">Catharus b. caniceps</emphasis>
</taxonomicName>
:
<collectingMunicipality id="6BD2ACB4836CFFCEB575499A24C0FB0C" box="[1063,1222,1165,1189]" pageId="22" pageNumber="250">Western Andes</collectingMunicipality>
in
<collectingCountry id="F31E765E836CFFCEB5BD499A2543FB0C" box="[1263,1349,1165,1189]" name="Ecuador" pageId="22" pageNumber="250">Ecuador</collectingCountry>
(El Oro), south to north-western
<collectingCountry id="F31E765E836CFFCEB54F49BA244BFB6D" box="[1053,1101,1197,1220]" name="Peru" pageId="22" pageNumber="250">Peru</collectingCountry>
(
<collectingRegion id="49CDF82C836CFFCEB50849BB24CAFB6D" box="[1114,1228,1196,1220]" country="Peru" name="Cajamarca" pageId="22" pageNumber="250">Cajamarca</collectingRegion>
,
<collectingRegion id="49CDF82C836CFFCEB58449BA2552FB6D" box="[1238,1364,1196,1220]" country="Peru" name="La Libertad" pageId="22" pageNumber="250">La Libertad</collectingRegion>
), and the Huancabamba and Chunchuca valleys, west of the
<location id="8ED66015836CFFCEB47449DB25B6FB4D" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB47449DB25B6FB4D" box="[1318,1456,1228,1252]" country="Ecuador" municipality="Western Andes" name="Rio Maranon" pageId="22" pageNumber="250" stateProvince="Cajamarca">Río Marañón</location>
.
</paragraph>
<paragraph id="8BB636CE836CFFCEB21749FC24FBFA29" blockId="22.[810,1460,1064,1408]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB21749FC240DFAAA" authorityName="Bonaparte" authorityYear="1850" box="[837,1035,1259,1283]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB21749FC240DFAAA" box="[837,1035,1259,1283]" italics="true" pageId="22" pageNumber="250">Catharus b. nebulus</emphasis>
</taxonomicName>
ssp. nov.:
<collectingCounty id="62D74E42836CFFCEB52E49FC24CFFAAB" box="[1148,1225,1259,1282]" pageId="22" pageNumber="250">Eastern</collectingCounty>
slope of the
<collectingMunicipality id="6BD2ACB4836CFFCEB40A49FC236DFA8B" pageId="22" pageNumber="250">Peruvian Andes</collectingMunicipality>
from
<collectingRegion id="49CDF82C836CFFCEB2FF481C2411FA8A" box="[941,1047,1291,1315]" country="Venezuela" name="Amazonas" pageId="22" pageNumber="250">Amazonas</collectingRegion>
(east of
<location id="8ED66015836CFFCEB524481D2502FA8B" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB524481D2502FA8B" box="[1142,1284,1290,1314]" county="Eastern" municipality="Peruvian Andes" name="Rio Maranon" pageId="22" pageNumber="250" stateProvince="Amazonas">Río Marañón</location>
) south to
<collectingRegion id="49CDF82C836CFFCEB420481D25B5FA8B" box="[1394,1459,1290,1314]" country="Peru" name="Cusco" pageId="22" pageNumber="250">Cusco</collectingRegion>
(north-west of
<location id="8ED66015836CFFCEB2BA483D240BFAEB" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB2BA483D240BFAEB" box="[1000,1037,1322,1346]" county="Eastern" municipality="Peruvian Andes" name="Rio" pageId="22" pageNumber="250" stateProvince="Amazonas">Río</location>
<collectingRegion id="49CDF82C836CFFCEB576483D2489FAEB" box="[1060,1167,1322,1346]" country="Peru" name="Apurimac" pageId="22" pageNumber="250">Apurímac</collectingRegion>
), including
<location id="8ED66015836CFFCEB478483D25B5FAEB" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB478483D25B5FAEB" box="[1322,1459,1322,1346]" county="Eastern" municipality="Peruvian Andes" name="Chachapoyas" pageId="22" pageNumber="250" stateProvince="Amazonas">Chachapoyas</location>
(
<collectingRegion id="49CDF82C836CFFCEB263485E23A5FAC8" box="[817,931,1353,1377]" country="Venezuela" name="Amazonas" pageId="22" pageNumber="250">Amazonas</collectingRegion>
),
<location id="8ED66015836CFFCEB2EA485E2413FAC8" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB2EA485E2413FAC8" box="[952,1045,1353,1377]" county="Eastern" municipality="Peruvian Andes" name="Tambillo" pageId="22" pageNumber="250" stateProvince="Amazonas">Tambillo</location>
(
<location id="8ED66015836CFFCEB574485E2498FAC8" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB574485E2498FAC8" box="[1062,1182,1353,1377]" county="Eastern" municipality="Peruvian Andes" name="Huamanga" pageId="22" pageNumber="250" stateProvince="Amazonas">Huamanga</location>
), and
<location id="8ED66015836CFFCEB5B3485E2536FAC8" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB5B3485E2536FAC8" box="[1249,1328,1353,1377]" county="Eastern" municipality="Peruvian Andes" name="Chilpes" pageId="22" pageNumber="250" stateProvince="Amazonas">Chilpes</location>
(
<collectingRegion id="49CDF82C836CFFCEB41A485E2584FAC8" box="[1352,1410,1353,1377]" country="Peru" name="Junin" pageId="22" pageNumber="250">Junín</collectingRegion>
) between 1770 and
<quantity id="4CF19B2B836CFFCEB280487F2426FA29" box="[978,1056,1384,1408]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="250" unit="m" value="2290.0">
<elevation id="0024D1FD836CFFCEB280487F2426FA29" box="[978,1056,1384,1408]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="22" pageNumber="250" unit="m" value="2290.0">2290 m</elevation>
</quantity>
(Taczanowski 1884).
</paragraph>
</subSubSection>
<subSubSection id="C3136545836CFFC2B2A648B420EBFE48" lastPageId="26" lastPageNumber="254" pageId="22" pageNumber="250" type="materials_examined">
<paragraph id="8BB636CE836CFFCEB2A648B424EFFA14" blockId="22.[1012,1257,1443,1469]" box="[1012,1257,1443,1469]" pageId="22" pageNumber="250">
<emphasis id="B97DEADC836CFFCEB2A648B424EFFA14" box="[1012,1257,1443,1469]" italics="true" pageId="22" pageNumber="250">Adult specimens examined</emphasis>
</paragraph>
<paragraph id="8BB636CE836CFFCEB27848DD253DF975" blockId="22.[810,1460,1482,1976]" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB27848DD2417FA4B" authorityName="Bonaparte" authorityYear="1850" box="[810,1041,1482,1506]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB27848DD2417FA4B" box="[810,1041,1482,1506]" italics="true" pageId="22" pageNumber="250">Catharus b. berlepschi</emphasis>
</taxonomicName>
(
<emphasis id="B97DEADC836CFFCEB57848DD243BFA48" box="[1066,1085,1482,1505]" italics="true" pageId="22" pageNumber="250">N</emphasis>
= 11):
<materialsCitation id="3B613C93836CFFCEB5C748DD23DDF9E9" ID-GBIF-Occurrence="4406001328" collectionCode="ANSP" collectorName="Maldonado-Tulcan Rd" country="Ecuador" location="Rio La Plata" pageId="22" pageNumber="250" specimenCode="ANSP 181213, ANSP 181212" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Carchi">
<collectingCountry id="F31E765E836CFFCEB5C748DD24EDFA4B" box="[1173,1259,1482,1506]" name="Ecuador" pageId="22" pageNumber="250">Ecuador</collectingCountry>
(
<emphasis id="B97DEADC836CFFCEB45648DD2511FA48" box="[1284,1303,1482,1505]" italics="true" pageId="22" pageNumber="250">N</emphasis>
= 11):
<collectingRegion id="49CDF82C836CFFCEB43C48DD25B5FA4B" box="[1390,1459,1482,1506]" country="Ecuador" name="Carchi" pageId="22" pageNumber="250">Carchi</collectingRegion>
(
<specimenCount id="9D0FFD47836CFFCEB26048FE23A4F9A8" box="[818,930,1513,1537]" pageId="22" pageNumber="250" type="male">two males</specimenCount>
,
<specimenCount id="9D0FFD47836CFFCEB2E248FD242AF9A8" box="[944,1068,1513,1537]" pageId="22" pageNumber="250" type="female">one female</specimenCount>
): west slope near
<collectorName id="26FC5318836CFFCEB5BD48FE234CF989" pageId="22" pageNumber="250">Maldonado-Tulcán Rd</collectorName>
, along
<location id="8ED66015836CFFCEB2C74B1F241CF989" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB2C74B1F241CF989" box="[917,1050,1544,1568]" country="Ecuador" name="Rio La Plata" pageId="22" pageNumber="250" stateProvince="Carchi">Río La Plata</location>
:
<specimenCode id="DBAF9EB5836CFFCEB5764B1F24BCF989" box="[1060,1210,1544,1568]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="250">ANSP 181213</specimenCode>
181214 (males),
<specimenCode id="DBAF9EB5836CFFCEB4204B1F237EF9E9" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="250">ANSP 181212</specimenCode>
(female)
</materialsCitation>
;
<materialsCitation id="3B613C93836CFFCEB2B44B3F241CF9F6" ID-GBIF-Occurrence="4406001457" collectionCode="ANSP, DMNH, AMNH" collectorName="Chiriboga &amp; Mindo" country="Ecuador" location="Chunchi" pageId="22" pageNumber="250" specimenCode="ANSP 59755-59757, DMNH 59252, 62883, AMNH 503903" specimenCount="2" specimenCount-male="2" stateProvince="Chimborazo">
<collectingRegion id="49CDF82C836CFFCEB2B44B3F2461F9E9" box="[998,1127,1576,1600]" country="Ecuador" name="Chimborazo" pageId="22" pageNumber="250">Chimborazo</collectingRegion>
(
<specimenCount id="9D0FFD47836CFFCEB5244B3F24F1F9E9" box="[1142,1271,1576,1600]" pageId="22" pageNumber="250" type="male">three males</specimenCount>
):
<location id="8ED66015836CFFCEB45B4B3F2561F9E9" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB45B4B3F2561F9E9" box="[1289,1383,1576,1600]" country="Ecuador" name="Chunchi" pageId="22" pageNumber="250" stateProvince="Chimborazo">Chunchi</location>
:
<specimenCode id="DBAF9EB5836CFFCEB4204B3F23BCF9F6" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="250">ANSP 5975559757</specimenCode>
(males);
</materialsCitation>
<materialsCitation id="3B613C93836CFFCEB5774B50246EF9D7" ID-GBIF-Occurrence="4406001413" collectionCode="ANSP, DMNH, AMNH" collectorName="Chiriboga &amp; Mindo" country="Ecuador" location="Chunchi" pageId="22" pageNumber="250" specimenCode="ANSP 59755-59757, DMNH 59252, 62883, AMNH 503903" specimenCount="2" specimenCount-male="2" stateProvince="Chimborazo">
Pinchincha (
<specimenCount id="9D0FFD47836CFFCEB5F84B502528F9F6" box="[1194,1326,1607,1631]" pageId="22" pageNumber="250" type="male">three males</specimenCount>
):
<collectorName id="26FC5318836CFFCEB4164B5025B6F9F6" box="[1348,1456,1607,1631]" pageId="22" pageNumber="250">Chiriboga</collectorName>
:
<specimenCode id="DBAF9EB5836CFFCEB2784B7023C3F9D7" box="[810,965,1638,1662]" collectionCode="DMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" lsid="urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="22" pageNumber="250" type="Museum">DMNH 59252</specimenCode>
,
<specimenCode id="DBAF9EB5836CFFCEB29F4B712408F9D7" box="[973,1038,1638,1662]" collectionCode="DMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" lsid="urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="22" pageNumber="250" type="Museum">62883</specimenCode>
(males);
</materialsCitation>
<materialsCitation id="3B613C93836CFFCEB53F4B7125A9F9D7" ID-GBIF-Occurrence="4406001366" box="[1133,1455,1638,1663]" collectionCode="ANSP, DMNH, AMNH" collectorName="Chiriboga &amp; Mindo" country="Ecuador" location="Chunchi" pageId="22" pageNumber="250" specimenCode="ANSP 59755-59757, DMNH 59252, 62883, AMNH 503903" specimenCount="2" specimenCount-male="2" stateProvince="Chimborazo">
<collectorName id="26FC5318836CFFCEB53F4B7124B1F9D7" box="[1133,1207,1638,1662]" pageId="22" pageNumber="250">Mindo</collectorName>
:
<specimenCode id="DBAF9EB5836CFFCEB5ED4B702564F9D7" box="[1215,1378,1638,1663]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 503903</specimenCode>
(male)
</materialsCitation>
;
<materialsCitation id="3B613C93836CFFCEB2784B912371F914" ID-GBIF-Occurrence="4406001428" collectionCode="AMNH" country="Ecuador" location="El Oro" pageId="22" pageNumber="250" specimenCode="AMNH 167890" specimenCount="1" specimenCount-male="1" stateProvince="El Oro">
<collectingRegion id="49CDF82C836CFFCEB2784B912370F937" box="[810,886,1670,1694]" country="Ecuador" name="El Oro" pageId="22" pageNumber="250">El Oro</collectingRegion>
(
<specimenCount id="9D0FFD47836CFFCEB2D94B9123F8F937" box="[907,1022,1670,1694]" pageId="22" pageNumber="250" type="male">two males</specimenCount>
,
<specimenCount id="9D0FFD47836CFFCEB55D4B912491F937" box="[1039,1175,1670,1694]" pageId="22" pageNumber="250" type="female">two females</specimenCount>
):
<collectorName id="26FC5318836CFFCEB5FC4B9124FEF937" box="[1198,1272,1670,1694]" pageId="22" pageNumber="250">Salvias</collectorName>
:
<specimenCode id="DBAF9EB5836CFFCEB4554B9125B5F937" box="[1287,1459,1670,1694]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 167890</specimenCode>
(male);
</materialsCitation>
<materialsCitation id="3B613C93836CFFCEB22C4BB223DFF975" ID-GBIF-Occurrence="4406001508" collectionCode="AMNH, ANSP" country="Ecuador" location="Zaruma" pageId="22" pageNumber="250" specimenCode="ANSP 83643, AMNH 130154, ANSP 83644" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectorName id="26FC5318836CFFCEB22C4BB223D2F915" box="[894,980,1701,1724]" pageId="22" pageNumber="250">Zaruma</collectorName>
:
<specimenCode id="DBAF9EB5836CFFCEB28C4BB22460F914" box="[990,1126,1701,1725]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="250">ANSP 83643</specimenCode>
(male),
<specimenCode id="DBAF9EB5836CFFCEB5ED4BB22561F914" box="[1215,1383,1701,1725]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 130154</specimenCode>
,
<specimenCode id="DBAF9EB5836CFFCEB4204BB2236DF975" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="250">ANSP 83644</specimenCode>
(females);
</materialsCitation>
<materialsCitation id="3B613C93836CFFCEB28C4BD32531F975" ID-GBIF-Occurrence="4406001395" box="[990,1335,1732,1756]" collectionCode="AMNH" country="Ecuador" location="El Chiral" pageId="22" pageNumber="250" specimenCode="AMNH 167888" specimenCount="1" specimenCount-male="1" stateProvince="El Oro">
<collectorName id="26FC5318836CFFCEB28C4BD32438F975" box="[990,1086,1732,1756]" pageId="22" pageNumber="250">El Chiral</collectorName>
:
<specimenCode id="DBAF9EB5836CFFCEB5144BD324ECF975" box="[1094,1258,1732,1756]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 167888</specimenCode>
(male)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636CE836CFFCFB2174BF32452F80C" blockId="22.[810,1460,1482,1976]" lastBlockId="23.[825,1475,1808,1957]" lastPageId="23" lastPageNumber="251" pageId="22" pageNumber="250">
<taxonomicName id="4C094D4D836CFFCEB2174BF32423F952" authorityName="Bonaparte" authorityYear="1850" box="[837,1061,1764,1788]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="22" pageNumber="250" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC836CFFCEB2174BF32423F952" box="[837,1061,1764,1788]" italics="true" pageId="22" pageNumber="250">Catharus b. caniceps</emphasis>
</taxonomicName>
(
<emphasis id="B97DEADC836CFFCEB5134BF32452F952" box="[1089,1108,1764,1787]" italics="true" pageId="22" pageNumber="250">N</emphasis>
= 33):
<materialsCitation id="3B613C93836CFFCEB5FD4BF32368F893" ID-GBIF-Occurrence="4406001503" collectionCode="AMNH" collectorName="San Bartolo" country="Ecuador" location="Celica" municipality="Alamor" pageId="22" pageNumber="250" specimenCode="AMNH 167894, AMNH 167891, 167893, AMNH 167892, AMNH 172116, AMNH 172117" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja">
<collectingCountry id="F31E765E836CFFCEB5FD4BF32501F955" box="[1199,1287,1764,1788]" name="Ecuador" pageId="22" pageNumber="250">Ecuador</collectingCountry>
(
<emphasis id="B97DEADC836CFFCEB4714BF32530F952" box="[1315,1334,1764,1787]" italics="true" pageId="22" pageNumber="250">N</emphasis>
= 6):
<collectingRegion id="49CDF82C836CFFCEB4D64BF325B5F955" box="[1412,1459,1764,1788]" country="Ecuador" name="Loja" pageId="22" pageNumber="250">Loja</collectingRegion>
(
<specimenCount id="9D0FFD47836CFFCEB2604A1423B3F8B2" box="[818,949,1795,1819]" pageId="22" pageNumber="250" type="male">three males</specimenCount>
,
<specimenCount id="9D0FFD47836CFFCEB2964A14245AF8B2" box="[964,1116,1795,1819]" pageId="22" pageNumber="250" type="female">three females</specimenCount>
):
<collectingMunicipality id="6BD2ACB4836CFFCEB5214A1424C0F8B2" box="[1139,1222,1795,1819]" pageId="22" pageNumber="250">Alamor</collectingMunicipality>
:
<specimenCode id="DBAF9EB5836CFFCEB5874A142587F8B2" box="[1237,1409,1795,1819]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 167894</specimenCode>
(female);
</materialsCitation>
<materialsCitation id="3B613C93836CFFCEB22C4A3523EBF8F3" ID-GBIF-Occurrence="4406001383" collectionCode="AMNH" collectorName="San Bartolo" country="Ecuador" location="Celica" municipality="Alamor" pageId="22" pageNumber="250" specimenCode="AMNH 167894, AMNH 167891, 167893, AMNH 167892, AMNH 172116, AMNH 172117" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja">
<location id="8ED66015836CFFCEB22C4A3523C2F893" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB22C4A3523C2F893" box="[894,964,1826,1850]" country="Ecuador" municipality="Alamor" name="Celica" pageId="22" pageNumber="250" stateProvince="Loja">Celica</location>
:
<specimenCode id="DBAF9EB5836CFFCEB2854A35248FF893" box="[983,1161,1826,1850]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 167891</specimenCode>
,
<specimenCode id="DBAF9EB5836CFFCEB5CF4A3524EAF893" box="[1181,1260,1826,1850]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">167893</specimenCode>
(males),
<specimenCode id="DBAF9EB5836CFFCEB4334A35237FF8F0" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 167892</specimenCode>
(female);
</materialsCitation>
<materialsCitation id="3B613C93836CFFCEB5524A55243FF8D0" ID-GBIF-Occurrence="4406001468" collectionCode="AMNH" collectorName="San Bartolo" country="Ecuador" location="Celica" municipality="Alamor" pageId="22" pageNumber="250" specimenCode="AMNH 167894, AMNH 167891, 167893, AMNH 167892, AMNH 172116, AMNH 172117" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Loja">
<collectorName id="26FC5318836CFFCEB5524A55248DF8F3" box="[1024,1163,1858,1882]" pageId="22" pageNumber="250">San Bartolo</collectorName>
:
<specimenCode id="DBAF9EB5836CFFCEB5F34A552552F8F0" box="[1185,1364,1857,1882]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 172116</specimenCode>
(male),
<specimenCode id="DBAF9EB5836CFFCEB2784A7623D2F8D0" box="[810,980,1889,1913]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 172117</specimenCode>
(female)
</materialsCitation>
.
<materialsCitation id="3B613C93836CFFCEB5144A7625A8F811" ID-GBIF-Occurrence="4406001350" collectionCode="AMNH, ANSP" country="Peru" location="Palambla" pageId="22" pageNumber="250" specimenCode="AMNH 175530-17533, ANSP 116437, 116438, AMNH 175534, 175535" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Piura">
<collectingCountry id="F31E765E836CFFCEB5144A762471F8D1" box="[1094,1143,1889,1912]" name="Peru" pageId="22" pageNumber="250">Peru</collectingCountry>
(
<emphasis id="B97DEADC836CFFCEB5D94A752498F8D0" box="[1163,1182,1890,1913]" italics="true" pageId="22" pageNumber="250">N</emphasis>
= 27):
<collectingRegion id="49CDF82C836CFFCEB5A04A76252FF8D0" box="[1266,1321,1889,1913]" country="Peru" name="Piura" pageId="22" pageNumber="250">Piura</collectingRegion>
(
<specimenCount id="9D0FFD47836CFFCEB4684A7625A8F8D0" box="[1338,1454,1889,1913]" pageId="22" pageNumber="250" type="male">four males</specimenCount>
,
<specimenCount id="9D0FFD47836CFFCEB2784A9723BDF831" box="[810,955,1920,1944]" pageId="22" pageNumber="250" type="female">four females</specimenCount>
):
<location id="8ED66015836CFFCEB2844A97243CF831" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836CFFCEB2844A97243CF831" box="[982,1082,1920,1944]" country="Peru" name="Palambla" pageId="22" pageNumber="250" stateProvince="Piura">Palambla</location>
:
<specimenCode id="DBAF9EB5836CFFCEB51E4A97254BF831" box="[1100,1357,1920,1944]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 17553017533</specimenCode>
(males),
<specimenCode id="DBAF9EB5836CFFCEB2784AB723C1F81E" box="[810,967,1951,1976]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="250">ANSP 116437</specimenCode>
,
<specimenCode id="DBAF9EB5836CFFCEB2874A882420F81E" box="[981,1062,1951,1975]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="22" pageNumber="250">116438</specimenCode>
,
<specimenCode id="DBAF9EB5836CFFCEB5664AB724E6F81E" box="[1076,1248,1951,1976]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">AMNH 175534</specimenCode>
,
<specimenCode id="DBAF9EB5836CFFCEB5BC4A88253BF81E" box="[1262,1341,1951,1975]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="22" pageNumber="250">175535</specimenCode>
(females)
</materialsCitation>
;
<materialsCitation id="3B613C93836DFFCFB1D34A072303F881" ID-GBIF-Occurrence="4406001371" box="[129,773,1808,1832]" collectionCode="ANSP" collectorName="Porculla" country="Peru" location="Lambayeque" pageId="23" pageNumber="251" specimenCode="ANSP 116441" specimenCount="1" specimenCount-female="1" stateProvince="Lambayeque">
<collectingRegion id="49CDF82C836DFFCFB1D34A072101F881" box="[129,263,1808,1832]" country="Peru" name="Lambayeque" pageId="23" pageNumber="251">Lambayeque</collectingRegion>
(
<specimenCount id="9D0FFD47836DFFCFB0474A072195F881" box="[277,403,1808,1832]" pageId="23" pageNumber="251" type="female">one female</specimenCount>
):
<collectorName id="26FC5318836DFFCFB0F64A0721F9F881" box="[420,511,1808,1832]" pageId="23" pageNumber="251">Porculla</collectorName>
:
<specimenCode id="DBAF9EB5836DFFCFB35B4A0722A6F881" box="[521,672,1808,1832]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="251">ANSP 116441</specimenCode>
(female)
</materialsCitation>
;
<materialsCitation id="3B613C93836DFFCFB1D34A38212BF8CF" ID-GBIF-Occurrence="4406001311" collectionCode="ANSP, LSUMZ, AMNH" collectorName="La Palma &amp; Cajabamba" country="Peru" location="Lanchal" municipality="Chira" pageId="23" pageNumber="251" specimenCode="ANSP 116442, 116443, LSUMZ 170101, 170102, LSUMZ 170103, LSUMZ 170104, AMNH 503909" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cajamarca">
<collectingRegion id="49CDF82C836DFFCFB1D34A3820E9F8EE" box="[129,239,1839,1863]" country="Peru" name="Cajamarca" pageId="23" pageNumber="251">Cajamarca</collectingRegion>
(
<specimenCount id="9D0FFD47836DFFCFB0514A38216BF8EE" box="[259,365,1839,1863]" pageId="23" pageNumber="251" type="male">six males</specimenCount>
,
<specimenCount id="9D0FFD47836DFFCFB02C4A2721F9F8EE" box="[382,511,1839,1863]" pageId="23" pageNumber="251" type="female">one female</specimenCount>
):
<collectingMunicipality id="6BD2ACB4836DFFCFB3444A382253F8EE" box="[534,597,1839,1863]" pageId="23" pageNumber="251">Chira</collectingMunicipality>
:
<specimenCode id="DBAF9EB5836DFFCFB3344A382303F8EE" box="[614,773,1839,1863]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="251">ANSP 116442</specimenCode>
,
<specimenCode id="DBAF9EB5836DFFCFB1D34A5920D6F8CF" box="[129,208,1870,1894]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="23" pageNumber="251">116443</specimenCode>
(males);
</materialsCitation>
<materialsCitation id="3B613C93836DFFCFB0614A592209F82F" ID-GBIF-Occurrence="4406001410" collectionCode="ANSP, LSUMZ, AMNH" collectorName="La Palma &amp; Cajabamba" country="Peru" location="Lanchal" municipality="Chira" pageId="23" pageNumber="251" specimenCode="ANSP 116442, 116443, LSUMZ 170101, 170102, LSUMZ 170103, LSUMZ 170104, AMNH 503909" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cajamarca">
Quebrada
<location id="8ED66015836DFFCFB0F24A5921FEF8CF" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836DFFCFB0F24A5921FEF8CF" box="[416,504,1870,1894]" country="Peru" municipality="Chira" name="Lanchal" pageId="23" pageNumber="251" stateProvince="Cajamarca">Lanchal</location>
:
<specimenCode id="DBAF9EB5836DFFCFB3534A5822B6F8CE" box="[513,688,1871,1895]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="23" pageNumber="251">LSUMZ 170101</specimenCode>
,
<specimenCode id="DBAF9EB5836DFFCFB3E84A58230FF8CE" box="[698,777,1871,1895]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="23" pageNumber="251">170102</specimenCode>
(males),
<specimenCode id="DBAF9EB5836DFFCFB1B44A79219BF82F" box="[230,413,1902,1926]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="23" pageNumber="251">LSUMZ 170103</specimenCode>
(female);
</materialsCitation>
<materialsCitation id="3B613C93836DFFCFB34D4A792300F80C" ID-GBIF-Occurrence="4406001381" collectionCode="ANSP, LSUMZ, AMNH" collectorName="La Palma &amp; Cajabamba" country="Peru" location="Lanchal" municipality="Chira" pageId="23" pageNumber="251" specimenCode="ANSP 116442, 116443, LSUMZ 170101, 170102, LSUMZ 170103, LSUMZ 170104, AMNH 503909" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cajamarca">
Quebrada
<collectorName id="26FC5318836DFFCFB3C54A792300F82F" box="[663,774,1902,1926]" pageId="23" pageNumber="251">La Palma</collectorName>
:
<specimenCode id="DBAF9EB5836DFFCFB1D34A9A2128F80C" box="[129,302,1933,1957]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="23" pageNumber="251">LSUMZ 170104</specimenCode>
(male);
<collectorName id="26FC5318836DFFCFB0D94A9A2203F80C" box="[395,517,1933,1957]" pageId="23" pageNumber="251">Cajabamba</collectorName>
:
<specimenCode id="DBAF9EB5836DFFCFB35C4A9A22B0F80C" box="[526,694,1933,1957]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">AMNH 503909</specimenCode>
(male)
</materialsCitation>
.
<materialsCitation id="3B613C93836DFFCFB26B4A072455F8CF" ID-GBIF-Occurrence="4406001456" collectionCode="AMNH" collectorName="Taulis &amp; Pacasmayo" country="Peru" location="Pacasmayo" municipality="Seques" pageId="23" pageNumber="251" specimenCode="AMNH 236066, 236068-236070, AMNH 236067, AMNH 236071, 236072, 236075, 236076, AMNH 236073, 236074" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="La Libertad">
<collectingRegion id="49CDF82C836DFFCFB26B4A0723B1F881" box="[825,951,1808,1832]" country="Peru" name="La Libertad" pageId="23" pageNumber="251">La Libertad</collectingRegion>
(
<specimenCount id="9D0FFD47836DFFCFB29A4A072441F881" box="[968,1095,1808,1832]" pageId="23" pageNumber="251" type="male">eight males</specimenCount>
,
<specimenCount id="9D0FFD47836DFFCFB5064A0724EFF881" box="[1108,1257,1808,1832]" pageId="23" pageNumber="251" type="female">three females</specimenCount>
):
<collectingMunicipality id="6BD2ACB4836DFFCFB5AF4A07254EF881" box="[1277,1352,1808,1832]" pageId="23" pageNumber="251">Seques</collectingMunicipality>
, north-east of
<location id="8ED66015836DFFCFB20C4A2723D3F8E1" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED66015836DFFCFB20C4A2723D3F8E1" box="[862,981,1840,1864]" country="Peru" municipality="Seques" name="Pacasmayo" pageId="23" pageNumber="251" stateProvince="La Libertad">Pacasmayo</location>
:
<specimenCode id="DBAF9EB5836DFFCFB2BA4A38249DF8EE" box="[1000,1179,1839,1863]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">AMNH 236066</specimenCode>
,
<specimenCode id="DBAF9EB5836DFFCFB5FD4A38255AF8EE" box="[1199,1372,1839,1863]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">236068236070</specimenCode>
(males),
<specimenCode id="DBAF9EB5836DFFCFB26B4A5823E0F8CF" box="[825,998,1870,1895]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">AMNH 236067</specimenCode>
(female);
</materialsCitation>
<materialsCitation id="3B613C93836DFFCFB50D4A592449F80C" ID-GBIF-Occurrence="4406001470" collectionCode="AMNH" collectorName="Taulis &amp; Pacasmayo" country="Peru" location="Pacasmayo" municipality="Seques" pageId="23" pageNumber="251" specimenCode="AMNH 236066, 236068-236070, AMNH 236067, AMNH 236071, 236072, 236075, 236076, AMNH 236073, 236074" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="La Libertad">
<collectorName id="26FC5318836DFFCFB50D4A5924A6F8CF" box="[1119,1184,1870,1894]" pageId="23" pageNumber="251">Taulis</collectorName>
, north-east of
<collectorName id="26FC5318836DFFCFB41A4A5825B9F8CE" box="[1352,1471,1871,1895]" pageId="23" pageNumber="251">Pacasmayo</collectorName>
:
<specimenCode id="DBAF9EB5836DFFCFB26B4A7923E1F82F" box="[825,999,1902,1926]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">AMNH 236071</specimenCode>
,
<specimenCode id="DBAF9EB5836DFFCFB2A44A792441F82F" box="[1014,1095,1902,1926]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">236072</specimenCode>
,
<specimenCode id="DBAF9EB5836DFFCFB5044A7924A1F82F" box="[1110,1191,1902,1926]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">236075</specimenCode>
,
<specimenCode id="DBAF9EB5836DFFCFB5E44A792503F82F" box="[1206,1285,1902,1926]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">236076</specimenCode>
(males),
<specimenCode id="DBAF9EB5836DFFCFB4224A79238CF80C" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">AMNH 236073</specimenCode>
,
<specimenCode id="DBAF9EB5836DFFCFB2C64A9A23E5F80C" box="[916,995,1933,1957]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="251">236074</specimenCode>
(females)
</materialsCitation>
.
</paragraph>
<caption id="DF766646836DFFCFB1D34BBB21CCF976" ID-DOI="http://doi.org/10.5281/zenodo.8326332" ID-Zenodo-Dep="8326332" httpUri="https://zenodo.org/record/8326332/files/figure.png" pageId="23" pageNumber="251" startId="23.[129,194,1708,1732]" targetBox="[257,1346,156,1680]" targetPageId="23" targetType="figure">
<paragraph id="8BB636CE836DFFCFB1D34BBB21CCF976" blockId="23.[129,1469,1708,1760]" pageId="23" pageNumber="251">
<emphasis id="B97DEADC836DFFCFB1D34BBB20E3F96D" bold="true" box="[129,229,1708,1732]" pageId="23" pageNumber="251">Figure 15.</emphasis>
Ventral and dorsal views of the adult male plumages in
<emphasis id="B97DEADC836DFFCFB3BE4BBB2374F96D" box="[748,882,1708,1732]" italics="true" pageId="23" pageNumber="251">
C. b.
<taxonomicName id="4C094D4D836DFFCFB2484BBB2374F96D" box="[794,882,1708,1732]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="23" pageNumber="251" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
,
<emphasis id="B97DEADC836DFFCFB2294BBB23F5F96D" box="[891,1011,1708,1732]" italics="true" pageId="23" pageNumber="251">C. b. caniceps</emphasis>
,
<emphasis id="B97DEADC836DFFCFB2AE4BBB2468F96D" box="[1020,1134,1708,1732]" italics="true" pageId="23" pageNumber="251">C. o. tenebris</emphasis>
ssp. nov.,
<emphasis id="B97DEADC836DFFCFB5994BBB253BF96D" box="[1227,1341,1708,1732]" italics="true" pageId="23" pageNumber="251">C. b. nebulus</emphasis>
ssp. nov., and
<taxonomicName id="4C094D4D836DFFCFB1D34BDF20E3F949" box="[129,229,1736,1760]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="23" pageNumber="251" phylum="Chordata" rank="species" species="mentalis">
<emphasis id="B97DEADC836DFFCFB1D34BDF20E3F949" box="[129,229,1736,1760]" italics="true" pageId="23" pageNumber="251">C. mentalis</emphasis>
</taxonomicName>
. Photograph by M.R.H.
</paragraph>
</caption>
<caption id="DF7666468362FFC0B1234BBB21E5F976" ID-DOI="http://doi.org/10.5281/zenodo.8326334" ID-Zenodo-Dep="8326334" httpUri="https://zenodo.org/record/8326334/files/figure.png" pageId="24" pageNumber="252" startId="24.[113,178,1708,1732]" targetBox="[285,1290,156,1680]" targetPageId="24" targetType="figure">
<paragraph id="8BB636CE8362FFC0B1234BBB21E5F976" blockId="24.[113,1431,1708,1760]" pageId="24" pageNumber="252">
<emphasis id="B97DEADC8362FFC0B1234BBB20D0F96D" bold="true" box="[113,214,1708,1732]" pageId="24" pageNumber="252">Figure 16.</emphasis>
Ventral and dorsal views of the adult female plumages in
<emphasis id="B97DEADC8362FFC0B3BC4BBB2372F96D" box="[750,884,1708,1732]" italics="true" pageId="24" pageNumber="252">
C. b.
<taxonomicName id="4C094D4D8362FFC0B24E4BBB2372F96D" box="[796,884,1708,1732]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="24" pageNumber="252" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
,
<emphasis id="B97DEADC8362FFC0B22F4BBB23F3F96D" box="[893,1013,1708,1732]" italics="true" pageId="24" pageNumber="252">C. b. caniceps</emphasis>
,
<emphasis id="B97DEADC8362FFC0B2AC4BBB2476F96D" box="[1022,1136,1708,1732]" italics="true" pageId="24" pageNumber="252">C. o. tenebris</emphasis>
ssp. nov.,
<emphasis id="B97DEADC8362FFC0B59F4BBB2539F96D" box="[1229,1343,1708,1732]" italics="true" pageId="24" pageNumber="252">C. b. nebulus</emphasis>
ssp. nov., and
<taxonomicName id="4C094D4D8362FFC0B1C84BDF20F8F949" box="[154,254,1736,1760]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="24" pageNumber="252" phylum="Chordata" rank="species" species="mentalis">
<emphasis id="B97DEADC8362FFC0B1C84BDF20F8F949" box="[154,254,1736,1760]" italics="true" pageId="24" pageNumber="252">C. mentalis</emphasis>
</taxonomicName>
. Photograph by M.R.H.
</paragraph>
</caption>
<paragraph id="8BB636CE8362FFC0B1DF4A07253EF86D" blockId="24.[113,763,1808,1989]" lastBlockId="24.[810,1461,1808,1989]" pageId="24" pageNumber="252">
<emphasis id="B97DEADC8362FFC0B1DF4A072146F881" box="[141,320,1808,1832]" italics="true" pageId="24" pageNumber="252">
<taxonomicName id="4C094D4D8362FFC0B1DF4A0720E7F881" authorityName="Bonaparte" authorityYear="1850" box="[141,225,1808,1832]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="24" pageNumber="252" phylum="Chordata" rank="genus">Catharus</taxonomicName>
b. nebulus
</emphasis>
<materialsCitation id="3B613C938362FFC0B0174A0720B6F8CF" ID-GBIF-Occurrence="4406001342" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
ssp. nov. (
<emphasis id="B97DEADC8362FFC0B0F64A0621B1F881" box="[420,439,1809,1832]" italics="true" pageId="24" pageNumber="252">N</emphasis>
= 19):
<collectingCountry id="F31E765E8362FFC0B0AD4A07222AF88E" box="[511,556,1808,1831]" name="Peru" pageId="24" pageNumber="252">Peru</collectingCountry>
(
<emphasis id="B97DEADC8362FFC0B36E4A062249F881" box="[572,591,1809,1832]" italics="true" pageId="24" pageNumber="252">N</emphasis>
= 19):
<collectingCounty id="62D74E428362FFC0B3C54A0722FDF881" box="[663,763,1808,1832]" pageId="24" pageNumber="252">Amazonas</collectingCounty>
(
<specimenCount id="9D0FFD478362FFC0B12B4A3820DAF8EE" box="[121,220,1839,1863]" pageId="24" pageNumber="252" type="male">two males</specimenCount>
,
<specimenCount id="9D0FFD478362FFC0B1B64A382158F8EE" box="[228,350,1839,1863]" pageId="24" pageNumber="252" type="female">four females</specimenCount>
):
<locationDeviation id="9A042FEC8362FFC0B03E4A38223EF8EE" box="[364,568,1839,1863]" location="Ingenio" pageId="24" pageNumber="252">
<quantity id="4CF19B2B8362FFC0B03E4A3821AFF8EE" box="[364,425,1839,1863]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="24" pageNumber="252" unit="km" value="33.0">33 km</quantity>
NE of Ingenio
</locationDeviation>
:
<specimenCode id="DBAF9EB58362FFC0B36D4A3822D2F8EE" box="[575,724,1839,1863]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 82159</specimenCode>
(female);
</materialsCitation>
<materialsCitation id="3B613C938362FFC0B1E14A5821D9F82F" ID-GBIF-Occurrence="4406001405" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
Cerro Montealegre:
<specimenCode id="DBAF9EB58362FFC0B0234A58220FF8CE" box="[369,521,1871,1895]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="252" type="Museum">FMNH 474413</specimenCode>
(male);
<collectorName id="26FC53188362FFC0B30A4A5822A0F8CF" box="[600,678,1870,1894]" pageId="24" pageNumber="252">
<collectingMunicipality id="6BD2ACB48362FFC0B30A4A582277F8CF" box="[600,625,1871,1894]" pageId="24" pageNumber="252">La</collectingMunicipality>
Lejia
</collectorName>
,north of
<location id="8ED660158362FFC0B1234A7920F2F82F" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED660158362FFC0B1234A7920F2F82F" box="[113,244,1902,1926]" country="Peru" county="Amazonas" municipality="La" name="Chachapoyas" pageId="24" pageNumber="252" stateProvince="Huanuco">Chachapoyas</location>
:
<specimenCode id="DBAF9EB58362FFC0B1A84A792193F82F" box="[250,405,1902,1926]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="252">AMNH 235051</specimenCode>
(male);
</materialsCitation>
<materialsCitation id="3B613C938362FFC0B0B14A7922FDF80C" ID-GBIF-Occurrence="4406001491" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
Leymebamba:
<specimenCode id="DBAF9EB58362FFC0B3224A7922FDF82F" box="[624,763,1902,1926]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="252">ANSP 109278</specimenCode>
(female);
<locationDeviation id="9A042FEC8362FFC0B1804A9A21A4F80C" box="[210,418,1933,1957]" location="La Peca" pageId="24" pageNumber="252">
<quantity id="4CF19B2B8362FFC0B1804A9A2115F80C" box="[210,275,1933,1957]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="252" unit="km" value="20.0">20 km</quantity>
E of La Peca
</locationDeviation>
:
<specimenCode id="DBAF9EB58362FFC0B0FC4A9A224EF80C" box="[430,584,1933,1957]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 88630</specimenCode>
,
<specimenCode id="DBAF9EB58362FFC0B3064A9A2297F80C" box="[596,657,1933,1957]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">88631</specimenCode>
(females);
</materialsCitation>
<materialsCitation id="3B613C938362FFC0B1234ABA22FDF86D" ID-GBIF-Occurrence="4406001387" box="[113,763,1964,1989]" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
<collectingRegion id="49CDF82C8362FFC0B1234ABA20CCF86C" box="[113,202,1965,1989]" country="Peru" name="Huanuco" pageId="24" pageNumber="252">Huánuco</collectingRegion>
(
<specimenCount id="9D0FFD478362FFC0B18E4ABB2153F86D" box="[220,341,1964,1988]" pageId="24" pageNumber="252" type="male">three males</specimenCount>
): Cayumba Alto:
<specimenCode id="DBAF9EB58362FFC0B3594ABA22AFF86C" box="[523,681,1965,1989]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="252" type="Museum">FMNH 299304</specimenCode>
(male);
</materialsCitation>
<materialsCitation id="3B613C938362FFC0B2784A07247DF8EE" ID-GBIF-Occurrence="4406001365" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
CordilleraCarpish:
<specimenCode id="DBAF9EB58362FFC0B2B04A072470F881" box="[994,1142,1808,1832]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 78972</specimenCode>
,
<specimenCode id="DBAF9EB58362FFC0B52E4A0724BFF881" box="[1148,1209,1808,1832]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">80744</specimenCode>
(males);
<collectingRegion id="49CDF82C8362FFC0B45C4A072547F881" box="[1294,1345,1808,1832]" country="Peru" name="Junin" pageId="24" pageNumber="252">Junín</collectingRegion>
(onemale): Rumicruz:
<specimenCode id="DBAF9EB58362FFC0B2C64A382436F8EE" box="[916,1072,1839,1863]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="252">AMNH 174319</specimenCode>
(male);
</materialsCitation>
<materialsCitation id="3B613C938362FFC0B52D4A272433F82F" ID-GBIF-Occurrence="4406001435" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
<collectingRegion id="49CDF82C8362FFC0B52D4A2724B0F8EE" box="[1151,1206,1840,1863]" country="Peru" name="Pasco" pageId="24" pageNumber="252">Pasco</collectingRegion>
(
<specimenCount id="9D0FFD478362FFC0B5934A38252EF8EE" box="[1217,1320,1839,1863]" pageId="24" pageNumber="252" type="male">four males</specimenCount>
,
<specimenCount id="9D0FFD478362FFC0B4624A3825ACF8EE" box="[1328,1450,1839,1863]" pageId="24" pageNumber="252" type="female">four females</specimenCount>
): Playa Pampa:
<specimenCode id="DBAF9EB58362FFC0B2E64A58245FF8CF" box="[948,1113,1870,1895]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 129006</specimenCode>
,
<specimenCode id="DBAF9EB58362FFC0B5364A5824A8F8CE" box="[1124,1198,1871,1895]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">129007</specimenCode>
,
<specimenCode id="DBAF9EB58362FFC0B5E84A582505F8CE" box="[1210,1283,1871,1895]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">129011</specimenCode>
(males),
<specimenCode id="DBAF9EB58362FFC0B4324A582372F82F" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 129002</specimenCode>
,
<specimenCode id="DBAF9EB58362FFC0B2D04A7923CDF82F" box="[898,971,1902,1926]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">129005</specimenCode>
(females);
</materialsCitation>
<materialsCitation id="3B613C938362FFC0B56D4A792487F80C" ID-GBIF-Occurrence="4406001462" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
Santa Cruz:
<specimenCode id="DBAF9EB58362FFC0B5EC4A792562F82F" box="[1214,1380,1902,1926]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 106496</specimenCode>
(male),
<specimenCode id="DBAF9EB58362FFC0B2784A9A23CDF80C" box="[810,971,1933,1957]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 106495</specimenCode>
,
<specimenCode id="DBAF9EB58362FFC0B2814A9A241AF80C" box="[979,1052,1933,1957]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">106497</specimenCode>
(females);
</materialsCitation>
<materialsCitation id="3B613C938362FFC0B5D64A9A2532F86D" ID-GBIF-Occurrence="4406001409" collectionCode="LSUMZ, FMNH, AMNH, ANSP" collectorName="La Lejia" country="Peru" county="Amazonas" location="Chachapoyas" municipality="La" pageId="24" pageNumber="252" specimenCode="LSUMZ 82159, FMNH 474413, AMNH 235051, ANSP 109278, LSUMZ 88630, 88631, FMNH 299304, LSUMZ 78972, 80744, AMNH 174319, LSUMZ 129006, 129007, 129011, LSUMZ 129002, 129005, LSUMZ 106496, LSUMZ 106495, 106497, LSUMZ 174203" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Huanuco">
<collectingRegion id="49CDF82C8362FFC0B5D64A9A24E9F80C" box="[1156,1263,1933,1957]" country="Peru" name="San Martin" pageId="24" pageNumber="252">San Martín</collectingRegion>
(
<specimenCount id="9D0FFD478362FFC0B5AB4A9A256FF80C" box="[1273,1385,1933,1957]" pageId="24" pageNumber="252" type="female">one female</specimenCount>
):
<locationDeviation id="9A042FEC8362FFC0B4244A9A243EF86D" location="Florida Pomacochas" pageId="24" pageNumber="252">
<quantity id="4CF19B2B8362FFC0B4244A9A25B2F80C" box="[1398,1460,1933,1957]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="252" unit="km" value="22.0">22 km</quantity>
ENE of Florida Pomacochas
</locationDeviation>
:
<specimenCode id="DBAF9EB58362FFC0B56C4ABA24DBF86C" box="[1086,1245,1965,1989]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="24" pageNumber="252">LSUMZ 174203</specimenCode>
(female)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636CE8363FFC1B0644D862252FF03" blockId="25.[129,778,144,269]" box="[310,596,144,170]" pageId="25" pageNumber="253">
<emphasis id="B97DEADC8363FFC1B0644D862252FF03" box="[310,596,144,170]" italics="true" pageId="25" pageNumber="253">Immature specimens examined</emphasis>
</paragraph>
<paragraph id="8BB636CE8363FFC1B1D34DA12428FF4E" blockId="25.[129,778,144,269]" lastBlockId="25.[825,1475,144,481]" pageId="25" pageNumber="253">
<emphasis id="B97DEADC8363FFC1B1D34DA1215BFF67" box="[129,349,182,206]" italics="true" pageId="25" pageNumber="253">
<taxonomicName id="4C094D4D8363FFC1B1D34DA120DAFF67" authorityName="Bonaparte" authorityYear="1850" box="[129,220,182,206]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="genus">Catharus</taxonomicName>
b.
<taxonomicName id="4C094D4D8363FFC1B1AC4DA1215BFF67" box="[254,349,182,206]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
(
<emphasis id="B97DEADC8363FFC1B0224DA02185FF67" box="[368,387,183,206]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 5):
<materialsCitation id="3B613C938363FFC1B09A4DA12213FEA4" ID-GBIF-Occurrence="4406001401" collectionCode="ANSP, DMNH, AMNH" country="Ecuador" location="Rio La Plata" municipality="Maldonado-Tulcan Rd." pageId="25" pageNumber="253" specimenCode="ANSP 181215, DMNH 62882, AMNH 180632" specimenCount="1" specimenCount-female="1" stateProvince="Carchi">
<collectingCountry id="F31E765E8363FFC1B09A4DA12218FF67" box="[456,542,182,206]" name="Ecuador" pageId="25" pageNumber="253">Ecuador</collectingCountry>
(
<emphasis id="B97DEADC8363FFC1B3604DA02243FF67" box="[562,581,183,206]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 5):
<collectingRegion id="49CDF82C8363FFC1B3D84DA122C9FF67" box="[650,719,182,206]" country="Ecuador" name="Carchi" pageId="25" pageNumber="253">Carchi</collectingRegion>
(
<specimenCount id="9D0FFD478363FFC1B38D4DA120D3FF47" pageId="25" pageNumber="253" type="female">two females</specimenCount>
): west slope near
<collectingMunicipality id="6BD2ACB48363FFC1B0D84DC1227CFF47" box="[394,634,214,238]" pageId="25" pageNumber="253">Maldonado-Tulcán Rd.</collectingMunicipality>
, along
<location id="8ED660158363FFC1B3914DC120BEFEA4" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED660158363FFC1B3914DC120BEFEA4" country="Ecuador" municipality="Maldonado-Tulcan Rd." name="Rio La Plata" pageId="25" pageNumber="253" stateProvince="Carchi">Río La Plata</location>
:
<specimenCode id="DBAF9EB58363FFC1B1924DE22152FEA4" box="[192,340,245,269]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="253">ANSP 181215</specimenCode>
181216 (females);
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B3494DE22421FF01" ID-GBIF-Occurrence="4406001305" box="[539,1063,144,269]" collectionCode="ANSP, DMNH, AMNH" country="Ecuador" location="Rio La Plata" municipality="Maldonado-Tulcan Rd." pageId="25" pageNumber="253" specimenCode="ANSP 181215, DMNH 62882, AMNH 180632" specimenCount="1" specimenCount-female="1" stateProvince="Carchi">
Pinchincha: Chiriboga:
<specimenCode id="DBAF9EB58363FFC1B26B4D8723D2FF01" box="[825,980,144,168]" collectionCode="DMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" lsid="urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="25" pageNumber="253" type="Museum">DMNH 62882</specimenCode>
(male);
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B57F4D8725B8FF01" ID-GBIF-Occurrence="4406001317" box="[1069,1470,144,168]" collectionCode="ANSP, DMNH, AMNH" country="Ecuador" location="Rio La Plata" municipality="Maldonado-Tulcan Rd." pageId="25" pageNumber="253" specimenCode="ANSP 181215, DMNH 62882, AMNH 180632" specimenCount="1" specimenCount-female="1" stateProvince="Carchi">
Mindo Arriba:
<specimenCode id="DBAF9EB58363FFC1B5984D872569FF01" box="[1226,1391,144,168]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="253">AMNH 180632</specimenCode>
(male)
</materialsCitation>
;
<materialsCitation id="3B613C938363FFC1B26B4DB82564FF6E" ID-GBIF-Occurrence="4406001330" box="[825,1378,175,200]" collectionCode="AMNH" country="Ecuador" location="Salvias" pageId="25" pageNumber="253" specimenCode="AMNH 167889, AMNH 130153" specimenCount="1" specimenCount-male="1" stateProvince="El Oro">
<collectingRegion id="49CDF82C8363FFC1B26B4DB82386FF6E" box="[825,896,175,199]" country="Ecuador" name="El Oro" pageId="25" pageNumber="253">El Oro</collectingRegion>
(
<specimenCount id="9D0FFD478363FFC1B2DD4DB82404FF6E" box="[911,1026,175,199]" pageId="25" pageNumber="253" type="male">two males</specimenCount>
):
<location id="8ED660158363FFC1B5464DB82458FF6E" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED660158363FFC1B5464DB82458FF6E" box="[1044,1118,175,199]" country="Ecuador" name="Salvias" pageId="25" pageNumber="253" stateProvince="El Oro">Salvias</location>
:
<specimenCode id="DBAF9EB58363FFC1B53A4DA72508FF6E" box="[1128,1294,175,200]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="253">AMNH 167889</specimenCode>
(male);
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B43B4DA7242CFF4E" ID-GBIF-Occurrence="4406001510" collectionCode="AMNH" country="Ecuador" location="Salvias" pageId="25" pageNumber="253" specimenCode="AMNH 167889, AMNH 130153" specimenCount="1" specimenCount-male="1" stateProvince="El Oro">
Zaruma:
<specimenCode id="DBAF9EB58363FFC1B26B4DD823DBFF4F" box="[825,989,206,231]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="253">AMNH 130153</specimenCode>
(male)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636CE8363FFC1B2074DF92441FE2A" blockId="25.[825,1475,144,481]" pageId="25" pageNumber="253">
<emphasis id="B97DEADC8363FFC1B2074DF9241DFEAF" box="[853,1051,238,262]" italics="true" pageId="25" pageNumber="253">
<taxonomicName id="4C094D4D8363FFC1B2074DF923B6FEAF" authorityName="Bonaparte" authorityYear="1850" box="[853,944,238,262]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="genus">Catharus</taxonomicName>
b. caniceps
</emphasis>
(
<emphasis id="B97DEADC8363FFC1B57F4DF82446FEAF" box="[1069,1088,239,262]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 3):
<materialsCitation id="3B613C938363FFC1B5D04DF92392FEEC" ID-GBIF-Occurrence="4406001432" collectionCode="ANSP, CM" country="Peru" location="Piura" pageId="25" pageNumber="253" specimenCode="ANSP 116439, CM 119947" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Piura">
<collectingCountry id="F31E765E8363FFC1B5D04DF924B4FEAC" box="[1154,1202,238,261]" name="Peru" pageId="25" pageNumber="253">Peru</collectingCountry>
(
<emphasis id="B97DEADC8363FFC1B5914DF824D0FEAF" box="[1219,1238,239,262]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 3):
<collectingRegion id="49CDF82C8363FFC1B44B4DF92549FEAF" box="[1305,1359,238,262]" country="Peru" name="Piura" pageId="25" pageNumber="253">Piura</collectingRegion>
(
<specimenCount id="9D0FFD478363FFC1B40F4DF925B9FEAF" box="[1373,1471,238,262]" pageId="25" pageNumber="253" type="male">one male</specimenCount>
,
<specimenCount id="9D0FFD478363FFC1B26B4C1923B1FE8C" box="[825,951,269,293]" pageId="25" pageNumber="253" type="female">one female</specimenCount>
): Palambla:
<specimenCode id="DBAF9EB58363FFC1B56D4C1A24DCFE8C" box="[1087,1242,269,293]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="253">ANSP 116439</specimenCode>
(male),
<specimenCode id="DBAF9EB58363FFC1B46C4C1A25C4FE8C" box="[1342,1474,269,293]" collectionCode="CM" country="China" name="Chongqing Museum" pageId="25" pageNumber="253">CM 119947</specimenCode>
(female)
</materialsCitation>
;
<materialsCitation id="3B613C938363FFC1B2F14C3A2348FECD" ID-GBIF-Occurrence="4406001346" country="Peru" location="Cajamarca" pageId="25" pageNumber="253" specimenCount="1" specimenCount-male="1" stateProvince="Cajamarca">
<collectingRegion id="49CDF82C8363FFC1B2F14C3A2409FEEC" box="[931,1039,301,325]" country="Peru" name="Cajamarca" pageId="25" pageNumber="253">Cajamarca</collectingRegion>
(
<specimenCount id="9D0FFD478363FFC1B5704C3A2488FEEC" box="[1058,1166,301,325]" pageId="25" pageNumber="253" type="male">one male</specimenCount>
): Chugur,
<quantity id="4CF19B2B8363FFC1B4504C3B254EFEEC" box="[1282,1352,300,325]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="25" pageNumber="253" unit="km" value="65.0">65 km</quantity>
north-west of
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B20E4C5B2445FE2A" ID-GBIF-Occurrence="4406001440" collectionCode="AMNH, LSUMZ" country="Peru" location="Cajamarca" pageId="25" pageNumber="253" specimenCode="AMNH 236065, LSUMZ 170100" specimenCount="1" stateProvince="Cajamarca">
<collectingRegion id="49CDF82C8363FFC1B20E4C5B23CBFECD" box="[860,973,332,356]" country="Peru" name="Cajamarca" pageId="25" pageNumber="253">Cajamarca</collectingRegion>
:
<specimenCode id="DBAF9EB58363FFC1B28C4C5B248DFECD" box="[990,1163,332,356]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="253">AMNH 236065</specimenCode>
(male); Quebrada Lanchal:
<specimenCode id="DBAF9EB58363FFC1B26B4C7C23E5FE2A" box="[825,995,363,387]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="25" pageNumber="253">LSUMZ 170100</specimenCode>
(female)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636CE8363FFC1B2074C9C23DEFE48" blockId="25.[825,1475,144,481]" pageId="25" pageNumber="253">
<taxonomicName id="4C094D4D8363FFC1B2074C9C2427FE0A" authorityName="Bonaparte" authorityYear="1850" box="[853,1057,395,419]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC8363FFC1B2074C9C2427FE0A" box="[853,1057,395,419]" italics="true" pageId="25" pageNumber="253">Catharus b. nebulus</emphasis>
</taxonomicName>
<materialsCitation id="3B613C938363FFC1B57C4C9C23D2FE48" ID-GBIF-Occurrence="4406001407" collectionCode="LSUMZ" country="Peru" location="Florida Pomacochas" pageId="25" pageNumber="253" specimenCode="LSUMZ 174204" specimenCount="1" specimenCount-male="1" stateProvince="San Martin">
ssp. nov. (
<emphasis id="B97DEADC8363FFC1B5CD4C9C24B4FE0B" box="[1183,1202,395,418]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 1):
<collectingCountry id="F31E765E8363FFC1B5A84C9C252CFE0B" box="[1274,1322,395,418]" name="Peru" pageId="25" pageNumber="253">Peru</collectingCountry>
(
<emphasis id="B97DEADC8363FFC1B4134C9C2552FE0B" box="[1345,1364,395,418]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 1):
<collectingRegion id="49CDF82C8363FFC1B4CE4C9C2387FE6B" country="Peru" name="San Martin" pageId="25" pageNumber="253">San Martín</collectingRegion>
(
<specimenCount id="9D0FFD478363FFC1B2DE4CBD23F7FE6B" box="[908,1009,426,450]" pageId="25" pageNumber="253" type="male">one male</specimenCount>
):
<quantity id="4CF19B2B8363FFC1B5524CBD2439FE6B" box="[1024,1087,426,450]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="25" pageNumber="253" unit="km" value="22.0">22 km</quantity>
ENE of
<location id="8ED660158363FFC1B5C34CBD2562FE6B" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED660158363FFC1B5C34CBD2562FE6B" box="[1169,1380,426,450]" country="Peru" name="Florida Pomacochas" pageId="25" pageNumber="253" stateProvince="San Martin">Florida Pomacochas</location>
:
<specimenCode id="DBAF9EB58363FFC1B4394CBD2381FE48" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="25" pageNumber="253">LSUMZ 174204</specimenCode>
(male)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB636CE8363FFC1B5524F1A24FDFD8E" blockId="25.[1024,1275,525,551]" box="[1024,1275,525,551]" pageId="25" pageNumber="253">
<emphasis id="B97DEADC8363FFC1B5524F1A24FDFD8E" box="[1024,1275,525,551]" italics="true" pageId="25" pageNumber="253">Audio recordings examined</emphasis>
</paragraph>
<paragraph id="8BB636CE8363FFC1B26B4F23255AFCEF" blockId="25.[825,1475,564,964]" pageId="25" pageNumber="253">
<taxonomicName id="4C094D4D8363FFC1B26B4F232426FDE5" authorityName="Bonaparte" authorityYear="1850" box="[825,1056,564,588]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC8363FFC1B26B4F232426FDE5" box="[825,1056,564,588]" italics="true" pageId="25" pageNumber="253">Catharus b. berlepschi</emphasis>
</taxonomicName>
(
<emphasis id="B97DEADC8363FFC1B56A4F23244DFDE2" box="[1080,1099,564,587]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 20):
<materialsCitation id="3B613C938363FFC1B5F04F23255DFDC2" ID-GBIF-Occurrence="4406001467" pageId="25" pageNumber="253">
<collectingCountry id="F31E765E8363FFC1B5F04F2324FEFDE5" box="[1186,1272,564,588]" name="Ecuador" pageId="25" pageNumber="253">Ecuador</collectingCountry>
(
<emphasis id="B97DEADC8363FFC1B4434F232522FDE2" box="[1297,1316,564,587]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 20):
<collectingRegion id="49CDF82C8363FFC1B4294F2325B9FDE5" box="[1403,1471,564,588]" country="Ecuador" name="Azuay" pageId="25" pageNumber="253">Azuay</collectingRegion>
: Cuenca: Molleturo, Corona de Oro Rd.: XC 602205;
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B4334F442550FD23" ID-GBIF-Occurrence="4406001312" pageId="25" pageNumber="253">Yunguilla Reserve: ML 135902, 135926, 76216901, XC 19999;</materialsCitation>
<materialsCitation id="3B613C938363FFC1B4094F6523F3FD03" ID-GBIF-Occurrence="4406001392" pageId="25" pageNumber="253">
<collectingRegion id="49CDF82C8363FFC1B4094F652588FD23" box="[1371,1422,626,650]" country="Ecuador" name="Loja" pageId="25" pageNumber="253">Loja</collectingRegion>
: near Celica: XC 67875;
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B2A84F8525C4FD03" ID-GBIF-Occurrence="4406001382" box="[1018,1474,658,682]" pageId="25" pageNumber="253">Reserva Utuana: ML 141950781, XC 67874;</materialsCitation>
<materialsCitation id="3B613C938363FFC1B26B4FA625C4FD60" ID-GBIF-Occurrence="4406001325" box="[825,1474,689,713]" pageId="25" pageNumber="253">
<collectingRegion id="49CDF82C8363FFC1B26B4FA623A5FD60" box="[825,931,689,713]" country="Ecuador" name="Pichincha" pageId="25" pageNumber="253">Pichincha</collectingRegion>
: Bellavista, Tandayapa: XC 128440, 13227, 4214;
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B26B4FC72466FD41" ID-GBIF-Occurrence="4406001471" box="[825,1120,720,744]" pageId="25" pageNumber="253">Mindo: XC 545139, 546525;</materialsCitation>
<materialsCitation id="3B613C938363FFC1B5374FC725C4FD41" ID-GBIF-Occurrence="4406001302" box="[1125,1474,720,744]" pageId="25" pageNumber="253">Mindo Cloud Forest: ML 139115;</materialsCitation>
<materialsCitation id="3B613C938363FFC1B26B4FE7256CFCAE" ID-GBIF-Occurrence="4406001391" box="[825,1386,751,776]" pageId="25" pageNumber="253">Old Mindo-Nono Rd., before Tandayapa: ML 63456;</materialsCitation>
<materialsCitation id="3B613C938363FFC1B4234FE724E0FC8E" ID-GBIF-Occurrence="4406001306" pageId="25" pageNumber="253">Recinto 23 de Junio: ML 216745971, XC 186913;</materialsCitation>
<materialsCitation id="3B613C938363FFC1B5BE4E1823AFFCEF" ID-GBIF-Occurrence="4406001314" pageId="25" pageNumber="253">Reserva Paz de Aves: XC 30933;</materialsCitation>
<materialsCitation id="3B613C938363FFC1B2FC4E39255AFCEF" ID-GBIF-Occurrence="4406001444" box="[942,1372,814,838]" pageId="25" pageNumber="253">Tandayapa Bird Lodge: XC 32386, 54902.</materialsCitation>
</paragraph>
<paragraph id="8BB636CE8363FFC1B2074E59244CFC6D" blockId="25.[825,1475,564,964]" pageId="25" pageNumber="253">
<taxonomicName id="4C094D4D8363FFC1B2074E59242BFCCC" authorityName="Bonaparte" authorityYear="1850" box="[853,1069,846,870]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC8363FFC1B2074E59242BFCCC" box="[853,1069,846,870]" italics="true" pageId="25" pageNumber="253">Catharus b. caniceps</emphasis>
</taxonomicName>
(
<emphasis id="B97DEADC8363FFC1B51A4E59245DFCCC" box="[1096,1115,846,869]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 4):
<materialsCitation id="3B613C938363FFC1B5F44E592559FC2C" ID-GBIF-Occurrence="4406001353" collectionCode="ML" country="Peru" county="Chotas" location="Sallique" municipality="Bosque Protector Pagaibamba" pageId="25" pageNumber="253" specimenCode="ML 515357, 515369" specimenCount="1" stateProvince="Cajamarca">
<collectingCountry id="F31E765E8363FFC1B5F44E5924D0FCCC" box="[1190,1238,846,869]" name="Peru" pageId="25" pageNumber="253">Peru</collectingCountry>
(
<emphasis id="B97DEADC8363FFC1B5A24E592505FCCC" box="[1264,1283,846,869]" italics="true" pageId="25" pageNumber="253">N</emphasis>
= 4):
<collectingRegion id="49CDF82C8363FFC1B41C4E5925B9FCCF" box="[1358,1471,846,870]" country="Peru" name="Cajamarca" pageId="25" pageNumber="253">Cajamarca</collectingRegion>
:
<collectingCounty id="62D74E428363FFC1B26B4E7A2382FC2C" box="[825,900,877,901]" pageId="25" pageNumber="253">Chotas</collectingCounty>
,
<collectingMunicipality id="6BD2ACB48363FFC1B2C04E7A24D7FC2C" box="[914,1233,877,901]" pageId="25" pageNumber="253">Bosque Protector Pagaibamba</collectingMunicipality>
: XC 108771;
</materialsCitation>
<materialsCitation id="3B613C938363FFC1B43B4E7A2500FC0D" ID-GBIF-Occurrence="4406001345" collectionCode="ML" country="Peru" county="Chotas" location="Sallique" municipality="Bosque Protector Pagaibamba" pageId="25" pageNumber="253" specimenCode="ML 515357, 515369" specimenCount="1" stateProvince="Cajamarca">
<location id="8ED660158363FFC1B43B4E7A25B9FC2C" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED660158363FFC1B43B4E7A25B9FC2C" box="[1385,1471,877,901]" country="Peru" county="Chotas" municipality="Bosque Protector Pagaibamba" name="Sallique" pageId="25" pageNumber="253" stateProvince="Cajamarca">Sallique</location>
: Quebrada
<location id="8ED660158363FFC1B2E24E9B2400FC0D" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED660158363FFC1B2E24E9B2400FC0D" box="[944,1030,908,932]" country="Peru" county="Chotas" municipality="Bosque Protector Pagaibamba" name="Lanchal" pageId="25" pageNumber="253" stateProvince="Cajamarca">Lanchal</location>
:
<specimenCode id="DBAF9EB58363FFC1B5484E9A24A6FC0D" box="[1050,1184,908,932]" collectionCode="ML" country="France" name="Musee de Lectoure" pageId="25" pageNumber="253">ML 515357</specimenCode>
,
<specimenCode id="DBAF9EB58363FFC1B5E64E9B2500FC0D" box="[1204,1286,908,932]" collectionCode="ML" country="France" name="Musee de Lectoure" pageId="25" pageNumber="253">515369</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B613C938363FFC1B4484E9B2440FC6D" ID-GBIF-Occurrence="4406001327" collectorName="de Cuyas" country="Peru" location="Bosque de Cuyas" pageId="25" pageNumber="253" specimenCount="1" stateProvince="Piura">
<collectingRegion id="49CDF82C8363FFC1B4484E9B2552FC0D" box="[1306,1364,908,932]" country="Peru" name="Piura" pageId="25" pageNumber="253">Piura</collectingRegion>
: Ayabaca,
<location id="8ED660158363FFC1B26B4EBB23EBFC6D" LSID="urn:lsid:plazi:treatment:03A087D8836CFFC3B1AD4EDD2140FA24:8ED660158363FFC1B26B4EBB23EBFC6D" box="[825,1005,940,964]" country="Peru" name="Bosque de Cuyas" pageId="25" pageNumber="253" stateProvince="Piura">
Bosque
<collectorName id="26FC53188363FFC1B2D94EBB23EBFC6D" box="[907,1005,940,964]" pageId="25" pageNumber="253">de Cuyas</collectorName>
</location>
: XC 231
</materialsCitation>
.
</paragraph>
<caption id="DF7666468363FFC1B1D34E9A215AFC68" ID-DOI="http://doi.org/10.5281/zenodo.8326336" ID-Zenodo-Dep="8326336" httpUri="https://zenodo.org/record/8326336/files/figure.png" pageId="25" pageNumber="253" startId="25.[129,194,909,933]" targetBox="[149,757,345,881]" targetPageId="25" targetType="figure">
<paragraph id="8BB636CE8363FFC1B1D34E9A215AFC68" blockId="25.[129,721,909,961]" pageId="25" pageNumber="253">
<emphasis id="B97DEADC8363FFC1B1D34E9A20E3FC0C" bold="true" box="[129,229,909,933]" pageId="25" pageNumber="253">Figure 17.</emphasis>
Side view of adult plumages in
<emphasis id="B97DEADC8363FFC1B3594E99227BFC0C" box="[523,637,909,933]" italics="true" pageId="25" pageNumber="253">C. b. nebulus</emphasis>
ssp. nov. Photograph by M.R.H.
</paragraph>
</caption>
<caption id="DF7666468363FFC1B1D34AB5258CF813" ID-DOI="http://doi.org/10.5281/zenodo.8326338" ID-Zenodo-Dep="8326338" box="[129,1418,1954,1979]" httpUri="https://zenodo.org/record/8326338/files/figure.png" pageId="25" pageNumber="253" startId="25.[129,194,1954,1978]" targetBox="[283,1360,1042,1926]" targetPageId="25" targetType="figure">
<paragraph id="8BB636CE8363FFC1B1D34AB5258CF813" blockId="25.[129,1418,1954,1979]" box="[129,1418,1954,1979]" pageId="25" pageNumber="253">
<emphasis id="B97DEADC8363FFC1B1D34AB520E3F812" bold="true" box="[129,229,1954,1979]" pageId="25" pageNumber="253">Figure 18.</emphasis>
Ventral and dorsal views of juvenile plumages in
<taxonomicName id="4C094D4D8363FFC1B3E34AB42319F812" box="[689,799,1955,1979]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="species" species="hellmayri">
<emphasis id="B97DEADC8363FFC1B3E34AB42319F812" box="[689,799,1955,1979]" italics="true" pageId="25" pageNumber="253">C. hellmayri</emphasis>
</taxonomicName>
,
<emphasis id="B97DEADC8363FFC1B27B4AB4239DF812" box="[809,923,1955,1979]" italics="true" pageId="25" pageNumber="253">C. b. nebulus</emphasis>
ssp. nov., and
<emphasis id="B97DEADC8363FFC1B5724AB424A3F812" box="[1056,1189,1955,1979]" italics="true" pageId="25" pageNumber="253">
C. b.
<taxonomicName id="4C094D4D8363FFC1B51F4AB424A3F812" box="[1101,1189,1955,1979]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="25" pageNumber="253" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
. Photograph by M.R.H.
</paragraph>
</caption>
<paragraph id="8BB636CE8360FFC2B1DF4D8720EBFE48" blockId="26.[113,763,144,481]" pageId="26" pageNumber="254">
<taxonomicName id="4C094D4D8360FFC2B1DF4D872166FF01" authorityName="Bonaparte" authorityYear="1850" box="[141,352,144,168]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="genus">
<emphasis id="B97DEADC8360FFC2B1DF4D872166FF01" box="[141,352,144,168]" italics="true" pageId="26" pageNumber="254">Catharus b. nebulus</emphasis>
</taxonomicName>
ssp. nov. (
<emphasis id="B97DEADC8360FFC2B0BA4D8621FDFF01" box="[488,507,145,168]" italics="true" pageId="26" pageNumber="254">N</emphasis>
= 18):
<materialsCitation id="3B613C938360FFC2B3064D872246FF6E" ID-GBIF-Occurrence="4406001364" pageId="26" pageNumber="254">
<collectingCountry id="F31E765E8360FFC2B3064D872282FF0E" box="[596,644,144,167]" name="Peru" pageId="26" pageNumber="254">Peru</collectingCountry>
(
<emphasis id="B97DEADC8360FFC2B3CC4D8622B7FF01" box="[670,689,145,168]" italics="true" pageId="26" pageNumber="254">N</emphasis>
= 18): Amazonas: Camino Atuen: ML 398834941;
</materialsCitation>
<materialsCitation id="3B613C938360FFC2B31A4DB8211CFF4E" ID-GBIF-Occurrence="4406001321" pageId="26" pageNumber="254">Camp Lowbrow: ML 304312761;</materialsCitation>
<materialsCitation id="3B613C938360FFC2B0724DD82286FF4F" ID-GBIF-Occurrence="4406001445" box="[288,640,206,231]" pageId="26" pageNumber="254">Fundo Alto Nieva: ML 72665721;</materialsCitation>
<materialsCitation id="3B613C938360FFC2B3D74DD821EBFEAF" ID-GBIF-Occurrence="4406001500" pageId="26" pageNumber="254">La Llantera cerca de Afluente: ML 272483101;</materialsCitation>
<materialsCitation id="3B613C938360FFC2B0AA4DF920C1FE8C" ID-GBIF-Occurrence="4406001394" pageId="26" pageNumber="254">above Leimebamba: XC 175873;</materialsCitation>
<materialsCitation id="3B613C938360FFC2B19D4C1A217AFEEC" ID-GBIF-Occurrence="4406001463" pageId="26" pageNumber="254">Utcubamba: Bagua Grande, ACP Bosque Berlín: ML 129649691, XC 123919;</materialsCitation>
<materialsCitation id="3B613C938360FFC2B0DA4C3A212DFECD" ID-GBIF-Occurrence="4406001372" pageId="26" pageNumber="254">
<collectingRegion id="49CDF82C8360FFC2B0DA4C3A21EDFEEC" box="[392,491,301,325]" country="Peru" name="Huanuco" pageId="26" pageNumber="254">Huánuco</collectingRegion>
: Paty Trail: XC 243435, 243436, 243437;
</materialsCitation>
<materialsCitation id="3B613C938360FFC2B0684C5B20E8FE2A" ID-GBIF-Occurrence="4406001498" pageId="26" pageNumber="254">
<collectingRegion id="49CDF82C8360FFC2B0684C5B217FFECA" box="[314,377,332,355]" country="Peru" name="Pasco" pageId="26" pageNumber="254">Pasco</collectingRegion>
: Bosque Shollet: ML 228217101, 289516811;
</materialsCitation>
<materialsCitation id="3B613C938360FFC2B1AC4C7C22C2FE2A" ID-GBIF-Occurrence="4406001474" box="[254,708,363,387]" pageId="26" pageNumber="254">Oxapampa: ML 269963401, 376022481;</materialsCitation>
<materialsCitation id="3B613C938360FFC2B3874C7C22FCFE0B" ID-GBIF-Occurrence="4406001505" pageId="26" pageNumber="254">
<collectingRegion id="49CDF82C8360FFC2B3874C7C20BBFE0A" country="Peru" name="San Martin" pageId="26" pageNumber="254">San Martín</collectingRegion>
: Abra Patricia: ECOAN Lodge: XC 30494, 144628;
</materialsCitation>
<materialsCitation id="3B613C938360FFC2B1234CBD21D2FE6B" ID-GBIF-Occurrence="4406001389" box="[113,468,426,450]" pageId="26" pageNumber="254">
<collectingRegion id="49CDF82C8360FFC2B1234CBD20C4FE6B" box="[113,194,426,450]" country="Peru" name="Ucayali" pageId="26" pageNumber="254">Ucayali</collectingRegion>
: Oventeni: ML 138773;
</materialsCitation>
<materialsCitation id="3B613C938360FFC2B0B14CBD20EBFE48" ID-GBIF-Occurrence="4406001351" pageId="26" pageNumber="254">Ulcumano Ecolodge: ML 287691191.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31365458360FFC2B0D54F192550FCBC" pageId="26" pageNumber="254" type="diagnosis">
<paragraph id="8BB636CE8360FFC2B0D54F1921E2FD8E" blockId="26.[391,484,526,551]" box="[391,484,526,551]" pageId="26" pageNumber="254">
<emphasis id="B97DEADC8360FFC2B0D54F1921E2FD8E" box="[391,484,526,551]" italics="true" pageId="26" pageNumber="254">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BB636CE8360FFC2B1234F2322FCF9FF" blockId="26.[113,763,564,1622]" pageId="26" pageNumber="254">
<emphasis id="B97DEADC8360FFC2B1234F2320CCFDE2" box="[113,202,564,587]" italics="true" pageId="26" pageNumber="254">Genetics:</emphasis>
In the UCE tree, samples from (i) western
<collectingCountry id="F31E765E8360FFC2B3F64F2322FCFDE5" box="[676,762,564,588]" name="Ecuador" pageId="26" pageNumber="254">Ecuador</collectingCountry>
and north-western
<collectingCountry id="F31E765E8360FFC2B0654F43216FFDC2" box="[311,361,596,619]" name="Peru" pageId="26" pageNumber="254">Peru</collectingCountry>
, encompassing the type localities of
<taxonomicName id="4C094D4D8360FFC2B3B64F442172FD23" authority="Lawrence 1888" authorityName="Lawrence" authorityYear="1888" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B3B64F4420D6FD23" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
<bibRefCitation id="EF984B3F8360FFC2B1844F642172FD23" author="Lawrence GN" box="[214,372,626,650]" pageId="26" pageNumber="254" pagination="503" refId="ref32168" refString="Lawrence GN. Description of a new species of bird of the genus Catharus, from Ecuador. Proceedings of the United States National Museum ' 1887 ' (= 1888); [1887]: 503." type="journal article" year="1888">Lawrence 1888</bibRefCitation>
</taxonomicName>
and
<emphasis id="B97DEADC8360FFC2B0F44F642225FD23" box="[422,547,626,650]" italics="true" pageId="26" pageNumber="254">C. f. caniceps</emphasis>
<bibRefCitation id="EF984B3F8360FFC2B37B4F6522CCFD23" author="Chapman FM" box="[553,714,626,650]" pageId="26" pageNumber="254" pagination="1 - 16" refId="ref30009" refString="Chapman FM. Descriptions of new birds from Ecuador, Colombia, Peru, and Bolivia. American Museum Novitates 1924; 138: 1 - 16." type="journal article" year="1924">Chapman 1924</bibRefCitation>
, and (ii) eastern
<collectingCountry id="F31E765E8360FFC2B1B54F852111FD00" box="[231,279,658,681]" name="Peru" pageId="26" pageNumber="254">Peru</collectingCountry>
from Amazonas south to the Río
<collectingRegion id="49CDF82C8360FFC2B33C4F8522D2FD03" box="[622,724,658,682]" country="Peru" name="Apurimac" pageId="26" pageNumber="254">Apurímac</collectingRegion>
(
<emphasis id="B97DEADC8360FFC2B3B64F8520D5FD60" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov.), formed reciprocally monophyletic clades. Together, these clades formed the sister-group of a clade composed of samples of
<emphasis id="B97DEADC8360FFC2B0184FE72167FCAE" box="[330,353,752,775]" italics="true" pageId="26" pageNumber="254">C.</emphasis>
[
<emphasis id="B97DEADC8360FFC2B0264FE72187FCA1" box="[372,385,752,776]" italics="true" pageId="26" pageNumber="254">f.</emphasis>
]
<taxonomicName id="4C094D4D8360FFC2B0C04FE721FDFCAE" box="[402,507,752,775]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="opertaneus">
<emphasis id="B97DEADC8360FFC2B0C04FE721FDFCAE" box="[402,507,752,775]" italics="true" pageId="26" pageNumber="254">opertaneus</emphasis>
</taxonomicName>
and Undescribed 2. We treat these major sister-groups as polytypic species:
<taxonomicName id="4C094D4D8360FFC2B3D24E1822FDFC8E" authorityName="Lawrence" authorityYear="1888" box="[640,763,783,807]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B3D24E1822FDFC8E" box="[640,763,783,807]" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
</taxonomicName>
(including
<emphasis id="B97DEADC8360FFC2B1BB4E38218FFCEF" box="[233,393,814,838]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B0784E39218FFCEF" authorityName="Lawrence" authorityYear="1888" box="[298,393,814,838]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
,
<emphasis id="B97DEADC8360FFC2B0C94E38222DFCEF" box="[411,555,814,838]" italics="true" pageId="26" pageNumber="254">C. b. caniceps</emphasis>
,
<emphasis id="B97DEADC8360FFC2B36C4E3922FCFCEF" box="[574,762,814,838]" italics="true" pageId="26" pageNumber="254">and C. b. nebulus</emphasis>
ssp. nov.) and
<taxonomicName id="4C094D4D8360FFC2B05D4E59219EFCCC" box="[271,408,846,869]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="opertaneus">
<emphasis id="B97DEADC8360FFC2B05D4E59219EFCCC" box="[271,408,846,869]" italics="true" pageId="26" pageNumber="254">C. opertaneus</emphasis>
</taxonomicName>
(including
<emphasis id="B97DEADC8360FFC2B3454E592228FCCC" box="[535,558,846,869]" italics="true" pageId="26" pageNumber="254">C.</emphasis>
[
<emphasis id="B97DEADC8360FFC2B3104E592249FCCF" box="[578,591,846,870]" italics="true" pageId="26" pageNumber="254">f.</emphasis>
]
<taxonomicName id="4C094D4D8360FFC2B3334E5922CCFCCC" box="[609,714,846,869]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="opertaneus">
<emphasis id="B97DEADC8360FFC2B3334E5922CCFCCC" box="[609,714,846,869]" italics="true" pageId="26" pageNumber="254">opertaneus</emphasis>
</taxonomicName>
and Undescribed 2, see below). The subspecies
<emphasis id="B97DEADC8360FFC2B3654E7A22CBFC2C" box="[567,717,877,901]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B33C4E7A22CBFC2C" authorityName="Lawrence" authorityYear="1888" box="[622,717,877,901]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
and
<emphasis id="B97DEADC8360FFC2B1234E9A20F8FC0D" box="[113,254,908,932]" italics="true" pageId="26" pageNumber="254">C. b. caniceps</emphasis>
were not genetically diagnosable, despite their plumage colour differences. A sample of
<emphasis id="B97DEADC8360FFC2B35E4EBB229BFC6D" box="[524,669,940,964]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B36C4EBB229BFC6D" authorityName="Lawrence" authorityYear="1888" box="[574,669,940,964]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
from the Yunguilla Valley,
<collectingRegion id="49CDF82C8360FFC2B0744EDC2160FC4A" box="[294,358,971,995]" country="Ecuador" name="Azuay" pageId="26" pageNumber="254">Azuay</collectingRegion>
,
<collectingCountry id="F31E765E8360FFC2B0204EDC21CEFC4A" box="[370,456,971,995]" name="Ecuador" pageId="26" pageNumber="254">Ecuador</collectingCountry>
(ZMUC 128360) was more similar to a
<typeStatus id="54B2886C8360FFC2B1BD4EFC214EFBAA" box="[239,328,1003,1027]" pageId="26" pageNumber="254" type="paratype">paratype</typeStatus>
of
<emphasis id="B97DEADC8360FFC2B03E4EFC21EBFBAB" box="[364,493,1002,1026]" italics="true" pageId="26" pageNumber="254">C. f. caniceps</emphasis>
(AMNH 175531) than a sample of
<emphasis id="B97DEADC8360FFC2B1B7491D2185FB8B" box="[229,387,1034,1058]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B076491D2185FB8B" authorityName="Lawrence" authorityYear="1888" box="[292,387,1034,1058]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
from
<collectingRegion id="49CDF82C8360FFC2B099491D2236FB8B" box="[459,560,1034,1058]" country="Ecuador" name="Imbabura" pageId="26" pageNumber="254">Imbabura</collectingRegion>
,
<collectingCountry id="F31E765E8360FFC2B312491D2290FB8B" box="[576,662,1034,1058]" name="Ecuador" pageId="26" pageNumber="254">Ecuador</collectingCountry>
(ZMUC 119570). In the
<emphasis id="B97DEADC8360FFC2B04A493E214DFBE8" box="[280,331,1065,1089]" italics="true" pageId="26" pageNumber="254">ND2</emphasis>
tree, these three samples, plus one sample each from the Huancabamba and Chunchuca valleys (i.e. east of the Abra de Porculla, an eastwest pass through the Western Cordillera at
<quantity id="4CF19B2B8360FFC2B05B49902164FB36" box="[265,354,1159,1183]" metricMagnitude="3" metricUnit="m" metricValue="2.145" pageId="26" pageNumber="254" unit="m" value="2145.0">2145 m</quantity>
) and a sample from
<collectingRegion id="49CDF82C8360FFC2B30F4990228BFB36" box="[605,653,1159,1183]" country="Ecuador" name="Loja" pageId="26" pageNumber="254">Loja</collectingRegion>
,
<collectingCountry id="F31E765E8360FFC2B3CD499022F0FB36" box="[671,758,1159,1183]" name="Ecuador" pageId="26" pageNumber="254">Ecuador</collectingCountry>
, formed a clade with low within-clade divergence (mean uncorrected
<emphasis id="B97DEADC8360FFC2B1B349D120E8FB74" box="[225,238,1222,1245]" italics="true" pageId="26" pageNumber="254">p</emphasis>
-distance = 0.005 ± 0.004). Samples from across the range of the eastern Andean group (
<emphasis id="B97DEADC8360FFC2B34149F22294FB54" box="[531,658,1253,1277]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov.) also formed a clade in the
<emphasis id="B97DEADC8360FFC2B0D8481321BBFAB5" box="[394,445,1284,1308]" italics="true" pageId="26" pageNumber="254">ND2</emphasis>
tree with low within-clade divergence (mean uncorrected
<emphasis id="B97DEADC8360FFC2B0F6483321B7FA92" box="[420,433,1316,1339]" italics="true" pageId="26" pageNumber="254">p</emphasis>
-distance = 0.002 ± 0.001). The divergence between these sister-clades (
<emphasis id="B97DEADC8360FFC2B345485422A8FAF2" box="[535,686,1347,1371]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B31D485422A8FAF2" authorityName="Lawrence" authorityYear="1888" box="[591,686,1347,1371]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
+
<emphasis id="B97DEADC8360FFC2B399485420C6FAD3" italics="true" pageId="26" pageNumber="254">C. b. caniceps</emphasis>
, vs.
<emphasis id="B97DEADC8360FFC2B1A348742176FAD3" box="[241,368,1378,1402]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov.) was an order of magnitude greater (mean uncorrected
<emphasis id="B97DEADC8360FFC2B0CC489521ADFA30" box="[414,427,1410,1433]" italics="true" pageId="26" pageNumber="254">p</emphasis>
-distance = 0.022 ± 0.002) than the mean divergence within either clade. ABGD analysis of
<emphasis id="B97DEADC8360FFC2B12348D720A2FA71" box="[113,164,1472,1496]" italics="true" pageId="26" pageNumber="254">ND2</emphasis>
data merged
<taxonomicName id="4C094D4D8360FFC2B07848D621A2FA71" authorityName="Lawrence" authorityYear="1888" box="[298,420,1472,1496]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B07848D621A2FA71" box="[298,420,1472,1496]" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
</taxonomicName>
with Undescribed 2 (uncorrected
<emphasis id="B97DEADC8360FFC2B12348F72078FA5E" box="[113,126,1504,1527]" italics="true" pageId="26" pageNumber="254">p</emphasis>
-distance = 0.01 ± &lt;0.01), although these were not sister-groups in the UCE tree, while treating
<emphasis id="B97DEADC8360FFC2B35748E8229FF9BE" box="[517,665,1535,1559]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B36848E8229FF9BE" authorityName="Lawrence" authorityYear="1888" box="[570,665,1535,1559]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
and
<emphasis id="B97DEADC8360FFC2B39E48E820BCF99F" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. as independent genetic clusters. ASAP analysis merged
<emphasis id="B97DEADC8360FFC2B1914B292157F9FC" box="[195,337,1597,1621]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B1A04B2A2157F9FC" authorityName="Lawrence" authorityYear="1888" box="[242,337,1597,1621]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
,
<emphasis id="B97DEADC8360FFC2B0094B2921D5F9FC" box="[347,467,1597,1621]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov., and Undescribed 2.
</paragraph>
<paragraph id="8BB636CE8360FFC2B1234B602454FE2A" blockId="26.[113,763,1655,1961]" lastBlockId="26.[810,1460,144,387]" pageId="26" pageNumber="254">
<emphasis id="B97DEADC8360FFC2B1234B6020F6F926" box="[113,240,1655,1679]" italics="true" pageId="26" pageNumber="254">Morphology:</emphasis>
Study skins from the range of
<taxonomicName id="4C094D4D8360FFC2B36A4B6022B3F926" authorityName="Lawrence" authorityYear="1888" box="[568,693,1655,1679]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B36A4B6022B3F926" box="[568,693,1655,1679]" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
</taxonomicName>
in the western Andes were sexually monochromatic in the adult plumage and exhibited a northsouth colour cline. At the northern end of the cline, adults had dark dorsal plumage (darker than Sepia, 119), medium-light ventral plumage (
<figureCitation id="13322A4B8360FFC2B32C4BE322CCF8A5" box="[638,714,1780,1804]" captionStart="Figure 15" captionStartId="23.[129,194,1708,1732]" captionTargetBox="[257,1346,156,1680]" captionTargetId="figure-75@23.[247,1356,144,1680]" captionTargetPageId="23" captionText="Figure 15. Ventral and dorsal views of the adult male plumages in C. b. berlepschi, C. b. caniceps, C. o.tenebris ssp. nov., C. b. nebulus ssp.nov., and C. mentalis. Photograph by M.R.H." figureDoi="http://doi.org/10.5281/zenodo.8326332" httpUri="https://zenodo.org/record/8326332/files/figure.png" pageId="26" pageNumber="254">Figs 15</figureCitation>
,
<figureCitation id="13322A4B8360FFC2B38B4BE322F5F8A5" box="[729,755,1780,1804]" captionStart="Figure 16" captionStartId="24.[113,178,1708,1732]" captionTargetBox="[285,1290,156,1680]" captionTargetId="figure-124@24.[282,1290,144,1680]" captionTargetPageId="24" captionText="Figure 16. Ventral and dorsal views of the adult female plumages in C. b. berlepschi, C. b. caniceps, C. o.tenebris ssp. nov., C. b. nebulus ssp.nov., and C. mentalis. Photograph by M.R.H." figureDoi="http://doi.org/10.5281/zenodo.8326334" httpUri="https://zenodo.org/record/8326334/files/figure.png" pageId="26" pageNumber="254">16</figureCitation>
;
<tableCitation id="C68B03758360FFC2B1234A0420C4F882" box="[113,194,1811,1835]" captionStart="Table 3" captionStartId="12.[115,170,143,167]" captionTargetPageId="12" captionText="Table 3. Adult plumage colour variation in the C.[fuscater] complex based on specimens in the ANSP, DMNH, FMNH, and LSUMZ. Numbers taken from Smithe (1975) denote colours on the crown, back, undertail coverts (tl-covs), flanks, breast, and throat. Chin is scored with two levels (0,1) referring to the absence or presence, respectively, of an unbroken (although sometimes faint) line of darker feathers connecting the malar regions across the chin.Specimens from southern Ecuador and north-western Peru (C. [f.] caniceps) were scored separately from north-western Ecuador (C. [f.] berlepschi), despite showing low genetic divergence, to illustrate plumage colour differences. Specimens from eastern Panama (C.[f.] mirabilis, Undescribed 1) were scored using non-fresh (faded) specimens (see text). Color scores from a brown type unsexed specimen of C. [f.] fuscater, the only study skin of that taxon available for the formal analysis, are included below as female." httpUri="http://table.plazi.org/id/DF7666468376FFD4B1214D9820BBFEC2" pageId="26" pageNumber="254" tableUuid="DF7666468376FFD4B1214D9820BBFEC2">Table 3</tableCitation>
), and completely black maxillae (
<tableCitation id="C68B03758360FFC2B3684A04228DF882" box="[570,651,1811,1835]" captionStart="Table 4" captionStartId="13.[129,184,1058,1082]" captionTargetPageId="13" captionText="Table 4. Taxonomic, sexual, and age-related variation in the extent of black on the maxilla, scored with five levels (see text) in a geographically broad sample of C. [fuscater] study skins (N = 295). The endpoints of the scale are no black (0) and completely black (4). Means (±SD) are shown for adults of each sex, and immatures (both sexes combined)" httpUri="http://table.plazi.org/id/DF7666468377FFD5B1D349352306FBDB" pageId="26" pageNumber="254" tableUuid="DF7666468377FFD5B1D349352306FBDB">Table 4</tableCitation>
), whereas southern adults (
<emphasis id="B97DEADC8360FFC2B0714A2421A3F8E3" box="[291,421,1843,1867]" italics="true" pageId="26" pageNumber="254">C. b. caniceps</emphasis>
), including in the Huancabamba and Chunchuca valleys, were paler on the flanks, breast, and throat (
<figureCitation id="13322A4B8360FFC2B1934A66210DF820" box="[193,267,1905,1929]" captionStart="Figure 15" captionStartId="23.[129,194,1708,1732]" captionTargetBox="[257,1346,156,1680]" captionTargetId="figure-75@23.[247,1356,144,1680]" captionTargetPageId="23" captionText="Figure 15. Ventral and dorsal views of the adult male plumages in C. b. berlepschi, C. b. caniceps, C. o.tenebris ssp. nov., C. b. nebulus ssp.nov., and C. mentalis. Photograph by M.R.H." figureDoi="http://doi.org/10.5281/zenodo.8326332" httpUri="https://zenodo.org/record/8326332/files/figure.png" pageId="26" pageNumber="254">Figs 15</figureCitation>
,
<figureCitation id="13322A4B8360FFC2B0454A662137F820" box="[279,305,1905,1929]" captionStart="Figure 16" captionStartId="24.[113,178,1708,1732]" captionTargetBox="[285,1290,156,1680]" captionTargetId="figure-124@24.[282,1290,144,1680]" captionTargetPageId="24" captionText="Figure 16. Ventral and dorsal views of the adult female plumages in C. b. berlepschi, C. b. caniceps, C. o.tenebris ssp. nov., C. b. nebulus ssp.nov., and C. mentalis. Photograph by M.R.H." figureDoi="http://doi.org/10.5281/zenodo.8326334" httpUri="https://zenodo.org/record/8326334/files/figure.png" pageId="26" pageNumber="254">16</figureCitation>
;
<tableCitation id="C68B03758360FFC2B06D4A66218DF820" box="[319,395,1905,1929]" captionStart="Table 3" captionStartId="12.[115,170,143,167]" captionTargetPageId="12" captionText="Table 3. Adult plumage colour variation in the C.[fuscater] complex based on specimens in the ANSP, DMNH, FMNH, and LSUMZ. Numbers taken from Smithe (1975) denote colours on the crown, back, undertail coverts (tl-covs), flanks, breast, and throat. Chin is scored with two levels (0,1) referring to the absence or presence, respectively, of an unbroken (although sometimes faint) line of darker feathers connecting the malar regions across the chin.Specimens from southern Ecuador and north-western Peru (C. [f.] caniceps) were scored separately from north-western Ecuador (C. [f.] berlepschi), despite showing low genetic divergence, to illustrate plumage colour differences. Specimens from eastern Panama (C.[f.] mirabilis, Undescribed 1) were scored using non-fresh (faded) specimens (see text). Color scores from a brown type unsexed specimen of C. [f.] fuscater, the only study skin of that taxon available for the formal analysis, are included below as female." httpUri="http://table.plazi.org/id/DF7666468376FFD4B1214D9820BBFEC2" pageId="26" pageNumber="254" tableUuid="DF7666468376FFD4B1214D9820BBFEC2">Table 3</tableCitation>
) and had dichromatic irides (
<tableCitation id="C68B03758360FFC2B3904A662078F801" captionStart="Table 5" captionStartId="14.[113,168,143,167]" captionTargetPageId="14" captionText="Table 5. Taxonomic variation in iris colour, scored with two levels (brown vs. white, see text) in a sample of C. [fuscater] study skins and digital photos (N = 340). Populations in which brown irides were detected in &lt;20% of specimens were categorized as monochromatic white (mono white). Populations in which white irides were detected in &lt;20% of specimens were categorized as monochromatic brown (mono brown).Populations in which brown and white irides were both detected at greater than 20% frequency are categorized as dichromatic. The complete list of specimens and digital photos is provided in the Supporting Information, Table S1." httpUri="http://table.plazi.org/id/DF7666468374FFD6B1234D9822E3FE0B" pageId="26" pageNumber="254" tableUuid="DF7666468374FFD6B1234D9822E3FE0B">Table 5</tableCitation>
). Specimens from Zaruma,
<collectingRegion id="49CDF82C8360FFC2B0E34A8621FDF800" box="[433,507,1937,1961]" country="Ecuador" name="El Oro" pageId="26" pageNumber="254">El Oro</collectingRegion>
(ANSP 83643, 83644) had somewhat intermediate characters between the endpoints of the plumage colour cline, and are here listed under the nominate form,
<emphasis id="B97DEADC8360FFC2B2CD4DD82437FF4F" box="[927,1073,206,230]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B2804DD92437FF4F" authorityName="Lawrence" authorityYear="1888" box="[978,1073,206,230]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
. Adult study skins of
<emphasis id="B97DEADC8360FFC2B45F4DD8258EFF4F" box="[1293,1416,206,230]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. were dichromatic in iris colour and more similar in plumage colour to
<emphasis id="B97DEADC8360FFC2B2DF4C1A2408FE8C" box="[909,1038,269,293]" italics="true" pageId="26" pageNumber="254">C. b. caniceps</emphasis>
than
<emphasis id="B97DEADC8360FFC2B51A4C1A24DFFE8C" box="[1096,1241,269,293]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B5284C1A24DFFE8C" authorityName="Lawrence" authorityYear="1888" box="[1146,1241,269,293]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
. A juvenile specimen identified as
<emphasis id="B97DEADC8360FFC2B2F84C3A2425FEED" box="[938,1059,300,324]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
was morphologically divergent from
<emphasis id="B97DEADC8360FFC2B4CF4C3A23A6FECD" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B2134C5B23A6FECD" authorityName="Lawrence" authorityYear="1888" box="[833,928,332,356]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
and
<emphasis id="B97DEADC8360FFC2B2804C5B2453FECA" box="[978,1109,332,356]" italics="true" pageId="26" pageNumber="254">C. b. caniceps</emphasis>
(see below), but this requires further investigation to confirm.
</paragraph>
<paragraph id="8BB636CE8360FFC2B2784CB22550FCBC" blockId="26.[810,1460,420,789]" pageId="26" pageNumber="254">
<emphasis id="B97DEADC8360FFC2B2784CB22362FE15" box="[810,868,421,444]" italics="true" pageId="26" pageNumber="254">Voice:</emphasis>
The blurred calls of
<emphasis id="B97DEADC8360FFC2B56C4CB224D5FE15" box="[1086,1235,420,444]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B5264CB324D5FE15" authorityName="Lawrence" authorityYear="1888" box="[1140,1235,420,444]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
and
<emphasis id="B97DEADC8360FFC2B45A4CB22580FE15" box="[1288,1414,420,444]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. had a check-like sonogram shape, which distinguished them from all other taxa in the complex, including Undescribed 2, which had a downward sloping blurred call that was narrower in bandwidth (
<figureCitation id="13322A4B8360FFC2B2964F36240CFD93" box="[964,1034,545,570]" captionStart="Figure 11" captionStartId="16.[113,178,1881,1905]" captionTargetBox="[305,1267,1451,1852]" captionTargetId="figure-38@16.[304,1267,1449,1853]" captionTargetPageId="16" captionText="Figure 11. Taxonomic variation in the structure of blurred calls, visualized in RavenPro 1.5 (Cornell University). The black line divides calls that modulate upward in frequency (left) and downward in frequency (right); both types were detected in C. [f.] berlepschi." figureDoi="http://doi.org/10.5281/zenodo.8326324" httpUri="https://zenodo.org/record/8326324/files/figure.png" pageId="26" pageNumber="254">Fig. 11</figureCitation>
). The subspecies
<emphasis id="B97DEADC8360FFC2B5904F352552FD90" box="[1218,1364,545,569]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B5A74F362552FD90" authorityName="Lawrence" authorityYear="1888" box="[1269,1364,545,569]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
and
<emphasis id="B97DEADC8360FFC2B4D44F352375FDF0" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. differed in punctuation call structure (
<typeStatus id="54B2886C8360FFC2B4184F562585FDF0" box="[1354,1411,577,601]" pageId="26" pageNumber="254">Type</typeStatus>
2 vs.
<typeStatus id="54B2886C8360FFC2B2784F762359FDD0" box="[810,863,609,633]" pageId="26" pageNumber="254">Type</typeStatus>
3, respectively). In song, of the five triadic contours (ABC, ACB, BCA, CAB, CBA) detected in
<emphasis id="B97DEADC8360FFC2B5F34F972532FD3E" box="[1185,1332,639,663]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B5874F682532FD3E" authorityName="Lawrence" authorityYear="1888" box="[1237,1332,639,663]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
, four (ABC, ACB, BAC, BCA) were shared with
<emphasis id="B97DEADC8360FFC2B5CA4F882514FD1E" box="[1176,1298,671,695]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov.; of the four tetradic contours (ACDB, BCDA, CABD, CDBA) detected in
<emphasis id="B97DEADC8360FFC2B2144FC923DCFD5C" box="[838,986,733,757]" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B2294FCA23DCFD5C" authorityName="Lawrence" authorityYear="1888" box="[891,986,733,757]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
, none was shared with
<emphasis id="B97DEADC8360FFC2B5824FC92549FD5C" box="[1232,1359,733,757]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov., and only one (BDAC) was shared with Undescribed 2.
</paragraph>
</subSubSection>
<subSubSection id="C31365458360FFC2B56E4E20237FFBBB" pageId="26" pageNumber="254" type="etymology">
<paragraph id="8BB636CE8360FFC2B56E4E2024A4FCF8" blockId="26.[810,1460,823,1042]" box="[1084,1186,823,849]" pageId="26" pageNumber="254">
<emphasis id="B97DEADC8360FFC2B56E4E2024A4FCF8" box="[1084,1186,823,849]" italics="true" pageId="26" pageNumber="254">Etymology</emphasis>
</paragraph>
<paragraph id="8BB636CE8360FFC2B2784E49237FFBBB" blockId="26.[810,1460,823,1042]" pageId="26" pageNumber="254">
We propose the English name Trans-Andean nightingale-thrush for the polytypic
<taxonomicName id="4C094D4D8360FFC2B5794E6A24A1FC3C" authorityName="Lawrence" authorityYear="1888" box="[1067,1191,893,917]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B5794E6A24A1FC3C" box="[1067,1191,893,917]" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
</taxonomicName>
because it is the only species in the complex that occurs on both slopes of the Andes. The scientific name
<emphasis id="B97DEADC8360FFC2B2AB4EAB2471FC7A" box="[1017,1143,955,979]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. is derived from Latin feminine noun
<emphasis id="B97DEADC8360FFC2B2974ECC2401FC5A" box="[965,1031,987,1011]" italics="true" pageId="26" pageNumber="254">nebula</emphasis>
(mist, fog), referring to its montane forest habitat.
</paragraph>
</subSubSection>
<subSubSection id="C31365458360FFC3B569492A2140FA24" lastPageId="27" lastPageNumber="255" pageId="26" pageNumber="254" type="discussion">
<paragraph id="8BB636CE8360FFC2B569492A24A7FBFE" blockId="26.[810,1460,1085,1962]" box="[1083,1185,1085,1111]" pageId="26" pageNumber="254">
<emphasis id="B97DEADC8360FFC2B569492A24A7FBFE" box="[1083,1185,1085,1111]" italics="true" pageId="26" pageNumber="254">Comments</emphasis>
</paragraph>
<paragraph id="8BB636CE8360FFC2B27849732395FA1C" blockId="26.[810,1460,1085,1962]" pageId="26" pageNumber="254">
We placed the name
<emphasis id="B97DEADC8360FFC2B5474973249DFBD2" box="[1045,1179,1123,1147]" italics="true" pageId="26" pageNumber="254">C. f. caniceps</emphasis>
into the synonymy of
<taxonomicName id="4C094D4D8360FFC2B4CE4973238AFB32" authorityName="Lawrence" authorityYear="1888" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B4CE4973238AFB32" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
</taxonomicName>
because, in both reconstructions, genetic samples from western
<collectingCountry id="F31E765E8360FFC2B2E949B52412FB13" box="[955,1044,1186,1210]" name="Ecuador" pageId="26" pageNumber="254">Ecuador</collectingCountry>
and north-western
<collectingCountry id="F31E765E8360FFC2B5B049B42515FB13" box="[1250,1299,1187,1210]" name="Peru" pageId="26" pageNumber="254">Peru</collectingCountry>
formed a clade that encompassed the
<typeStatus id="54B2886C8360FFC2B57649D52455FB73" box="[1060,1107,1218,1242]" pageId="26" pageNumber="254">type</typeStatus>
localities of both taxa, and the older name has priority. Based on the shallow
<emphasis id="B97DEADC8360FFC2B45A49F6253DFB50" box="[1288,1339,1249,1273]" italics="true" pageId="26" pageNumber="254">ND2</emphasis>
divergence between samples from the western Andes, on one hand, and the eastern side of the Abra de Porculla in the Huancabamba (LSUMZ 170103) and upper Chunchuca valleys (LSUMZ 170104) on the other, we hypothesize that
<taxonomicName id="4C094D4D8360FFC2B479484925B5FADF" authorityName="Lawrence" authorityYear="1888" box="[1323,1459,1374,1398]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B479484925B5FADF" box="[1323,1459,1374,1398]" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
</taxonomicName>
crossed the Abra de Porculla, from the west, sometime after the LGM.
</paragraph>
<paragraph id="8BB636CE8360FFC3B21748AB20BAFE8C" blockId="26.[810,1460,1085,1962]" lastBlockId="27.[129,779,144,1421]" lastPageId="27" lastPageNumber="255" pageId="26" pageNumber="254">
Despite its monophyly in both genetic datasets, we treat the taxon from the eastern Andes as a subspecies of
<taxonomicName id="4C094D4D8360FFC2B44048CB258AFA5A" authorityName="Lawrence" authorityYear="1888" box="[1298,1420,1499,1523]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8360FFC2B44048CB258AFA5A" box="[1298,1420,1499,1523]" italics="true" pageId="26" pageNumber="254">C. berlepschi</emphasis>
</taxonomicName>
, instead of its sister-species, because additional sampling is needed to assess the degree of vocal and morphological divergence. We note that
<emphasis id="B97DEADC8360FFC2B2DD4B2D240BF9F8" box="[911,1037,1593,1617]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. is apparently divergent from
<emphasis id="B97DEADC8360FFC2B4CF4B2D23A6F9D8" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B2134B4E23A6F9D8" authorityName="Lawrence" authorityYear="1888" box="[833,928,1625,1649]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
in juvenile plumage and punctuation call structure, but this may be an artefact of low sample size. Study skins of
<emphasis id="B97DEADC8360FFC2B2784B8F23ACF906" box="[810,938,1687,1711]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. also exhibit subtle sexual dichromatism, unlike the sexually monochromatic (but clinally variable)
<emphasis id="B97DEADC8360FFC2B4D74BA0238FF947" italics="true" pageId="26" pageNumber="254">
C. b.
<taxonomicName id="4C094D4D8360FFC2B2784BC1238FF947" authorityName="Lawrence" authorityYear="1888" box="[810,905,1750,1774]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="26" pageNumber="254" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
+
<emphasis id="B97DEADC8360FFC2B2F74BC1242CF947" box="[933,1066,1750,1774]" italics="true" pageId="26" pageNumber="254">C. b. caniceps</emphasis>
clade. Males of
<emphasis id="B97DEADC8360FFC2B5874BC12552F947" box="[1237,1364,1750,1774]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. (e.g. FMNH 474413, LSUMZ 129006) were darker and less brown on the dorsal surface than adult females, including on the crown, and some adult males of
<emphasis id="B97DEADC8360FFC2B53A4A2324E6F8E5" box="[1128,1248,1844,1868]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. approached the brownish plumage of the adult female (e.g. LSUMZ 106496, with enlarged testes: 9 ×
<quantity id="4CF19B2B8360FFC2B57A4A652475F822" box="[1064,1139,1906,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="254" unit="mm" value="15.0">15 mm</quantity>
). This suggests that
<emphasis id="B97DEADC8360FFC2B46B4A6425B5F823" box="[1337,1459,1906,1930]" italics="true" pageId="26" pageNumber="254">C. b. nebulus</emphasis>
ssp. nov. may exhibit polychromatic plumage types like
<emphasis id="B97DEADC8360FFC2B4274A85258AF800" box="[1397,1420,1938,1961]" italics="true" pageId="26" pageNumber="254">C.</emphasis>
[
<emphasis id="B97DEADC8360FFC2B4CF4A8525ADF803" box="[1437,1451,1938,1962]" italics="true" pageId="26" pageNumber="254">f.</emphasis>
]
<taxonomicName id="4C094D4D8361FFC3B1D34D8720D4FF01" box="[129,210,144,168]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="mentalis">
<emphasis id="B97DEADC8361FFC3B1D34D8720D4FF01" box="[129,210,144,168]" italics="true" pageId="27" pageNumber="255">mentalis</emphasis>
</taxonomicName>
(see below). Notably, the strongest colour differences between
<emphasis id="B97DEADC8361FFC3B1B24DA72164FF6E" box="[224,354,175,199]" italics="true" pageId="27" pageNumber="255">C. b. nebulus</emphasis>
ssp. nov. and the
<emphasis id="B97DEADC8361FFC3B3774DA722BBFF6E" box="[549,701,175,199]" italics="true" pageId="27" pageNumber="255">
C. b.
<taxonomicName id="4C094D4D8361FFC3B30C4DB822BBFF6E" authorityName="Lawrence" authorityYear="1888" box="[606,701,175,199]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
+
<emphasis id="B97DEADC8361FFC3B3884DA720D6FF4F" italics="true" pageId="27" pageNumber="255">C. b. caniceps</emphasis>
clade occur where their ranges approach each other in northern
<collectingCountry id="F31E765E8361FFC3B1B74DF92111FEAC" box="[229,279,238,261]" name="Peru" pageId="27" pageNumber="255">Peru</collectingCountry>
, which may be indicative of character displacement.
</paragraph>
<paragraph id="8BB636CE8361FFC3B1CE4C3A20F1FC31" blockId="27.[129,779,144,1421]" pageId="27" pageNumber="255">
The plumage of a juvenile from central
<collectingCountry id="F31E765E8361FFC3B3354C3A229FFEED" box="[615,665,301,324]" name="Peru" pageId="27" pageNumber="255">Peru</collectingCountry>
(LSUMZ 174204), identified as
<emphasis id="B97DEADC8361FFC3B0D44C5B2213FECD" box="[390,533,332,356]" italics="true" pageId="27" pageNumber="255">C. b. nebulus</emphasis>
ssp. nov., was boldly marked relative to a juvenile
<emphasis id="B97DEADC8361FFC3B0E64C7B224AFE2A" box="[436,588,363,387]" italics="true" pageId="27" pageNumber="255">
C. b.
<taxonomicName id="4C094D4D8361FFC3B0B54C7C224AFE2A" authorityName="Lawrence" authorityYear="1888" box="[487,588,363,387]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
(ANSP 181216). LSUMZ 174204 has tawny streaks on the mantle, tawny spots on the breast, a whitish throat and abdomen, and a pale mandible (
<figureCitation id="13322A4B8361FFC3B1AB4CDE2145FE48" box="[249,323,457,481]" captionStart="Figure 19" captionStartId="27.[825,890,733,757]" captionTargetBox="[846,1454,166,705]" captionTargetId="figure-830@27.[846,1454,144,705]" captionTargetPageId="27" captionText="Figure 19. Side view of adult plumages in C. o. tenebris ssp.nov. Photograph by M.R.H." figureDoi="http://doi.org/10.5281/zenodo.8326340" httpUri="https://zenodo.org/record/8326340/files/figure.png" pageId="27" pageNumber="255">Fig. 19</figureCitation>
). At first glance, this specimen resembles the juvenile of
<taxonomicName id="4C094D4D8361FFC3B04D4CFE21A2FDA8" box="[287,420,489,513]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B97DEADC8361FFC3B04D4CFE21A2FDA8" box="[287,420,489,513]" italics="true" pageId="27" pageNumber="255">C. maculatus</emphasis>
</taxonomicName>
, but there are multiple reasons to doubt that identification. CM 85107, a genuine
<taxonomicName id="4C094D4D8361FFC3B3D14F1F230FFD89" box="[643,777,520,544]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B97DEADC8361FFC3B3D14F1F230FFD89" box="[643,777,520,544]" italics="true" pageId="27" pageNumber="255">C. maculatus</emphasis>
</taxonomicName>
juvenile from
<collectingRegion id="49CDF82C8361FFC3B0444F3021A0FD96" box="[278,422,551,575]" country="Bolivia" name="Cochabamba" pageId="27" pageNumber="255">Cochabamba</collectingRegion>
,
<collectingCountry id="F31E765E8361FFC3B0E24F3021F8FD96" box="[432,510,551,575]" name="Bolivia" pageId="27" pageNumber="255">Bolivia</collectingCountry>
, has a dark mandible (vs. pale with dusky tip in LSUMZ 174204), a darker and more heavily marked belly (vs. white), and a brown collar across the upper breast (vs. no collar). LSU 174204 was also collected at
<quantity id="4CF19B2B8361FFC3B1BA4FB22139FD15" box="[232,319,677,701]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="255" unit="m" value="2300.0">2300 m</quantity>
, at the same site as an adult female
<emphasis id="B97DEADC8361FFC3B3854FB220C9FD75" italics="true" pageId="27" pageNumber="255">C. b. nebulus</emphasis>
ssp. nov. (LSUMZ 174203), and about
<quantity id="4CF19B2B8361FFC3B3C64FD322DDFD72" box="[660,731,708,732]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="255" unit="m" value="500.0">500 m</quantity>
upslope of the known upper range limit of
<taxonomicName id="4C094D4D8361FFC3B3604FF322BFFD52" box="[562,697,739,763]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B97DEADC8361FFC3B3604FF322BFFD52" box="[562,697,739,763]" italics="true" pageId="27" pageNumber="255">C. maculatus</emphasis>
</taxonomicName>
in central
<collectingCountry id="F31E765E8361FFC3B1F94E1420DBFCB3" box="[171,221,771,794]" name="Peru" pageId="27" pageNumber="255">Peru</collectingCountry>
(Schulenberg
<emphasis id="B97DEADC8361FFC3B02E4E1421ABFCB3" box="[380,429,770,794]" italics="true" pageId="27" pageNumber="255">et al.</emphasis>
2007). As noted by Ridgely and Tudor (1989: 110),
<taxonomicName id="4C094D4D8361FFC3B0364E3521EBFC93" box="[356,493,802,826]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B97DEADC8361FFC3B0364E3521EBFC93" box="[356,493,802,826]" italics="true" pageId="27" pageNumber="255">C. maculatus</emphasis>
</taxonomicName>
and
<emphasis id="B97DEADC8361FFC3B3754E352239FC90" box="[551,575,802,825]" italics="true" pageId="27" pageNumber="255">C.</emphasis>
[
<taxonomicName id="4C094D4D8361FFC3B3014E3522A4FC93" authorityName="Taczanowski" authorityYear="1874" box="[595,674,802,826]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="fuscater">
<emphasis id="B97DEADC8361FFC3B3014E3522A4FC93" box="[595,674,802,826]" italics="true" pageId="27" pageNumber="255">fuscater</emphasis>
</taxonomicName>
] populations in South America seem rarely if ever actually to occur together, rather, replacing each other in an as yet not understood way.
</paragraph>
<caption id="DF7666468361FFC3B26B4FCA2412FCB9" ID-DOI="http://doi.org/10.5281/zenodo.8326340" ID-Zenodo-Dep="8326340" httpUri="https://zenodo.org/record/8326340/files/figure.png" pageId="27" pageNumber="255" startId="27.[825,890,733,757]" targetBox="[846,1454,166,705]" targetPageId="27" targetType="figure">
<paragraph id="8BB636CE8361FFC3B26B4FCA2412FCB9" blockId="27.[825,1418,733,785]" pageId="27" pageNumber="255">
<emphasis id="B97DEADC8361FFC3B26B4FCA2398FD5C" bold="true" box="[825,926,733,757]" pageId="27" pageNumber="255">Figure 19.</emphasis>
Side view of adult plumages in
<emphasis id="B97DEADC8361FFC3B5964FCA2530FD5C" box="[1220,1334,733,757]" italics="true" pageId="27" pageNumber="255">C. o. tenebris</emphasis>
ssp. nov. Photograph by M.R.H.
</paragraph>
</caption>
<paragraph id="8BB636CE8361FFC3B1CE4EB72140FA24" blockId="27.[129,779,144,1421]" pageId="27" pageNumber="255">
<bibRefCitation id="EF984B3F8361FFC3B1CE4EB72179FC1E" author="Freeman BG &amp; Rolland J &amp; Montgomery GA" box="[156,383,927,951]" pageId="27" pageNumber="255" pagination="20211514" refId="ref30792" refString="Freeman BG, Rolland J, Montgomery GA, et al. Faster evolution of a premating reproductive barrier is not associated with faster speciation rates in New World passerine birds. Proceedings of the Royal Society B 2022; 289: 20211514." type="journal article" year="2022">
Freeman
<emphasis id="B97DEADC8361FFC3B1AC4EB7212EFC1E" box="[254,296,927,951]" italics="true" pageId="27" pageNumber="255">et al</emphasis>
. (2022)
</bibRefCitation>
played recordings of
<emphasis id="B97DEADC8361FFC3B30D4EB722D8FC1E" box="[607,734,927,951]" italics="true" pageId="27" pageNumber="255">C. b. nebulus</emphasis>
ssp. nov. in territories of
<emphasis id="B97DEADC8361FFC3B00C4EA821F7FC7F" box="[350,497,958,982]" italics="true" pageId="27" pageNumber="255">
C. b.
<taxonomicName id="4C094D4D8361FFC3B0C24EA921F7FC7F" authorityName="Lawrence" authorityYear="1888" box="[400,497,958,982]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
and recorded the response of the territorial bird (i.e. how closely the playback target approached the speaker, and whether and how much it vocalized). In each of eight different experiments, one recording was played in the vicinity of a singing individual, and three different stimuli were used overall (XC 175873 for two experiments, XC 30494 for three experiments, XC 144268 for three experiments). We reiterate that interpreting the results of these experiments (and sympatric experiments using audio playback of
<emphasis id="B97DEADC8361FFC3B1CE49CE2135FB59" box="[156,307,1240,1264]" italics="true" pageId="27" pageNumber="255">
C. b.
<taxonomicName id="4C094D4D8361FFC3B18049CF2135FB59" authorityName="Lawrence" authorityYear="1888" box="[210,307,1240,1264]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="berlepschi">berlepschi</taxonomicName>
</emphasis>
recordings in the range of
<taxonomicName id="4C094D4D8361FFC3B30249CE22E0FB59" authorityName="Lawrence" authorityYear="1888" box="[592,742,1240,1264]" class="Aves" family="Turdidae" genus="Catharus" kingdom="Animalia" order="Passeriformes" pageId="27" pageNumber="255" phylum="Chordata" rank="species" species="berlepschi">
<emphasis id="B97DEADC8361FFC3B30249CE22E0FB59" box="[592,742,1240,1264]" italics="true" pageId="27" pageNumber="255">C. b. berlepschi</emphasis>
</taxonomicName>
) is not straightforward, because the function of song (and, therefore, any response to it) is not clearly defined in species that exhibit cooperative parental care and/or facultative territoriality (e.g.
<bibRefCitation id="EF984B3F8361FFC3B1BB4841219AFAC7" author="Goetz JE &amp; McFarland KP &amp; Rimmer CC" box="[233,412,1366,1390]" pageId="27" pageNumber="255" pagination="1044 - 53" refId="ref30922" refString="Goetz JE, McFarland KP, Rimmer CC. Multiple paternity and multiple male feeders in Bicknell's Thrush (Catharus bicknelli). The Auk 2003; 120: 1044 - 53." type="journal article" year="2003">
Goetz
<emphasis id="B97DEADC8361FFC3B06248412167FAC7" box="[304,353,1366,1390]" italics="true" pageId="27" pageNumber="255">et al.</emphasis>
2003
</bibRefCitation>
,
<bibRefCitation id="EF984B3F8361FFC3B0FB4841222CFAC7" author="Halley MR" box="[425,554,1366,1390]" pageId="27" pageNumber="255" refId="ref31052" refString="Halley MR. Kin structure and mating system of the Veery (Catharus fuscescens) in the Mid-Atlantic Piedmont. M. S. Thesis, Delaware State University 2014. hups: // doi. org / 10.13140 / 2.1.1775.9369" type="book" year="2014">Halley 2014</bibRefCitation>
,
<bibRefCitation id="EF984B3F8361FFC3B36548412302FAC4" author="Greeney HF &amp; Dyrcz A &amp; Mikusek R" box="[567,772,1365,1390]" pageId="27" pageNumber="255" pagination="323 - 5" refId="ref30977" refString="Greeney HF, Dyrcz A, Mikusek R, et al. Cooperative breeding at a nest of Slaty-backed Nightingale-Thrushes (Catharus fuscater). Wilson Journal of Ornithology 2015; 127: 323 - 5." type="journal article" year="2015">
Greeney
<emphasis id="B97DEADC8361FFC3B3C5484122CEFAC7" box="[663,712,1366,1390]" italics="true" pageId="27" pageNumber="255">et al.</emphasis>
2015
</bibRefCitation>
,
<bibRefCitation id="EF984B3F8361FFC3B1D348622133FA24" author="Halley MR &amp; Heckscher CM &amp; Kalavacharla V" box="[129,309,1397,1421]" pageId="27" pageNumber="255" pagination="0157051" refId="ref31250" refString="Halley MR, Heckscher CM, Kalavacharla V. Multi-generational kinship, multiple mating, and flexible modes of parental care in a breeding population of the Veery (Catharus fuscescens), a trans-hemispheric migratory songbird. PLoS One 2016; 11: e 0157051." type="journal article" year="2016">
Halley
<emphasis id="B97DEADC8361FFC3B199486120FDFA24" box="[203,251,1397,1421]" italics="true" pageId="27" pageNumber="255">et al.</emphasis>
2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>