treatments-xml/data/03/8A/87/038A87D6FFB5FFDCFF7FF8FC4004F9F1.xml

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<mods:title id="DC613B1E475C7BECA0EF1C2E67FA913F">Phylogeny and revision of Schraderiellus Rider, 1998 with the description of five new species (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini)</mods:title>
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<mods:affiliation id="5DB63AFAD38D04EEEFC42F5643D2859F">Programa de Pós Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, RS, Brazil 91501 - 970.</mods:affiliation>
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<taxonomicName id="4C234D43FFB5FFDBFF7FF8FC4321F830" authority="(Ruckes, 1959)" baseAuthorityName="Ruckes" baseAuthorityYear="1959" box="[151,611,1903,1930]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis id="B957EAD2FFB5FFDBFF7FF8FC40E6F833" bold="true" box="[151,420,1903,1929]" italics="true" pageId="11" pageNumber="58">Schraderiellus cinctus</emphasis>
(
<bibRefCitation id="EFB24B31FFB5FFDBFE5CF8E34319F830" author="Ruckes, H." box="[436,603,1904,1930]" journalOrPublisher="American Museum Novitates" pageId="11" pageNumber="58" pagination="1 - 18" part="139" refId="ref12672" refString="Ruckes, H. (1959) New genera and species of Pentatomids from Panama and Costa Rica (Heteroptera, Pentatomidae). American Museum Novitates, 139, 1 - 18." title="New genera and species of Pentatomids from Panama and Costa Rica (Heteroptera, Pentatomidae)" type="journal article" year="1959">Ruckes, 1959</bibRefCitation>
)
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<paragraph id="8B9C36C0FFB5FFDBFF7FF8014373F811" blockId="11.[151,611,1903,1963]" box="[151,561,1938,1963]" pageId="11" pageNumber="58">
(
<figureCitation id="13182A45FFB5FFDBFF77F8014061F811" box="[159,291,1938,1963]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="11" pageNumber="58">Figs. 3C–D</figureCitation>
, 
<figureCitation id="13182A45FFB5FFDBFEC7F801402DF811" box="[303,367,1938,1963]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. A–C, S. hughesae; D–F, S. cinctus; G–I, S. luridomaculatus sp. nv.; J–L, S. rufilineatus sp. nv.; M–O, S. luteipedis sp. nv.; P–R, S. brevicolis sp. nv.; S–U, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="11" pageNumber="58">5D–F</figureCitation>
, 
<figureCitation id="13182A45FFB5FFDBFE93F80140DCF811" box="[379,414,1938,1963]" captionStart="FIGURE 6" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[315,1253,194,1929]" captionTargetId="figure-68@7.[315,1260,193,1943]" captionTargetPageId="7" captionText="FIGURE 6. Parameres; dorsal, posterior, inner lateral and external lateral respectively. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983236" httpUri="https://zenodo.org/record/4983236/files/figure.png" pageId="11" pageNumber="58">6B</figureCitation>
, 
<figureCitation id="13182A45FFB5FFDBFE41F800408EF810" box="[425,460,1939,1962]" captionStart="FIGURE 7" captionStartId="8.[151,250,854,876]" captionTargetBox="[159,1400,198,833]" captionTargetId="figure-898@8.[151,1436,193,833]" captionTargetPageId="8" captionText="FIGURE 7. Male internal genitalia (phallus), lateral view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.4983238" httpUri="https://zenodo.org/record/4983238/files/figure.png" pageId="11" pageNumber="58">7B</figureCitation>
, 
<figureCitation id="13182A45FFB5FFDBFE30F80140B9F811" box="[472,507,1938,1963]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="11" pageNumber="58">8B</figureCitation>
, 
<figureCitation id="13182A45FFB5FFDBFDEFF801436AF811" box="[519,552,1938,1963]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1357,233,1748]" captionTargetId="figure-88@10.[150,1436,193,1781]" captionTargetPageId="10" captionText="FIGURE 9. Female internal genitalia, posterior view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.. Abbreviations: cs, capsula seminalis; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner ducts; laIX, laterotergites IX; md, medium ducts; od, outer ducts; pi, pars intermedialis; rs, ring sclerites; vi, vaginal intima. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983242" httpUri="https://zenodo.org/record/4983242/files/figure.png" pageId="11" pageNumber="58">9B</figureCitation>
)
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<paragraph id="8B9C36C0FFB5FFDCFF7FF84945FEFF14" blockId="11.[151,709,2010,2032]" lastBlockId="12.[151,1212,152,174]" lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58">
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<taxonomicName id="4C234D43FFB5FFDBFF7FF8494017F84A" baseAuthorityName="Ruckes" baseAuthorityYear="1959" box="[151,341,2010,2032]" class="Malacostraca" family="Pentatomidae" genus="Schraderia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis id="B957EAD2FFB5FFDBFF7FF8494017F84A" box="[151,341,2010,2032]" italics="true" pageId="11" pageNumber="58">Schraderia cinctus</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFB24B31FFB5FFDBFE8BF84940B2F84A" author="Ruckes, H." box="[355,496,2010,2032]" journalOrPublisher="American Museum Novitates" pageId="11" pageNumber="58" pagination="1 - 18" part="139" refId="ref12672" refString="Ruckes, H. (1959) New genera and species of Pentatomids from Panama and Costa Rica (Heteroptera, Pentatomidae). American Museum Novitates, 139, 1 - 18." title="New genera and species of Pentatomids from Panama and Costa Rica (Heteroptera, Pentatomidae)" type="journal article" year="1959">Ruckes, 1959</bibRefCitation>
; 
<treatmentCitation id="0A8210D1FFB5FFDBFE12F8494387F84A" author="Schrader, F." box="[506,709,2010,2032]" page="499" pageId="11" pageNumber="58" year="1960">
<bibRefCitation id="EFB24B31FFB5FFDBFE12F8494387F84A" author="Schrader, F." box="[506,709,2010,2032]" pageId="11" pageNumber="58" pagination="498 - 508" refId="ref12806" refString="Schrader, F. (1960) Evolutionary aspects of aberrant meiosis in some Pentatominae (Heteroptera). Evolution, 14, 498 - 508. https: // doi. org / 10.1111 / j. 1558 - 5646.1960. tb 03116. x" type="journal article" year="1960">Schrader, 1960: 499</bibRefCitation>
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<treatmentCitation id="0A8210D1FFB2FFDCFF7FFF0B436EFF14" author="Rider, D. A." box="[151,556,152,174]" page="505" pageId="12" pageNumber="59" year="1998">
<taxonomicName id="4C234D43FFB2FFDCFF7FFF0B436EFF14" authority="Rider, 1998: 505" authorityName="Rider" authorityPageNumber="505" authorityYear="1998" box="[151,556,152,174]" class="Insecta" family="Pentatomidae" genus="Schraderiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis id="B957EAD2FFB2FFDCFF7FFF0B403BFF14" box="[151,377,152,174]" italics="true" pageId="12" pageNumber="59">Schraderiellus cinctus</emphasis>
<bibRefCitation id="EFB24B31FFB2FFDCFE97FF0B436EFF14" author="Rider, D. A." box="[383,556,152,174]" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="12" pageNumber="59" pagination="504 - 510" part="100" refId="ref12493" refString="Rider, D. A. (1998) Nomenclatural changes in the Pentatomoidea (Hemiptera - Heteroptera: Pentatomidae, Tessaratomidae). III. Generic level changes. Proceedings of the Entomological Society of Washington, 100 (3), 504 - 510." title="Nomenclatural changes in the Pentatomoidea (Hemiptera - Heteroptera: Pentatomidae, Tessaratomidae). III. Generic level changes" type="journal article" year="1998">Rider, 1998: 505</bibRefCitation>
</taxonomicName>
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; 
<treatmentCitation id="0A8210D1FFB2FFDCFDDEFF0B4220FF14" author="Campos, L. A. &amp; Grazia, J." box="[566,866,152,174]" page="154" pageId="12" pageNumber="59" year="2006">
<bibRefCitation id="EFB24B31FFB2FFDCFDDEFF0B4220FF14" author="Campos, L. A. &amp; Grazia, J." box="[566,866,152,174]" journalOrPublisher="Iheringia Serie Zoologia" pageId="12" pageNumber="59" pagination="147 - 163" part="96" refId="ref11580" refString="Campos, L. A. &amp; Grazia, J. (2006) Analise cladistica e biogeografia de Ochlerini (Heteroptera, Pentatomidae, Discocephalinae). Iheringia Serie Zoologia, 96, 147 - 163. https: // doi. org / 10.1590 / s 0073 - 47212006000200004" title="Analise cladistica e biogeografia de Ochlerini (Heteroptera, Pentatomidae, Discocephalinae)" type="journal article" year="2006">Campos &amp; Grazia, 2006: 154</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0A8210D1FFB2FFDCFC85FF0B45FEFF14" author="Roell, T. &amp; Campos, L. A." box="[877,1212,152,174]" page="575" pageId="12" pageNumber="59" year="2015">
<bibRefCitation id="EFB24B31FFB2FFDCFC85FF0B45FEFF14" author="Roell, T. &amp; Campos, L. A." box="[877,1212,152,174]" journalOrPublisher="Zootaxa" pageId="12" pageNumber="59" pagination="573 - 583" part="4018" refId="ref12540" refString="Roell, T. &amp; Campos, L. A. (2015) Candeocoris bistillatus, new genus and new species of Ochlerini from Ecuador (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 4018 (4), 573 - 583. https: // doi. org / 10.11646 / zootaxa. 4018.4.6" title="Candeocoris bistillatus, new genus and new species of Ochlerini from Ecuador (Hemiptera: Heteroptera: Pentatomidae)" type="journal article" year="2015">Roell &amp; Campos, 2015: 575–576</bibRefCitation>
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</paragraph>
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<paragraph id="8B9C36C0FFB2FFDCFF7FFF4940EFFF49" blockId="12.[151,1437,218,1611]" box="[151,429,218,243]" pageId="12" pageNumber="59">
<emphasis id="B957EAD2FFB2FFDCFF7FFF49404FFF49" bold="true" box="[151,269,218,243]" pageId="12" pageNumber="59">
<typeStatus id="54988862FFB2FFDCFF7FFF49404AFF49" box="[151,264,218,243]" pageId="12" pageNumber="59" type="holotype">Holotype</typeStatus>
:
</emphasis>
male, 
<collectionCode id="ED32AE05FFB2FFDCFEB0FF4940EFFF49" box="[344,429,218,243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="12" pageNumber="59">AMNH</collectionCode>
</paragraph>
<paragraph id="8B9C36C0FFB2FFDCFF2FFF6D43CAFEAD" blockId="12.[151,1437,218,1611]" box="[199,648,254,279]" pageId="12" pageNumber="59">
<emphasis id="B957EAD2FFB2FFDCFF2FFF6D402BFEAD" bold="true" box="[199,361,254,279]" pageId="12" pageNumber="59">Type locality:</emphasis>
<collectingCountry id="F3347650FFB2FFDCFE87FF6D4354FEAD" box="[367,534,254,279]" name="Costa Rica" pageId="12" pageNumber="59">COSTA RICA</collectingCountry>
, Turrialba
</paragraph>
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<paragraph id="8B9C36C0FFB2FFDCFF2FFEB1445EFE39" blockId="12.[151,1437,218,1611]" pageId="12" pageNumber="59">
<emphasis id="B957EAD2FFB2FFDCFF2FFEB1401BFE81" bold="true" box="[199,345,290,315]" inLineHeading="true" pageId="12" pageNumber="59" reason="1">Description.</emphasis>
Dark brown to black. Lateral margins of pronotum, hemelytra and connexivum usually outlined in yellow or orange. Often with a yellow spot at apex of each radial vein. Pronotum, scutellum and hemelytra may present sparse yellow spots. Coxae, trochanter, femora and tibiae dark brown, tarsi yellowish (
<figureCitation id="13182A45FFB2FFDCFB5FFEF9444DFE39" box="[1207,1295,362,387]" captionStart="FIGURE 3" captionStartId="4.[151,250,1955,1977]" captionTargetBox="[174,1428,196,1930]" captionTargetId="figure-86@4.[150,1436,193,1933]" captionTargetPageId="4" captionText="FIGURE 3. Species of Schraderiellus, dorsal and ventral habitus. A, B, S. hughesae; C, D, S. cinctus; E, F, S. luridomaculatus sp. nv.; G, H, S. rufilineatus sp. nv.; I, J, S. luteipedis sp. nv.; K, L, S. brevicolis sp. nv.; M, N, S. falcatus sp. nv.. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983228" httpUri="https://zenodo.org/record/4983228/files/figure.png" pageId="12" pageNumber="59">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="8B9C36C0FFB2FFDCFF2FFE1D4422FDE1" blockId="12.[151,1437,218,1611]" pageId="12" pageNumber="59">Male. Measurements (n=8): head length 2.65 ± 0.15 (2.41–2.80); width 3.11 ± 0.08 (3.04–3.24); pronotum length 3.37 ± 0.14 (3.25–3.60); width 6.17 ± 0.13 (6.03–6.39); scutellum length 5.12 ± 0.20 (4.80–5.35); width 4.27 ± 0.32 (3.90–4.72); abdomen length 4.92 ± 0.42 (4.26–5.46); width 6.22 ± 0.20 (6.03–6.60); length of antennal segments: I 1.23 ± 0.05 (1.20–1.32); II 1.17 ± 0.15 (1.00–1.40); III 2.09 ± 0.14 (2.04–2.40); VI 2.23 ± 0.15 (2.04–2.40); V 2.56 ± 0.17 (2.44–2.76); length of labial segments: I 1.64 ± 0.09 (1.55–1.79); II 2.99 ± 0.13 (2.80–3.16); III 2.30 ± 0.22 (1.92–2.48); IV 2.72 ± 0.10 (2.60–2.84); total length 13.17 ± 0.58 (12.48–14.00).</paragraph>
<paragraph id="8B9C36C0FFB2FFDCFF2FFDF54576FCB5" blockId="12.[151,1437,218,1611]" pageId="12" pageNumber="59">
Genitalia. Exposed surface of pygophore castaneous; posterolateral angles moderately divergent, stout, posteriorly depressed, apices rounded (
<figureCitation id="13182A45FFB2FFDCFD8CFD1943A0FD19" box="[612,738,650,675]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. A–C, S. hughesae; D–F, S. cinctus; G–I, S. luridomaculatus sp. nv.; J–L, S. rufilineatus sp. nv.; M–O, S. luteipedis sp. nv.; P–R, S. brevicolis sp. nv.; S–U, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="12" pageNumber="59">Fig. 5D–F</figureCitation>
). Parameres yellowish, subparallel to posterolateral angles, attaining apex of segment Xor slightly shorter, cylindrical, apices flattened with subapical denticle (
<figureCitation id="13182A45FFB2FFDCFAFEFD3D44DAFD7D" box="[1302,1432,686,711]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. A–C, S. hughesae; D–F, S. cinctus; G–I, S. luridomaculatus sp. nv.; J–L, S. rufilineatus sp. nv.; M–O, S. luteipedis sp. nv.; P–R, S. brevicolis sp. nv.; S–U, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="12" pageNumber="59">Figs. 5D–F</figureCitation>
, 
<figureCitation id="13182A45FFB2FFDCFF7FFD4141F8FD51" box="[151,186,722,747]" captionStart="FIGURE 6" captionStartId="7.[151,250,1967,1989]" captionTargetBox="[315,1253,194,1929]" captionTargetId="figure-68@7.[315,1260,193,1943]" captionTargetPageId="7" captionText="FIGURE 6. Parameres; dorsal, posterior, inner lateral and external lateral respectively. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bar: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983236" httpUri="https://zenodo.org/record/4983236/files/figure.png" pageId="12" pageNumber="59">6B</figureCitation>
). Segment X subcylindrical, basal portion membranous forming a line at the limit of sclerotized area (
<figureCitation id="13182A45FFB2FFDCFADFFD4141E8FCB5" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[314,1259,195,1859]" captionTargetId="figure-110@6.[314,1260,193,1868]" captionTargetPageId="6" captionText="FIGURE 5. Pygophores in posterior, dorsal and lateral views. A–C, S. hughesae; D–F, S. cinctus; G–I, S. luridomaculatus sp. nv.; J–L, S. rufilineatus sp. nv.; M–O, S. luteipedis sp. nv.; P–R, S. brevicolis sp. nv.; S–U, S. falcatus sp. nv.. Abbreviations: dr, dorsal rim; il, inferior layer; p1, dorsal projection of the posterolateral angles; p2, basal projection of the posterolateral angles; pa, posterolateral angles; par, parameres; vr, ventral rim; X, segment X. Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983234" httpUri="https://zenodo.org/record/4983234/files/figure.png" pageId="12" pageNumber="59">Fig. 5D– F</figureCitation>
). Phallotheca globose, vesica sinuous, slightly longer than phallotheca (
<figureCitation id="13182A45FFB2FFDCFC26FD654565FCB5" box="[974,1063,758,783]" captionStart="FIGURE 7" captionStartId="8.[151,250,854,876]" captionTargetBox="[159,1400,198,833]" captionTargetId="figure-898@8.[151,1436,193,833]" captionTargetPageId="8" captionText="FIGURE 7. Male internal genitalia (phallus), lateral view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.; E, S. brevicolis sp. nv.; F, S. falcatus sp. nv.. Scale bars = 0.5mm" figureDoi="http://doi.org/10.5281/zenodo.4983238" httpUri="https://zenodo.org/record/4983238/files/figure.png" pageId="12" pageNumber="59">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8B9C36C0FFB2FFDCFF2FFC89458CFC5D" blockId="12.[151,1437,218,1611]" pageId="12" pageNumber="59">Female. Measurements (n=11): head length 2.40 ± 0.19 (2.80–3.20); width 3.16 ± 0.08 (3.03–3.28); pronotum length 3.66 ± 0.17 (3.30–3.78); width 6.60 ± 0.23 (6.10–6.81); scutellum length 5.73 ± 0.24 (5.35–5.98); width 4.23 ± 0.15 (4.04–4.47); abdomen length 6.27 ± 0.31 (5.68–6.53); width 6.88 ± 0.20 (6.53–7.1); length of antennal segments: I 1.17 ± 0.12 (1.00–1.26); II 1.14 ± 0.09 (1.05–1.26); III 2.18 ± 0.10 (2.04–2.32); IV 2.31 ± 0.11 (2.17– 2.45); V 2.64 ± 0.12 (2.51–2.75); length of labial segments: I 1.60 ± 0.16 (1.42–1.85); II 3.00 ± 0.22 (2.62–3.30); III 2.40 ± 0.21 (2.13–2.75); IV 2.58 ± 0.27 (2.13–2.85); total length 14.75 ± 0.74 (13.57–15.44).</paragraph>
<paragraph id="8B9C36C0FFB2FFDCFF2FFC6142EDFBCD" blockId="12.[151,1437,218,1611]" pageId="12" pageNumber="59">
Genitalia. Gonocoxites VIII longer than wide covering the base of laterotergites IX. Posterior margins of gonocoxites VIII over gonocoxites IX rectilinear. Laterotergites IX subrectangular, diagonally directed to apex (
<figureCitation id="13182A45FFB2FFDCFF77FBA941BBFBE9" box="[159,249,1082,1107]" captionStart="FIGURE 8" captionStartId="9.[151,250,1935,1957]" captionTargetBox="[180,1393,531,1897]" captionTargetId="figure-449@9.[180,1394,525,1898]" captionTargetPageId="9" captionText="FIGURE 8. Female genital plates, posterior view. A, S. hughesae; B, S. cinctus; C, S. luridomaculatus sp. nv.; D, S. rufilineatus sp. nv.; E, S. luteipedis sp. nv.. Abbreviations. gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; laVIII, laterotergites VIII; laIX, laterotergites IX; X, segment X. Scale bars: 1mm." figureDoi="http://doi.org/10.5281/zenodo.4983240" httpUri="https://zenodo.org/record/4983240/files/figure.png" pageId="12" pageNumber="59">Fig. 8B</figureCitation>
). Arms of gonocoxites IX widening to apex, sclerotized. Vesicular area about six times longer than the combined length of pars intermedialis and capsula seminalis (
<figureCitation id="13182A45FFB2FFDCFCA2FBCD42E1FBCD" box="[842,931,1118,1143]" captionStart="FIGURE 9" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[151,1357,233,1748]" captionTargetId="figure-88@10.[150,1436,193,1781]" captionTargetPageId="10" captionText="FIGURE 9. Female internal genitalia, posterior view. A, S. hughesae; B, S. cinctus; C, S. rufilineatus sp. nv.; D, S. luteipedis sp. nv.. Abbreviations: cs, capsula seminalis; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner ducts; laIX, laterotergites IX; md, medium ducts; od, outer ducts; pi, pars intermedialis; rs, ring sclerites; vi, vaginal intima. Scale bars: 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.4983242" httpUri="https://zenodo.org/record/4983242/files/figure.png" pageId="12" pageNumber="59">Fig. 9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C339654BFFB2FFDCFF2FFB1142EAFB20" box="[199,936,1154,1179]" pageId="12" pageNumber="59" type="distribution">
<paragraph id="8B9C36C0FFB2FFDCFF2FFB1142EAFB20" blockId="12.[151,1437,218,1611]" box="[199,936,1154,1179]" pageId="12" pageNumber="59">
<emphasis id="B957EAD2FFB2FFDCFF2FFB11401BFB21" bold="true" box="[199,345,1154,1179]" pageId="12" pageNumber="59">Distribution</emphasis>
: 
<collectingCountry id="F3347650FFB2FFDCFE80FB1140A4FB21" box="[360,486,1154,1179]" name="Costa Rica" pageId="12" pageNumber="59">Costa Rica</collectingCountry>
. NEW RECORDS: 
<collectingCountry id="F3347650FFB2FFDCFD2EFB11427EFB21" box="[710,828,1154,1179]" name="Nicaragua" pageId="12" pageNumber="59">Nicaragua</collectingCountry>
, 
<collectingCountry id="F3347650FFB2FFDCFCA0FB1042E6FB20" box="[840,932,1155,1178]" name="Panama" pageId="12" pageNumber="59">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C339654BFFB2FFDCFF2FFB354004F9F1" pageId="12" pageNumber="59" type="materials_examined">
<paragraph id="8B9C36C0FFB2FFDCFF2FFB354004F9F1" blockId="12.[151,1437,218,1611]" pageId="12" pageNumber="59">
<emphasis id="B957EAD2FFB2FFDCFF2FFB3540F6FB05" bold="true" box="[199,436,1190,1215]" inLineHeading="true" pageId="12" pageNumber="59" reason="1">Material examined.</emphasis>
<collectingCountry id="F3347650FFB2FFDCFE55FB35432AFB05" box="[445,616,1190,1215]" name="Nicaragua" pageId="12" pageNumber="59">NICARAGUA</collectingCountry>
, 
<emphasis id="B957EAD2FFB2FFDCFD9FFB35438EFB05" box="[631,716,1190,1215]" italics="true" pageId="12" pageNumber="59">Zelaya:</emphasis>
(Sulum, 
<geoCoordinate id="EE175007FFB2FFDCFCD2FB3542E0FB04" box="[826,930,1190,1215]" degrees="14" direction="north" minutes="15" orientation="latitude" pageId="12" pageNumber="59" precision="925" value="14.25">14º15’ N</geoCoordinate>
<geoCoordinate id="EE175007FFB2FFDCFC45FB35455DFB05" box="[941,1055,1190,1215]" degrees="84" direction="west" minutes="36" orientation="longitude" pageId="12" pageNumber="59" precision="925" value="-84.6">84º36’ W</geoCoordinate>
), 
<specimenCount id="9D25FD49FFB2FFDCFBDEFB35451BFB05" box="[1078,1113,1190,1215]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFB83FB3545F1FB05" box="[1131,1203,1190,1215]" pageId="12" pageNumber="59" value="1996-01">I.1996</date>
Maes &amp; Hernandes (
<collectionCode id="ED32AE05FFB2FFDCFF77FB5941B2FB59" box="[159,240,1226,1251]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="12" pageNumber="59">UFRG</collectionCode>
). 
<collectingCountry id="F3347650FFB2FFDCFEEBFB5940E8FB59" box="[259,426,1226,1251]" name="Costa Rica" pageId="12" pageNumber="59">COSTA RICA</collectingCountry>
, 
<emphasis id="B957EAD2FFB2FFDCFE5EFB58437CFB59" box="[438,574,1226,1251]" italics="true" pageId="12" pageNumber="59">San Carlos:</emphasis>
<specimenCount id="9D25FD49FFB2FFDCFDADFB59432BFB59" box="[581,617,1226,1251]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
, Schild &amp; Burgdorf (
<collectionCode id="ED32AE05FFB2FFDCFCB2FB5942F3FB59" box="[858,945,1226,1251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="59" type="Museum">USNM</collectionCode>
); 
<emphasis id="B957EAD2FFB2FFDCFC2DFB58452CFB58" box="[965,1134,1227,1250]" italics="true" pageId="12" pageNumber="59">Los Diamantes</emphasis>
: 
<specimenCount id="9D25FD49FFB2FFDCFB95FB5945E2FB59" box="[1149,1184,1226,1251]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
and 
<specimenCount id="9D25FD49FFB2FFDCFB31FB5945BEFB59" box="[1241,1276,1226,1251]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFAE3FB594439FB59" box="[1291,1403,1226,1251]" pageId="12" pageNumber="59" value="1948-05-31">31.v.1948</date>
, F. Schrader (
<collectionCode id="ED32AE05FFB2FFDCFEE5FB7D4026FABD" box="[269,356,1262,1287]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="59" type="Museum">USNM</collectionCode>
); 
<emphasis id="B957EAD2FFB2FFDCFE93FB7D40A4FABD" box="[379,486,1262,1287]" italics="true" pageId="12" pageNumber="59">Turrialba</emphasis>
: 
<specimenCount id="9D25FD49FFB2FFDCFE11FB7D435EFABD" box="[505,540,1262,1287]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFDC5FB7D43DCFABD" box="[557,670,1262,1287]" pageId="12" pageNumber="59" value="1951-05-22">22.v.1951</date>
, OL Cartwright (
<collectionCode id="ED32AE05FFB2FFDCFC81FB7D4282FABD" box="[873,960,1262,1287]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="59" type="Museum">USNM</collectionCode>
); 
<specimenCount id="9D25FD49FFB2FFDCFC3FFB7D42B8FABD" box="[983,1018,1262,1287]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFBE3FB7D453AFABD" box="[1035,1144,1262,1287]" pageId="12" pageNumber="59" value="1965-01-21">21.i.1965</date>
, J.A Slater &amp; N. 
<collectionCode id="ED32AE05FFB2FFDCFAD6FB7C4413FABC" box="[1342,1361,1263,1286]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="59">T</collectionCode>
Davis (
<collectionCode id="ED32AE05FFB2FFDCFF77FA8141B1FA91" box="[159,243,1298,1323]" pageId="12" pageNumber="59">DARC</collectionCode>
); 
<specimenCount id="9D25FD49FFB2FFDCFEE4FA81406DFA91" box="[268,303,1298,1323]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFEABFA8140ABFA91" box="[323,489,1298,1323]" pageId="12" pageNumber="59" value="1965-03-13" valueMax="1965-03-17" valueMin="1965-03-13">13–17.iii.1965</date>
, SS &amp; Duckworth (
<collectionCode id="ED32AE05FFB2FFDCFD35FA814276FA91" box="[733,820,1298,1323]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="59" type="Museum">USNM</collectionCode>
); (
<collectingRegion id="49E7F822FFB2FFDCFCBEFA8142F1FA91" box="[854,947,1298,1323]" country="Costa Rica" name="Cartago" pageId="12" pageNumber="59">Cartago</collectingRegion>
, 
<quantity id="4CDB9B25FFB2FFDCFC2CFA81455CFA91" box="[964,1054,1298,1323]" metricMagnitude="-1" metricUnit="m" metricValue="6.459999999999999" pageId="12" pageNumber="59" unit="mm" value="646.0">646mm</quantity>
), 
<specimenCount id="9D25FD49FFB2FFDCFBD0FA814519FA91" box="[1080,1115,1298,1323]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFB87FA8145A3FA91" box="[1135,1249,1298,1323]" pageId="12" pageNumber="59" value="1965-07-09">9.vii.1965</date>
M.G Naumann (
<collectionCode id="ED32AE05FFB2FFDCFF77FAA541B2FAF5" box="[159,240,1334,1359]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="12" pageNumber="59">UFRG</collectionCode>
); 
<emphasis id="B957EAD2FFB2FFDCFEEBFAA5401CFAF5" box="[259,350,1334,1359]" italics="true" pageId="12" pageNumber="59">
<collectingRegion id="49E7F822FFB2FFDCFEEBFAA5401CFAF5" box="[259,350,1334,1359]" country="Costa Rica" name="Heredia" pageId="12" pageNumber="59">Heredia</collectingRegion>
</emphasis>
: (La Selva 
<quantity id="4CDB9B25FFB2FFDCFE08FAA54350FAF5" box="[480,530,1334,1359]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="59" unit="km" value="3.0">3km</quantity>
S Pto. Viejo 
<geoCoordinate id="EE175007FFB2FFDCFD40FAA5424FFAF4" box="[680,781,1334,1359]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="12" pageNumber="59" precision="925" value="10.433333">10º26’ N</geoCoordinate>
<geoCoordinate id="EE175007FFB2FFDCFCFDFAA542C7FAF5" box="[789,901,1334,1359]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="12" pageNumber="59" precision="925" value="-84.01667">84º01’ W</geoCoordinate>
), 
<specimenCount id="9D25FD49FFB2FFDCFC70FAA542F9FAF5" box="[920,955,1334,1359]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 1983, HA. Hespenheide (
<collectionCode id="ED32AE05FFB2FFDCFB0FFAA54479FAF5" box="[1255,1339,1334,1359]" pageId="12" pageNumber="59">DARC</collectionCode>
); 
<specimenCount id="9D25FD49FFB2FFDCFAA7FAA54431FAF5" box="[1359,1395,1334,1359]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFA68FAA54041FAC9" pageId="12" pageNumber="59" value="1984-06-03" valueMax="1984-06-05" valueMin="1984-06-03">3– 5.vi.1984</date>
, Riley, Rider &amp; LeDoux (
<collectionCode id="ED32AE05FFB2FFDCFDCCFAC94337FAC9" box="[548,629,1370,1395]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="12" pageNumber="59">UFRG</collectionCode>
); 
<specimenCount id="9D25FD49FFB2FFDCFD60FAC943EEFAC9" box="[648,684,1370,1395]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFD51FAC9421FFAC9" box="[697,861,1370,1395]" pageId="12" pageNumber="59" value="1986-06-11" valueMax="1986-06-17" valueMin="1986-06-11">11–17.vi.1986</date>
, W. Hanson &amp; G. Bohart (
<collectionCode id="ED32AE05FFB2FFDCFB69FAC945F6FAC9" box="[1153,1204,1370,1395]" pageId="12" pageNumber="59">JEE</collectionCode>
); (Est. Biol La Selva 
<quantity id="4CDB9B25FFB2FFDCFF7FFAED4198FA2D" box="[151,218,1406,1431]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="59" unit="m" value="150.0">150m</quantity>
), 
<specimenCount id="9D25FD49FFB2FFDCFF07FAED4051FA2D" box="[239,275,1406,1431]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFEC9FAED4096FA2D" box="[289,468,1406,1431]" pageId="12" pageNumber="59" value="1993-04-20" valueMax="1993-05-30" valueMin="1993-04-20">20.iv-30.v.1993</date>
, Paul M. Freytag &amp; Thomas 
<collectionCode id="ED32AE05FFB2FFDCFCC8FAEC4276FA2C" box="[800,820,1407,1430]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="59" type="Museum">V</collectionCode>
. Myers (
<collectionCode id="ED32AE05FFB2FFDCFC70FAED4289FA2D" box="[920,971,1406,1431]" pageId="12" pageNumber="59">JEE</collectionCode>
); (Est. Biol La Selva), 
<specimenCount id="9D25FD49FFB2FFDCFB38FAED45B1FA2D" box="[1232,1267,1406,1431]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFAEAFAED44D5FA2D" box="[1282,1431,1406,1431]" pageId="12" pageNumber="59" value="1995-01-11" valueMax="1995-01-13" valueMin="1995-01-11">11–13.i.1995</date>
, E.G. Riley (
<collectionCode id="ED32AE05FFB2FFDCFEF2FA31400FFA01" box="[282,333,1442,1467]" pageId="12" pageNumber="59">JEE</collectionCode>
); (Est. Biol La Selva 
<geoCoordinate id="EE175007FFB2FFDCFDD6FA3143E1FA00" box="[574,675,1442,1467]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="12" pageNumber="59" precision="925" value="10.433333">10º26’ N</geoCoordinate>
<geoCoordinate id="EE175007FFB2FFDCFD42FA314254FA00" box="[682,790,1442,1467]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="12" pageNumber="59" precision="925" value="-84.01667">84º01’ W</geoCoordinate>
<quantity id="4CDB9B25FFB2FFDCFCF5FA3142C4FA01" box="[797,902,1442,1467]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="12" pageNumber="59" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50-150m</quantity>
), 
<specimenCount id="9D25FD49FFB2FFDCFC72FA3142FFFA01" box="[922,957,1442,1467]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
and 
<specimenCount id="9D25FD49FFB2FFDCFC1CFA314555FA01" box="[1012,1047,1442,1467]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFBCDFA31459AFA01" box="[1061,1240,1442,1467]" pageId="12" pageNumber="59" value="1999-08-19" valueMax="1999-08-23" valueMin="1999-08-19">19–23.viii.1999</date>
, 
<collectionCode id="ED32AE05FFB2FFDCFB0DFA304454FA00" box="[1253,1302,1443,1466]" country="New Zealand" name="Dominion Museum" pageId="12" pageNumber="59">DM</collectionCode>
. &amp; S. Davis (
<collectionCode id="ED32AE05FFB2FFDCFF77FA5541B4FA65" box="[159,246,1478,1503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="59" type="Museum">USNM</collectionCode>
); 
<emphasis id="B957EAD2FFB2FFDCFEF9FA554035FA65" box="[273,375,1478,1503]" italics="true" pageId="12" pageNumber="59">
<collectingRegion id="49E7F822FFB2FFDCFEF9FA554031FA65" box="[273,371,1478,1503]" country="Costa Rica" name="Alajuela" pageId="12" pageNumber="59">Alajuela</collectingRegion>
,
</emphasis>
(
<quantity id="4CDB9B25FFB2FFDCFE97FA554080FA65" box="[383,450,1478,1503]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="59" unit="m" value="750.0">750m</quantity>
) 
<specimenCount id="9D25FD49FFB2FFDCFE31FA5540BFFA65" box="[473,509,1478,1503]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFDFBFA5543C9FA65" box="[531,651,1478,1503]" pageId="12" pageNumber="59" value="1988-04-19">19.iv.1988</date>
, J. Soto (
<collectionCode id="ED32AE05FFB2FFDCFCE3FA55421DFA65" box="[779,863,1478,1503]" pageId="12" pageNumber="59">DARC</collectionCode>
); [no state], 
<specimenCount id="9D25FD49FFB2FFDCFBEAFA554567FA65" box="[1026,1061,1478,1503]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, G. 
<collectionCode id="ED32AE05FFB2FFDCFBBBFA554524FA65" box="[1107,1126,1478,1503]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="59">R</collectionCode>
Wilson Collector (
<collectionCode id="ED32AE05FFB2FFDCFABCFA5544D2FA65" box="[1364,1424,1478,1503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="12" pageNumber="59" type="Museum">CAS</collectionCode>
). 
<collectingCountry id="F3347650FFB2FFDCFF7FFA794053F9B9" box="[151,273,1514,1539]" name="Panama" pageId="12" pageNumber="59">PANAMA</collectingCountry>
, 
<emphasis id="B957EAD2FFB2FFDCFEF6FA784029F9B8" box="[286,363,1515,1538]" italics="true" pageId="12" pageNumber="59">Bocas:</emphasis>
(
<collectionCode id="ED32AE05FFB2FFDCFE95FA7940E6F9B9" box="[381,420,1514,1539]" pageId="12" pageNumber="59">DT</collectionCode>
Miramar 
<geoCoordinate id="EE175007FFB2FFDCFDF2FA794313F9B8" box="[538,593,1514,1539]" degrees="9" direction="north" orientation="latitude" pageId="12" pageNumber="59" precision="55555" value="9.0">9º N</geoCoordinate>
, 
<geoCoordinate id="EE175007FFB2FFDCFDB7FA794393F9B9" box="[607,721,1514,1539]" degrees="82" direction="west" minutes="15" orientation="longitude" pageId="12" pageNumber="59" precision="925" value="-82.25">82º15’ W</geoCoordinate>
), 
<specimenCount id="9D25FD49FFB2FFDCFD0FFA794248F9B9" box="[743,778,1514,1539]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFCF3FA7942D3F9B9" box="[795,913,1514,1539]" pageId="12" pageNumber="59" value="1979-04-25">25.iv.1979</date>
(
<collectionCode id="ED32AE05FFB2FFDCFC4BFA7942B5F9B9" box="[931,1015,1514,1539]" pageId="12" pageNumber="59">DARC</collectionCode>
); (
<collectionCode id="ED32AE05FFB2FFDCFBFDFA79457EF9B9" box="[1045,1084,1514,1539]" pageId="12" pageNumber="59">DT</collectionCode>
Corriente Grande 
<geoCoordinate id="EE175007FFB2FFDCFAFDFA7944D9F9B8" box="[1301,1435,1514,1539]" degrees="9" direction="north" minutes="17" orientation="latitude" pageId="12" pageNumber="59" precision="15" seconds="30" value="9.291667">9º17’30’’ N</geoCoordinate>
<geoCoordinate id="EE175007FFB2FFDCFF7FF99D4072F99C" box="[151,304,1550,1575]" degrees="82" direction="west" minutes="32" orientation="longitude" pageId="12" pageNumber="59" precision="15" seconds="41" value="-82.54472">82º32’41’’ W</geoCoordinate>
, 
<quantity id="4CDB9B25FFB2FFDCFED3F99D40D7F99D" box="[315,405,1550,1575]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="59" unit="mm" value="100.0">100mm</quantity>
), 
<specimenCount id="9D25FD49FFB2FFDCFE40F99D4089F99D" box="[424,459,1550,1575]" pageId="12" pageNumber="59" type="female">1♀</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFE32F99D4317F99D" box="[474,597,1550,1575]" pageId="12" pageNumber="59" value="1980-03-24">24.iii.1980</date>
, Henk Wolda (
<collectionCode id="ED32AE05FFB2FFDCFD15F99D4213F99D" box="[765,849,1550,1575]" pageId="12" pageNumber="59">DARC</collectionCode>
); 
<emphasis id="B957EAD2FFB2FFDCFC8CF99C4566F99C" box="[868,1060,1551,1574]" italics="true" pageId="12" pageNumber="59">Cerro Campana:</emphasis>
<specimenCount id="9D25FD49FFB2FFDCFBC4F99D4512F99D" box="[1068,1104,1550,1575]" pageId="12" pageNumber="59" type="male">1♂</specimenCount>
, 
<date id="FF9D1000FFB2FFDCFBB5F99D458BF99D" box="[1117,1225,1550,1575]" pageId="12" pageNumber="59" value="1995-06-05">5.vi.1995</date>
, B. Ratcliffe &amp; M. Jameson (
<collectionCode id="ED32AE05FFB2FFDCFEEEF9A1407BF9F1" box="[262,313,1586,1611]" pageId="12" pageNumber="59">JEE</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>