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<document id="798F10555E7E5A5FE216C7ABBD2D9535" ID-CLB-Dataset="2961" ID-DOI="10.11646/palaeoentomology.4.6.5" ID-GBIF-Dataset="73a0820d-7503-472f-8c32-3836cf5fe857" ID-ISSN="2624-2834" ID-Zenodo-Dep="5778576" ID-ZooBank="95AA3380-CA95-46E8-B233-E9F0E8D60603" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639469484134" checkinUser="plazi" docAuthor="Béthoux, Olivier &amp; Anderson, John M." docDate="2021" docId="039DFA6CFFD9FFD86EC7FB4DCA79FE10" docLanguage="en" docName="palaeoentomology.4.6.5.pdf" docOrigin="Palaeoentomology 4 (6)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Moltenojurina parva Béthoux &amp; Anderson 2021, sp. nov." docType="treatment" docUuid="BFCFDAB4-295D-4C86-A9AB-7441215855A4" docUuidSource="ZooBank" docVersion="6" lastPageNumber="557" masterDocId="FFA48214FFDFFFDF6E4FFF9CC82EFFF8" masterDocTitle="The Polycytellidae viewed as Gondwanan Glosselytrodea" masterLastPageNumber="558" masterPageNumber="550" pageNumber="556" updateTime="1699472406386" updateUser="ExternalLinkService">
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<mods:titleInfo id="A9DBB5D01282879F2FB053762FDA0C74">
<mods:title id="03052F27D5F05788DF6510BE44CA4061">The Polycytellidae viewed as Gondwanan Glosselytrodea</mods:title>
</mods:titleInfo>
<mods:name id="DE9F13E9AE56F6BFDAD1C8EC7F94D7DF" type="personal">
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<mods:namePart id="C90A9C174DBDA7242078D579CE87B18F">Béthoux, Olivier</mods:namePart>
<mods:affiliation id="E6F5EDBEA4B02D48698DC593CF9BE98C">Centre de Recherche sur la Paléontologie - Paris (CR 2 P), Sorbonne Université, MNHN, CNRS, Paris, France</mods:affiliation>
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<mods:namePart id="58DED438998A503739E850BB61C4AAD6">Anderson, John M.</mods:namePart>
<mods:affiliation id="E11AE72CA1039AFE7AEB17A22055B328">Evolutionary Studies Institute, Witwatersrand University, Johannesburg, South Africa</mods:affiliation>
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<mods:title id="97E646D34E7EC47FD7AE2AFC03FBDE2B">Palaeoentomology</mods:title>
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<mods:date id="369A73C7F8CD57767557ACA5E9CD07D1">2021</mods:date>
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<mods:number id="D76FE0BC3F05C47C2A7C363FE0AA658C">2021-12-14</mods:number>
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<mods:identifier id="F88D0CB288ECC2805118242D5164BE97" type="DOI">10.11646/palaeoentomology.4.6.5</mods:identifier>
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<treatment id="039DFA6CFFD9FFD86EC7FB4DCA79FE10" ID-DOI="http://doi.org/10.5281/zenodo.5778590" ID-GBIF-Taxon="191571265" ID-Zenodo-Dep="5778590" LSID="urn:lsid:zoobank.org:act:BFCFDAB4-295D-4C86-A9AB-7441215855A4" httpUri="http://treatment.plazi.org/id/039DFA6CFFD9FFD86EC7FB4DCA79FE10" lastPageId="7" lastPageNumber="557" pageId="6" pageNumber="556">
<subSubSection id="C32E18F1FFD9FFD96EC7FB4DC9F7FB14" box="[136,473,1233,1260]" pageId="6" pageNumber="556" type="nomenclature">
<paragraph id="8B8B4B7AFFD9FFD96EC7FB4DC9F7FB14" blockId="6.[136,777,1233,1365]" box="[136,473,1233,1260]" pageId="6" pageNumber="556">
<heading id="D0C3FC16FFD9FFD96EC7FB4DC9F7FB14" bold="true" box="[136,473,1233,1260]" fontSize="10" level="2" pageId="6" pageNumber="556" reason="3">
<emphasis id="B9409768FFD9FFD96EC7FB4DC9F7FB14" bold="true" box="[136,473,1233,1260]" pageId="6" pageNumber="556">
<taxonomicName id="4C3430F9FFD9FFD96EC7FB4DC956FB14" authority="Béthoux &amp; Anderson, 2021" authorityName="Béthoux &amp; Anderson" authorityYear="2021" box="[136,376,1233,1260]" class="Insecta" family="Polycytellidae" genus="Moltenojurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="B9409768FFD9FFD96EC7FB4DC956FB14" bold="true" box="[136,376,1233,1260]" italics="true" pageId="6" pageNumber="556">Moltenojurina parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2732A13FFD9FFD96F30FB4EC9F7FB14" box="[383,473,1234,1260]" pageId="6" pageNumber="556" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32E18F1FFD9FFD96EC7FB68C9A6FAAD" pageId="6" pageNumber="556" type="description">
<paragraph id="8B8B4B7AFFD9FFD96EC7FB68C9A6FAAD" blockId="6.[136,777,1233,1365]" pageId="6" pageNumber="556">
(
<figureCitation id="130F57FFFFD9FFD96EDFFB68C8FAFAF7" box="[144,212,1268,1295]" captionStart="FIGURE 4" captionStartId="6.[810,909,1974,1998]" captionTargetBox="[809,1452,1435,1950]" captionTargetId="figure-692@6.[809,1452,1430,1950]" captionTargetPageId="6" captionText="FIGURE 4. Moltenojurina parva gen. et sp. nov., holotype specimen (PRE/F/20922), right forewing. A, Interpretative drawing. B, Photograph (light-mirrored, flipped horizontally)." figureDoi="http://doi.org/10.5281/zenodo.5778588" httpUri="https://zenodo.org/record/5778588/files/figure.png" pageId="6" pageNumber="556">Fig. 4</figureCitation>
)
<uri id="FFA54778FFD9FFD96EC7FA8BC9A6FAAD" pageId="6" pageNumber="556">
urn:lsid:zoobank.org:act:
<uuid id="FF9271AFFFD9FFD96FF4FA8BC9A6FAAD" pageId="6" pageNumber="556">BFCFDAB4-295D-4C86- A9AB-7441215855A4</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32E18F1FFD9FFD96EC7FAE3C8F9F9F9" pageId="6" pageNumber="556" type="materials_examined">
<paragraph id="8B8B4B7AFFD9FFD96EC7FAE3C8F9F9F9" blockId="6.[136,778,1407,2071]" pageId="6" pageNumber="556">
<materialsCitation id="3B5C4127FFD9FFD96EC7FAE3C8F9F9F9" ID-GBIF-Occurrence="3418318302" country="South Africa" county="Johannesburg" location="Specimen" municipality="Witwatersrand" pageId="6" pageNumber="556" specimenCount="1" typeStatus="holotype">
<emphasis id="B9409768FFD9FFD96EC7FAE3C8D2FA61" bold="true" box="[136,252,1407,1433]" pageId="6" pageNumber="556">
<typeStatus id="548FF5D8FFD9FFD96EC7FAE3C8D9FA61" box="[136,247,1407,1433]" pageId="6" pageNumber="556" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EEB1DA1FFD9FFD96F45FAE3C956FA61" LSID="urn:lsid:plazi:treatment:039DFA6CFFD9FFD86EC7FB4DCA79FE10:8EEB1DA1FFD9FFD96F45FAE3C956FA61" box="[266,376,1407,1433]" country="South Africa" county="Johannesburg" municipality="Witwatersrand" name="Specimen" pageId="6" pageNumber="556">Specimen</location>
PRE/F/20922 (part, preserving a wing in in ventral aspect),
<location id="8EEB1DA1FFD9FFD96FFBFA3ECB28FA44" LSID="urn:lsid:plazi:treatment:039DFA6CFFD9FFD86EC7FB4DCA79FE10:8EEB1DA1FFD9FFD96FFBFA3ECB28FA44" box="[436,774,1442,1468]" country="South Africa" county="Johannesburg" municipality="Witwatersrand" name="Evolutionary Studies Institute" pageId="6" pageNumber="556">Evolutionary Studies Institute</location>
,
<location id="8EEB1DA1FFD9FFD96EC7FA58C8D0FA26" LSID="urn:lsid:plazi:treatment:039DFA6CFFD9FFD86EC7FB4DCA79FE10:8EEB1DA1FFD9FFD96EC7FA58C8D0FA26" box="[136,254,1476,1502]" country="South Africa" county="Johannesburg" municipality="Witwatersrand" name="University" pageId="6" pageNumber="556">University</location>
of the
<collectingMunicipality id="6BEFD100FFD9FFD96F2EFA58CA26FA26" box="[353,520,1476,1502]" pageId="6" pageNumber="556">Witwatersrand</collectingMunicipality>
,
<collectingCounty id="62EA33F6FFD9FFD96C55FA58CA98FA26" box="[538,694,1476,1502]" pageId="6" pageNumber="556">Johannesburg</collectingCounty>
,
<collectingCountry id="F3230BEAFFD9FFD96C87FA58C8FDF9F9" name="South Africa" pageId="6" pageNumber="556">South Africa</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32E18F1FFD9FFD96EF2F996CA58F9BF" pageId="6" pageNumber="556" type="etymology">
<paragraph id="8B8B4B7AFFD9FFD96EF2F996CA58F9BF" blockId="6.[136,778,1407,2071]" pageId="6" pageNumber="556">
<emphasis id="B9409768FFD9FFD96EF2F996C96AF9DC" bold="true" box="[189,324,1546,1572]" pageId="6" pageNumber="556">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C3430F9FFD9FFD96C0BF997CAABF9DC" authorityName="Béthoux &amp; Anderson" authorityYear="2021" box="[580,645,1547,1572]" class="Insecta" family="Polycytellidae" genus="Moltenojurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B9409768FFD9FFD96C0BF997CAABF9DC" box="[580,645,1547,1572]" italics="true" pageId="6" pageNumber="556">parva</emphasis>
</taxonomicName>
(small in Latin) refers to the small size of the species.
</paragraph>
</subSubSection>
<subSubSection id="C32E18F1FFD9FFD96EF2F9D3CAD2F92A" pageId="6" pageNumber="556" type="diagnosis">
<paragraph id="8B8B4B7AFFD9FFD96EF2F9D3CAD2F92A" blockId="6.[136,778,1407,2071]" pageId="6" pageNumber="556">
<emphasis id="B9409768FFD9FFD96EF2F9D3C916F991" bold="true" box="[189,312,1615,1641]" pageId="6" pageNumber="556">Diagnosis.</emphasis>
Forewing: marginal band broad (opposite the 3
<superScript id="7C41E632FFD9FFD96EF1F9EEC8E2F978" attach="left" box="[190,204,1650,1664]" fontSize="6" pageId="6" pageNumber="556">rd</superScript>
quarter of wing length, accounting for about a third of total wing width); apex not deflected posteriorly; small size (forewing length about
<quantity id="4CCCE69FFFD9FFD96F8DF924CA36F92A" box="[450,536,1720,1746]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="556" unit="mm" value="3.9">3.9 mm</quantity>
in known material).
</paragraph>
</subSubSection>
<subSubSection id="C32E18F1FFD9FFD96EF2F940C9BAF89A" pageId="6" pageNumber="556" type="materials_examined">
<paragraph id="8B8B4B7AFFD9FFD96EF2F940C9BAF89A" blockId="6.[136,778,1407,2071]" pageId="6" pageNumber="556">
<emphasis id="B9409768FFD9FFD96EF2F940CA26F90D" bold="true" box="[189,520,1755,1782]" pageId="6" pageNumber="556">Type locality and horizon.</emphasis>
Birds River (locality code Bir 111; see
<bibRefCitation id="EFA5368BFFD9FFD96F27F963CA9DF8E2" author="Anderson, J. M. &amp; Anderson, H. M." box="[360,691,1791,1818]" pageId="6" pageNumber="556" pagination="39 - 59" refId="ref5642" refString="Anderson, J. M. &amp; Anderson, H. M. (1984) The fossil content of the Upper Triassic Molteno Formation, South Africa. Palaeontologia Africana, 25, 39 - 59." type="journal article" year="1984">Anderson &amp; Anderson, 1984</bibRefCitation>
),
<collectingCountry id="F3230BEAFFD9FFD96C87F963C8FDF8C5" name="South Africa" pageId="6" pageNumber="556">South Africa</collectingCountry>
; Molteno Formation; lower Carnian, Upper Triassic (
<bibRefCitation id="EFA5368BFFD9FFD96EC0F8DBC9A9F89A" author="Anderson, J. M. &amp; Anderson, H. M. &amp; Cruickshank, A. R. I." box="[143,391,1863,1890]" pageId="6" pageNumber="556" pagination="387 - 421" refId="ref5754" refString="Anderson, J. M., Anderson, H. M. &amp; Cruickshank, A. R. I. (1998) Late Triassic ecosystems of the Molteno / Lower Elliot Biome of Southern Africa. Palaeontology, 41 (3), 387 - 421." type="journal article" year="1998">
Anderson
<emphasis id="B9409768FFD9FFD96F4AF8D4C918F899" box="[261,310,1863,1889]" italics="true" pageId="6" pageNumber="556">et al</emphasis>
., 1998
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E18F1FFD9FFD96EF2F8F7CB5FFE68" pageId="6" pageNumber="556" type="description">
<paragraph id="8B8B4B7AFFD9FFD96EF2F8F7CB5FFE68" blockId="6.[136,778,1407,2071]" lastBlockId="6.[810,1452,158,1373]" pageId="6" pageNumber="556">
<emphasis id="B9409768FFD9FFD96EF2F8F7C961F87D" bold="true" box="[189,335,1899,1925]" pageId="6" pageNumber="556">Description.</emphasis>
A right forewing, posterior area incomplete, clavus missing, and, according to the proposed reconstruction, area between anterior wing margin and ScP missing; length
<quantity id="4CCCE69FFFD9FFD96F3EF844C9E7F80A" box="[369,457,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="556" unit="mm" value="3.3">3.3 mm</quantity>
as preserved (
<emphasis id="B9409768FFD9FFD96C20F844CAA7F809" box="[623,649,2008,2033]" italics="true" pageId="6" pageNumber="556">ca</emphasis>
.
<quantity id="4CCCE69FFFD9FFD96CD6F844CADFF80A" box="[665,753,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="556" unit="mm" value="3.9">3.9 mm</quantity>
if complete), width about
<quantity id="4CCCE69FFFD9FFD96FDDF860C9C6F7EE" box="[402,488,2044,2070]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="556" unit="mm" value="1.1">1.1 mm</quantity>
if complete; area anterior to RA broad (
<quantity id="4CCCE69FFFD9FFD96D97FF02CC18FF41" box="[984,1078,158,185]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="556" unit="mm" value="0.3">0.3 mm</quantity>
opposite the presumed end of ScP), with 4 rows of cells at best, and retaining 2 rows of cells until shortly before the apex; area between RA and MP+CuA
<emphasis id="B9409768FFD9FFD96DD8FE97CB95FEDC" box="[919,955,267,292]" italics="true" pageId="6" pageNumber="556">ant</emphasis>
with 2 veins, zigzagging and without clear origin; area between MP+CuA
<emphasis id="B9409768FFD9FFD96AC0FEB3CC9DFEB0" box="[1167,1203,303,328]" italics="true" pageId="6" pageNumber="556">ant</emphasis>
and CuA
<emphasis id="B9409768FFD9FFD96B69FEB3CD7BFEB0" box="[1318,1365,303,328]" italics="true" pageId="6" pageNumber="556">post</emphasis>
with 2 zigzagging veins; in its posterior section, marginal band broad.
</paragraph>
</subSubSection>
<subSubSection id="C32E18F1FFD9FFD86D10FE06CA79FE10" lastPageId="7" lastPageNumber="557" pageId="6" pageNumber="556" type="discussion">
<paragraph id="8B8B4B7AFFD9FFD96D10FE06CBCDFB98" blockId="6.[810,1452,158,1373]" pageId="6" pageNumber="556">
<emphasis id="B9409768FFD9FFD96D10FE06CBF0FE4C" bold="true" box="[863,990,410,436]" pageId="6" pageNumber="556">Discussion.</emphasis>
Because of its incompleteness and the symmetry characterizing
<taxonomicName id="4C3430F9FFD9FFD96A0EFE22CCCFFE20" authorityName="Martynov" authorityYear="1938" box="[1089,1249,446,472]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
forewings, it was not straightforward to orientate the specimen. We rested on the fact that the clavus is commonly delimited by a series of rectilinear, close and parallel veins in
<taxonomicName id="4C3430F9FFD9FFD96B4AFDB6CD86FDBC" authorityName="Martynov" authorityYear="1938" box="[1285,1448,554,580]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
. Provided that it is not the case in the preserved area which could have been interpreted as the clavus, we interpreted it as the area anterior to RA instead. Additionally, veinlike elements preserved in the corresponding area display a zigzagging course, known to be the case for similar elements preserved in the area anterior to RA in
<taxonomicName id="4C3430F9FFD9FFD96B7BFC9ECD82FCE4" authorityName="Tillyard" authorityYear="1922" box="[1332,1452,770,796]" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="B9409768FFD9FFD96B7BFC9ECD82FCE4" box="[1332,1452,770,796]" italics="true" pageId="6" pageNumber="556">Polycytella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3430F9FFD9FFD96D14FCBACC60FCB8" authorityName="Martins-Neto &amp; Gallego" authorityYear="2001" box="[859,1102,806,832]" class="Insecta" family="Glosselytridae" genus="Argentinoglosselytrina" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="B9409768FFD9FFD96D14FCBACC60FCB8" box="[859,1102,806,832]" italics="true" pageId="6" pageNumber="556">Argentinoglosselytrina</emphasis>
</taxonomicName>
. In a similar line of reasoning, the lack of a rectilinear vein interpretable as ScP suggests that the specimen was broken along this vein, a damage not uncommon in fossil wings of
<taxonomicName id="4C3430F9FFD9FFD96A9FFC0ECD5EFC54" authorityName="Martynov" authorityYear="1938" box="[1232,1392,914,940]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
(see
<bibRefCitation id="EFA5368BFFD9FFD96D65FC2BCB85FC28" author="Hong, Y. C." box="[810,939,950,977]" pageId="6" pageNumber="556" pagination="271 - 280" refId="ref6256" refString="Hong, Y. C. (2007) Discovery of the fossil glosselytrids (Insecta: Glosselytrodea) from Shaanxi, China. Acta Entomologica Sinica, 50 (3), 271 - 280." type="journal article" year="2007">Hong, 2007</bibRefCitation>
; Rasnitsyn &amp; Aristov in
<bibRefCitation id="EFA5368BFFD9FFD96AFBFC2ACDA7FC28" author="Aristov, D. S. &amp; Bashkuev, A. &amp; Golubev, V. K. &amp; Gorochov, A. V. &amp; Karasev, E. V. &amp; Kopylov, D. S. &amp; Ponomarenko, A. B. &amp; Rasnitsyn, A. P. &amp; Rasnitsyn, D. A. &amp; Sinitshenkova, N. D. &amp; Sukatsheva, I. D. &amp; Vassilenko, D. V." box="[1204,1417,950,977]" pageId="6" pageNumber="556" pagination="641 - 832" refId="ref5804" refString="Aristov, D. S., Bashkuev, A., Golubev, V. K., Gorochov, A. V., Karasev, E. V., Kopylov, D. S., Ponomarenko, A. B., Rasnitsyn, A. P., Rasnitsyn, D. A., Sinitshenkova, N. D., Sukatsheva, I. D. &amp; Vassilenko, D. V. (2013) Fossil insects of the Middle and Upper Permian of European Russia. Paleontological Journal, 47 (7), 641 - 832. https: // doi. org / 10.1134 / S 0031030113070010" type="journal article" year="2013">
Aristov
<emphasis id="B9409768FFD9FFD96B44FC2BCD15FC28" box="[1291,1339,950,976]" italics="true" pageId="6" pageNumber="556">et al</emphasis>
., 2013
</bibRefCitation>
). A tentative reconstruction was elaborated accordingly (
<figureCitation id="130F57FFFFD9FFD96B12FC46CDB0FC0C" box="[1373,1438,986,1012]" captionStart="FIGURE 4" captionStartId="6.[810,909,1974,1998]" captionTargetBox="[809,1452,1435,1950]" captionTargetId="figure-692@6.[809,1452,1430,1950]" captionTargetPageId="6" captionText="FIGURE 4. Moltenojurina parva gen. et sp. nov., holotype specimen (PRE/F/20922), right forewing. A, Interpretative drawing. B, Photograph (light-mirrored, flipped horizontally)." figureDoi="http://doi.org/10.5281/zenodo.5778588" httpUri="https://zenodo.org/record/5778588/files/figure.png" pageId="6" pageNumber="556">Fig. 4</figureCitation>
). It implies that the area anterior to RA is very broad, ensuring the assignment of the specimen to the
<taxonomicName id="4C3430F9FFD9FFD96ABFFBBECDA6FBC4" authorityName="Martynova" authorityYear="1952" box="[1264,1416,1058,1084]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
as diagnosed above.
</paragraph>
<paragraph id="8B8B4B7AFFD9FFD86D10FBF6CA21FEF7" blockId="6.[810,1452,158,1373]" lastBlockId="7.[136,778,173,488]" lastPageId="7" lastPageNumber="557" pageId="6" pageNumber="556">
The new material differs from any known
<taxonomicName id="4C3430F9FFD9FFD96D65FB12CBE4FB50" authorityName="Martynova" authorityYear="1952" box="[810,970,1166,1192]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
by its small size. As a matter of fact, to our knowledge, it is the smallest
<taxonomicName id="4C3430F9FFD9FFD96AE9FB2ECD60FB34" authorityName="Martynov" authorityYear="1938" box="[1190,1358,1202,1228]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
to date. More decisively, the marginal band is broader than in other
<taxonomicName id="4C3430F9FFD9FFD96D3EFB66CC33FAEC" authorityName="Martynov" authorityYear="1938" box="[881,1053,1274,1300]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="order">Glosselytrodea</taxonomicName>
: opposite the 3
<superScript id="7C41E632FFD9FFD96A98FB66CCCBFAF0" attach="left" box="[1239,1253,1274,1288]" fontSize="6" pageId="6" pageNumber="556">rd</superScript>
quarter of wing length, the marginal band (anterior and posterior sections) accounts for about a quarter of wing width in
<taxonomicName id="4C3430F9FFD9FFD86B62FADEC896FF3F" class="Insecta" family="Glosselytridae" genus="Polycytella" kingdom="Animalia" lastPageId="7" lastPageNumber="557" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9409768FFD9FFD96B62FADECD85FAA4" box="[1325,1451,1346,1372]" italics="true" pageId="6" pageNumber="556">Polycytella</emphasis>
spp.
</taxonomicName>
, while it accounts for more than a third in the new material. These traits justify the erection of a new genus and, incidentally, of a new species.
</paragraph>
<caption id="DF4B1BF2FFD9FFD96D65F82ACDB1F7EE" ID-DOI="http://doi.org/10.5281/zenodo.5778588" ID-Zenodo-Dep="5778588" httpUri="https://zenodo.org/record/5778588/files/figure.png" pageId="6" pageNumber="556" startId="6.[810,909,1974,1998]" targetBox="[809,1452,1435,1950]" targetPageId="6">
<paragraph id="8B8B4B7AFFD9FFD96D65F82ACDB1F7EE" blockId="6.[810,1452,1974,2070]" pageId="6" pageNumber="556">
<emphasis id="B9409768FFD9FFD96D65F82ACB84F836" bold="true" box="[810,938,1974,1998]" pageId="6" pageNumber="556">FIGURE 4.</emphasis>
<taxonomicName id="4C3430F9FFD9FFD96DFBF82ACCA2F836" authority="Béthoux &amp; Anderson, 2021" authorityName="Béthoux &amp; Anderson" authorityYear="2021" box="[948,1164,1974,1998]" class="Insecta" family="Polycytellidae" genus="Moltenojurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="parva" status="gen. et sp. nov.">
<emphasis id="B9409768FFD9FFD96DFBF82ACCA2F836" box="[948,1164,1974,1998]" italics="true" pageId="6" pageNumber="556">Moltenojurina parva</emphasis>
</taxonomicName>
<emphasis id="B9409768FFD9FFD96AD8F82ACD6FF836" bold="true" box="[1175,1345,1974,1998]" pageId="6" pageNumber="556">
<taxonomicNameLabel id="A2732A13FFD9FFD96AD8F82ACD6FF836" box="[1175,1345,1974,1998]" pageId="6" pageNumber="556" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype specimen (PRE/F/20922), right forewing.
<emphasis id="B9409768FFD9FFD96AB5F846CD22F80A" bold="true" box="[1274,1292,2010,2034]" pageId="6" pageNumber="556">A</emphasis>
, Interpretative drawing.
<emphasis id="B9409768FFD9FFD96DC6F862CBB4F7EE" bold="true" box="[905,922,2046,2070]" pageId="6" pageNumber="556">B</emphasis>
, Photograph (light-mirrored, flipped horizontally).
</paragraph>
</caption>
<paragraph id="8B8B4B7AFFD8FFD86EF2FE85CA79FE10" blockId="7.[136,778,173,488]" pageId="7" pageNumber="557">
General similarity between the new material and
<taxonomicName id="4C3430F9FFD8FFD86EC7FEA2CA5DFEAF" authority="Martynova, 1943" authorityName="Martynova" authorityYear="1943" box="[136,627,317,344]" class="Insecta" family="Jurinidae" genus="Mesojurina" kingdom="Animalia" order="Glosselytrodea" pageId="7" pageNumber="557" phylum="Arthropoda" rank="species" species="sogjutensis">
<emphasis id="B9409768FFD8FFD86EC7FEA2C9B8FEAF" box="[136,406,318,343]" italics="true" pageId="7" pageNumber="557">Mesojurina sogjutensis</emphasis>
<bibRefCitation id="EFA5368BFFD8FFD86FE7FEA2CA5DFEAF" author="Martynova, O. M." box="[424,627,317,344]" pageId="7" pageNumber="557" pagination="312 - 313" refId="ref6646" refString="Martynova, O. M. (1943) Glosselytrodea from the Jurassic shales of the coal bed Sogjuta. Comptes Rendus de l'Academie des Sciences de l'URSS, 39 (7), 312 - 313." type="journal article" year="1943">Martynova, 1943</bibRefCitation>
</taxonomicName>
is regarded as due to the small size characterizing both species. Notably,
<taxonomicName id="4C3430F9FFD8FFD86EBDFE1AC9D4FE67" authorityName="Martynova" authorityYear="1943" box="[242,506,390,415]" class="Insecta" family="Jurinidae" genus="Mesojurina" kingdom="Animalia" order="Glosselytrodea" pageId="7" pageNumber="557" phylum="Arthropoda" rank="species" species="sogjutensis">
<emphasis id="B9409768FFD8FFD86EBDFE1AC9D4FE67" box="[242,506,390,415]" italics="true" pageId="7" pageNumber="557">Mesojurina sogjutensis</emphasis>
</taxonomicName>
has an area anterior to RA comparatively narrow, this precluding its assignment to the
<taxonomicName id="4C3430F9FFD8FFD86E81FE52C940FE10" authorityName="Martynova" authorityYear="1952" box="[206,366,462,488]" class="Insecta" family="Polycytellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Glosselytrodea" pageId="7" pageNumber="557" phylum="Arthropoda" rank="family">Polycytellidae</taxonomicName>
as diagnosed above.
</paragraph>
</subSubSection>
</treatment>
</document>