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<document id="EE872E799F22173C6CED73F2211A3BF3" ID-CLB-Dataset="55750" ID-DOI="10.5281/zenodo.5385198" ID-GBIF-Dataset="c7f95b94-53f8-420d-97c5-4e27d7c211bf" ID-ISSN="2345-7600" ID-Zenodo-Dep="5385198" ID-ZooBank="AC7F1491-CD89-4F10-9D61-55354B55E368" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630300948061" checkinUser="valdenar" docAuthor="Hájek, Jiří &amp; Brancucci, Michel" docDate="2015" docId="038A87C4FFA0FF99F09393A4FEBCBDF9" docLanguage="en" docName="RafflesBZool.63.309-326.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Laccophilus ceylonicus Zimmermann 1919" docType="treatment" docVersion="9" lastPageNumber="314" masterDocId="FFB3FFBCFFA5FF9CF02A9279FFD5BB5C" masterDocTitle="A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)" masterLastPageNumber="326" masterPageNumber="309" pageNumber="314" updateTime="1699240741915" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="97D116495B16B726926F6467F1D5AE55">A taxonomic review of the Oriental Laccophilus javanicus species group (Coleoptera: Dytiscidae)</mods:title>
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<mods:affiliation id="2D3A9658BE7CB25721A5E35A9EA554D5">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 Horní Počernice, Czech Republic</mods:affiliation>
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<mods:affiliation id="B324F40B96D1E97588EBB850A2C81B46">Naturhistorisches Museum, Basel, Switzerland</mods:affiliation>
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<treatment id="038A87C4FFA0FF99F09393A4FEBCBDF9" ID-DOI="http://doi.org/10.5281/zenodo.5466883" ID-GBIF-Taxon="194955238" ID-Zenodo-Dep="5466883" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9" httpUri="http://treatment.plazi.org/id/038A87C4FFA0FF99F09393A4FEBCBDF9" lastPageNumber="314" pageId="5" pageNumber="314">
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<paragraph id="8B9C36D2FFA0FF99F09393A4FD7FBAA9" blockId="5.[185,682,477,533]" box="[185,682,477,501]" pageId="5" pageNumber="314">
<heading id="D0D481BEFFA0FF99F09393A4FD7FBAA9" bold="true" box="[185,682,477,501]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="314" reason="2">
<taxonomicName id="4C234D51FFA0FF99F09393A4FD7FBAA9" ID-CoL="3RM5X" authority="Zimmermann, 1919" authorityName="Zimmermann" authorityYear="1919" box="[185,682,477,501]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B957EAC0FFA0FF99F09393A4FD7FBAA9" bold="true" box="[185,682,477,501]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F09393A4FE68BAA9" bold="true" box="[185,445,477,501]" italics="true" pageId="5" pageNumber="314">Laccophilus ceylonicus</emphasis>
<bibRefCitation id="EFB24B23FFA0FF99F1ED93A4FD7FBAA9" author="Zimmermann A" box="[455,682,477,501]" firstAuthor="Zimmermann" journalOrPublisher="Archiv fur Naturgeschichte" pageId="5" pageNumber="314" pagination="68 - 249" part="83" refId="ref16045" refString="Zimmermann A (1919) Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem. Archiv fur Naturgeschichte, 83 (1917) (A 12): 68 - 249." title="Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem" type="journal article" year="1919">Zimmermann, 1919</bibRefCitation>
</emphasis>
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<subSubSection id="C3396559FFA0FF99F1449384FE21B949" box="[366,500,509,533]" pageId="5" pageNumber="314" type="description">
<paragraph id="8B9C36D2FFA0FF99F1449384FE21B949" blockId="5.[185,682,477,533]" box="[366,500,509,533]" pageId="5" pageNumber="314">
(
<figureCitation id="13182A57FFA0FF99F15F9384FE16B949" box="[373,451,509,533]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 19. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="5" pageNumber="314">Figs. 3</figureCitation>
,
<figureCitation id="13182A57FFA0FF99F1FA9384FE39B949" box="[464,492,509,533]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 1835. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="5" pageNumber="314">20</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="AB3311FCFFA0FF99F05B9045FE25B932" pageId="5" pageNumber="314">
<treatmentCitation id="0A8210C3FFA0FF99F05B9045FDD9B90E" author="Regimbart M" box="[113,524,572,594]" page="468" pageId="5" pageNumber="314" year="1902">
<taxonomicName id="4C234D51FFA0FF99F05B9045FDD9B90E" ID-CoL="3RM7R" authority="Regimbart, 1902: 468" authorityName="Regimbart" authorityPageNumber="468" authorityYear="1902" box="[113,524,572,594]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="horni">
<emphasis id="B957EAC0FFA0FF99F05B9045FEF2B90D" box="[113,295,572,593]" italics="true" pageId="5" pageNumber="314">Laccophilus horni</emphasis>
<bibRefCitation id="EFB24B23FFA0FF99F11B9045FDD9B90E" author="Regimbart M" box="[305,524,572,594]" firstAuthor="Regimbart" journalOrPublisher="Annales de la Societe Entomologique de France" page="468" pageId="5" pageNumber="314" pagination="468 - 475" part="71" refId="ref15820" refString="Regimbart M (1902) Voyage du Dr W. Horn a Ceylan. Coleopteres aquatiques. Annales de la Societe Entomologique de France, 71: 468 - 475." title="Voyage du Dr W. Horn a Ceylan. Coleopteres aquatiques" type="journal article" year="1902">Régimbart, 1902: 468</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description); preoccupied by
<treatmentCitation id="0A8210C3FFA0FF99F11E9020FE38B932" author="Branden C van den" box="[308,493,601,623]" page="21" pageId="5" pageNumber="314" year="1885">
<bibRefCitation id="EFB24B23FFA0FF99F11E9020FE38B932" author="Branden C van den" box="[308,493,601,623]" firstAuthor="Branden" journalOrPublisher="Annales de la Societe Entomologique de Belgique" page="21" pageId="5" pageNumber="314" pagination="5 - 116" part="29" refId="ref15174" refString="Branden C van den (1885) Catalogue des coleopteres carnassiers aquatiques (Haliplidae, Amphizoidae, Pelobiidae et Dytiscidae). Annales de la Societe Entomologique de Belgique, 29 (1): 5 - 116." title="Catalogue des coleopteres carnassiers aquatiques (Haliplidae, Amphizoidae, Pelobiidae et Dytiscidae)" type="journal article" year="1885">Branden, 1885: 21</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B9C36D2FFA0FF99F05B900CFDD2B999" blockId="5.[113,753,572,709]" pageId="5" pageNumber="314">
<treatmentCitationGroup id="AB3311FCFFA0FF99F05B900CFDD2B999" pageId="5" pageNumber="314">
<treatmentCitation id="0A8210C3FFA0FF99F05B900CFDB0B9D7" author="Zimmermann A" box="[113,613,629,651]" page="123" pageId="5" pageNumber="314" year="1919">
<taxonomicName id="4C234D51FFA0FF99F05B900CFDB0B9D7" ID-CoL="3RM5X" authority="Zimmermann, 1919: 123" authorityName="Zimmermann" authorityPageNumber="123" authorityYear="1919" box="[113,613,629,651]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="314" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B957EAC0FFA0FF99F05B900CFE8BB9D6" box="[113,350,629,650]" italics="true" pageId="5" pageNumber="314">Laccophilus ceylonicus</emphasis>
<bibRefCitation id="EFB24B23FFA0FF99F14D900CFDB0B9D7" author="Zimmermann A" box="[359,613,629,651]" firstAuthor="Zimmermann" journalOrPublisher="Archiv fur Naturgeschichte" page="123" pageId="5" pageNumber="314" pagination="68 - 249" part="83" refId="ref16045" refString="Zimmermann A (1919) Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem. Archiv fur Naturgeschichte, 83 (1917) (A 12): 68 - 249." title="Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem" type="journal article" year="1919">Zimmermann, 1919: 123</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(replacement name);
<treatmentCitation id="0A8210C3FFA0FF99F0F490EBFE48B9FB" author="Vazirani TG" box="[222,413,658,680]" page="119" pageId="5" pageNumber="314" year="1972">
<bibRefCitation id="EFB24B23FFA0FF99F0F490EBFE48B9FB" author="Vazirani TG" box="[222,413,658,680]" firstAuthor="Vazirani" journalOrPublisher="Proceedings of the Zoological Society of Calcutta" page="119" pageId="5" pageNumber="314" pagination="117 - 122" part="25" refId="ref15934" refString="Vazirani TG (1972) Contribution to the study of aquatic beetles. 13. A collection of Dytiscidae from Nilgiri Hills, South India, with the description of a new species. Proceedings of the Zoological Society of Calcutta, 25: 117 - 122." title="Contribution to the study of aquatic beetles. 13. A collection of Dytiscidae from Nilgiri Hills, South India, with the description of a new species" type="journal article" year="1972">Vazirani, 1972: 119</bibRefCitation>
</treatmentCitation>
: (
<typeStatus id="54988870FFA0FF99F18690EBFDD0B9F4" box="[428,517,658,680]" pageId="5" pageNumber="314" type="lectotype">lectotype</typeStatus>
designation);
<treatmentCitation id="0A8210C3FFA0FF99F2A090EBFF2FB999" author="Brancucci M" page="253" pageId="5" pageNumber="314" year="1983">
<bibRefCitation id="EFB24B23FFA0FF99F2A090EBFF2FB999" author="Brancucci M" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" page="253" pageId="5" pageNumber="314" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983: 253</bibRefCitation>
</treatmentCitation>
(description, new records).
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</paragraph>
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<subSubSection id="C3396559FFA0FF99F05B9094FDFDBE79" pageId="5" pageNumber="314" type="materials_examined">
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<emphasis id="B957EAC0FFA0FF99F05B9094FEC0B859" bold="true" box="[113,277,749,773]" pageId="5" pageNumber="314">
<typeStatus id="54988870FFA0FF99F05B9094FF79B859" box="[113,172,749,773]" pageId="5" pageNumber="314">Type</typeStatus>
locality.
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<location id="8EFC6009FFA0FF99F10A9094FEA9B859" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F10A9094FEA9B859" box="[288,380,749,773]" country="Sri Lanka" name="Nalanda" pageId="5" pageNumber="314" stateProvince="Central">Nalanda</location>
[=
<collectingCountry id="F3347642FFA0FF99F1839094FDC9B859" box="[425,540,749,773]" name="Sri Lanka" pageId="5" pageNumber="314">Srí Lanka</collectingCountry>
,
<collectingRegion id="49E7F830FFA0FF99F2019094FD39B859" box="[555,748,749,773]" country="Sri Lanka" name="Central" pageId="5" pageNumber="314">Central Province</collectingRegion>
, Nalanda].
</paragraph>
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<emphasis id="B957EAC0FFA0FF99F05B9134FE85B839" bold="true" box="[113,336,845,869]" pageId="5" pageNumber="314">Material examined.</emphasis>
<materialsCitation id="3B4B3C8FFFA0FF99F17C9134FD3CB839" ID-GBIF-Occurrence="3760488383" box="[342,745,845,869]" country="India" location="Maharashtra" pageId="5" pageNumber="314" specimenCount="28" stateProvince="Maharashtra">
<specimenCount id="9D25FD5BFFA0FF99F17C9134FE3FB839" box="[342,490,845,869]" pageId="5" pageNumber="314" type="generic">28 specimens</specimenCount>
.
<emphasis id="B957EAC0FFA0FF99F1DE9134FD3CB839" bold="true" box="[500,745,845,869]" pageId="5" pageNumber="314">
<collectingCountry id="F3347642FFA0FF99F1DE9134FD92B839" box="[500,583,845,869]" name="India" pageId="5" pageNumber="314">INDIA</collectingCountry>
:
<collectingRegion id="49E7F830FFA0FF99F2789134FD3CB839" box="[594,745,845,869]" country="India" name="Maharashtra" pageId="5" pageNumber="314">Maharashtra</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B4B3C8FFFA0FF99F05B9114FD39B8F9" ID-GBIF-Occurrence="3760488372" collectingDate="1991-09-16" collectionCode="GWCV, NHMW" collectorName="R. Schuh" country="India" elevation="1300" location="Mahabaleshwar" pageId="5" pageNumber="314" specimenCount="7" specimenCount-female="6" specimenCount-male="1">
<specimenCount id="9D25FD5BFFA0FF99F05B9114FF6CB8D9" box="[113,185,877,901]" pageId="5" pageNumber="314" type="male">1 male</specimenCount>
<specimenCount id="9D25FD5BFFA0FF99F0959114FEFDB8D9" box="[191,296,877,901]" pageId="5" pageNumber="314" type="female">6 females</specimenCount>
,
<locationDeviation id="9A2E2FF0FFA0FF99F1189114FE2EB8D9" box="[306,507,877,901]" location="Poona" pageId="5" pageNumber="314">
<quantity id="4CDB9B37FFA0FF99F1189114FE57B8D9" box="[306,386,877,901]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="314" unit="km" value="100.0">100 km</quantity>
SW Poona
</locationDeviation>
,
<location id="8EFC6009FFA0FF99F22F9114FD64B8D9" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F22F9114FD64B8D9" box="[517,689,877,901]" country="India" name="Mahabaleshwar" pageId="5" pageNumber="314">Mahabaleshwar</location>
,
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</quantity>
,
<date id="FF9D1012FFA0FF99F0BC91F4FEB8B8F9" box="[150,365,909,933]" pageId="5" pageNumber="314" value="1991-09-16">
<collectingDate id="EFD9E9FAFFA0FF99F0BC91F4FEB8B8F9" box="[150,365,909,933]" pageId="5" pageNumber="314" value="1991-09-16">16 September 1991</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F19B91F7FDC3B8F9" box="[433,534,909,933]" pageId="5" pageNumber="314">R. Schuh</collectorName>
(
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,
<collectionCode id="ED32AE17FFA0FF99F2A291F4FD31B8F9" box="[648,740,909,933]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="314" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA0FF99F05B91D4FDCCB8B9" ID-GBIF-Occurrence="3760488342" collectingDate="2005-09-30" collectingDateMax="2005-10-02" collectingDateMin="2005-09-30" collectionCode="NMPC, PMCB" collectorName="J. Bezdek" country="India" elevation="1400" location="Mahabaleshwar" pageId="5" pageNumber="314" specimenCount="12">
<specimenCount id="9D25FD5BFFA0FF99F05B91D4FF60B899" box="[113,181,941,965]" pageId="5" pageNumber="314" type="generic">12 ex.</specimenCount>
, same locality, but
<quantity id="4CDB9B37FFA0FF99F1AC91D4FE0CB899" box="[390,473,941,965]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="314" unit="m" value="1400.0">
<elevation id="000ED1E1FFA0FF99F1AC91D4FE0CB899" box="[390,473,941,965]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="314" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="FF9D1012FFA0FF99F1CF91D4FF7CB8B9" pageId="5" pageNumber="314" value="2005-09-30" valueMax="2005-10-02" valueMin="2005-09-30">
<collectingDate id="EFD9E9FAFFA0FF99F1CF91D4FF7CB8B9" pageId="5" pageNumber="314" value="2005-09-30" valueMax="2005-10-02" valueMin="2005-09-30">30 September2 October 2005</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F0C791B7FE82B8B9" box="[237,343,973,997]" pageId="5" pageNumber="314">J. Bezděk</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F14C91B4FE63B8B9" box="[358,438,973,997]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="314">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFA0FF99F1E991B4FDC7B8B9" box="[451,530,973,997]" pageId="5" pageNumber="314">PMCB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA0FF99F20C91B4FD40BF79" ID-GBIF-Occurrence="3760488409" collectingDate="2002-08-01" collectingDateMax="2002-08-12" collectingDateMin="2002-08-01" collectionCode="NMPC" collectorName="P. Sipek &amp; M. Fikacek" country="India" location="Igatpuri" pageId="5" pageNumber="314" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D25FD5BFFA0FF99F20C91B4FD44B8B9" box="[550,657,973,997]" pageId="5" pageNumber="314" type="female">2 females</specimenCount>
,
<locationDeviation id="9A2E2FF0FFA0FF99F2B491B4FED4BF59" location="Mumbai" pageId="5" pageNumber="314">
<quantity id="4CDB9B37FFA0FF99F2B491B4FD25B8B9" box="[670,752,973,997]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="314" unit="km" value="120.0">120 km</quantity>
NE Mumbai
</locationDeviation>
,
<location id="8EFC6009FFA0FF99F13A9194FEBDBF59" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F13A9194FEBDBF59" box="[272,360,1005,1029]" country="India" name="Igatpuri" pageId="5" pageNumber="314">Igatpuri</location>
env., 19.42,17N 73.33,06E,
<date id="FF9D1012FFA0FF99F2939194FF28BF79" pageId="5" pageNumber="314" value="2002-08-01" valueMax="2002-08-12" valueMin="2002-08-01">
<collectingDate id="EFD9E9FAFFA0FF99F2939194FF28BF79" pageId="5" pageNumber="314" value="2002-08-01" valueMax="2002-08-12" valueMin="2002-08-01">112 August 2002</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F1169677FE40BF74" box="[316,405,1038,1064]" pageId="5" pageNumber="314">P. Šípek</collectorName>
&amp;
<collectorName id="26D65304FFA0FF99F19F9677FDFBBF79" box="[437,558,1037,1061]" pageId="5" pageNumber="314">M. Fikáček</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F2169674FD5BBF79" box="[572,654,1037,1061]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="314">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4B3C8FFFA0FF99F2B59674FD6ABF39" ID-GBIF-Occurrence="3760488413" collectionCode="NHMB" collectorName="G. Topal" location="Jajpur - Keonjahr" municipality="Daitari Distr." pageId="5" pageNumber="314" specimenCount="1" specimenCount-male="1" stateProvince="Orissa">
<emphasis id="B957EAC0FFA0FF99F2B59674FD3CBF79" bold="true" box="[671,745,1037,1061]" pageId="5" pageNumber="314">
<collectingRegion id="49E7F830FFA0FF99F2B59674FD3CBF79" box="[671,745,1037,1061]" country="India" name="Odisha" pageId="5" pageNumber="314">Orissa</collectingRegion>
</emphasis>
:
<specimenCount id="9D25FD5BFFA0FF99F05B9654FF15BF19" box="[113,192,1069,1093]" pageId="5" pageNumber="314" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFA0FF99F0E49654FEB6BF19" box="[206,355,1069,1093]" pageId="5" pageNumber="314">Daitari Distr.</collectingMunicipality>
,
<location id="8EFC6009FFA0FF99F15B9654FDECBF19" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F15B9654FDECBF19" box="[369,569,1069,1093]" country="India" municipality="Daitari Distr." name="Jajpur - Keonjahr" pageId="5" pageNumber="314" stateProvince="Orissa">Jajpur Keonjahr</location>
[
<location id="8EFC6009FFA0FF99F2639654FD33BF19" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F2639654FD33BF19" box="[585,742,1069,1093]" municipality="Daitari Distr." name="Keonjhargarh" pageId="5" pageNumber="314" stateProvince="Orissa">Keonjhargarh</location>
], trapped during monsoon 1967, coll.
<collectorName id="26D65304FFA0FF99F1DB9634FD9ABF39" box="[497,591,1101,1125]" pageId="5" pageNumber="314">G. Topál</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F2779634FD66BF39" box="[605,691,1101,1125]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="5" pageNumber="314">NHMB</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4B3C8FFFA0FF99F2EE9634FD0CBFD9" ID-GBIF-Occurrence="3760488364" collectionCode="ZSMG" collectorName="W. Horn" country="Sri Lanka" location="Nalanda" pageId="5" pageNumber="314" specimenCount="1" specimenCount-female="1">
<emphasis id="B957EAC0FFA0FF99F2EE9634FF04BFD9" bold="true" pageId="5" pageNumber="314">
<collectingCountry id="F3347642FFA0FF99F2EE9634FF04BFD9" name="Sri Lanka" pageId="5" pageNumber="314">SRI LANKA</collectingCountry>
</emphasis>
:
<specimenCount id="9D25FD5BFFA0FF99F0F79614FEEEBFD9" box="[221,315,1133,1157]" pageId="5" pageNumber="314" type="female">1 female</specimenCount>
,
<location id="8EFC6009FFA0FF99F1699614FE4BBFD9" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F1699614FE4BBFD9" box="[323,414,1133,1157]" country="Sri Lanka" name="Nalanda" pageId="5" pageNumber="314">Nalanda</location>
, 1899, coll.
<collectorName id="26D65304FFA0FF99F2319617FDA3BFD9" box="[539,630,1134,1157]" pageId="5" pageNumber="314">W. Horn</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F2A99614FD07BFD9" box="[643,722,1133,1157]" pageId="5" pageNumber="314">ZSMG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA0FF99F2C99614FF0FBF99" ID-GBIF-Occurrence="3760488404" collectionCode="GWCV" collectorName="E. Bugnion" country="Sri Lanka" location="Puwakpitya" pageId="5" pageNumber="314" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D25FD5BFFA0FF99F2C99614FF73BFF9" pageId="5" pageNumber="314" type="male">1 male</specimenCount>
,
<location id="8EFC6009FFA0FF99F08596F4FEF8BFF9" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F08596F4FEF8BFF9" box="[175,301,1165,1189]" country="Sri Lanka" name="Puwakpitya" pageId="5" pageNumber="314">Puwakpitya</location>
, hiver [winter] 19061907, coll.
<collectorName id="26D65304FFA0FF99F25096F7FD25BFF9" box="[634,752,1165,1189]" pageId="5" pageNumber="314">E. Bugnion</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F05396D4FF07BF99" box="[121,210,1197,1221]" pageId="5" pageNumber="314">GWCV</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA0FF99F0C296D4FD39BFB9" ID-GBIF-Occurrence="3760488425" collectingDate="1970-12-08" collectionCode="GWCV" collectorName="Starmulluer" country="Sri Lanka" location="Veli Oya Belihuoya" pageId="5" pageNumber="314" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D25FD5BFFA0FF99F0C296D4FE98BF99" box="[232,333,1197,1221]" pageId="5" pageNumber="314" type="female">1 female</specimenCount>
,
<location id="8EFC6009FFA0FF99F17196D4FDE9BF99" LSID="urn:lsid:plazi:treatment:038A87C4FFA0FF99F09393A4FEBCBDF9:8EFC6009FFA0FF99F17196D4FDE9BF99" box="[347,572,1197,1221]" country="Sri Lanka" name="Veli Oya Belihuoya" pageId="5" pageNumber="314">Veli Oya Belihuoya</location>
[most probably Belihul Oya],
<date id="FF9D1012FFA0FF99F12D96B4FE10BFB9" box="[263,453,1229,1253]" pageId="5" pageNumber="314" value="1970-12-08">
<collectingDate id="EFD9E9FAFFA0FF99F12D96B4FE10BFB9" box="[263,453,1229,1253]" pageId="5" pageNumber="314" value="1970-12-08">8 December 1970</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F22896B4FD55BFB9" box="[514,640,1229,1253]" pageId="5" pageNumber="314">Starmülluer</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F2A796B4FD30BFB9" box="[653,741,1229,1253]" pageId="5" pageNumber="314">GWCV</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3C8FFFA0FF99F05B9694FDFDBE79" ID-GBIF-Occurrence="3760488313" collectingDate="1991-04-19" collectingDateMax="1991-05-09" collectingDateMin="1991-04-19" collectionCode="NMPC, NHMB" collectorName="J. Kolibac" elevation="300" location="Dambulla" municipality="Dambulla" pageId="5" pageNumber="314" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D25FD5BFFA0FF99F05B9694FF10BE59" box="[113,197,1261,1285]" pageId="5" pageNumber="314" type="male">2 males</specimenCount>
<specimenCount id="9D25FD5BFFA0FF99F0E49694FEE5BE59" box="[206,304,1261,1285]" pageId="5" pageNumber="314" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BF8ACA8FFA0FF99F1179694FE73BE59" box="[317,422,1261,1285]" country="Sri Lanka" name="Dambulla" pageId="5" pageNumber="314">Dambulla</collectingMunicipality>
env.,
<quantity id="4CDB9B37FFA0FF99F1C19694FDE6BE59" box="[491,563,1261,1285]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="314" unit="m" value="300.0">
<elevation id="000ED1E1FFA0FF99F1C19694FDE6BE59" box="[491,563,1261,1285]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="314" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FF9D1012FFA0FF99F26A9694FF7CBE79" pageId="5" pageNumber="314" value="1991-04-19" valueMax="1991-05-09" valueMin="1991-04-19">
<collectingDate id="EFD9E9FAFFA0FF99F26A9694FF7CBE79" pageId="5" pageNumber="314" value="1991-04-19" valueMax="1991-05-09" valueMin="1991-04-19">19 April9 May 1991</collectingDate>
</date>
, coll.
<collectorName id="26D65304FFA0FF99F0C79777FE8EBE79" box="[237,347,1293,1317]" pageId="5" pageNumber="314">J. Kolibáč</collectorName>
(
<collectionCode id="ED32AE17FFA0FF99F1419774FE6EBE79" box="[363,443,1293,1317]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="314">NMPC</collectionCode>
,
<collectionCode id="ED32AE17FFA0FF99F1E29774FDC8BE79" box="[456,541,1293,1317]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="5" pageNumber="314">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F05B9734FDD6BEB9" pageId="5" pageNumber="314" type="diagnosis">
<paragraph id="8B9C36D2FFA0FF99F05B9734FDD6BEB9" blockId="5.[113,752,1357,1509]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F05B9734FF30BE39" bold="true" box="[113,229,1357,1381]" pageId="5" pageNumber="314">Diagnosis.</emphasis>
Medium sized, elongate oval species (body length
<quantity id="4CDB9B37FFA0FF99F05B9714FF23BED9" box="[113,246,1389,1413]" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="5" pageNumber="314" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.84.1 mm</quantity>
). Elytra with subbasal testaceous band sinuous, narrow, well separated from elytral base (
<figureCitation id="13182A57FFA0FF99F20D97F4FDB2BEF9" box="[551,615,1421,1445]" captionStart="Figs" captionStartId="3.[113,157,2051,2073]" captionTargetBox="[115,1452,190,2035]" captionTargetId="figure-10@3.[97,1463,180,2039]" captionTargetPageId="3" captionText="Figs. 19. Habitus of Laccophilus: 1, L. aurofasciatus; 2, L. auropictus; 3, L. ceylonicus; 4, L. elegans; 5, L. girardi; 6, L. hendrichi, new species; 7, L. jaechi, new species; 8, L. javanicus; 9, L. komareki, new species. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5385200" httpUri="https://zenodo.org/record/5385200/files/figure.png" pageId="5" pageNumber="314">Fig. 3</figureCitation>
). Apical part of median lobe in lateral view with distinct bulge on dorsal side; apex with pointed tip (
<figureCitation id="13182A57FFA0FF99F18F97B4FE20BEB9" box="[421,501,1485,1509]" captionStart="Figs" captionStartId="7.[113,157,2006,2028]" captionTargetBox="[120,1445,189,1990]" captionTargetId="figure-10@7.[110,1454,186,1997]" captionTargetPageId="7" captionText="Figs. 1835. Median lobe in lateral view: 18, Laccophilus aurofasciatus; 19, L. auropictus; 20, L. ceylonicus; 21, L. elegans; 22, L. girardi; 23, L. hendrichi, new species; 24, L. jaechi, new species; 25, L. javanicus; 26, L. kalimantanensis, new species; 27, L. komareki, new species. 28, L. mahakamensis; 29, L. mazzoldii, new species; 30, L. nusatenggaraensis, new species; 31, L. schillhammeri, new species; 32, L. stastnyi, new species; 33, L. tobaensis; 34, L. tonkinensis; 35, L. wittmeri. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5385204" httpUri="https://zenodo.org/record/5385204/files/figure.png" pageId="5" pageNumber="314">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396559FFA0FF99F05B9474FEBCBDF9" pageId="5" pageNumber="314" type="distribution">
<paragraph id="8B9C36D2FFA0FF99F05B9474FEBCBDF9" blockId="5.[113,753,1549,1701]" pageId="5" pageNumber="314">
<emphasis id="B957EAC0FFA0FF99F05B9474FEC7BD79" bold="true" box="[113,274,1549,1573]" pageId="5" pageNumber="314">Distribution.</emphasis>
The species was described and known previously only from
<collectingCountry id="F3347642FFA0FF99F1459454FE0ABD19" box="[367,479,1581,1605]" name="Sri Lanka" pageId="5" pageNumber="314">Sri Lanka</collectingCountry>
(
<bibRefCitation id="EFB24B23FFA0FF99F1DB9454FD65BD19" author="Brancucci M" box="[497,688,1581,1605]" firstAuthor="Brancucci" journalOrPublisher="Entomologische Arbeiten aus dem Museum G. Frey" pageId="5" pageNumber="314" pagination="241 - 426" part="31 / 32" refId="ref15135" refString="Brancucci M (1983) Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum G. Frey, 31 / 32: 241 - 426." title="Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae)" type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
). The present records from Maharashtra and Orissa allowed us to expect that is widely distributed also in the Indian subcontinent (
<figureCitation id="13182A57FFA0FF99F12194F4FE88BDF9" box="[267,349,1677,1701]" captionStart="Figs" captionStartId="9.[113,157,2093,2115]" captionTargetBox="[141,1427,190,2077]" captionTargetPageId="9" captionText="Figs. 4041. Distribution of the species of the Laccophilus javanicus group" figureDoi="http://doi.org/10.5281/zenodo.5385208" httpUri="https://zenodo.org/record/5385208/files/figure.png" pageId="5" pageNumber="314">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>