159 lines
21 KiB
XML
159 lines
21 KiB
XML
<document id="47869A9C6178A7D30176E8E1CCD6F70B" ID-CLB-Dataset="26068" ID-DOI="10.11646/zootaxa.4652.1.13" ID-GBIF-Dataset="b79add95-a479-45e5-ba55-60409cfac833" ID-ISSN="1175-5326" ID-Zenodo-Dep="3363525" ID-ZooBank="87176577-EF1F-4A8A-918C-2775AE9D2425" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565270295833" checkinUser="plazi" docAuthor="Parisi, Francesco & Fanti, Fabrizio" docDate="2019" docId="038A87B76864FFA1FF45FCD4AE598CFC" docLanguage="en" docName="zootaxa.4652.1.13.pdf" docOrigin="Zootaxa 4652 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malthodes (Malthodes) tognettii Parisi & Fanti 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="190" masterDocId="FFB3FFCF6865FFA3FFD2FFF5AB778C07" masterDocTitle="A new fossil Malthodes Kiesenwetter, 1852 from the Eocene Baltic amber (Coleoptera Cantharidae)" masterLastPageNumber="195" masterPageNumber="189" pageNumber="190" updateTime="1698749081420" updateUser="ExternalLinkService">
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<mods:title id="E62C37602C9004DAA6044FA509C1A622">A new fossil Malthodes Kiesenwetter, 1852 from the Eocene Baltic amber (Coleoptera Cantharidae)</mods:title>
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<mods:namePart id="41800B250DB105409D54439E234B6D31">Parisi, Francesco</mods:namePart>
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<mods:namePart id="68ECC24F900AE49E7BBDAB1B5AD61DDE">Fanti, Fabrizio</mods:namePart>
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<mods:title id="1879C83E71AC7C64869A8AE97EBA760B">Zootaxa</mods:title>
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<mods:date id="0C23606A908E393CEA9CE278FCFC6B07">2019</mods:date>
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<treatment id="038A87B76864FFA1FF45FCD4AE598CFC" ID-DOI="http://doi.org/10.5281/zenodo.5934506" ID-GBIF-Taxon="159043955" ID-Zenodo-Dep="5934506" LSID="urn:lsid:plazi:treatment:038A87B76864FFA1FF45FCD4AE598CFC" httpUri="http://treatment.plazi.org/id/038A87B76864FFA1FF45FCD4AE598CFC" lastPageId="2" lastPageNumber="190" pageId="1" pageNumber="190">
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<subSubSection id="C339652A6864FFA2FF45FCD4A9F78F3C" box="[151,640,801,828]" pageId="1" pageNumber="190" type="nomenclature">
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<paragraph id="8B9C36A16864FFA2FF45FCD4A9F78F3C" blockId="1.[151,640,801,863]" box="[151,640,801,828]" pageId="1" pageNumber="190">
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<heading id="D0D481CD6864FFA2FF45FCD4A9F78F3C" bold="true" box="[151,640,801,828]" fontSize="11" level="1" pageId="1" pageNumber="190" reason="1">
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<emphasis id="B957EAB36864FFA2FF45FCD4A9F78F3C" bold="true" box="[151,640,801,828]" pageId="1" pageNumber="190">
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<taxonomicName id="4C234D226864FFA2FF45FCD4A96C8F3C" authority="Parisi & Fanti, 2019" authorityName="Parisi & Fanti" authorityYear="2019" box="[151,539,801,828]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="tognettii" status="sp. nov." subGenus="Malthodes">
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<emphasis id="B957EAB36864FFA2FF45FCD4AA618F3B" bold="true" box="[151,278,801,828]" italics="true" pageId="1" pageNumber="190">Malthodes</emphasis>
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(
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<emphasis id="B957EAB36864FFA2FEF5FCD4AAD18F3B" bold="true" box="[295,422,801,828]" italics="true" pageId="1" pageNumber="190">Malthodes</emphasis>
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)
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<emphasis id="B957EAB36864FFA2FE6AFCD4A96C8F3C" bold="true" box="[440,539,801,827]" italics="true" pageId="1" pageNumber="190">tognettii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26457C86864FFA2FDF0FCD4A9F78F3C" box="[546,640,801,827]" pageId="1" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C339652A6864FFA2FF45FCB0AA658F58" box="[151,274,837,863]" pageId="1" pageNumber="190" type="description">
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<paragraph id="8B9C36A16864FFA2FF45FCB0AA658F58" blockId="1.[151,640,801,863]" box="[151,274,837,863]" pageId="1" pageNumber="190">
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(
