206 lines
29 KiB
XML
206 lines
29 KiB
XML
<document id="E79E1DCA28771E011EF7E4D349A61FF0" ID-CLB-Dataset="9183" ID-DOI="10.11646/zootaxa.4820.1.2" ID-GBIF-Dataset="8f361ffe-9e56-41f8-ab77-fae3b124798d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4397238" ID-ZooBank="ED703EC8-3124-413F-8B17-3C1695B789C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609205927210" checkinUser="plazi" docAuthor="Numberger-Thuy, Lea D. & Thuy, Ben" docDate="2020" docId="039FFC4D5958FFC1C3C6C3B9794A73D1" docLanguage="en" docName="zootaxa.4820.1.2.pdf" docOrigin="Zootaxa 4820 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophiacantha oceani Numberger-Thuy & Thuy 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="24" masterDocId="FFA68435595BFFC4C351C32E7A0D7304" masterDocTitle="A new bathyal ophiacanthid brittle star (Ophiuroidea: Ophiacanthidae) with Caribbean affinities from the Plio-Pleistocene of the Mediterranean" masterLastPageNumber="30" masterPageNumber="19" pageNumber="22" updateTime="1698848199796" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="100F52A86F3D90B8C5D0A4509C1A37F1">
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<mods:title id="8049D158DB2FC2198CCFDFAEB78E1ACB">A new bathyal ophiacanthid brittle star (Ophiuroidea: Ophiacanthidae) with Caribbean affinities from the Plio-Pleistocene of the Mediterranean</mods:title>
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<mods:namePart id="3BC6AC0FFED29E16D55F3E849BC69A83">Numberger-Thuy, Lea D.</mods:namePart>
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<mods:roleTerm id="7DFFD33FA7FF088F6A2BE334F0230F3D">Author</mods:roleTerm>
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<mods:namePart id="8EED2A5391E28ECBFFCA0BB34C23BF82">Thuy, Ben</mods:namePart>
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<mods:typeOfResource id="29DD8553CD1387DC47BC4C85C377817C">text</mods:typeOfResource>
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<mods:title id="5E841A22CB07A0AE4F0072F5353C9E09">Zootaxa</mods:title>
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<mods:part id="851BE9B70C013EDB67A61E7FFDF823B3">
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<mods:date id="0F2EA942374848532D7050B64E9C60D9">2020</mods:date>
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<mods:number id="E88E886E573FC8C452020E6A13231F45">2020-07-27</mods:number>
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<mods:number id="39306C5449E6EAC3FC50338F34660A45">4820</mods:number>
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<mods:number id="7F94FB9F0F34F7F2C9CA073E625AB7BE">1</mods:number>
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<mods:identifier id="D1CEDA9AB10222C511742CEEC45C9095" type="CLB-Dataset">9183</mods:identifier>
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<mods:identifier id="D37E92E8C5A53E4B3CA11CFECBADC0DE" type="DOI">10.11646/zootaxa.4820.1.2</mods:identifier>
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<mods:identifier id="84E6CE9F99A4673F8527004151AC6483" type="GBIF-Dataset">8f361ffe-9e56-41f8-ab77-fae3b124798d</mods:identifier>
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<treatment id="039FFC4D5958FFC1C3C6C3B9794A73D1" ID-DOI="http://doi.org/10.5281/zenodo.4397217" ID-GBIF-Taxon="176342298" ID-Zenodo-Dep="4397217" LSID="urn:lsid:plazi:treatment:039FFC4D5958FFC1C3C6C3B9794A73D1" httpUri="http://treatment.plazi.org/id/039FFC4D5958FFC1C3C6C3B9794A73D1" lastPageId="5" lastPageNumber="24" pageId="3" pageNumber="22">
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<subSubSection id="C32C1ED05958FFC7C3C6C3B97BFC73B5" box="[151,497,151,178]" pageId="3" pageNumber="22" type="nomenclature">
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<paragraph id="8B894D5B5958FFC7C3C6C3B97BFC73B5" blockId="3.[151,961,151,250]" box="[151,497,151,178]" pageId="3" pageNumber="22">
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<heading id="D0C1FA375958FFC7C3C6C3B97BFC73B5" bold="true" box="[151,497,151,178]" fontSize="11" level="1" pageId="3" pageNumber="22" reason="1">
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<emphasis id="B94291495958FFC7C3C6C3B97BFC73B5" bold="true" box="[151,497,151,178]" pageId="3" pageNumber="22">
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<taxonomicName id="4C3636D85958FFC7C3C6C3B97B8073B5" authority="Numberger-Thuy & Thuy, 2020" authorityName="Numberger-Thuy & Thuy" authorityYear="2020" box="[151,397,151,178]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="22" phylum="Echinodermata" rank="species" species="oceani" status="sp. nov.">
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<emphasis id="B94291495958FFC7C3C6C3B97B8073B5" bold="true" box="[151,397,151,178]" italics="true" pageId="3" pageNumber="22">Ophiacantha oceani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2712C325958FFC7C2C5C3B97BFC73B5" box="[404,497,151,177]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C1ED05958FFC7C3C6C39579CC73FE" pageId="3" pageNumber="22" type="description">
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<paragraph id="8B894D5B5958FFC7C3C6C3957AF873D1" blockId="3.[151,961,151,250]" box="[151,245,187,213]" pageId="3" pageNumber="22">
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<figureCitation id="130D51DE5958FFC7C3C6C3957AF873D1" box="[151,245,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1534,1558]" captionTargetBox="[164,1427,181,1510]" captionTargetId="figure-25@4.[151,1436,181,1510]" captionTargetPageId="4" captionText="FIGURE 2. Ophiacantha oceani sp. nov. (A–F) from the Piacenzian to Gelasian, latest Pliocene to earliest Pleistocene (approximately 2.6 mya) of Punta Mazza, Capo Milazzo, NE Sicily, Italy, A–B: MnhnL OPH074, holotype, proximal lateral arm plate in external (A) and internal (B) views; C–D: MnhnL OPH075, paratype, proximal lateral arm plate in external (C) and internal (D) views; E–F: MnhnL OPH076, paratype, distal lateral arm plate in external (E) and internal (F) views. Recent Ophiacantha stellata (G–J) from the tropical NW-Atlantic as closest living relative of Ophiacantha oceani sp. nov., G: complete skeleton in ventral view; H: detail of arm in lateral view; I–J: lateral arm plate in external (I) and internal (J) views. Abbreviations: ckv: central knob of the vertebral articular structures; di: distal; do: dorsal; pr: proximal; prv: proximal ridge of the vertebral articular structures; sa: spine articulation; sp: spur; ve: ventral." figureDoi="http://doi.org/10.5281/zenodo.4397242" httpUri="https://zenodo.org/record/4397242/files/figure.png" pageId="3" pageNumber="22">Figure 2</figureCitation>
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</paragraph>
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<paragraph id="8B894D5B5958FFC7C3C6C3F179CC73FE" blockId="3.[151,961,151,250]" box="[151,961,223,250]" pageId="3" pageNumber="22">
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<uri id="FFA741595958FFC7C3C6C3F179CC73FE" box="[151,961,223,250]" pageId="3" pageNumber="22">
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urn:lsid:zoobank.org:act:
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<uuid id="FF90778E5958FFC7C2FDC3F179CC73FE" box="[428,961,223,250]" pageId="3" pageNumber="22">F80B0B8E-7E60-4666-B682-E59E0B50A7A6</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C1ED05958FFC7C3C6C209785F72D5" pageId="3" pageNumber="22" type="etymology">
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<paragraph id="8B894D5B5958FFC7C3C6C209785F72D5" blockId="3.[151,1437,295,2013]" pageId="3" pageNumber="22">
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<emphasis id="B94291495958FFC7C3C6C2097B137245" bold="true" box="[151,286,295,321]" pageId="3" pageNumber="22">Etymology.</emphasis>
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Species named in honour of progressive metal band ‘The Ocean’. Musicians who so skillfully combine arts and science, composing albums like ‘Precambrian’ (with songs named after the periods of the Precambrian), ‘Pelagial’ (with songs named after the bathymetric subdivisions of the water column) and ‘Phanerozoic’ as well as the song ‘Turritopsis dohrnii’ referring to the immortal jellyfish from the Mediterranean, are more than deserving of being immortalized in the fossil record.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C1ED05958FFC7C396C2F57E0D7139" pageId="3" pageNumber="22" type="materials_examined">
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<paragraph id="8B894D5B5958FFC7C396C2F57F8172F1" blockId="3.[151,1437,295,2013]" box="[199,1420,475,502]" pageId="3" pageNumber="22">
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<materialsCitation id="3B5E47065958FFC7C396C2F57F8172F1" ID-GBIF-Occurrence="3005587302" box="[199,1420,475,502]" country="Luxembourg" location="Musee" pageId="3" pageNumber="22" specimenCount="1" typeStatus="holotype">
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<emphasis id="B94291495958FFC7C396C2F57B3672F1" bold="true" box="[199,315,475,501]" pageId="3" pageNumber="22">
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<typeStatus id="548DF3F95958FFC7C396C2F57B3B72F1" box="[199,310,475,501]" pageId="3" pageNumber="22" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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MnhnL (
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<location id="8EE91B805958FFC7C2F5C2F57BFD72F1" LSID="urn:lsid:plazi:treatment:039FFC4D5958FFC1C3C6C3B9794A73D1:8EE91B805958FFC7C2F5C2F57BFD72F1" box="[420,496,475,501]" country="Luxembourg" name="Musee" pageId="3" pageNumber="22">Musée</location>
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national d’histoire naturelle,
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<collectingCountry id="F3210DCB5958FFC7C06AC2F579C072F2" box="[827,973,475,502]" name="Luxembourg" pageId="3" pageNumber="22">Luxembourg</collectingCountry>
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) OPH074, dissociated lateral arm plate.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B894D5B5958FFC7C396C2D17E21711D" blockId="3.[151,1437,295,2013]" box="[199,1068,511,538]" pageId="3" pageNumber="22">
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<emphasis id="B94291495958FFC7C396C2D17B4B711D" bold="true" box="[199,326,511,537]" pageId="3" pageNumber="22">
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<typeStatus id="548DF3F95958FFC7C396C2D17B4C711D" box="[199,321,511,537]" pageId="3" pageNumber="22" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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MnhnL OPH075 and OPH076, two dissociated lateral arm plates.
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</paragraph>
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<paragraph id="8B894D5B5958FFC7C396C10D7E0D7139" blockId="3.[151,1437,295,2013]" box="[199,1024,547,574]" pageId="3" pageNumber="22">
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<emphasis id="B94291495958FFC7C396C10D7BB77139" bold="true" box="[199,442,547,573]" pageId="3" pageNumber="22">Additional material.</emphasis>
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MnhnL OPH077, ten dissociated lateral arm plates.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C1ED05958FFC7C396C16978227086" pageId="3" pageNumber="22" type="diagnosis">
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<paragraph id="8B894D5B5958FFC7C396C1697E0E7011" blockId="3.[151,1437,295,2013]" pageId="3" pageNumber="22">
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<emphasis id="B94291495958FFC7C396C1697B4F7165" bold="true" box="[199,322,583,609]" pageId="3" pageNumber="22">Diagnosis.</emphasis>
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Species of
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<taxonomicName id="4C3636D85958FFC7C29CC16978527165" authorityName="Muller & Troschel" authorityYear="1842" box="[461,607,583,609]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="22" phylum="Echinodermata" rank="genus">
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<emphasis id="B94291495958FFC7C29CC16978527165" box="[461,607,583,609]" italics="true" pageId="3" pageNumber="22">Ophiacantha</emphasis>
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</taxonomicName>
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with lateral arm plates showing a very strong constriction; weak vertical striation close to the row of spine articulations; large, poorly defined, proximalwards protruding and slightly prominent spur on the outer proximal edge; up to seven dorso-ventrally compressed spine articulations composed of massive dorsal and ventral ridges fully separated at their proximal tips; inner side of lateral arm plates with vertebral articular structures composed of a well-defined prominent triangular central knob and a similarly well-defined oblique, slightly arched proximal ridge with a ventralward-pointing dorsal tip.
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</paragraph>
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<paragraph id="8B894D5B5958FFC7C396C0317ED5703E" blockId="3.[151,1437,295,2013]" box="[199,1240,799,826]" pageId="3" pageNumber="22">
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<emphasis id="B94291495958FFC7C396C0317B7F703D" bold="true" box="[199,370,799,825]" pageId="3" pageNumber="22">Locus typicus.</emphasis>
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<materialsCitation id="3B5E47065958FFC7C228C00E7ED9703E" ID-GBIF-Occurrence="3005587301" box="[377,1236,799,826]" collectorName="Capo Milazzo, NE" country="Italy" latitude="38.270695" location="Punta Mazza" longLatPrecision="1" longitude="15.239373" pageId="3" pageNumber="22" specimenCount="1" stateProvince="Sicilia">
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<location id="8EE91B805958FFC7C228C00E7800703E" LSID="urn:lsid:plazi:treatment:039FFC4D5958FFC1C3C6C3B9794A73D1:8EE91B805958FFC7C228C00E7800703E" box="[377,525,800,826]" country="Italy" latitude="38.270695" longLatPrecision="1" longitude="15.239373" name="Punta Mazza" pageId="3" pageNumber="22" stateProvince="Sicilia">Punta Mazza</location>
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,
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<collectorName id="26C3288D5958FFC7C148C03178E7703E" box="[537,746,799,826]" pageId="3" pageNumber="22">Capo Milazzo, NE</collectorName>
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<collectingRegion id="49F283B95958FFC7C1A0C031793E703D" box="[753,819,799,825]" country="Italy" name="Sicilia" pageId="3" pageNumber="22">Sicily</collectingRegion>
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,
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<collectingCountry id="F3210DCB5958FFC7C06FC031797C703D" box="[830,881,799,825]" name="Italy" pageId="3" pageNumber="22">Italy</collectingCountry>
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(
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<geoCoordinate id="EE022B9C5958FFC7C0D1C0317E2F703E" box="[896,1058,799,826]" degrees="38.270694" direction="north" orientation="latitude" pageId="3" pageNumber="22" precision="1" value="38.270695">38.270694° N</geoCoordinate>
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,
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<geoCoordinate id="EE022B9C5958FFC7C77CC0317EC1703E" box="[1069,1228,799,826]" degrees="15.239373" direction="east" orientation="longitude" pageId="3" pageNumber="22" precision="1" value="15.239373">15.239373° E</geoCoordinate>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B894D5B5958FFC7C396C06D78227086" blockId="3.[151,1437,295,2013]" pageId="3" pageNumber="22">
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<emphasis id="B94291495958FFC7C396C06D7B937059" bold="true" box="[199,414,835,861]" pageId="3" pageNumber="22">Stratum typicum.</emphasis>
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Yellowish marls of the MPL5–MPL6, Piacenzian to Gelasian, latest Pliocene to earliest Pleistocene, approximately 2.6 mya.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C1ED05958FFC1C396C0A5794A73D1" lastPageId="5" lastPageNumber="24" pageId="3" pageNumber="22" type="description">
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<paragraph id="8B894D5B5958FFC7C396C0A57F8D7699" blockId="3.[151,1437,295,2013]" pageId="3" pageNumber="22">
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<emphasis id="B94291495958FFC7C396C0A5781470A1" bold="true" box="[199,537,907,933]" pageId="3" pageNumber="22">
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Description of the
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<typeStatus id="548DF3F95958FFC7C2FAC0A5781970A1" box="[427,532,907,933]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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MnhnL OPH074 (
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<figureCitation id="130D51DE5958FFC7C1A3C0A5793A70A1" box="[754,823,907,933]" captionStart="FIGURE 2" captionStartId="4.[151,250,1534,1558]" captionTargetBox="[164,1427,181,1510]" captionTargetId="figure-25@4.[151,1436,181,1510]" captionTargetPageId="4" captionText="FIGURE 2. Ophiacantha oceani sp. nov. (A–F) from the Piacenzian to Gelasian, latest Pliocene to earliest Pleistocene (approximately 2.6 mya) of Punta Mazza, Capo Milazzo, NE Sicily, Italy, A–B: MnhnL OPH074, holotype, proximal lateral arm plate in external (A) and internal (B) views; C–D: MnhnL OPH075, paratype, proximal lateral arm plate in external (C) and internal (D) views; E–F: MnhnL OPH076, paratype, distal lateral arm plate in external (E) and internal (F) views. Recent Ophiacantha stellata (G–J) from the tropical NW-Atlantic as closest living relative of Ophiacantha oceani sp. nov., G: complete skeleton in ventral view; H: detail of arm in lateral view; I–J: lateral arm plate in external (I) and internal (J) views. Abbreviations: ckv: central knob of the vertebral articular structures; di: distal; do: dorsal; pr: proximal; prv: proximal ridge of the vertebral articular structures; sa: spine articulation; sp: spur; ve: ventral." figureDoi="http://doi.org/10.5281/zenodo.4397242" httpUri="https://zenodo.org/record/4397242/files/figure.png" pageId="3" pageNumber="22">Fig. 2</figureCitation>
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A–B) is a dissociated proximal to median lateral arm plate approximately 1.3 times longer than high with deeply concave dorsal edge as a result of an extreme constriction; ventral edge weakly convex; outer proximal edge convex, with a large, smooth, protruding, slightly prominent and poorly defined spur proximally fading into the crest of the outer surface curvature; outer surface with moderately coarsely meshed stereom with very weak vertical striation in a narrow line along the proximal edge of a welldefined, slightly wavy ridge bordering the row of spine articulations; outer surface stereom slightly more densely meshed on crest of curvature in distal continuation of the spur on the outer proximal edge; seven spine articulations on strongly bulging distal portion of lateral arm plate, forming a row from the ventral to the dorsal edge of the lateral arm plate and probably forming a continuous to near-continuous fan of arm spines in the intact individual; size of spine articulations and of gaps separating them increasing towards the dorsal edge of the lateral arm plate; spine articulations dorso-ventrally compressed, consisting of a large, comma-shaped, near-horizontal dorsal lobe composed of compact stereom and a much shorter, straight, horizontal ventral lobe composed of compact to weakly perforate stereom; dorsal and ventral lobes proximally and distally completely separate, with weak sigmoidal fold distally; spine articulations distally bordered by brink composed of slightly more finely meshed stereom than on outer surface, with brink starting at second ventralmost spine articulation, widest at second dorsalmost spine articulation.
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</paragraph>
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<paragraph id="8B894D5B5958FFC7C396C6867F9B7401" blockId="3.[151,1437,295,2013]" pageId="3" pageNumber="22">
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Inner side of lateral arm plate (
|
||
<figureCitation id="130D51DE5958FFC7C177C689787076C5" box="[550,637,1447,1473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1534,1558]" captionTargetBox="[164,1427,181,1510]" captionTargetId="figure-25@4.[151,1436,181,1510]" captionTargetPageId="4" captionText="FIGURE 2. Ophiacantha oceani sp. nov. (A–F) from the Piacenzian to Gelasian, latest Pliocene to earliest Pleistocene (approximately 2.6 mya) of Punta Mazza, Capo Milazzo, NE Sicily, Italy, A–B: MnhnL OPH074, holotype, proximal lateral arm plate in external (A) and internal (B) views; C–D: MnhnL OPH075, paratype, proximal lateral arm plate in external (C) and internal (D) views; E–F: MnhnL OPH076, paratype, distal lateral arm plate in external (E) and internal (F) views. Recent Ophiacantha stellata (G–J) from the tropical NW-Atlantic as closest living relative of Ophiacantha oceani sp. nov., G: complete skeleton in ventral view; H: detail of arm in lateral view; I–J: lateral arm plate in external (I) and internal (J) views. Abbreviations: ckv: central knob of the vertebral articular structures; di: distal; do: dorsal; pr: proximal; prv: proximal ridge of the vertebral articular structures; sa: spine articulation; sp: spur; ve: ventral." figureDoi="http://doi.org/10.5281/zenodo.4397242" httpUri="https://zenodo.org/record/4397242/files/figure.png" pageId="3" pageNumber="22">Fig. 2B</figureCitation>
|
||
) with thick ventral edge showing a large contact surface with opposite lateral arm plate suggesting small ventral arm plates widely separated by lateral arm plates; dorsal edge much thinner but also showing extensive contact surface with opposite lateral arm plate, suggesting small dorsal arm plates restricted to the distal quarter of the arm segment; vertebral articular structures consisting of a well-defined, prominent central knob composed of slightly more finely meshed stereom than remaining inner side of lateral arm plate, proximally bordered and seemingly merged with a much smaller but similarly prominent and well-defined knob; additional vertebral articular structure consisting of a similarly well-defined, boomerang-shaped proximal ridge; weakly developed, narrow vertical ridge in distal third of the lateral arm plate, distally bordered by a very weak vertical furrow without clearly discernible perforations; inner distal edge with a large but poorly defined area of compact to very dense stereom corresponding to spur on outer proximal edge; tentacle notch small and inconspicuous.
|
||
</paragraph>
|
||
<paragraph id="8B894D5B5958FFC7C396C4217F8B7491" blockId="3.[151,1437,295,2013]" pageId="3" pageNumber="22">
|
||
<emphasis id="B94291495958FFC7C396C4217BDC742D" bold="true" box="[199,465,1807,1833]" pageId="3" pageNumber="22">
|
||
<typeStatus id="548DF3F95958FFC7C396C4217B3F742D" box="[199,306,1807,1833]" pageId="3" pageNumber="22" type="paratype">Paratype</typeStatus>
|
||
supplements.
|
||
</emphasis>
|
||
MnhnL OPH075 (
|
||
<figureCitation id="130D51DE5958FFC7C1CCC42178D1742D" box="[669,732,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[151,250,1534,1558]" captionTargetBox="[164,1427,181,1510]" captionTargetId="figure-25@4.[151,1436,181,1510]" captionTargetPageId="4" captionText="FIGURE 2. Ophiacantha oceani sp. nov. (A–F) from the Piacenzian to Gelasian, latest Pliocene to earliest Pleistocene (approximately 2.6 mya) of Punta Mazza, Capo Milazzo, NE Sicily, Italy, A–B: MnhnL OPH074, holotype, proximal lateral arm plate in external (A) and internal (B) views; C–D: MnhnL OPH075, paratype, proximal lateral arm plate in external (C) and internal (D) views; E–F: MnhnL OPH076, paratype, distal lateral arm plate in external (E) and internal (F) views. Recent Ophiacantha stellata (G–J) from the tropical NW-Atlantic as closest living relative of Ophiacantha oceani sp. nov., G: complete skeleton in ventral view; H: detail of arm in lateral view; I–J: lateral arm plate in external (I) and internal (J) views. Abbreviations: ckv: central knob of the vertebral articular structures; di: distal; do: dorsal; pr: proximal; prv: proximal ridge of the vertebral articular structures; sa: spine articulation; sp: spur; ve: ventral." figureDoi="http://doi.org/10.5281/zenodo.4397242" httpUri="https://zenodo.org/record/4397242/files/figure.png" pageId="3" pageNumber="22">Fig. 2</figureCitation>
|
||
C–D) is a dissociated proximal to proximalmost lateral arm plate slightly higher than long; dorso-proximal tip missing, otherwise well preserved, with respect to finer stereom structures even better preserved than
|
||
<typeStatus id="548DF3F95958FFC7C295C47978297475" box="[452,548,1879,1905]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
specimen; outer proximal edge and outer surface ornamentation as in
|
||
<typeStatus id="548DF3F95958FFC7C664C4797F957475" box="[1333,1432,1879,1905]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
; seven spine articulations as in
|
||
<typeStatus id="548DF3F95958FFC7C2B5C45578497491" box="[484,580,1915,1941]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
except for enlarged and distalwards displaced dorsalmost spine articulation.
|
||
</paragraph>
|
||
<paragraph id="8B894D5B5958FFC7C396C48E7B3874D9" blockId="3.[151,1437,295,2013]" pageId="3" pageNumber="22">
|
||
Inner side with vertebral articular structures as in
|
||
<typeStatus id="548DF3F95958FFC7C051C4B1796F74BD" box="[768,866,1951,1977]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
except for more clearly separated large and small central knobs.
|
||
</paragraph>
|
||
<caption id="DF491DD3595FFFC0C3C6C6D078BB7416" ID-DOI="http://doi.org/10.5281/zenodo.4397242" ID-Zenodo-Dep="4397242" httpUri="https://zenodo.org/record/4397242/files/figure.png" pageId="4" pageNumber="23" startId="4.[151,250,1534,1558]" targetBox="[164,1427,181,1510]" targetPageId="4">
|
||
<paragraph id="8B894D5B595FFFC0C3C6C6D078BB7416" blockId="4.[151,1437,1534,1810]" pageId="4" pageNumber="23">
|
||
<emphasis id="B9429149595FFFC0C3C6C6D07B1F7512" bold="true" box="[151,274,1534,1558]" pageId="4" pageNumber="23">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C3636D8595FFFC0C246C6D07BE97512" authority="Numberger-Thuy & Thuy, 2020" authorityName="Numberger-Thuy & Thuy" authorityYear="2020" box="[279,484,1534,1558]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="23" phylum="Echinodermata" rank="species" species="oceani" status="sp. nov.">
|
||
<emphasis id="B9429149595FFFC0C246C6D07BE97512" box="[279,484,1534,1558]" italics="true" pageId="4" pageNumber="23">Ophiacantha oceani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2712C32595FFFC0C2B8C6D1783B7512" box="[489,566,1535,1558]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A–F) from the Piacenzian to Gelasian, latest Pliocene to earliest Pleistocene (approximately 2.6 mya) of Punta Mazza, Capo Milazzo, NE Sicily, Italy, A–B: MnhnL OPH074, holotype, proximal lateral arm plate in external (A) and internal (B) views; C–D: MnhnL OPH075, paratype, proximal lateral arm plate in external (C) and internal (D) views; E–F: MnhnL OPH076, paratype, distal lateral arm plate in external (E) and internal (F) views. Recent
|
||
<taxonomicName id="4C3636D8595FFFC0C649C5447AEF75A2" authorityName="Lyman" authorityYear="1875" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="23" phylum="Echinodermata" rank="species" species="stellata">
|
||
<emphasis id="B9429149595FFFC0C649C5447AEF75A2" italics="true" pageId="4" pageNumber="23">Ophiacantha stellata</emphasis>
|
||
</taxonomicName>
|
||
(G–J) from the tropical NW-Atlantic as closest living relative of
|
||
<taxonomicName id="4C3636D8595FFFC0C02BC5A07E4475A2" authority="Numberger-Thuy & Thuy, 2020" authorityName="Numberger-Thuy & Thuy" authorityYear="2020" box="[890,1097,1678,1702]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="23" phylum="Echinodermata" rank="species" species="oceani" status="sp. nov.">
|
||
<emphasis id="B9429149595FFFC0C02BC5A07E4475A2" box="[890,1097,1678,1702]" italics="true" pageId="4" pageNumber="23">Ophiacantha oceani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2712C32595FFFC0C71EC5A17EAD75A2" box="[1103,1184,1679,1702]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, G: complete skeleton in ventral view; H: detail of arm in lateral view; I–J: lateral arm plate in external (I) and internal (J) views. Abbreviations: ckv: central knob of the vertebral articular structures; di: distal; do: dorsal; pr: proximal; prv: proximal ridge of the vertebral articular structures; sa: spine articulation; sp: spur; ve: ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B894D5B595FFFC0C396C46E7B2F74EE" blockId="4.[151,1436,1856,2026]" pageId="4" pageNumber="23">
|
||
MnhnL OPH076 (
|
||
<figureCitation id="130D51DE595FFFC0C2C2C46E7BD9745E" box="[403,468,1856,1882]" captionStart="FIGURE 2" captionStartId="4.[151,250,1534,1558]" captionTargetBox="[164,1427,181,1510]" captionTargetId="figure-25@4.[151,1436,181,1510]" captionTargetPageId="4" captionText="FIGURE 2. Ophiacantha oceani sp. nov. (A–F) from the Piacenzian to Gelasian, latest Pliocene to earliest Pleistocene (approximately 2.6 mya) of Punta Mazza, Capo Milazzo, NE Sicily, Italy, A–B: MnhnL OPH074, holotype, proximal lateral arm plate in external (A) and internal (B) views; C–D: MnhnL OPH075, paratype, proximal lateral arm plate in external (C) and internal (D) views; E–F: MnhnL OPH076, paratype, distal lateral arm plate in external (E) and internal (F) views. Recent Ophiacantha stellata (G–J) from the tropical NW-Atlantic as closest living relative of Ophiacantha oceani sp. nov., G: complete skeleton in ventral view; H: detail of arm in lateral view; I–J: lateral arm plate in external (I) and internal (J) views. Abbreviations: ckv: central knob of the vertebral articular structures; di: distal; do: dorsal; pr: proximal; prv: proximal ridge of the vertebral articular structures; sa: spine articulation; sp: spur; ve: ventral." figureDoi="http://doi.org/10.5281/zenodo.4397242" httpUri="https://zenodo.org/record/4397242/files/figure.png" pageId="4" pageNumber="23">Fig. 2</figureCitation>
|
||
E–F) is a dissociated distal lateral arm plate approximately 1.5 times longer than high; general plate outline, outer surface ornamentation and outer proximal edge as in
|
||
<typeStatus id="548DF3F9595FFFC0C749C44A7E77747A" box="[1048,1146,1892,1918]" pageId="4" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
but with weaker constriction and thus less strongly concave dorsal edge; dorso-proximal tip of lateral arm plate missing; six spine articulations as in
|
||
<typeStatus id="548DF3F9595FFFC0C25CC4827B7E74C2" box="[269,371,1964,1990]" pageId="4" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
; size of gaps separating spine articulations increasing dorsalwards but size of spine articulations unchanging.
|
||
</paragraph>
|
||
<paragraph id="8B894D5B595EFFC1C396C3B9794A73D1" blockId="5.[151,1436,151,213]" pageId="5" pageNumber="24">
|
||
Inner side as in
|
||
<typeStatus id="548DF3F9595EFFC1C22AC3B97BD073B5" box="[379,477,151,177]" pageId="5" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
but with widely separated large and small central knobs; proximal vertebral articular structure roughly boomerang shaped but very poorly defined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |