307 lines
39 KiB
XML
307 lines
39 KiB
XML
<document id="259E142476BE040DEEF6035541FFACE8" ID-CLB-Dataset="5285" ID-DOI="10.11646/zootaxa.3835.4.3" ID-GBIF-Dataset="dbf8de20-9c7a-4b66-9130-615b6b42e84e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921623" ID-ZooBank="AD845CF3-CB19-4924-891F-330AAE283D07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623283883149" checkinUser="felipe" docAuthor="Yoshizawa, Kazunori, Lienhard, Charles & Abd, Idris" docDate="2014" docId="039F5A5FFFEDFFE9FF20FAB2FB803D43" docLanguage="en" docName="zt03835p500.pdf" docOrigin="Zootaxa 3835 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichadenotecnum sibolangitense Endang & New" docType="treatment" docVersion="7" lastPageNumber="488" masterDocId="FFA62227FFFFFFFAFFB7FF9FFF893E1E" masterDocTitle="Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)" masterLastPageNumber="500" masterPageNumber="469" pageNumber="487" updateTime="1699009618324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="E8040B7AE0A0C637095D6035BAA9FDCA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7AAAD794E558A463EDD8F7E9BC73CA8D">
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<mods:title id="1008514DFFEEE7E61D584AD4811B4530">Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)</mods:title>
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<mods:name id="531BF8A14247455EAD78DC0CE785467E" type="personal">
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<mods:role id="08B75FE45FE4F9D98EF162C2018B56C0">
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<mods:roleTerm id="EFCF7EDFC1BA45A28C9CEBB96130AD7A">Author</mods:roleTerm>
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<mods:namePart id="CE9CA8DC0EFD0C8BC350141834A95B22">Yoshizawa, Kazunori</mods:namePart>
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<mods:affiliation id="C9907EE777F8CBBE3666C8D8F5FB7C05">Systematic Entomology, School of Agriculture, Hokkaido University, Sapporo 060 - 8589, Japan.</mods:affiliation>
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</mods:name>
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<mods:name id="F8715DBBD94B456E2FAE4083902347C5" type="personal">
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<mods:roleTerm id="06E5ECF1E98E4E3C7975CE776CA5AE46">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3C10395B731AAB5A90DAD6E96B38FCD4">Lienhard, Charles</mods:namePart>
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<mods:affiliation id="7BA0080026A76C001CD4AFC8195E6BD2">Muséum d'histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name id="6715FB9A5F7BBE02674F5FC0565FE66D" type="personal">
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<mods:role id="EC71C3CB0A58B6EB1EC400A92A254199">
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<mods:roleTerm id="B75AFC287E6766B6E94D1C0DC18EA957">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="87B7A405EE5C2BEA8F86200CCD289F35">Abd, Idris</mods:namePart>
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<mods:affiliation id="7B47EEC8636683BDDB68A67C73719BA4">Center for Insect Systematics, Faculty of Sciences and Technology, Universiti Kebangsaan Malaysia, 43600 Bangi, Malaysia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="BEF9A2C10CD19102E28FE26835382825">text</mods:typeOfResource>
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<mods:title id="A14F88B4B18E63552532A7B03CC2F6D9">Zootaxa</mods:title>
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<mods:part id="0A6B0E7F554656763B251ECCECA11335">
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<mods:date id="E6574E8C7CC8C94E9B5EFCA89DF42617">2014</mods:date>
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<mods:number id="C88C4BD17244CDF6EC7CB44908A40CA2">2014-07-14</mods:number>
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<mods:detail id="B3F1244ACDF490DC79836FB37A94E185" type="volume">
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<mods:number id="7175260ABAB3D31605FBDC81A7C01E7B">3835</mods:number>
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<mods:detail id="3A8430A31089ADFE40C655F6B2A45AF3" type="issue">
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<mods:number id="A2E69705A064979F328B63E9C3A78013">4</mods:number>
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<mods:start id="69C8DDB2E02E2F5D8B66C076C3133AC2">469</mods:start>
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<mods:end id="73F78FF711D6301B9139F79A0B8FC5A7">500</mods:end>
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<mods:identifier id="BAB9C14AB272D31800A2A3E67435BA56" type="DOI">10.11646/zootaxa.3835.4.3</mods:identifier>
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<treatment id="039F5A5FFFEDFFE9FF20FAB2FB803D43" ID-DOI="http://doi.org/10.5281/zenodo.4921961" ID-GBIF-Taxon="183731224" ID-Zenodo-Dep="4921961" LSID="urn:lsid:plazi:treatment:039F5A5FFFEDFFE9FF20FAB2FB803D43" httpUri="http://treatment.plazi.org/id/039F5A5FFFEDFFE9FF20FAB2FB803D43" lastPageId="19" lastPageNumber="488" pageId="18" pageNumber="487">
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<subSubSection id="C32CB8C2FFEDFFE8FF20FAB2FD7F3B59" box="[151,758,1325,1351]" pageId="18" pageNumber="487" type="nomenclature">