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<figureCitation id="13182A246864FFA2FF4DFCB0AA7E8F58" box="[159,265,837,863]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1721,1746]" captionStartId-1="4.[151,250,1793,1818]" captionStartId-2="5.[151,250,1801,1826]" captionTargetBox-0="[151,1436,273,1702]" captionTargetBox-1="[151,1436,345,1774]" captionTargetBox-2="[229,1358,273,1782]" captionTargetId-0="figure@2.[151,1436,273,1702]" captionTargetId-1="figure@4.[151,1436,345,1774]" captionTargetId-2="figure@5.[229,1358,273,1782]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Malthodes (Malthodes) tognettii sp. nov. in Baltic amber, holotype. A: dorsolateral view, scale bar = 0.5 mm. B: reconstruction, scale bar = 0.5 mm." captionText-1="FIGURE 2. Malthodes (Malthodes) tognettii sp. nov. in Baltic amber, holotype. A: detail of antennae, scale bar = 0.5 mm. B: detail of the head, pronotum and elytra (lateral view), scale bar = 0.3 mm." captionText-2="FIGURE 3. Malthodes (Malthodes) tognettii sp. nov. in Baltic amber, holotype. A: detail of the last abdominal segments, scale bar = 0.1 mm. B: line drawing of the last abdominal segments, scale bar = 0.1 mm." httpUri-0="https://zenodo.org/record/3363527/files/figure.png" httpUri-1="https://zenodo.org/record/3363529/files/figure.png" httpUri-2="https://zenodo.org/record/3363531/files/figure.png" pageId="1" pageNumber="190">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C339652A6864FFA1FF45FC79AE598CFC" lastPageId="2" lastPageNumber="191" pageId="1" pageNumber="190" type="materials_examined">
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<paragraph id="8B9C36A16864FFA2FF45FC79AE618A70" blockId="1.[151,1437,908,2015]" pageId="1" pageNumber="190">
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<emphasis id="B957EAB36864FFA2FF45FC79AA5E8FA0" bold="true" box="[151,297,908,935]" pageId="1" pageNumber="190">Description.</emphasis>
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Adult, winged, slender, male defined on the basis of the last urites strongly modified. Body length:
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<quantity id="4CDB9B446864FFA2FAABFC78ABB28FCC" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="190" unit="mm" value="2.1">2.1 mm</quantity>
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, antennae: about 2.0 mm. Entirely dark brown without yellow spots on elytra. Head exposed, slightly elongate, rounded behind the eyes, with shallow punctation and small hairs. Eyes very large, rounded, prominent, inserted in the lateral-upper part of the head. Maxillary palpi 4-segmented, first palpomere short, second palpomere triangular, third robust and elongate, last palpomere globular and distally pointed. Labial palpi 3-segmented, last palpomere globular and distally pointed. Antennae filiform, 11-segmented, elongate, surpassing apex of elytra and as long as abdomen; antennomere I elongate, club-shaped; antennomere II about 1.7 times shorter than antennomere I; antennomere III longer than antennomere II; antennomeres IV-VIII subequal in length, slightly longer than antennomere III; antennomere IX shorter than previous; antennomere X as long as antennomere III; antennomere XI elongate, rounded at apex; all antennomeres covered by long setae. Pronotum strongly transverse, narrower than the head, surface scarcely punctate and with short setae, sides almost straight, posterior margin slightly bordered. Elytra slightly narrower than pronotum, covered with several short and erected setae, parallel-sided, reaching the base of the eighth abdominal segment, rounded at the apexes. Hind wings infuscate, surpassing elytra and almost reaching last abdominal segment. Legs rather robust, pubescent; coxae elongate, trochanters with rounded apex; femora enlarged; tibiae longer than the femora, thin, cylindrical, with a spur at the apex. Tarsi 5-segmented, pubescent; tarsomeres I thin, elongate; tarsomeres II shorter and stouter than tarsomere I; tarsomeres III shorter than second; tarsomeres IV bilobed; tarsomeres
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<collectionCode id="ED32AE646864FFA2FE70FA38AAC189E0" box="[418,438,1485,1511]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="190" type="Museum">V</collectionCode>
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elongate and slender; claws simple. Metasternum elongate. Sternites are transverse and slightly pubescent. Last tergite (tg10) elongate, almost flat and slightly cylindrical, slightly bent downwards from half length, apically weakly forked; last sternite (st9) elongate, strongly curved, folded backwards and weakly forked apically, the initial part of the last sternite is cylindrical while the backward bending is flat with the sides bent upwards in a kind of small wings or lobes and concave inside. Aedeagus not visible. Female unknown.
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</paragraph>
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<paragraph id="8B9C36A16864FFA2FF15F974AEFE8A9C" blockId="1.[151,1437,908,2015]" box="[199,1417,1665,1692]" pageId="1" pageNumber="190">
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<emphasis id="B957EAB36864FFA2FF15F974AA398A9C" bold="true" box="[199,334,1665,1691]" pageId="1" pageNumber="190">Etymology.</emphasis>
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Named in honor of Roberto Tognetti (University of
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<collectingRegion id="49E7F8436864FFA2FC48F974A89C8A9C" box="[922,1003,1665,1691]" country="Italy" name="Molise" pageId="1" pageNumber="190">Molise</collectingRegion>
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,
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<collectingCountry id="F33476316864FFA2FC27F974AF5B8A9B" box="[1013,1068,1665,1692]" name="Italy" pageId="1" pageNumber="190">Italy</collectingCountry>
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), the mentor of the first author.
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</paragraph>
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<paragraph id="8B9C36A16864FFA2FF15F950AA558AE4" blockId="1.[151,1437,908,2015]" pageId="1" pageNumber="190">
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<materialsCitation id="3B4B3CFC6864FFA2FF15F950AA558AE4" ID-GBIF-Occurrence="2313474987" collectorName="No. Unimol" location="Molise" pageId="1" pageNumber="190" specimenCount="1" stateProvince="Molise" typeStatus="holotype">
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<emphasis id="B957EAB36864FFA2FF15F950AA4C8AB8" bold="true" box="[199,315,1701,1727]" pageId="1" pageNumber="190">
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<typeStatus id="549888036864FFA2FF15F950AA418AB8" box="[199,310,1701,1727]" pageId="1" pageNumber="190" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male, in Baltic amber, deposited at the University of
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<collectingRegion id="49E7F8436864FFA2FC73F950A8988AB8" box="[929,1007,1701,1727]" country="Italy" name="Molise" pageId="1" pageNumber="190">Molise</collectingRegion>
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(Unimol) with accession
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<collectorName id="26D653776864FFA2FAC7F950AEEA8AB8" box="[1301,1437,1701,1727]" pageId="1" pageNumber="190">No. Unimol</collectorName>
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AAA002FP.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9C36A16864FFA2FF15F918AF418B00" blockId="1.[151,1437,908,2015]" box="[199,1078,1772,1799]" pageId="1" pageNumber="190">
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<emphasis id="B957EAB36864FFA2FF15F918AA118B00" bold="true" box="[199,358,1772,1799]" pageId="1" pageNumber="190">Type locality.</emphasis>
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Yantarny mine, Sambian Peninsula,
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<collectingRegion id="49E7F8436864FFA2FCD0F918A8AF8B00" box="[770,984,1773,1799]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="190">Kaliningrad region</collectingRegion>
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,
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<collectingCountry id="F33476316864FFA2FC36F918AF458B00" box="[996,1074,1773,1799]" name="Russia" pageId="1" pageNumber="190">Russia</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B9C36A16864FFA2FF15F8E4AF398B2C" blockId="1.[151,1437,908,2015]" box="[199,1102,1808,1836]" pageId="1" pageNumber="190">
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<emphasis id="B957EAB36864FFA2FF15F8E4AA1B8B2C" bold="true" box="[199,364,1808,1835]" pageId="1" pageNumber="190">Type horizon.</emphasis>
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Upper Eocene (Priabonian), Prussian Formation: 37.8–33.9 Mya.
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</paragraph>
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<paragraph id="8B9C36A16864FFA2FF15F8C1A8068B48" blockId="1.[151,1437,908,2015]" box="[199,881,1844,1871]" pageId="1" pageNumber="190">
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<emphasis id="B957EAB36864FFA2FF15F8C1AA058B48" bold="true" box="[199,370,1844,1871]" pageId="1" pageNumber="190">Syninclusions.</emphasis>
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Air bubbles, wood remains and stellate hairs.
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</paragraph>
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<paragraph id="8B9C36A16864FFA1FF15F8ADAE598CFC" blockId="1.[151,1437,908,2015]" lastBlockId="2.[151,1436,152,251]" lastPageId="2" lastPageNumber="191" pageId="1" pageNumber="190">
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<emphasis id="B957EAB36864FFA2FF15F8ADAAB98B74" bold="true" box="[199,462,1880,1907]" pageId="1" pageNumber="190">Differential diagnosis.</emphasis>
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No living species of
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<taxonomicName id="4C234D226864FFA2FD6DF8ACA8428B74" authorityName="Kiesenwetter" authorityYear="1852" box="[703,821,1881,1907]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="190" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAB36864FFA2FD6DF8ACA8428B74" box="[703,821,1881,1907]" italics="true" pageId="1" pageNumber="190">Malthodes</emphasis>
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</taxonomicName>
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from the Baltic area and Central Europe or the Alps is phylogenetically related to the new species.
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<taxonomicName id="4C234D226864FFA2FD5EF888AF008B90" authority="Kiesenwetter, 1852" authorityName="Kiesenwetter" authorityYear="1852" box="[652,1143,1916,1943]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="hexacanthus">
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<emphasis id="B957EAB36864FFA2FD5EF888A8E18B90" box="[652,918,1917,1943]" italics="true" pageId="1" pageNumber="190">Malthodes hexacanthus</emphasis>
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<bibRefCitation id="EFB24B506864FFA2FC4CF888AF008B90" author="Kiesenwetter, E. A. H. von" box="[926,1143,1916,1943]" pageId="1" pageNumber="190" pagination="239 - 324" refId="ref5036" refString="Kiesenwetter, E. A. H. von (1852) Beitrage zu einer Monographie der Malthinen. Linnaea entomologica, 7, 239 - 324, tables 1 - 2." type="journal article" year="1852">Kiesenwetter, 1852</bibRefCitation>
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</taxonomicName>
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has similarities only with regard to the last sternite (st9) which however in the latter species does not have lateral extensions near the apex and it is deeply forked (
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<bibRefCitation id="EFB24B506864FFA2FEA3F830A93E8BD8" author="Kiesenwetter, E. A. H. von" box="[369,585,1988,2015]" pageId="1" pageNumber="190" pagination="239 - 324" refId="ref5036" refString="Kiesenwetter, E. A. H. von (1852) Beitrage zu einer Monographie der Malthinen. Linnaea entomologica, 7, 239 - 324, tables 1 - 2." type="journal article" year="1852">Kiesenwetter 1852</bibRefCitation>
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;
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<bibRefCitation id="EFB24B506864FFA2FD87F830A9968BD8" author="Liberti, G." box="[597,737,1989,2015]" pageId="1" pageNumber="190" pagination="147 - 246" refId="ref5199" refString="Liberti, G. (2011) Le specie di Malthodes Kiesenwetter, 1852 delle Alpi Marittime e Liguri (Coleoptera, Cantharidae). Annali del Museo Civico di Storia Naturale " G. Doria ", Genova, 103, 147 - 246." type="journal article" year="2011">Liberti 2011</bibRefCitation>
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) and not weakly as in
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<taxonomicName id="4C234D226864FFA2FC0FF833AF288BD8" authority="Parisi & Fanti, 2019" authorityName="Parisi & Fanti" authorityYear="2019" box="[989,1119,1990,2015]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="190" phylum="Arthropoda" rank="species" species="tognettii" status="sp. nov.">
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<emphasis id="B957EAB36864FFA2FC0FF833AF288BD8" box="[989,1119,1990,2015]" italics="true" pageId="1" pageNumber="190">M. tognettii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26457C86864FFA2FBB5F830AFB68BD8" box="[1127,1217,1989,2015]" pageId="1" pageNumber="190" rank="species">
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<emphasis id="B957EAB36864FFA2FBB5F830AFB68BD8" bold="true" box="[1127,1217,1989,2015]" pageId="1" pageNumber="190">sp. nov.</emphasis>
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</taxonomicNameLabel>
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No fossil is similar to the new species, except for
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<taxonomicName id="4C234D226867FFA1FE22FF6CAF5A8CB4" authority="Kuska & Kupryjanowicz, 2005" authorityName="Kuska & Kupryjanowicz" authorityYear="2005" box="[496,1069,152,179]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="serafini">
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<emphasis id="B957EAB36867FFA1FE22FF6CA9B38CB4" box="[496,708,153,179]" italics="true" pageId="2" pageNumber="191">Malthodes serafini</emphasis>
|
||
<bibRefCitation id="EFB24B506867FFA1FD1EFF6CAF5A8CB4" author="Kuska, A. & Kupryjanowicz, J." box="[716,1069,152,179]" pageId="2" pageNumber="191" pagination="309 - 316" refId="ref5166" refString="Kuska, A. & Kupryjanowicz, J. (2005) Soldier beetles (Coleoptera: Cantharidae) from Baltic amber. Polskie Pismo Entomologiczne, 74, 309 - 316." type="journal article" year="2005">Kuśka & Kupryjanowicz, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
which has the last tergite (tg10) very similar though less bent and (as for
|
||
<taxonomicName id="4C234D226867FFA1FD88FF4BA87C8CD0" authorityName="Kiesenwetter" authorityYear="1852" box="[602,779,189,215]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="191" phylum="Arthropoda" rank="species" species="hexacanthus">
|
||
<emphasis id="B957EAB36867FFA1FD88FF4BA87C8CD0" box="[602,779,189,215]" italics="true" pageId="2" pageNumber="191">M. hexacanthus</emphasis>
|
||
</taxonomicName>
|
||
) does not have lateral expansions on the last sternite which is also strongly bifurcated at the apex (
|
||
<bibRefCitation id="EFB24B506867FFA1FD9AFF14A8D38CFC" author="Kuska, A. & Kupryjanowicz, J." box="[584,932,224,251]" pageId="2" pageNumber="191" pagination="309 - 316" refId="ref5166" refString="Kuska, A. & Kupryjanowicz, J. (2005) Soldier beetles (Coleoptera: Cantharidae) from Baltic amber. Polskie Pismo Entomologiczne, 74, 309 - 316." type="journal article" year="2005">Kuśka & Kupryjanowicz 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB24B506867FFA1FC62FF14AF4E8CFC" author="Fanti, F." box="[944,1081,225,251]" pageId="2" pageNumber="191" pagination="1 - 18" refId="ref4327" refString="Fanti, F. (2017 a) Catalogo Cantharidae fossili del mondo. Fossils & Minerals Review, 2, 1 - 18 [abbreviated Italian version, available: 12 March 2017] / world catalog of fossil Cantharidae. Fossils & Minerals Review, 2 (Special Issue), 1 - 52. [extended English version, available: 25 May 2017]" type="journal article" year="2017">Fanti 2017a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB24B506867FFA1FB91FF14AE558CFC" author="Fanti, F. & Vitali, F." box="[1091,1314,225,251]" pageId="2" pageNumber="191" pagination="19 - 27" refId="ref4647" refString="Fanti, F. & Vitali, F. (2017) Key to fossil Malthininae, with description of two new species in Baltic amber (Coleoptera Canthari- dae). Baltic Journal of Coleopterology, 17 (1), 19 - 27." type="journal article" year="2017">Fanti & Vitali 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |