treatments-xml/data/03/8A/87/038A87DAA77CFF9D0788FC4FFC7D0B86.xml

<document id="FEB05E89C7C41358175148858635EF9F" ID-CLB-Dataset="20416" ID-DOI="10.5852/ejt.2022.798.1671" ID-GBIF-Dataset="dc185c1a-2bc2-48f1-bbdd-e0ae42f852f1" ID-ISSN="2118-9773" ID-Zenodo-Dep="6323040" ID-ZooBank="F136E044-62C8-4FB3-8160-7DAE663D9600" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646225490206" checkinUser="felipe" docAuthor="Houben, Albrecht M., Monnens, Marlies, Proesmans, Willem &amp; Artois, Tom J." docDate="2022" docId="038A87DAA77CFF9D0788FC4FFC7D0B86" docLanguage="en" docName="ejt-798_70-102.pdf" docOrigin="European Journal of Taxonomy 798" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1671" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Faunulus nielsi Houben, Proesmans &amp; Artois 2022, gen. et sp. nov." docType="treatment" docUuid="E8B0D042-259A-427D-8F29-5DB6ED8C8411" docUuidSource="ZooBank" docVersion="9" lastPageNumber="77" masterDocId="FFB3FFA2A779FF9A0626FF8AFFA50F38" masterDocTitle="Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus" masterLastPageNumber="102" masterPageNumber="70" pageNumber="75" updateTime="1699316596528" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B5AE858CB33B9713760E87AFE8039E8D">Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus</mods:title>
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<mods:namePart id="FBED98E133EADE13D905DF27E4F04820">Houben, Albrecht M.</mods:namePart>
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<mods:affiliation id="D8259E2E60F75B639924077F524C3AAA">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
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<mods:namePart id="D2E6C93C948360B5E981DFA3EBF8291B">Monnens, Marlies</mods:namePart>
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<mods:affiliation id="33B0DD442395AEB58725B2325F58B4C9">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
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<mods:namePart id="779FAD7402431BCB16A1652D30557679">Proesmans, Willem</mods:namePart>
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<mods:affiliation id="631D5177E4E9AF152417AD98830378F3">Agroécologie, INRAE, 17 Rue Sully, F- 21065 Dijon, France.</mods:affiliation>
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<mods:namePart id="7FFCCECCA69D55FCD355D2F18A84A142">Artois, Tom J.</mods:namePart>
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<mods:affiliation id="62C451FF128E62CC54B09F097057232E">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
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<mods:date id="73B80DA04812F23DCF32605AFAF7D2C7">2022</mods:date>
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<taxonomicName id="4C234D4FA77CFF9F0788FC4FFC6B0CD8" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[430,974,965,992]" family="Typhloplanidae" genus="Faunulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="5" pageNumber="75" phylum="Platyhelminthes" rank="species" species="nielsi" status="gen. et sp. nov.">
<emphasis id="B957EADEA77CFF9F0788FC4FFDCA0CD8" bold="true" box="[430,623,965,992]" italics="true" pageId="5" pageNumber="75">Faunulus nielsi</emphasis>
Houben, Proesmans &amp; Artois
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urn:lsid:zoobank.org:act: 
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<figureCitation id="13182A49A77CFF9F04D1FB86FC990B1E" box="[759,828,1036,1062]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="5" pageNumber="75">Fig. 2</figureCitation>
</paragraph>
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<heading id="D0D481A0A77CFF9F069BFBC7FE920B5F" bold="true" box="[189,311,1101,1127]" fontSize="11" level="3" pageId="5" pageNumber="75" reason="3">
<emphasis id="B957EADEA77CFF9F069BFBC7FE920B5F" bold="true" box="[189,311,1101,1127]" pageId="5" pageNumber="75">Diagnosis</emphasis>
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<paragraph id="8B9C36CCA77CFF9F069BFBF6FD780BAF" blockId="5.[189,733,1148,1175]" box="[189,733,1148,1175]" pageId="5" pageNumber="75">Provisionally the same diagnosis as the genus.</paragraph>
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<subSubSection id="C3396547A77CFF9F069BFB34FB630A3F" box="[189,1222,1214,1287]" pageId="5" pageNumber="75" type="etymology">
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<heading id="D0D481A0A77CFF9F069BFB34FEE00BE0" bold="true" box="[189,325,1214,1240]" fontSize="11" level="3" pageId="5" pageNumber="75" reason="3">
<emphasis id="B957EADEA77CFF9F069BFB34FEE00BE0" bold="true" box="[189,325,1214,1240]" pageId="5" pageNumber="75">Etymology</emphasis>
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<paragraph id="8B9C36CCA77CFF9F069BFB66FB630A3F" blockId="5.[189,1222,1260,1287]" box="[189,1222,1260,1287]" pageId="5" pageNumber="75">The species epithet is dedicated to Niels Houben, son of the first author (A.M. Houben).</paragraph>
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<emphasis id="B957EADEA77CFF9F069BFAA7FE080A7F" bold="true" box="[189,429,1325,1351]" pageId="5" pageNumber="75">Material examined</emphasis>
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<materialsCitation id="3B4B3C91A77CFF9F06CCFAD6FD2D0AD8" ID-GBIF-Occurrence="3695141321" collectedFrom="forest soil of Pinus, Sambucus, Corylus, and Cornus" collectingDate="2011-08-29" collectingMethod="live observations and sagittal sections" collectionCode="FMNH" collectorName="A. M. Houben &amp; W. Proesmans" country="Austria" latitude="47.859997" location="AUSTRIA" longLatPrecision="18" longitude="14.652499" municipality="Weyer" pageId="5" pageNumber="75" specimenCode="KV.686" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5498886EA77CFF9F06CCFAD6FEF80A4E" box="[234,349,1372,1398]" pageId="5" pageNumber="75" type="holotype">Holotype</typeStatus>
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• 
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, 
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; 
<locationDeviation id="9A2E2FEEA77CFF9F0589FAF5FB630AA2" box="[943,1222,1407,1434]" pageId="5" pageNumber="75">Kreuzberg near Weyer</locationDeviation>
; 
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, 
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; 
<date id="FF9D100CA77CFF9F0742FA28FDAB0A84" box="[356,526,1442,1469]" pageId="5" pageNumber="75" value="2011-08-29">
<collectingDate id="EFD9E9E4A77CFF9F0742FA28FDAB0A84" box="[356,526,1442,1469]" pageId="5" pageNumber="75" value="2011-08-29">29 Aug. 2011</collectingDate>
</date>
; 
<collectorName id="26D6531AA77CFF9F043AFA29FD660A84" box="[540,707,1442,1469]" pageId="5" pageNumber="75">A.M. Houben</collectorName>
and 
<collectorName id="26D6531AA77CFF9F0525FA29FC0B0A85" box="[771,942,1443,1469]" pageId="5" pageNumber="75">W. Proesmans</collectorName>
leg.; 
<collectedFrom id="383F7531A77CFF9F05DFFA28FE170AD8" pageId="5" pageNumber="75">
forest soil of 
<emphasis id="B957EADEA77CFF9F0287FA29FB410A84" box="[1185,1252,1443,1468]" italics="true" pageId="5" pageNumber="75">Pinus</emphasis>
, 
<emphasis id="B957EADEA77CFF9F02D1FA28FACA0A84" box="[1271,1391,1442,1468]" italics="true" pageId="5" pageNumber="75">Sambucus</emphasis>
, 
<emphasis id="B957EADEA77CFF9F069BFA4FFEBE0AE7" box="[189,283,1477,1503]" italics="true" pageId="5" pageNumber="75">Corylus</emphasis>
, and 
<emphasis id="B957EADEA77CFF9F077DFA4CFE170AD8" box="[347,434,1478,1504]" italics="true" pageId="5" pageNumber="75">Cornus</emphasis>
</collectedFrom>
; 
<specimenCode id="DB859EB7A77CFF9F07E4FA4FFD850AE7" box="[450,544,1477,1503]" pageId="5" pageNumber="75">KV.686</specimenCode>
; 
<collectionCode id="ED32AE09A77CFF9F040DFA4CFD210AD8" box="[555,644,1478,1504]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="75" type="Museum">FMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9C36CCA77CFF9F06CCF98DFD9C0950" blockId="5.[189,1398,1543,1640]" pageId="5" pageNumber="75">
<materialsCitation id="3B4B3C91A77CFF9F06CCF98DFD9C0950" ID-GBIF-Occurrence="3695141315" collectedFrom="forest soil of Pinus, Sambucus, Corylus, and Cornus" collectingDate="2011-08-29" collectingMethod="live observations, one of which whole mounted" collectionCode="HU" collectorName="A. M. Houben &amp; W. Proesmans" country="Austria" latitude="47.859997" location="AUSTRIA" longLatPrecision="18" longitude="14.652499" municipality="Weyer" pageId="5" pageNumber="75" specimenCode="821" specimenCount="2" typeStatus="paratype">
<emphasis id="B957EADEA77CFF9F06CCF98DFECC0919" bold="true" box="[234,361,1543,1569]" pageId="5" pageNumber="75">
<typeStatus id="5498886EA77CFF9F06CCF98DFECC0919" box="[234,361,1543,1569]" pageId="5" pageNumber="75" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F334765CA77CFF9F069BF9A0FE9F097C" box="[189,314,1578,1604]" name="Austria" pageId="5" pageNumber="75">AUSTRIA</collectingCountry>
• 
<specimenCount id="9D25FD45A77CFF9F077BF9A0FE1F097D" box="[349,442,1578,1605]" pageId="5" pageNumber="75">2 specs</specimenCount>
, 
<collectingMethod id="52624EDBA77CFF9F07EDF9A0FBBF097C" box="[459,1050,1578,1605]" pageId="5" pageNumber="75">live observations, one of which whole mounted</collectingMethod>
; same collection data as for holotype; paratype no. 
<specimenCode id="DB859EB7A77CFF9F07EDF9C4FE5F0950" box="[459,506,1614,1640]" pageId="5" pageNumber="75">821</specimenCode>
; 
<collectionCode id="ED32AE09A77CFF9F0421F9C4FD900950" box="[519,565,1614,1640]" pageId="5" pageNumber="75">HU</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
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<paragraph id="8B9C36CCA77CFF9F069BF905FEEA0991" blockId="5.[189,335,1679,1705]" box="[189,335,1679,1705]" pageId="5" pageNumber="75">
<heading id="D0D481A0A77CFF9F069BF905FEEA0991" bold="true" box="[189,335,1679,1705]" fontSize="11" level="3" pageId="5" pageNumber="75" reason="3">
<emphasis id="B957EADEA77CFF9F069BF905FEEA0991" bold="true" box="[189,335,1679,1705]" pageId="5" pageNumber="75">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA77CFF9F069BF937FE8C08F6" blockId="5.[189,1401,1725,1998]" pageId="5" pageNumber="75">
Specimens about 
<quantity id="4CDB9B29A77CFF9F07B4F937FE7C09E0" box="[402,473,1725,1752]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="75" unit="mm" value="1.0">1 mm</quantity>
long. The body shape is pointed anteriorly and blunt to rounded posteriorly (see 
<figureCitation id="13182A49A77CFF9F06D3F96AFEF609C3" box="[245,339,1760,1787]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="5" pageNumber="75">Fig. 2A</figureCitation>
). On free-swimming specimens, a small tail can be observed. Both anterior and posterior ends possess very typical, strongly eosinophilic glands that appear rod-shaped, like big rhabdites, and are situated more or less parallel to each other. Rostrally, rhabdite glands (
<figureCitation id="13182A49A77CFF9F0273F8ADFB1D0879" box="[1109,1208,1831,1857]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="5" pageNumber="75">Fig. 2A</figureCitation>
: ar) are present and arranged in two groups. The distal part of the epidermis is completely filled with coarse, strongly eosinophilic secretions, which probably are dermal rhabdites (
<figureCitation id="13182A49A77CFF9F0583F8E7FBA008BF" box="[933,1029,1901,1927]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="5" pageNumber="75">Fig. 2B</figureCitation>
: dr?). The protonephridiopores were not observed with certainty. The slightly forward slanted rosulate pharynx (
<figureCitation id="13182A49A77CFF9F0257F81AFB750893" box="[1137,1232,1936,1963]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="5" pageNumber="75">Fig. 2A</figureCitation>
: ph) is located centrally.
</paragraph>
<paragraph id="8B9C36CCA77FFF9C069BFE83FAFE0E7E" blockId="6.[189,1398,265,327]" pageId="6" pageNumber="76">
The gonopore (
<figureCitation id="13182A49A77FFF9C074AFE83FE4E0E1B" box="[364,491,265,291]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="6" pageNumber="76">Fig. 2A–B</figureCitation>
: gp) is situated at ±65% of the body and connected to a genital atrium (
<figureCitation id="13182A49A77FFF9C0329FE83FF710E7F" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="6" pageNumber="76">Fig. 2A– B</figureCitation>
: ga) that is surrounded by muscles, although their orientation could not be observed with certainty.
</paragraph>
<paragraph id="8B9C36CCA77FFF9D069BFEE4FF4D0ECF" blockId="6.[189,1398,366,428]" lastBlockId="7.[188,1400,265,503]" lastPageId="7" lastPageNumber="77" pageId="6" pageNumber="76">
The paired, differently sized testes (
<figureCitation id="13182A49A77FFF9C0455FEE4FD700EB0" box="[627,725,366,392]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="6" pageNumber="76">Fig. 2A</figureCitation>
: t) lie in front of the pharynx and ventral to the paired vitellaria (
<figureCitation id="13182A49A77FFF9C0713FE1BFE100E93" box="[309,437,401,427]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="6" pageNumber="76">Fig. 2A–B</figureCitation>
: vi). One testis is relatively large, while the other is barely visible. The paired vasa deferentia (
<figureCitation id="13182A49A77EFF9D0764FE83FE3A0E1B" box="[322,415,265,291]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2B</figureCitation>
: vde) enter the copulatory organ laterally (
<figureCitation id="13182A49A77EFF9D05ABFE83FC4E0E1B" box="[909,1003,265,291]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2A</figureCitation>
: co). Two layers of spiral muscles surround the 50 µm long copulatory organ, which bears an intracapsular seminal vesicle (
<figureCitation id="13182A49A77EFF9D02D7FEA6FAD60E7E" box="[1265,1395,300,326]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2B–C</figureCitation>
: vs) and a cirrus (
<figureCitation id="13182A49A77EFF9D07A7FEC5FDA40E51" box="[385,513,335,361]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2B–C</figureCitation>
: cir). Large, coarse-grained, extracapsular eosinophilic glands (
<figureCitation id="13182A49A77EFF9D02C1FEC5FAE10E51" box="[1255,1348,335,361]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2B</figureCitation>
: gg) are associated with the copulatory organ, although the point of entry remains uncertain. The 30 µm long cirrus is more or less straight and covered with spines in its distal part (see 
<figureCitation id="13182A49A77EFF9D0263FE1CFB670E88" box="[1093,1218,406,432]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2B–F</figureCitation>
: cir). The most distal part of the cirrus bears the larger spines and is enveloped in a conical, sclerotised pouch (
<figureCitation id="13182A49A77EFF9D0331FE33FAD10EEB" box="[1303,1396,441,467]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2C</figureCitation>
: sp).
</paragraph>
<caption id="DF5C6644A77FFF9C069AF802FC0F08D0" ID-DOI="http://doi.org/10.5281/zenodo.6323046" ID-Zenodo-Dep="6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="6" pageNumber="76" startId="6.[188,231,1928,1954]" targetBox="[208,1398,526,1886]" targetPageId="6">
<paragraph id="8B9C36CCA77FFF9C069AF802FC0F08D0" blockId="6.[188,1399,1928,2025]" pageId="6" pageNumber="76">
<emphasis id="B957EADEA77FFF9C069AF802FEA9089A" bold="true" box="[188,268,1928,1954]" pageId="6" pageNumber="76">Fig. 2.</emphasis>
<taxonomicName id="4C234D4FA77FFF9C0733F802FC95089A" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[277,816,1928,1954]" family="Typhloplanidae" genus="Faunulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="6" pageNumber="76" phylum="Platyhelminthes" rank="species" species="nielsi" status="gen. et sp. nov.">
<emphasis id="B957EADEA77FFF9C0733F802FE68089A" box="[277,461,1928,1954]" italics="true" pageId="6" pageNumber="76">Faunulus nielsi</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A77FFF9C051FF802FC49089A" box="[825,1004,1928,1954]" pageId="6" pageNumber="76" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="B957EADEA77FFF9C05D2F802FBAD089A" bold="true" box="[1012,1032,1928,1954]" pageId="6" pageNumber="76">A</emphasis>
. Internal organisation of a live specimen, 
<emphasis id="B957EADEA77FFF9C071FF821FEE808FD" bold="true" box="[313,333,1963,1989]" pageId="6" pageNumber="76">B</emphasis>
. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). 
<emphasis id="B957EADEA77FFF9C069AF844FF5708D0" bold="true" box="[188,242,1998,2024]" pageId="6" pageNumber="76">C–F</emphasis>
. Male copulatory organ of the paratype (paratype no. 821).
</paragraph>
</caption>
<paragraph id="8B9C36CCA77EFF9D069BFD94FB440D63" blockId="7.[189,1398,542,603]" pageId="7" pageNumber="77">
The female duct (
<figureCitation id="13182A49A77EFF9D07AAFD94FE4F0D00" box="[396,490,542,568]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2B</figureCitation>
: fd) is relatively long and lined with a high, nuclear epithelium. Proximally it widens into a seminal receptacle (
<figureCitation id="13182A49A77EFF9D046CFDCBFD0D0D63" box="[586,680,577,603]" captionStart="Fig" captionStartId="6.[188,231,1928,1954]" captionTargetBox="[208,1398,526,1886]" captionTargetId="figure-84@6.[189,1398,493,1889]" captionTargetPageId="6" captionText="Fig. 2. Faunulus nielsi Houben, Proesmans &amp; Artois gen. et sp. nov. A. Internal organisation of a live specimen, B. Reconstruction of reproductive organs based on sagittal sections of the holotype (KV.686). C–F. Male copulatory organ of the paratype (paratype no. 821)." figureDoi="http://doi.org/10.5281/zenodo.6323046" httpUri="https://zenodo.org/record/6323046/files/figure.png" pageId="7" pageNumber="77">Fig. 2B</figureCitation>
: rs) that receives the oviduct and the vitelloduct.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A77EFF9D069BFD08FC7D0B86" pageId="7" pageNumber="77" type="discussion">
<paragraph id="8B9C36CCA77EFF9D069BFD08FEE70DA4" blockId="7.[189,322,642,668]" box="[189,322,642,668]" pageId="7" pageNumber="77">
<heading id="D0D481A0A77EFF9D069BFD08FEE70DA4" bold="true" box="[189,322,642,668]" fontSize="11" level="3" pageId="7" pageNumber="77" reason="3">
<emphasis id="B957EADEA77EFF9D069BFD08FEE70DA4" bold="true" box="[189,322,642,668]" pageId="7" pageNumber="77">Discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA77EFF9D069BFD3BFAC90B76" blockId="7.[187,1400,689,1103]" pageId="7" pageNumber="77">
<taxonomicName id="4C234D4FA77EFF9D069BFD3BFD740DF3" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[189,721,689,715]" family="Typhloplanidae" genus="Faunulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="7" pageNumber="77" phylum="Platyhelminthes" rank="species" species="nielsi" status="gen. et sp. nov.">
<emphasis id="B957EADEA77EFF9D069BFD3BFED10DF3" box="[189,372,689,715]" italics="true" pageId="7" pageNumber="77">Faunulus nielsi</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A77EFF9D04FEFD3BFC230DF3" box="[728,902,689,715]" pageId="7" pageNumber="77" rank="species">gen. et sp. nov.</taxonomicNameLabel>
strongly resembles species of 
<emphasis id="B957EADEA77EFF9D02CCFD3BFAD30DF3" box="[1258,1398,689,715]" italics="true" pageId="7" pageNumber="77">Adenocerca</emphasis>
<bibRefCitation id="EFB24B3DA77EFF9D069BFD5EFED10DD6" author="Reisinger E." box="[189,372,724,750]" pageId="7" pageNumber="77" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
because of the combination of following features: centrally positioned pharynx, presence of tail glands and a spiny cirrus, and absence of a bursa (
<bibRefCitation id="EFB24B3DA77EFF9D0577FD7DFB280C29" author="Van Steenkiste N. &amp; Davison P. &amp; Artois T." box="[849,1165,759,785]" pageId="7" pageNumber="77" pagination="285 - 291" refId="ref16235" refString="Van Steenkiste N., Davison P. &amp; Artois T. 2010. Bryoplana xerophila n. g. n. sp., a new limnoterrestrial microturbellarian (Platyhelminthes, Typhloplanidae, Protoplanellinae) from epilithic mosses, with notes on its ecology. Zoological Science 27 (3): 285 - 291. https: // doi. org / 10.2108 / zsj. 27.285" type="journal article" year="2010">
Van Steenkiste 
<emphasis id="B957EADEA77EFF9D022CFD72FBE30C29" box="[1034,1094,759,785]" italics="true" pageId="7" pageNumber="77">et al.</emphasis>
2010
</bibRefCitation>
). However, 
<taxonomicName id="4C234D4FA77EFF9D033FFD72FDBE0C0C" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" family="Typhloplanidae" genus="Faunulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="7" pageNumber="77" phylum="Platyhelminthes" rank="species" species="nielsi" status="gen. et sp. nov.">
<emphasis id="B957EADEA77EFF9D033FFD72FAD30C29" box="[1305,1398,759,785]" italics="true" pageId="7" pageNumber="77">F. nielsi</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A77EFF9D0403FC91FD790C0D" box="[549,732,795,821]" pageId="7" pageNumber="77" rank="species">gen. et sp. nov.</taxonomicNameLabel>
differs from all species of 
<emphasis id="B957EADEA77EFF9D020EFC90FB110C0C" box="[1064,1204,794,820]" italics="true" pageId="7" pageNumber="77">Adenocerca</emphasis>
in having testes that lie rostral to the pharynx. In all species of 
<emphasis id="B957EADEA77EFF9D04D7FCB7FCD90C6F" box="[753,892,829,855]" italics="true" pageId="7" pageNumber="77">Adenocerca</emphasis>
, they are situated posterior to the pharynx and lie next to the copulatory organ. Moreover, there is a difference in size between the two testes of 
<taxonomicName id="4C234D4FA77EFF9D0699FC0FFDD90CA6" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[191,636,900,927]" family="Typhloplanidae" genus="Faunulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="7" pageNumber="77" phylum="Platyhelminthes" rank="species" species="nielsi" status="gen. et sp. nov.">
<emphasis id="B957EADEA77EFF9D0699FC0FFEB90CA6" box="[191,284,900,926]" italics="true" pageId="7" pageNumber="77">F. nielsi</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A77EFF9D04A3FC0FFC9C0CA7" box="[645,825,901,927]" pageId="7" pageNumber="77" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, which was never reported for 
<emphasis id="B957EADEA77EFF9D028EFC0EFA910CA6" box="[1192,1332,900,926]" italics="true" pageId="7" pageNumber="77">Adenocerca</emphasis>
. This size difference is consistent for all observed specimens. All species of 
<emphasis id="B957EADEA77EFF9D05C1FC2DFBD60CF9" box="[999,1139,935,961]" italics="true" pageId="7" pageNumber="77">Adenocerca</emphasis>
either have no dermal rhabdites (
<emphasis id="B957EADEA77EFF9D071BFC40FDE30CDC" box="[317,582,970,996]" italics="true" pageId="7" pageNumber="77">Adenocerca teshirogii</emphasis>
Kolasa, 1981) or dermal rhabdites occurring throughout the whole epidermis. Conversely, what we assume are dermal rhabdites occur solely in the caudal body part of 
<taxonomicName id="4C234D4FA77EFF9D0699FB98FD270B13" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" box="[191,642,1041,1067]" family="Typhloplanidae" genus="Faunulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabdocoela" pageId="7" pageNumber="77" phylum="Platyhelminthes" rank="species" species="nielsi" status="gen. et sp. nov.">
<emphasis id="B957EADEA77EFF9D0699FB98FEBB0B13" box="[191,286,1041,1067]" italics="true" pageId="7" pageNumber="77">F. nielsi</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A77EFF9D04AAFB9BFCE40B13" box="[652,833,1041,1067]" pageId="7" pageNumber="77" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Because of the unique combination of features present in the new species described, we find it justified to erect a new genus within ‘Typhloplanidae’.
</paragraph>
<paragraph id="8B9C36CCA77EFF9D069BFBFCFE950BA8" blockId="7.[189,304,1142,1168]" box="[189,304,1142,1168]" pageId="7" pageNumber="77">
<heading id="D0D481A0A77EFF9D069BFBFCFE950BA8" bold="true" box="[189,304,1142,1168]" fontSize="11" level="3" pageId="7" pageNumber="77" reason="3">
<emphasis id="B957EADEA77EFF9D069BFBFCFE950BA8" bold="true" box="[189,304,1142,1168]" pageId="7" pageNumber="77">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA77EFF9D069BFB2EFC7D0B86" blockId="7.[189,984,1188,1214]" box="[189,984,1188,1214]" pageId="7" pageNumber="77">Animals seem to feed on rotifers since trophi were found in the gut.</paragraph>
</subSubSection>
</treatment>
</document>