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<paragraph id="8B89EB49FFEDFFE8FF20FAB2FD7F3B59" blockId="18.[151,758,1325,1384]" box="[151,758,1325,1351]" pageId="18" pageNumber="487">
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<heading id="D0C15C25FFEDFFE8FF20FAB2FD7F3B59" bold="true" box="[151,758,1325,1351]" fontSize="11" level="1" pageId="18" pageNumber="487" reason="1">
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<taxonomicName id="4C3690CAFFEDFFE8FF20FAB2FD7F3B59" ID-CoL="587H9" authority="Endang & New" authorityName="Endang & New" box="[151,758,1325,1351]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="487" phylum="Arthropoda" rank="species" species="sibolangitense">
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<emphasis id="B942375BFFEDFFE8FF20FAB2FD7F3B59" bold="true" box="[151,758,1325,1351]" pageId="18" pageNumber="487">
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<emphasis id="B942375BFFEDFFE8FF20FAB2FDB93B59" bold="true" box="[151,560,1325,1351]" italics="true" pageId="18" pageNumber="487">Trichadenotecnum sibolangitense</emphasis>
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Endang & New
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32CB8C2FFEDFFE8FF20FAD0FEDE3B76" box="[151,343,1359,1384]" pageId="18" pageNumber="487" type="description">
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<paragraph id="8B89EB49FFEDFFE8FF20FAD0FEDE3B76" blockId="18.[151,758,1325,1384]" box="[151,343,1359,1384]" pageId="18" pageNumber="487">
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(
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<figureCitation id="130DF7CCFFEDFFE8FF28FAD0FF753B76" box="[159,252,1359,1384]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="18" pageNumber="487">Figs 1N</figureCitation>
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,
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<figureCitation id="130DF7CCFFEDFFE8FEBFFAD0FEAF3B76" box="[264,294,1359,1384]" captionStart="FIGURE 18" captionStartId="19.[151,250,1832,1854]" captionTargetBox="[159,1432,922,1804]" captionTargetId="figure-398@19.[151,1435,916,1811]" captionTargetPageId="19" captionText="FIGURE 18. Male terminalia of Trichadenotecnum sibolangitense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921662" httpUri="https://zenodo.org/record/4921662/files/figure.png" pageId="18" pageNumber="487">18</figureCitation>
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,
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<figureCitation id="130DF7CCFFEDFFE8FE85FAD0FEC63B76" box="[306,335,1359,1384]" captionStart="FIGURE 19" captionStartId="20.[151,250,1045,1067]" captionTargetBox="[154,1429,196,1019]" captionTargetId="figure-253@20.[151,1435,193,1024]" captionTargetPageId="20" captionText="FIGURE 19. Female genitalia of Trichadenotecnum sibolangitense, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921664" httpUri="https://zenodo.org/record/4921664/files/figure.png" pageId="18" pageNumber="487">19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32CB8C2FFEDFFE8FF20FA07FC973825" pageId="18" pageNumber="487" type="reference_group">
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<paragraph id="8B89EB49FFEDFFE8FF20FA07FD723BB0" blockId="18.[151,763,1432,1454]" box="[151,763,1432,1454]" pageId="18" pageNumber="487">
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<treatmentCitationGroup id="AB26CC67FFEDFFE8FF20FA07FD723BB0" box="[151,763,1432,1454]" pageId="18" pageNumber="487">
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<treatmentCitation id="0A97CD58FFEDFFE8FF20FA07FD7F3BB0" author="Endang, S. K. & New, T. R." box="[151,758,1432,1454]" page="27" pageId="18" pageNumber="487" year="2005">
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<taxonomicName id="4C3690CAFFEDFFE8FF20FA07FD7F3BB0" ID-CoL="587H9" authority="Endang & New, 2005: 27" authorityName="Endang & New" authorityPageNumber="27" authorityYear="2005" box="[151,758,1432,1454]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="18" pageNumber="487" phylum="Arthropoda" rank="species" species="sibolangitense">
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<emphasis id="B942375BFFEDFFE8FF20FA07FE653BB0" box="[151,492,1432,1454]" italics="true" pageId="18" pageNumber="487">Trichadenotecnum sibolangitense</emphasis>
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<bibRefCitation id="EFA796B8FFEDFFE8FE45FA07FD7F3BB0" author="Endang, S. K. & New, T. R." box="[498,758,1432,1454]" pageId="18" pageNumber="487" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. & New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang & New, 2005: 27</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B89EB49FFEDFFE8FF20FA45FC973825" blockId="18.[151,1437,1498,2027]" pageId="18" pageNumber="487">
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<emphasis id="B942375BFFEDFFE8FF20FA45FE1E3BEC" box="[151,407,1498,1523]" inLineHeading="true" pageId="18" pageNumber="487" reason="1">
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<emphasis id="B942375BFFEDFFE8FF20FA45FE193BED" bold="true" box="[151,400,1498,1523]" pageId="18" pageNumber="487">Specimens examined</emphasis>
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.
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</emphasis>
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[
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<collectingCountry id="F321ABD9FFEDFFE8FE11FA45FD9B3BED" box="[422,530,1498,1523]" name="Malaysia" pageId="18" pageNumber="487">Malaysia</collectingCountry>
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]
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<specimenCount id="9D3020C0FFEDFFE8FD95FA45FD0E3BED" box="[546,647,1498,1523]" pageId="18" pageNumber="487" type="female">1 female</specimenCount>
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, Gunung Jasar, Cameron Highlands,
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<collectingRegion id="49F225ABFFEDFFE8FB99FA45FB0A3BED" box="[1070,1155,1498,1523]" country="Malaysia" name="Pahang" pageId="18" pageNumber="487">Pahang</collectingRegion>
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,
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<date id="FF88CD89FFEDFFE8FB27FA45FA853BED" box="[1168,1292,1498,1523]" pageId="18" pageNumber="487" value="2003-03-14">14.iii.2003</date>
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, KY;
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<materialsCitation id="3B5EE114FFEDFFE8FAF8FA45FB3C3809" ID-GBIF-Occurrence="3326724336" collectingDate="2005-03-10" country="Singapore" location="Gunung Berembun" municipality="Cameron Highlands" pageId="18" pageNumber="487" specimenCount="1" specimenCount-male="1" stateProvince="Pahang">
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<specimenCount id="9D3020C0FFEDFFE8FAF8FA45FA153BED" box="[1359,1436,1498,1523]" pageId="18" pageNumber="487" type="male">1 male</specimenCount>
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(KY345)
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<location id="8EE9BD92FFEDFFE8FEBCFA61FE643809" LSID="urn:lsid:plazi:treatment:039F5A5FFFEDFFE9FF20FAB2FB803D43:8EE9BD92FFEDFFE8FEBCFA61FE643809" box="[267,493,1534,1559]" country="Singapore" municipality="Cameron Highlands" name="Gunung Berembun" pageId="18" pageNumber="487" stateProvince="Pahang">Gunung Berembun</location>
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,
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<collectingMunicipality id="6BED7133FFEDFFE8FDB7FA61FD653809" box="[512,748,1534,1559]" pageId="18" pageNumber="487">Cameron Highlands</collectingMunicipality>
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,
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<collectingRegion id="49F225ABFFEDFFE8FCB7FA61FCDC3809" box="[768,853,1534,1559]" country="Malaysia" name="Pahang" pageId="18" pageNumber="487">Pahang</collectingRegion>
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,
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<date id="FF88CD89FFEDFFE8FCDFFA61FC6D3809" box="[872,996,1534,1559]" pageId="18" pageNumber="487" value="2005-03-10">
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<collectingDate id="EFCC3461FFEDFFE8FCDFFA61FC6D3809" box="[872,996,1534,1559]" pageId="18" pageNumber="487" value="2005-03-10">10.iii.2005</collectingDate>
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</date>
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, NT. [
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<collectingCountry id="F321ABD9FFEDFFE8FB8FFA61FB263809" box="[1080,1199,1534,1559]" name="Singapore" pageId="18" pageNumber="487">Singapore</collectingCountry>
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]
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</materialsCitation>
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<materialsCitation id="3B5EE114FFEDFFE8FB72FA61FC923825" ID-GBIF-Occurrence="3326724302" collectingDate="1981-11-06" collectionCode="MHNG" country="Singapore" location="Botanic Gardens" municipality="Forest" pageId="18" pageNumber="487" specimenCount="1" specimenCount-female="1" stateProvince="Pahang">
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<specimenCount id="9D3020C0FFEDFFE8FB72FA61FAB83809" box="[1221,1329,1534,1559]" pageId="18" pageNumber="487" type="female">1 female</specimenCount>
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,
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<location id="8EE9BD92FFEDFFE8FAF2FA61FF7E3825" LSID="urn:lsid:plazi:treatment:039F5A5FFFEDFFE9FF20FAB2FB803D43:8EE9BD92FFEDFFE8FAF2FA61FF7E3825" country="Singapore" municipality="Forest" name="Botanic Gardens" pageId="18" pageNumber="487" stateProvince="Pahang">Botanic Gardens</location>
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,
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<collectingMunicipality id="6BED7133FFEDFFE8FEB5F9BCFEC13824" box="[258,328,1571,1594]" pageId="18" pageNumber="487">Forest</collectingMunicipality>
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(low foliage),
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<date id="FF88CD89FFEDFFE8FE59F9BDFDD23825" box="[494,603,1570,1595]" pageId="18" pageNumber="487" value="1981-11-06">
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<collectingDate id="EFCC3461FFEDFFE8FE59F9BDFDD23825" box="[494,603,1570,1595]" pageId="18" pageNumber="487" value="1981-11-06">6.xi.1981</collectingDate>
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</date>
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, DHM (
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<collectionCode id="ED27738CFFEDFFE8FD01F9BDFC9B3825" box="[694,786,1570,1595]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="18" pageNumber="487" type="Museum">MHNG</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32CB8C2FFEDFFE8FF70F9D9FD053928" pageId="18" pageNumber="487" type="description">
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<paragraph id="8B89EB49FFEDFFE8FF70F9D9FD053928" blockId="18.[151,1437,1498,2027]" pageId="18" pageNumber="487">
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<emphasis id="B942375BFFEDFFE8FF70F9D9FDB53840" box="[199,572,1606,1631]" inLineHeading="true" pageId="18" pageNumber="487" reason="1">
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<emphasis id="B942375BFFEDFFE8FF70F9D9FDBC3841" bold="true" box="[199,565,1606,1631]" pageId="18" pageNumber="487">Description of female genitalia</emphasis>
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.
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</emphasis>
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Egg guide (
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<figureCitation id="130DF7CCFFEDFFE8FD71F9D9FCA43841" box="[710,813,1606,1631]" captionStart="FIGURE 19" captionStartId="20.[151,250,1045,1067]" captionTargetBox="[154,1429,196,1019]" captionTargetId="figure-253@20.[151,1435,193,1024]" captionTargetPageId="20" captionText="FIGURE 19. Female genitalia of Trichadenotecnum sibolangitense, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921664" httpUri="https://zenodo.org/record/4921664/files/figure.png" pageId="18" pageNumber="487">Fig. 19A</figureCitation>
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) short, shorter than basal width, ventrally membranous and dorsally widely sclerotized, distal margin concave; body of subgenital plate broadly sclerotized, anteromedially with wide membranous region, posterior margin with pair of long, narrow and sharply pointed lateral processes. Gonapophyses (
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<figureCitation id="130DF7CCFFEDFFE8FDA1F92DFDF738D5" box="[534,638,1714,1739]" captionStart="FIGURE 19" captionStartId="20.[151,250,1045,1067]" captionTargetBox="[154,1429,196,1019]" captionTargetId="figure-253@20.[151,1435,193,1024]" captionTargetPageId="20" captionText="FIGURE 19. Female genitalia of Trichadenotecnum sibolangitense, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921664" httpUri="https://zenodo.org/record/4921664/files/figure.png" pageId="18" pageNumber="487">Fig. 19B</figureCitation>
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). Ventral valve not exceeding external valve; dorsal valve broad, with deep pouch posteromedially, distal process long; external valve with broad membranous region anteriorly and with flap-like expansion posteriorly, posterior lobe broad, internal lobe narrow. Spermapore plate (
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<figureCitation id="130DF7CCFFEDFFE8FA92F965FA1D390D" box="[1317,1428,1786,1811]" captionStart="FIGURE 19" captionStartId="20.[151,250,1045,1067]" captionTargetBox="[154,1429,196,1019]" captionTargetId="figure-253@20.[151,1435,193,1024]" captionTargetPageId="20" captionText="FIGURE 19. Female genitalia of Trichadenotecnum sibolangitense, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921664" httpUri="https://zenodo.org/record/4921664/files/figure.png" pageId="18" pageNumber="487">Fig. 19C</figureCitation>
|
|
) symmetrical, pigmented around spermapore.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C32CB8C2FFEDFFE9FF70F8DDFB803D43" lastPageId="19" lastPageNumber="488" pageId="18" pageNumber="487" type="discussion">
|
|
<paragraph id="8B89EB49FFEDFFE9FF70F8DDFA0C3C25" blockId="18.[151,1437,1498,2027]" lastBlockId="19.[151,1438,151,861]" lastPageId="19" lastPageNumber="488" pageId="18" pageNumber="487">
|
|
<emphasis id="B942375BFFEDFFE8FF70F8DDFEB23944" box="[199,315,1858,1883]" inLineHeading="true" pageId="18" pageNumber="487" reason="1">
|
|
<emphasis id="B942375BFFEDFFE8FF70F8DDFEBD3945" bold="true" box="[199,308,1858,1883]" pageId="18" pageNumber="487">Remarks</emphasis>
|
|
.
|
|
</emphasis>
|
|
This species was originally described from
|
|
<collectingRegion id="49F225ABFFEDFFE8FC85F8DDFC1B3945" box="[818,914,1858,1883]" country="Indonesia" name="Sumatera Utara" pageId="18" pageNumber="487">Sumatra</collectingRegion>
|
|
,
|
|
<collectingCountry id="F321ABD9FFEDFFE8FC28F8DDFB853945" box="[927,1036,1858,1883]" name="Indonesia" pageId="18" pageNumber="487">Indonesia</collectingCountry>
|
|
based on a single male and is here recorded from Peninsula
|
|
<collectingCountry id="F321ABD9FFEDFFE8FE0EF8F9FD963961" box="[441,543,1894,1919]" name="Malaysia" pageId="18" pageNumber="487">Malaysia</collectingCountry>
|
|
for the first time. The male specimens examined here agree exactly with the original description except for the hypandrial structures being in mirror image (
|
|
<figureCitation id="130DF7CCFFEDFFE8FBEFF815FB4C39BD" box="[1112,1221,1930,1955]" captionStart="FIGURE 18" captionStartId="19.[151,250,1832,1854]" captionTargetBox="[159,1432,922,1804]" captionTargetId="figure-398@19.[151,1435,916,1811]" captionTargetPageId="19" captionText="FIGURE 18. Male terminalia of Trichadenotecnum sibolangitense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921662" httpUri="https://zenodo.org/record/4921662/files/figure.png" pageId="18" pageNumber="487">Fig. 18C</figureCitation>
|
|
). Given the exact similarity of each structure, we concluded that the illustration of the hypandrium of this species presented in
|
|
<bibRefCitation id="EFA796B8FFEDFFE8FF20F84DFE2C39F5" author="Endang, S. K. & New, T. R." box="[151,421,2002,2027]" pageId="18" pageNumber="487" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. & New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang & New (2005)</bibRefCitation>
|
|
is drawn in dorsal (internal) view or reproduced in reversed position. This species is apparently closely related to
|
|
<taxonomicName id="4C3690CAFFECFFE9FE6FFF06FC433EAE" authority="Endang, Thornton & New, 2005" authorityName="Endang, Thornton & New" authorityYear="2005" box="[472,970,151,176]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="alinguum">
|
|
<emphasis id="B942375BFFECFFE9FE6FFF06FDD43EAE" box="[472,605,151,176]" italics="true" pageId="19" pageNumber="488">T. alinguum</emphasis>
|
|
Endang, Thornton & New, 2005
|
|
</taxonomicName>
|
|
and, in
|
|
<taxonomicName id="4C3690CAFFECFFE9FB94FF06FB693EAE" box="[1059,1248,151,176]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="sibolangitense">
|
|
<emphasis id="B942375BFFECFFE9FB94FF06FB693EAE" box="[1059,1248,151,176]" italics="true" pageId="19" pageNumber="488">T. sibolangitense</emphasis>
|
|
</taxonomicName>
|
|
, the right side of the hypandrium is strongly expanded, as illustrated in
|
|
<figureCitation id="130DF7CCFFECFFE9FCB2FF23FCE53ECB" box="[773,876,188,213]" captionStart="FIGURE 18" captionStartId="19.[151,250,1832,1854]" captionTargetBox="[159,1432,922,1804]" captionTargetId="figure-398@19.[151,1435,916,1811]" captionTargetPageId="19" captionText="FIGURE 18. Male terminalia of Trichadenotecnum sibolangitense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921662" httpUri="https://zenodo.org/record/4921662/files/figure.png" pageId="19" pageNumber="488">Fig. 18C</figureCitation>
|
|
. Given the lack of the hypandrial median tongue and the triangular shape of the hypandrium, this species is considered to be closely related to
|
|
<taxonomicName id="4C3690CAFFECFFE9FABAFF7EFA123EE6" authorityName=": New & Lee" authorityYear="1992" box="[1293,1435,223,248]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">
|
|
<emphasis id="B942375BFFECFFE9FABAFF7EFA123EE6" box="[1293,1435,223,248]" italics="true" pageId="19" pageNumber="488">T. arciforme</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="EFA796B8FFECFFE9FF29FE9BFEDB3F03" author="Thornton, I. W. B." box="[158,338,260,285]" pageId="19" pageNumber="488" pagination="1 - 24" refId="ref15445" refString="Thornton, I. W. B. (1961) The Trichadenotecnum group (Psocoptera: Psocidae) in Hong Kong, with descriptions of new species. Transactions of the Royal Entomological Society, London, 113, 1 - 24. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1961. tb 00797. x" type="journal article" year="1961">Thornton, 1961</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFA796B8FFECFFE9FEE8FE9BFD393F03" author="Yoshizawa, K. & Lienhard, C." box="[351,688,260,285]" pageId="19" pageNumber="488" pagination="121 - 149" refId="ref15885" refString="Yoshizawa, K. & Lienhard, C. (2004) Systematics of Trichadenotecnum (Psocoptera: Psocidae) in Hong Kong. Publicaciones Especiales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 20, 121 - 149." type="journal article" year="2004">Yoshizawa & Lienhard, 2004</bibRefCitation>
|
|
). In contrast, these species are clearly different in the condition of the epiproct: strongly expanded over the clunium in
|
|
<taxonomicName id="4C3690CAFFECFFE9FD5EFEB6FC213F5E" box="[745,936,295,320]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="sibolangitense">
|
|
<emphasis id="B942375BFFECFFE9FD5EFEB6FC213F5E" box="[745,936,295,320]" italics="true" pageId="19" pageNumber="488">T. sibolangitense</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="130DF7CCFFECFFE9FC0FFEB8FB973F5E" box="[952,1054,295,320]" captionStart="FIGURE 18" captionStartId="19.[151,250,1832,1854]" captionTargetBox="[159,1432,922,1804]" captionTargetId="figure-398@19.[151,1435,916,1811]" captionTargetPageId="19" captionText="FIGURE 18. Male terminalia of Trichadenotecnum sibolangitense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921662" httpUri="https://zenodo.org/record/4921662/files/figure.png" pageId="19" pageNumber="488">Fig. 18A</figureCitation>
|
|
) whereas almost flat and covered by the clunial flap in
|
|
<emphasis id="B942375BFFECFFE9FE31FED2FD9E3F7B" box="[390,535,332,357]" italics="true" pageId="19" pageNumber="488">
|
|
<taxonomicName id="4C3690CAFFECFFE9FE31FED2FD9B3F7B" authorityName=": New & Lee" authorityYear="1992" box="[390,530,332,357]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">T. arciforme</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
The state as observed in
|
|
<taxonomicName id="4C3690CAFFECFFE9FC86FED2FC333F7B" authorityName=": New & Lee" authorityYear="1992" box="[817,954,332,357]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">
|
|
<emphasis id="B942375BFFECFFE9FC86FED2FC333F7B" box="[817,954,332,357]" italics="true" pageId="19" pageNumber="488">T. arciforme</emphasis>
|
|
</taxonomicName>
|
|
is unique even among the tribe
|
|
<taxonomicName id="4C3690CAFFECFFE9FA96FED3FAF23F7B" authorityName="Mockford" authorityYear="1993" box="[1313,1403,332,357]" class="Insecta" family="Psocidae" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="tribe" tribe="Ptyctini">Ptyctini</taxonomicName>
|
|
, in which
|
|
<taxonomicName id="4C3690CAFFECFFE9FF53FEF0FE3D3F96" authorityName="Enderlein" authorityYear="1909" box="[228,436,367,392]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B942375BFFECFFE9FF53FEF0FE3D3F96" box="[228,436,367,392]" italics="true" pageId="19" pageNumber="488">Trichadenotecnum</emphasis>
|
|
</taxonomicName>
|
|
is classified. Because of this highly unusual feature,
|
|
<bibRefCitation id="EFA796B8FFECFFE9FBA1FEF0FAF93F96" author="Yoshizawa, K. & Lienhard, C." box="[1046,1392,367,392]" pageId="19" pageNumber="488" pagination="121 - 149" refId="ref15885" refString="Yoshizawa, K. & Lienhard, C. (2004) Systematics of Trichadenotecnum (Psocoptera: Psocidae) in Hong Kong. Publicaciones Especiales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 20, 121 - 149." type="journal article" year="2004">Yoshizawa & Lienhard (2004)</bibRefCitation>
|
|
did not assign
|
|
<taxonomicName id="4C3690CAFFECFFE9FEB8FE0AFE113FB3" authorityName=": New & Lee" authorityYear="1992" box="[271,408,404,429]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">
|
|
<emphasis id="B942375BFFECFFE9FEB8FE0AFE113FB3" box="[271,408,404,429]" italics="true" pageId="19" pageNumber="488">T. arciforme</emphasis>
|
|
</taxonomicName>
|
|
to any species group, although they pointed out that
|
|
<taxonomicName id="4C3690CAFFECFFE9FC5CFE0AFBFA3FB3" authorityName=": New & Lee" authorityYear="1992" box="[1003,1139,404,429]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">
|
|
<emphasis id="B942375BFFECFFE9FC5CFE0AFBFA3FB3" box="[1003,1139,404,429]" italics="true" pageId="19" pageNumber="488">T. arciforme</emphasis>
|
|
</taxonomicName>
|
|
is possibly closely related to
|
|
<taxonomicName id="4C3690CAFFECFFE9FF02FE26FEA63FCE" authorityName="Thornton" authorityYear="1961" box="[181,303,441,464]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="apertum">
|
|
<emphasis id="B942375BFFECFFE9FF02FE26FEA63FCE" box="[181,303,441,464]" italics="true" pageId="19" pageNumber="488">T. apertum</emphasis>
|
|
</taxonomicName>
|
|
. The strongly expanded epiproct as observed in
|
|
<taxonomicName id="4C3690CAFFECFFE9FCEBFE26FB923FCE" box="[860,1051,439,464]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="sibolangitense">
|
|
<emphasis id="B942375BFFECFFE9FCEBFE26FB923FCE" box="[860,1051,439,464]" italics="true" pageId="19" pageNumber="488">T. sibolangitense</emphasis>
|
|
</taxonomicName>
|
|
and similarities of the hypandrial structure between this species and
|
|
<taxonomicName id="4C3690CAFFECFFE9FD93FE42FD273FEB" authorityName=": New & Lee" authorityYear="1992" box="[548,686,476,501]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">
|
|
<emphasis id="B942375BFFECFFE9FD93FE42FD273FEB" box="[548,686,476,501]" italics="true" pageId="19" pageNumber="488">T. arciforme</emphasis>
|
|
</taxonomicName>
|
|
corroborate the idea that
|
|
<taxonomicName id="4C3690CAFFECFFE9FC60FE42FBE83FEB" authorityName=": New & Lee" authorityYear="1992" box="[983,1121,476,501]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">
|
|
<emphasis id="B942375BFFECFFE9FC60FE42FBE83FEB" box="[983,1121,476,501]" italics="true" pageId="19" pageNumber="488">T. arciforme</emphasis>
|
|
</taxonomicName>
|
|
(hypandrial median tongue absent and ventral valve of gonapophyses short) is closely related to
|
|
<taxonomicName id="4C3690CAFFECFFE9FC25FD9EFB833C06" authorityName="Thornton" authorityYear="1961" box="[914,1034,513,536]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="apertum">
|
|
<emphasis id="B942375BFFECFFE9FC25FD9EFB833C06" box="[914,1034,513,536]" italics="true" pageId="19" pageNumber="488">T. apertum</emphasis>
|
|
</taxonomicName>
|
|
and its relatives, and that the highly specialized condition of the epiproct-clunium interface has to be considered as an autapomorphy of
|
|
<taxonomicName id="4C3690CAFFECFFE9FB42FDBAFAF43C23" authorityName=": New & Lee" authorityYear="1992" box="[1269,1405,548,573]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="arciforme">
|
|
<emphasis id="B942375BFFECFFE9FB42FDBAFAF43C23" box="[1269,1405,548,573]" italics="true" pageId="19" pageNumber="488">T. arciforme</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8B89EB49FFECFFE9FF70FDD8FB803D43" blockId="19.[151,1438,151,861]" pageId="19" pageNumber="488">
|
|
The female of
|
|
<taxonomicName id="4C3690CAFFECFFE9FEDDFDD6FDAE3C7E" box="[362,551,583,608]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="sibolangitense">
|
|
<emphasis id="B942375BFFECFFE9FEDDFDD6FDAE3C7E" box="[362,551,583,608]" italics="true" pageId="19" pageNumber="488">T. sibolangitense</emphasis>
|
|
</taxonomicName>
|
|
is described here for the first time. As shown in
|
|
<figureCitation id="130DF7CCFFECFFE9FBF3FDD8FB233C7E" box="[1092,1194,583,608]" captionStart="FIGURE 19" captionStartId="20.[151,250,1045,1067]" captionTargetBox="[154,1429,196,1019]" captionTargetId="figure-253@20.[151,1435,193,1024]" captionTargetPageId="20" captionText="FIGURE 19. Female genitalia of Trichadenotecnum sibolangitense, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921664" httpUri="https://zenodo.org/record/4921664/files/figure.png" pageId="19" pageNumber="488">Fig. 19A</figureCitation>
|
|
, it is characterized by possession of a pair of sharp processes arising from the posterior margin of the subgenital plate. Similar structures have been reported from some species such as
|
|
<taxonomicName id="4C3690CAFFECFFE9FD2AFD0EFBCF3CB6" authority="Yoshizawa & Lienhard, 2007" authorityName="Yoshizawa & Lienhard" authorityYear="2007" box="[669,1094,655,680]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="suwai">
|
|
<emphasis id="B942375BFFECFFE9FD2AFD0EFD703CB9" box="[669,761,656,680]" italics="true" pageId="19" pageNumber="488">T. suwai</emphasis>
|
|
<bibRefCitation id="EFA796B8FFECFFE9FCB7FD10FBCF3CB6" author="Yoshizawa, K. & Lienhard, C. & Thapa, V. K." box="[768,1094,655,680]" pageId="19" pageNumber="488" pagination="1 - 33" refId="ref15928" refString="Yoshizawa, K., Lienhard, C. & Thapa, V. K. (2007) Systematic study of the genus Trichadenotecnum in Nepal (Psocodea: ' Psocoptera': Psocidae). Insecta matsumurana, new series, 63, 1 - 33." type="journal article" year="2007">Yoshizawa & Lienhard, 2007</bibRefCitation>
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F321ABD9FFECFFE9FB3DFD10FB443CB6" box="[1162,1229,655,680]" name="Nepal" pageId="19" pageNumber="488">Nepal</collectingCountry>
|
|
or
|
|
<taxonomicName id="4C3690CAFFECFFE9FB47FD0EFE053CD3" authority="Endang et al., 2002" authorityName="Endang" authorityYear="2002" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="laticornutum">
|
|
<emphasis id="B942375BFFECFFE9FB47FD0EFA153CB6" box="[1264,1436,655,680]" italics="true" pageId="19" pageNumber="488">T. laticornutum</emphasis>
|
|
<bibRefCitation id="EFA796B8FFECFFE9FF20FD2BFE053CD3" author="Endang, S. K. & Thornton, I. W. B. & New, T. R." box="[151,396,692,717]" pageId="19" pageNumber="488" pagination="107 - 176" refId="ref15093" refString="Endang, S. K., Thornton, I. W. B. & New, T. R. (2002) The Psocidae (Insecta: Psocoptera) of Java and the eastern islands of Indonesia. Invertebrate Systematics, 16, 107 - 176." type="journal article" year="2002">
|
|
Endang
|
|
<emphasis id="B942375BFFECFFE9FF4BFD2AFEB43CD3" box="[252,317,692,717]" italics="true" pageId="19" pageNumber="488">et al.</emphasis>
|
|
, 2002
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C3690CAFFECFFE9FE63FD2AFC313CD3" authority="Endang & New, 2005" authorityName="Endang & New" authorityYear="2005" box="[468,952,692,717]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="waykananense">
|
|
<emphasis id="B942375BFFECFFE9FE63FD2AFD123CD3" box="[468,667,692,717]" italics="true" pageId="19" pageNumber="488">T. waykananense</emphasis>
|
|
<bibRefCitation id="EFA796B8FFECFFE9FD1CFD2BFC313CD3" author="Endang, S. K. & New, T. R." box="[683,952,692,717]" pageId="19" pageNumber="488" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. & New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang & New, 2005</bibRefCitation>
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="F321ABD9FFECFFE9FBB9FD2BFBF63CD3" box="[1038,1151,692,717]" name="Indonesia" pageId="19" pageNumber="488">Indonesia</collectingCountry>
|
|
. The former species is phylogenetically distant (belonging to the
|
|
<emphasis id="B942375BFFECFFE9FDCAFD48FC903CEE" box="[637,793,727,752]" italics="true" pageId="19" pageNumber="488">spiniserrulum</emphasis>
|
|
group) from
|
|
<taxonomicName id="4C3690CAFFECFFE9FC01FD46FBFF3CEE" box="[950,1142,727,752]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="sibolangitense">
|
|
<emphasis id="B942375BFFECFFE9FC01FD46FBFF3CEE" box="[950,1142,727,752]" italics="true" pageId="19" pageNumber="488">T. sibolangitense</emphasis>
|
|
</taxonomicName>
|
|
so that presence of these processes can be recognized as convergence. In contrast, although the species-group assignments of
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<taxonomicName id="4C3690CAFFECFFE9FA30FD62FEA13D26" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="laticornutum">
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<emphasis id="B942375BFFECFFE9FA30FD62FEA13D26" italics="true" pageId="19" pageNumber="488">T. laticornutum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3690CAFFECFFE9FEDFFCBEFDA23D26" authorityName="Endang & New" authorityYear="2005" box="[360,555,799,824]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="waykananense">
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<emphasis id="B942375BFFECFFE9FEDFFCBEFDA23D26" box="[360,555,799,824]" italics="true" pageId="19" pageNumber="488">T. waykananense</emphasis>
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</taxonomicName>
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have not been proposed in the original descriptions (
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<bibRefCitation id="EFA796B8FFECFFE9FB1EFC80FA1E3D26" author="Endang, S. K. & Thornton, I. W. B. & New, T. R." box="[1193,1431,799,824]" pageId="19" pageNumber="488" pagination="107 - 176" refId="ref15093" refString="Endang, S. K., Thornton, I. W. B. & New, T. R. (2002) The Psocidae (Insecta: Psocoptera) of Java and the eastern islands of Indonesia. Invertebrate Systematics, 16, 107 - 176." type="journal article" year="2002">
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Endang
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|
<emphasis id="B942375BFFECFFE9FABBFCBEFAC83D26" box="[1292,1345,799,824]" italics="true" pageId="19" pageNumber="488">et al</emphasis>
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|
., 2002
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|
</bibRefCitation>
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|
;
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|
<bibRefCitation id="EFA796B8FFECFFE9FF20FCDBFE043D43" author="Endang, S. K. & New, T. R." box="[151,397,836,861]" pageId="19" pageNumber="488" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. & New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang & New, 2005</bibRefCitation>
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|
), their close affinity with
|
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<taxonomicName id="4C3690CAFFECFFE9FD1AFCDAFCE33D43" box="[685,874,836,861]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="19" pageNumber="488" phylum="Arthropoda" rank="species" species="sibolangitense">
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|
<emphasis id="B942375BFFECFFE9FD1AFCDAFCE33D43" box="[685,874,836,861]" italics="true" pageId="19" pageNumber="488">T. sibolangitense</emphasis>
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|
</taxonomicName>
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is very likely.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |