320 lines
36 KiB
XML
320 lines
36 KiB
XML
<document id="CD89FF63A6DB9F6A17ABC298C03E79E3" ID="10.11646/zootaxa.4243.1.7SLASH10478" ID-CLB-Dataset="36283" ID-DOI="10.11646/zootaxa.4243.1.7" ID-GBIF-Dataset="1facafb7-179b-4200-ac2f-9d00375a7ce4" ID-ISSN="1175-5326" ID-Zenodo-Dep="398752" ID-ZooBank="96C38B34-BD4C-438B-BC73-FAD6CCBDE212" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489483589214" checkinUser="plazi" docAuthor="Schädel, Mario & Lechner, Thomas S." docDate="2017" docId="03A087A4FFBAFFBAFF42376EFED814C7" docLanguage="en" docName="zootaxa.4243.1.7SLASH10478.pdf" docOrigin="Zootaxa 4243 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gomphaeschna carinthiae Schädel & Lechner, 2017, sp. nov." docType="treatment" docVersion="8" lastPageNumber="162" masterDocId="FF99FFDCFFBCFFB3FFD5316EFFDB1533" masterDocTitle="Two new dragonflies (Odonata: Anisoptera) from the Miocene of Carinthia (Austria)" masterLastPageNumber="164" masterPageNumber="153" pageNumber="159" updateTime="1698437928032" updateUser="ExternalLinkService">
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<mods:mods id="2C172F41A3695FBDF0D1AD2826FAD2F4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FB8F1C55B7CAB1806851CEB22F586C7B">
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<mods:title id="15B0A894F67D4A6C28A61449840852C2">Two new dragonflies (Odonata: Anisoptera) from the Miocene of Carinthia (Austria)</mods:title>
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<mods:name id="0C5B9D483D497AC784222C5E05B554CF" type="personal">
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<mods:roleTerm id="873EA70FE6162D1D1B2193EB58FAF64E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="62AF47CD1079104E0445B13D6098CC0F">Schädel, Mario</mods:namePart>
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</mods:name>
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<mods:name id="DE151ED5825F17AD493F47B1FEDA18F1" type="personal">
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<mods:roleTerm id="6DEB3CDAD41C07AC8FF2F7F78EFF4ABA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C2EA155BE2219C761C76E210024CCAD8">Lechner, Thomas S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="38A77DD96A0D88E19502BCDAF5D92A36">text</mods:typeOfResource>
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<mods:title id="325B56726E8A8EB35B790510A6F435D7">Zootaxa</mods:title>
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<mods:part id="AFDDD9A2D1E2F1040CE1C8F9506EE60C">
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<mods:date id="6EA3EABC3D0C68E48EC5086C241BC498">2017</mods:date>
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<mods:number id="20698409C070E9B6D3EE901A2D15B589">4243</mods:number>
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<mods:detail id="2B95F6522F8D9BEF751A27EE6484B85D" type="issue">
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<mods:number id="6C0E5F22D874DEE94689DF3021FEB497">1</mods:number>
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<mods:classification id="268FA61E7503259B1A65C3E95AA87F4A">journal article</mods:classification>
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<mods:identifier id="03623CAE9EA9A91377BC6A08BEAC8E2E" type="CLB-Dataset">36283</mods:identifier>
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<mods:identifier id="19B756A17671870026DCEE8BE8902F2B" type="DOI">10.11646/zootaxa.4243.1.7</mods:identifier>
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<mods:identifier id="8D2C0D8C6AC64A30C75B5ED132D3DF35" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2D8D02481CCC88072300E40C37AA21A1" type="Zenodo-Dep">398752</mods:identifier>
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<treatment id="03A087A4FFBAFFBAFF42376EFED814C7" ID-DOI="http://doi.org/10.5281/zenodo.6033075" ID-GBIF-Taxon="127843938" ID-Zenodo-Dep="6033075" LSID="urn:lsid:plazi:treatment:03A087A4FFBAFFBAFF42376EFED814C7" httpUri="http://treatment.plazi.org/id/03A087A4FFBAFFBAFF42376EFED814C7" lastPageId="9" lastPageNumber="162" pageId="6" pageNumber="159">
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<subSubSection id="C3136539FFBAFFB5FF42376EFEC7130E" pageId="6" pageNumber="159" type="nomenclature">
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<paragraph id="8BB636B2FFBAFFB5FF42376EFDE01328" blockId="6.[151,571,1536,1597]" box="[151,571,1536,1563]" pageId="6" pageNumber="159">
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<heading id="D0FE81DEFFBAFFB5FF42376EFDE01328" bold="true" box="[151,571,1536,1563]" fontSize="11" level="1" pageId="6" pageNumber="159" reason="1">
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<emphasis id="B97DEAA0FFBAFFB5FF42376EFDE01328" bold="true" box="[151,571,1536,1563]" pageId="6" pageNumber="159">
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<taxonomicName id="4C094D31FFBAFFB5FF42376EFE0C1329" box="[151,471,1536,1562]" class="Insecta" family="Aeshnidae" genus="Gomphaeschna" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="159" phylum="Arthropoda" rank="species" species="carinthiae" status="sp. nov.">
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<emphasis id="B97DEAA0FFBAFFB5FF42376EFE0C1329" bold="true" box="[151,471,1536,1562]" italics="true" pageId="6" pageNumber="159">Gomphaeschna carinthiae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E57DBFFBAFFB5FE0B376FFDE01328" box="[478,571,1537,1563]" pageId="6" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB636B2FFBAFFB5FF42374AFEC7130E" blockId="6.[151,571,1536,1597]" box="[151,284,1572,1597]" pageId="6" pageNumber="159">
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<figureCitation id="13322A37FFBAFFB5FF42374AFEC7130E" box="[151,284,1572,1597]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="7.[151,250,893,915]" captionStartId-1="7.[151,250,1719,1741]" captionStartId-2="8.[151,250,730,752]" captionTargetBox-0="[151,1436,193,872]" captionTargetBox-1="[151,1436,996,1698]" captionTargetBox-2="[151,1436,193,709]" captionTargetId-0="figure@7.[151,1436,193,872]" captionTargetId-1="figure@7.[151,1436,996,1698]" captionTargetId-2="figure@8.[151,1436,193,709]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 3. Photograph of the hind wing fragment of Gomphaeschna carinthiae sp. nov., holotype UMJG & P 211472, plate A. The wingtip is covered with sediment." captionText-1="FIGURE 4. Photograph of the hind wing fragment of Gomphaeschna carinthiae sp. nov., holotype UMJG & P 211472, plate B. The posterobasal area of the wing is covered with sediment." captionText-2="FIGURE 5. Drawing of the hind wing fragment of Gomphaeschna carinthiae sp. nov., holotype UMJG & P 211472. The drawing involves features of both plates." httpUri-0="https://zenodo.org/record/398755/files/figure.png" httpUri-1="https://zenodo.org/record/398756/files/figure.png" httpUri-2="https://zenodo.org/record/398757/files/figure.png" pageId="6" pageNumber="159">Figures 3–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136539FFBAFFB5FF423702FB4213B6" box="[151,1177,1644,1669]" pageId="6" pageNumber="159" type="materials_examined">
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<paragraph id="8BB636B2FFBAFFB5FF423702FB4213B6" blockId="6.[151,1437,1644,2029]" box="[151,1177,1644,1669]" pageId="6" pageNumber="159">
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<materialsCitation id="3B613CEFFFBAFFB5FF423702FB4213B6" ID-GBIF-Occurrence="1439650070" box="[151,1177,1644,1669]" collectionCode="UMJG" country="Austria" location="Universalmuseum Joanneum" municipality="Graz" pageId="6" pageNumber="159" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97DEAA0FFBAFFB5FF423702FED013B6" bold="true" box="[151,267,1644,1669]" pageId="6" pageNumber="159">
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<typeStatus id="54B28810FFBAFFB5FF423702FEDD13B6" box="[151,262,1644,1669]" pageId="6" pageNumber="159" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED18AE77FFBAFFB5FEC73702FEBB13B7" box="[274,352,1644,1668]" pageId="6" pageNumber="159">UMJG</collectionCode>
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&P 211472, collection of the
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<location id="8ED66069FFBAFFB5FD7D3702FC3413B7" LSID="urn:lsid:plazi:treatment:03A087A4FFBAFFBAFF42376EFED814C7:8ED66069FFBAFFB5FD7D3702FC3413B7" box="[680,1007,1644,1669]" country="Austria" municipality="Graz" name="Universalmuseum Joanneum" pageId="6" pageNumber="159">Universalmuseum Joanneum</location>
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,
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<collectingMunicipality id="6BD2ACC8FFBAFFB5FC2F3702FBE813B7" box="[1018,1075,1644,1668]" pageId="6" pageNumber="159">Graz</collectingMunicipality>
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,
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<collectingCountry id="F31E7622FFBAFFB5FBEA3702FB4E13B6" box="[1087,1173,1644,1669]" name="Austria" pageId="6" pageNumber="159">Austria</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136539FFBAFFB5FF1237E1FC2513FE" pageId="6" pageNumber="159" type="diagnosis">
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<paragraph id="8BB636B2FFBAFFB5FF1237E1FC2513FE" blockId="6.[151,1437,1644,2029]" pageId="6" pageNumber="159">
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<emphasis id="B97DEAA0FFBAFFB5FF1237E1FE9A139B" bold="true" box="[199,321,1679,1704]" pageId="6" pageNumber="159">Diagnosis.</emphasis>
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The specimen differs from all Recent and fossil gomphaeschnid species in the broad space between Rspl and IR2 and the distinct oblique crossvein that connects these two veins.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136539FFBAFFBBFF1237B9FB761311" lastPageId="8" lastPageNumber="161" pageId="6" pageNumber="159" type="description">
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<paragraph id="8BB636B2FFBAFFB5FF1237B9FCAF126F" blockId="6.[151,1437,1644,2029]" pageId="6" pageNumber="159">
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<emphasis id="B97DEAA0FFBAFFB5FF1237B9FE4513C3" bold="true" box="[199,414,1751,1776]" pageId="6" pageNumber="159">Description Graz.</emphasis>
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The fossil was split during its excavation and is now present in two halves. Both halves show mostly the same but mirrored parts of the venation. Almost the complete wing is preserved and visible. The preservation lacks only the proximal posterior part of the wing. The tip of the wing dips into the sediment of plate A whereas the proximal anterior edge of the wing dips in plate B.
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</paragraph>
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<paragraph id="8BB636B2FFBAFFB5FF123609FA9A12DE" blockId="6.[151,1437,1644,2029]" pageId="6" pageNumber="159">
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Total length
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<quantity id="4CF19B57FFBAFFB5FE823606FE1B124C" box="[343,448,1896,1920]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="6" pageNumber="159" unit="mm" value="28.4">28.4 mm</quantity>
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; total width
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<quantity id="4CF19B57FFBAFFB5FD853609FD71124C" box="[592,682,1895,1920]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="6" pageNumber="159" unit="mm" value="9.6">9.6 mm</quantity>
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. Secondary antenodal crossveins not aligned. Pterostigma roughly parallel sided;
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<quantity id="4CF19B57FFBAFFB5FEEB36E2FE4E1297" box="[318,405,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="159" unit="mm" value="3.1">3.1 mm</quantity>
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in proximodistal dimension and maximum
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<quantity id="4CF19B57FFBAFFB5FC5036E2FC071297" box="[901,988,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="159" unit="mm" value="0.9">0.9 mm</quantity>
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in anteroposterior dimension; proximal and distal margin oblique with an angle of 134° between RA and proximal margin and 135° between posterior margin and distal margin. Pterostigmal brace vein oblique in extension of proximal margin of pterostigma.
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</paragraph>
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<caption id="DF76663AFFBBFFB4FF423213FDC71681" httpUri="https://zenodo.org/record/398755/files/figure.png" pageId="7" pageNumber="160" targetBox="[151,1436,193,872]" targetPageId="7">
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<paragraph id="8BB636B2FFBBFFB4FF423213FDC71681" blockId="7.[151,1436,893,946]" pageId="7" pageNumber="160">
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<emphasis id="B97DEAA0FFBBFFB4FF423213FEC916A0" bold="true" box="[151,274,893,915]" pageId="7" pageNumber="160">FIGURE 3.</emphasis>
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Photograph of the hind wing fragment of
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<taxonomicName id="4C094D31FFBBFFB4FD6E3213FC1D16A0" box="[699,966,893,915]" class="Insecta" family="Aeshnidae" genus="Gomphaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="160" phylum="Arthropoda" rank="species" species="carinthiae" status="sp. nov.">
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<emphasis id="B97DEAA0FFBBFFB4FD6E3213FC1D16A0" box="[699,966,893,915]" italics="true" pageId="7" pageNumber="160">Gomphaeschna carinthiae</emphasis>
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</taxonomicName>
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<emphasis id="B97DEAA0FFBBFFB4FC193210FBC616A0" bold="true" box="[972,1053,894,915]" pageId="7" pageNumber="160">
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<taxonomicNameLabel id="A24E57DBFFBBFFB4FC193210FBC616A0" box="[972,1053,894,915]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype UMJG&P 211472, plate A. The wingtip is covered with sediment.
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</paragraph>
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</caption>
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<caption id="DF76663AFFBBFFB4FF4237D9FD2C13DF" httpUri="https://zenodo.org/record/398756/files/figure.png" pageId="7" pageNumber="160" targetBox="[151,1436,996,1698]" targetPageId="7">
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<paragraph id="8BB636B2FFBBFFB4FF4237D9FD2C13DF" blockId="7.[151,1436,1719,1772]" pageId="7" pageNumber="160">
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<emphasis id="B97DEAA0FFBBFFB4FF4237D9FEC913FF" bold="true" box="[151,274,1719,1741]" pageId="7" pageNumber="160">FIGURE 4.</emphasis>
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Photograph of the hind wing fragment of
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<taxonomicName id="4C094D31FFBBFFB4FD6937D9FC1C13FE" box="[700,967,1719,1741]" class="Insecta" family="Aeshnidae" genus="Gomphaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="160" phylum="Arthropoda" rank="species" species="carinthiae" status="sp. nov.">
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<emphasis id="B97DEAA0FFBBFFB4FD6937D9FC1C13FE" box="[700,967,1719,1741]" italics="true" pageId="7" pageNumber="160">Gomphaeschna carinthiae</emphasis>
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</taxonomicName>
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<emphasis id="B97DEAA0FFBBFFB4FC1837D9FBC513FF" bold="true" box="[973,1054,1719,1740]" pageId="7" pageNumber="160">
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<taxonomicNameLabel id="A24E57DBFFBBFFB4FC1837D9FBC513FF" box="[973,1054,1719,1740]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype UMJG&P 211472, plate B. The posterobasal area of the wing is covered with sediment.
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</paragraph>
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</caption>
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<paragraph id="8BB636B2FFBBFFB4FF12367DFC95121F" blockId="7.[151,1436,1811,2016]" box="[199,846,1811,1836]" pageId="7" pageNumber="160">Preradial and interradial fields both with one row of cells.</paragraph>
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<paragraph id="8BB636B2FFBBFFB4FF123659FE4C12D3" blockId="7.[151,1436,1811,2016]" pageId="7" pageNumber="160">
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Distalmost crossvein in antesubnodal space at level of junction between RP´and IR2. Distance between RP´–RP´´furca and subnodus 4.0 mm. Distinct
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<quantity id="4CF19B57FFBBFFB4FC7C3632FC241240" box="[937,1023,1884,1908]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="160" unit="mm" value="1.3">1.3 mm</quantity>
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long oblique crossvein at distance of
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<quantity id="4CF19B57FFBBFFB4FF4236EEFF3712A4" box="[151,236,1920,1944]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="7" pageNumber="160" unit="mm" value="9.8">9.8 mm</quantity>
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from RP´–RP´´furca between IR2 and Rspl with an angle of 127° to proximal side of IR2. The part of Rspl proximal to the distinct IR2-Rspl crossvein is not preserved. At least five straight veins running from Rspl towards posterior wing margin.
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</paragraph>
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<caption id="DF76663AFFB4FFBBFF4233B4FDEF163D" httpUri="https://zenodo.org/record/398757/files/figure.png" pageId="8" pageNumber="161" targetBox="[151,1436,193,709]" targetPageId="8">
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||
<paragraph id="8BB636B2FFB4FFBBFF4233B4FDEF163D" blockId="8.[151,1435,730,782]" pageId="8" pageNumber="161">
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<emphasis id="B97DEAA0FFB4FFBBFF4233B4FECD17DC" bold="true" box="[151,278,730,752]" pageId="8" pageNumber="161">FIGURE 5.</emphasis>
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Drawing of the hind wing fragment of
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<taxonomicName id="4C094D31FFB4FFBBFD1033B4FC0E17C3" box="[709,981,730,752]" class="Insecta" family="Aeshnidae" genus="Gomphaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="161" phylum="Arthropoda" rank="species" species="carinthiae" status="sp. nov.">
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||
<emphasis id="B97DEAA0FFB4FFBBFD1033B4FC0E17C3" box="[709,981,730,752]" italics="true" pageId="8" pageNumber="161">Gomphaeschna carinthiae</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B97DEAA0FFB4FFBBFC3533B4FBEF17DC" bold="true" box="[992,1076,730,751]" pageId="8" pageNumber="161">
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||
<taxonomicNameLabel id="A24E57DBFFB4FFBBFC3533B4FBEF17DC" box="[992,1076,730,751]" pageId="8" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype UMJG&P 211472. The drawing involves features of both plates.
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</paragraph>
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</caption>
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<paragraph id="8BB636B2FFB4FFBBFF123252FEE516A8" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
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RP2 slightly undulating with least distance to RP1 at level of the pterostigmal brace vein. One row of cells between RP1 and RP
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<quantity id="4CF19B57FFB4FFBBFE52320EFE68164B" box="[391,435,863,888]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="161" unit="in" value="2.0">2 in</quantity>
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proximal part; two rows of cells from
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<quantity id="4CF19B57FFB4FFBBFCBE3231FC0B1644" box="[875,976,863,888]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="161" unit="mm" value="13.5">13.5 mm</quantity>
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distance to RP´-RP´´furca to distalmost preserved part.
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</paragraph>
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<paragraph id="8BB636B2FFB4FFBBFF1232C9FE1116D0" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">Premedial field with one row of cells and with parallel boundaries; posterodistal part of premedial field not distinctly broader than rest.</paragraph>
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<paragraph id="8BB636B2FFB4FFBBFF12329EFC85113B" blockId="8.[151,1436,827,2031]" box="[199,862,1007,1032]" pageId="8" pageNumber="161">At least four antefurcal crossveins between RP - and MA+.</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF12357AFEF4117C" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
Htc at least
|
||
<quantity id="4CF19B57FFB4FFBBFE9B357AFE7F1118" box="[334,420,1044,1068]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="161" unit="mm" value="3.8">3.8 mm</quantity>
|
||
long and
|
||
<quantity id="4CF19B57FFB4FFBBFDC0357AFDB71118" box="[533,620,1044,1068]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="161" unit="mm" value="0.4">0.4 mm</quantity>
|
||
wide at half length of tcv; MA almost parallel to tcv and only curved in its distal part.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF123535FC8E118F" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
Discoidal triangle longitudinally stretched and divided by at least one secondary crossvein; basal side
|
||
<quantity id="4CF19B57FFB4FFBBFAEA3532FA431140" box="[1343,1432,1116,1140]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="161" unit="mm" value="1.7">1.7 mm</quantity>
|
||
; costal side
|
||
<quantity id="4CF19B57FFB4FFBBFECC35EEFEAE11A4" box="[281,373,1152,1176]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="161" unit="mm" value="3.3">3.3 mm</quantity>
|
||
; posterodistal side
|
||
<quantity id="4CF19B57FFB4FFBBFD8635EEFD7411A4" box="[595,687,1152,1176]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="161" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; posterodistal side (ddv) almost straight with a slight kink at the origin of the distinct Tspl. Tspl meets MP almost orthogonally.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF1235A9FCA41037" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">Three rows of cells in basal part of postdiscoidal space. One intercalated vein in medial field. At least one intercalated vein or secondary branch of CuA´in antecubital field.</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF123461FE281043" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
Preanal field with 3 cells on anterior side; anterior side width
|
||
<quantity id="4CF19B57FFB4FFBBFCA8347EFC0D1014" box="[893,982,1296,1320]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="161" unit="mm" value="3.1">3.1 mm</quantity>
|
||
. Anal field
|
||
<quantity id="4CF19B57FFB4FFBBFB8F347EFB6B1014" box="[1114,1200,1296,1320]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="161" unit="mm" value="1.2">1.2 mm</quantity>
|
||
wide and unicellular on anterior side, narrowing towards posterior wing margin. (AA´´& AP) concave on proximal side and approaching AA2b posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF123412FEC210E8" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
CuP crossing
|
||
<quantity id="4CF19B57FFB4FFBBFEBE3412FE1E10A0" box="[363,453,1404,1428]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="161" unit="mm" value="0.7">0.7 mm</quantity>
|
||
long and
|
||
<quantity id="4CF19B57FFB4FFBBFD953412FD4110A0" box="[576,666,1404,1428]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="161" unit="mm" value="1.4">1.4 mm</quantity>
|
||
proximal to anteroproximal angle of triangulum. Anterior side of subdiscoidal cell
|
||
<quantity id="4CF19B57FFB4FFBBFE8C34F1FE6B1084" box="[345,432,1439,1463]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="161" unit="mm" value="1.4">1.4 mm</quantity>
|
||
long. PsA joins proximal side of triangulum
|
||
<quantity id="4CF19B57FFB4FFBBFC7F34F1FBDA1084" box="[938,1025,1439,1463]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="161" unit="mm" value="0.2">0.2 mm</quantity>
|
||
posterior to anteroproximal angle of triangulum.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF12348BFB761311" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
Subtriangular cell with posterior side
|
||
<quantity id="4CF19B57FFB4FFBBFDA7348BFD1710CD" box="[626,716,1509,1534]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="161" unit="mm" value="1.6">1.6 mm</quantity>
|
||
; anterior side (PsA) 1.0 mm; distal side
|
||
<quantity id="4CF19B57FFB4FFBBFB48348BFB2C10CD" box="[1181,1271,1509,1534]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="161" unit="mm" value="1.5">1.5 mm</quantity>
|
||
. Posterior side of subtriangulum not joining posterior angle of triangulum; sdv distinct but short (>
|
||
<quantity id="4CF19B57FFB4FFBBFB923764FB7B1311" box="[1095,1184,1546,1570]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="161" unit="mm" value="0.1">0.1 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFB4FFBBFF123743FE9F13BF" pageId="8" pageNumber="161" type="materials_examined">
|
||
<paragraph id="8BB636B2FFB4FFBBFF123743FE9F13BF" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
<materialsCitation id="3B613CEFFFB4FFBBFF123743FE9F13BF" ID-GBIF-Occurrence="1439650069" country="Austria" county="Laminated" latitude="46.798145" location="St. Andra" longLatPrecision="1" longitude="14.806603" municipality="Wolfsberg" pageId="8" pageNumber="161" specimenCount="1" stateProvince="Carinthia" typeStatus="holotype">
|
||
<emphasis id="B97DEAA0FFB4FFBBFF123743FDC31375" bold="true" box="[199,536,1581,1606]" pageId="8" pageNumber="161">
|
||
<typeStatus id="54B28810FFB4FFBBFF123743FED91375" box="[199,258,1581,1606]" pageId="8" pageNumber="161">Type</typeStatus>
|
||
locality and horizon.
|
||
</emphasis>
|
||
Claypit Schaßbach (
|
||
<geoCoordinate id="EE3D5075FFB4FFBBFCCE3743FC1D1375" box="[795,966,1581,1606]" direction="north" orientation="latitude" pageId="8" pageNumber="161" precision="1" value="46.798145">46°47'53.32"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3D5075FFB4FFBBFC0F3743FB591376" box="[986,1154,1581,1605]" direction="east" orientation="longitude" pageId="8" pageNumber="161" precision="1" value="14.806603">14°48'23.77"E</geoCoordinate>
|
||
),
|
||
<location id="8ED66069FFB4FFBBFB4B3743FACE1375" LSID="urn:lsid:plazi:treatment:03A087A4FFBAFFBAFF42376EFED814C7:8ED66069FFB4FFBBFB4B3743FACE1375" box="[1182,1301,1581,1606]" country="Austria" county="Laminated" latitude="46.798145" longLatPrecision="1" longitude="14.806603" municipality="Wolfsberg" name="St. Andra" pageId="8" pageNumber="161" stateProvince="Carinthia">St. Andrä</location>
|
||
,
|
||
<collectingMunicipality id="6BD2ACC8FFB4FFBBFAFD3743FA471375" box="[1320,1436,1581,1606]" pageId="8" pageNumber="161">Wolfsberg</collectingMunicipality>
|
||
(district),
|
||
<collectingRegion id="49CDF850FFB4FFBBFED3373EFEAA135A" box="[262,369,1616,1641]" country="Austria" name="Karnten" pageId="8" pageNumber="161">Carinthia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31E7622FFB4FFBBFEA8373EFE08135A" box="[381,467,1616,1641]" name="Austria" pageId="8" pageNumber="161">Austria</collectingCountry>
|
||
.
|
||
<collectingCounty id="62D74E3EFFB4FFBBFE0A373EFD8C135A" box="[479,599,1616,1641]" pageId="8" pageNumber="161">Laminated</collectingCounty>
|
||
siltstone of the lacustrine lower part of the Mühldorf Formation. Age late Early Miocene.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFB4FFBBFF1237F9FE40120D" pageId="8" pageNumber="161" type="etymology">
|
||
<paragraph id="8BB636B2FFB4FFBBFF1237F9FA4E1383" blockId="8.[151,1436,827,2031]" box="[199,1429,1687,1712]" pageId="8" pageNumber="161">
|
||
<emphasis id="B97DEAA0FFB4FFBBFF1237F9FE951383" bold="true" box="[199,334,1687,1712]" pageId="8" pageNumber="161">Etymology.</emphasis>
|
||
The species name refers to the state of
|
||
<collectingRegion id="49CDF850FFB4FFBBFCD237F9FCB41383" box="[775,879,1687,1712]" country="Austria" name="Karnten" pageId="8" pageNumber="161">Carinthia</collectingRegion>
|
||
(
|
||
<collectingRegion id="49CDF850FFB4FFBBFCAB37F9FC07139C" box="[894,988,1687,1711]" country="Austria" name="Karnten" pageId="8" pageNumber="161">Kärnten</collectingRegion>
|
||
) in
|
||
<collectingCountry id="F31E7622FFB4FFBBFBD237F9FB811383" box="[1031,1114,1687,1712]" name="Austria" pageId="8" pageNumber="161">Austria</collectingCountry>
|
||
where the fossil was found.
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF1237D4FE40120D" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
<emphasis id="B97DEAA0FFB4FFBBFF1237D4FE7213E0" bold="true" box="[199,425,1722,1747]" pageId="8" pageNumber="161">Sex determination.</emphasis>
|
||
The
|
||
<typeStatus id="54B28810FFB4FFBBFE3737D4FD9F13E0" box="[482,580,1722,1747]" pageId="8" pageNumber="161" type="holotype">holotype</typeStatus>
|
||
is presumably the hind wing of a male specimen. The anal field is relatively narrow and unicellular in the basal part. Also (AA´´& AP) is concave on its proximal side and approaching the straight AA2b posteriorly. This could indicate an anal angle characteristic for male anisopterans that is not preserved in this fossil.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFB4FFBAFF123626FBF415E7" lastPageId="9" lastPageNumber="162" pageId="8" pageNumber="161" type="description">
|
||
<paragraph id="8BB636B2FFB4FFBBFF123626FE18129B" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
<emphasis id="B97DEAA0FFB4FFBBFF123626FE0D1252" bold="true" box="[199,470,1864,1889]" pageId="8" pageNumber="161">Phylogenetic affinities.</emphasis>
|
||
The specimen can be clearly attributed to the suborder
|
||
<taxonomicName id="4C094D31FFB4FFBBFB913626FA971252" authority="Selys, 1854" authorityName="Selys" authorityYear="1854" box="[1092,1356,1864,1889]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Anisoptera" kingdom="Animalia" order="Malvales" pageId="8" pageNumber="161" phylum="Tracheophyta" rank="genus">Anisoptera Selys, 1854</taxonomicName>
|
||
, by the strong expansion of the cubito-anal field and the presence of a discoidal triangle as well as a hypertriangle and a subtriangle (
|
||
<bibRefCitation id="EF984B43FFB4FFBBFEF736E1FE6C129B" author="Bechly" box="[290,439,1935,1960]" pageId="8" pageNumber="161" refString="Bechly, G. (2007) Chapter 11.5 Odonata: damselflies and dragonflies. In: Martill, D. M., Bechly, G. & Loveridge, R. F. (Eds.), The Crato fossil beds of Brazil: window into an ancient world. Cambridge University Press, Cambridge, pp. 184 - 222." type="book chapter" year="2007">Bechly 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB4FFBBFF1236DDFE0A12DC" blockId="8.[151,1436,827,2031]" pageId="8" pageNumber="161">
|
||
Aeshnoptera
|
||
<bibRefCitation id="EF984B43FFB4FFBBFE8836DDFE2E12FF" author="Bechly" box="[349,501,1971,1996]" pageId="8" pageNumber="161" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der * Odonata. Petalura, special Volume 2, 1 - 402." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
: space between RP1 and RP2 narrowed with only one row of cells to pterostigma; Rspl present (
|
||
<bibRefCitation id="EF984B43FFB4FFBBFEE436B8FE1D12DC" author="Bechly" box="[305,454,2006,2031]" pageId="8" pageNumber="161" refString="Bechly, G. (2003) Phylogenetic systematics of Odonata. In: Schorr, M. & Lindeboom, M. (Eds.), Dragonfly Research, Folume 1. Zerf-Tubingen (CD-ROM)." type="book chapter" year="2003">Bechly 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB5FFBAFF1231F9FC0E1583" blockId="9.[151,1437,151,500]" box="[199,981,151,176]" pageId="9" pageNumber="162">
|
||
Aeshnomorpha
|
||
<bibRefCitation id="EF984B43FFB5FFBAFEAC31F9FDCA1583" author="Bechly" box="[377,529,151,176]" pageId="9" pageNumber="162" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">Bechly, 2001</bibRefCitation>
|
||
: RP2 slightly undulating (
|
||
<bibRefCitation id="EF984B43FFB5FFBAFCE131F9FC121583" author="Bechly" box="[820,969,151,176]" pageId="9" pageNumber="162" refString="Bechly, G. (2003) Phylogenetic systematics of Odonata. In: Schorr, M. & Lindeboom, M. (Eds.), Dragonfly Research, Folume 1. Zerf-Tubingen (CD-ROM)." type="book chapter" year="2003">Bechly 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB5FFBAFF1231D2FBF415E7" blockId="9.[151,1437,151,500]" box="[199,1071,188,213]" pageId="9" pageNumber="162">
|
||
Euaeshnida
|
||
<bibRefCitation id="EF984B43FFB5FFBAFE9A31D2FE3D15E6" author="Bechly" box="[335,486,188,213]" pageId="9" pageNumber="162" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der * Odonata. Petalura, special Volume 2, 1 - 402." type="journal article" year="1996">Bechly, 1996</bibRefCitation>
|
||
: RP2 and IR2 distinctly non-parallel (
|
||
<bibRefCitation id="EF984B43FFB5FFBAFC5B31D2FBF815E7" author="Bechly" box="[910,1059,188,213]" pageId="9" pageNumber="162" refString="Bechly, G. (2003) Phylogenetic systematics of Odonata. In: Schorr, M. & Lindeboom, M. (Eds.), Dragonfly Research, Folume 1. Zerf-Tubingen (CD-ROM)." type="book chapter" year="2003">Bechly 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFB5FFBAFF1231B1FD1F149E" pageId="9" pageNumber="162" type="reference_group">
|
||
<paragraph id="8BB636B2FFB5FFBAFF1231B1FB871473" blockId="9.[151,1437,151,500]" pageId="9" pageNumber="162">
|
||
<taxonomicName id="4C094D31FFB5FFBAFF1231B1FD6015CB" authority="Tillyard & Fraser, 1940" authorityName="Tillyard & Fraser" authorityYear="1940" box="[199,699,223,248]" class="Insecta" family="Gomphaeschnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="162" phylum="Arthropoda" rank="family">Gomphaeschnidae Tillyard & Fraser, 1940</taxonomicName>
|
||
: ‘cordulegastrid gap’ (distalmost part of the antesubnodal area without crossveins); no accessory crossveins between CuP-crossing and PsA in subdiscoidal cell; triangulum divided by one crossvein; hypertriangle unicellular (
|
||
<bibRefCitation id="EF984B43FFB5FFBAFD0A3049FC6A1473" author="Bechly" box="[735,945,295,320]" pageId="9" pageNumber="162" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B97DEAA0FFB5FFBAFCE23047FCAB1473" box="[823,880,296,320]" italics="true" pageId="9" pageNumber="162">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF984B43FFB5FFBAFC683049FB8A1473" author="Bechly" box="[957,1105,295,320]" pageId="9" pageNumber="162" refString="Bechly, G. (2003) Phylogenetic systematics of Odonata. In: Schorr, M. & Lindeboom, M. (Eds.), Dragonfly Research, Folume 1. Zerf-Tubingen (CD-ROM)." type="book chapter" year="2003">Bechly 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB636B2FFB5FFBAFF123022FD1F149E" blockId="9.[151,1437,151,500]" pageId="9" pageNumber="162">
|
||
<taxonomicName id="4C094D31FFB5FFBAFF123022FDD91457" ID-CoL="4PVG" authority="Selys, 1871" authorityName="Selys" authorityYear="1871" box="[199,514,332,357]" class="Insecta" family="Aeshnidae" genus="Gomphaeschna" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="162" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97DEAA0FFB5FFBAFF123022FEAE1457" box="[199,373,332,356]" italics="true" pageId="9" pageNumber="162">Gomphaeschna</emphasis>
|
||
Selys, 1871
|
||
</taxonomicName>
|
||
: IR2 and Rspl more or less parallel; RP2 undulating; hind wing triangle with basal side longer than 1/2 costal side; cubito-anal space with 1 crossvein (CuP-crossing); Rspl with no distinct connection to wing margin (
|
||
<bibRefCitation id="EF984B43FFB5FFBAFE0730FAFD62149E" author="Garrison" box="[466,697,404,429]" pageId="9" pageNumber="162" refString="Garrison, R. W., von Ellenrieder, N. & Louton, J. A. (2006) Dragonfly genera of the New World: an illustrated and annotated key to the Anisoptera. The Johns Hopkins University Press, Baltimore, 368 pp." type="book" year="2006">
|
||
Garrison
|
||
<emphasis id="B97DEAA0FFB5FFBAFDE930FBFDAE149F" box="[572,629,404,428]" italics="true" pageId="9" pageNumber="162">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3136539FFB5FFBAFF1230D6FED814C7" pageId="9" pageNumber="162" type="description">
|
||
<paragraph id="8BB636B2FFB5FFBAFF1230D6FED814C7" blockId="9.[151,1437,151,500]" pageId="9" pageNumber="162">
|
||
No elongated paranal cell basal to anal loop in hind wing as in
|
||
<taxonomicName id="4C094D31FFB5FFBAFC4030D9FAC414E3" authority="Bechly, 2001" authorityName="Bechly" authorityYear="2001" box="[917,1311,439,464]" class="Insecta" family="Gomphaeschnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="162" phylum="Arthropoda" rank="tribe" tribe="Gomphaeschnaoidini">
|
||
Gomphaeschnaoidini
|
||
<bibRefCitation id="EF984B43FFB5FFBAFB5E30D9FAC414E3" author="Bechly" box="[1163,1311,439,464]" pageId="9" pageNumber="162" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">Bechly, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF984B43FFB5FFBAFAFA30D9FF2C14C7" author="Bechly" pageId="9" pageNumber="162" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 1 - 219." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B97DEAA0FFB5FFBAFA5D30D7FF6F14C7" italics="true" pageId="9" pageNumber="162">et al.</emphasis>
|
||
2001
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |