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<document id="4B877B5FA0F85824CEC491817D99E39A" ID-CLB-Dataset="68523" ID-DOI="10.11646/zootaxa.5150.2.1" ID-GBIF-Dataset="1e8c5179-1951-45cd-b1eb-3dedbc03b1b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="6621158" ID-ZooBank="99F57360-DC86-48D5-8527-CBAE787DFFEA" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="admin" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1654625336811" checkinUser="plazi" docAuthor="Philips, T. Keith, Mudge, Alan D., Orozco, Jesus, Neidlinger, Robert, Beinhundner, Gerhard &amp; Joly, Claude" docDate="2022" docId="038A87B6FF9FF815FF13FA7DDA6A97A0" docLanguage="en" docName="zootaxa.5150.2.1.pdf" docOrigin="Zootaxa 5150 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Charadronota Burmeister 1842" docType="treatment" docVersion="8" lastPageNumber="177" masterDocId="FFB3FFCEFF85F80EFF84FFA2DF589759" masterDocTitle="A rich and diverse fauna: an illustrated guide to the cetoniines of Ghana with comparisons to Ivory Coast and Benin (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="188" masterPageNumber="151" pageNumber="177" updateTime="1699334540354" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="11722381C9E5653DE5452C80F25E6CB5">A rich and diverse fauna: an illustrated guide to the cetoniines of Ghana with comparisons to Ivory Coast and Benin (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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<mods:namePart id="85157958906A6BC3F2EA6D989F055A81">Philips, T. Keith</mods:namePart>
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<mods:affiliation id="14E69A4BA783A677E71A28FD844EB4F1">Systematics and Evolution Laboratory, Department of Biology, Western Kentucky University, Bowling Green, KY 42101 - 3576 USA. Keith. Philips @ wku. edu; https: // orcid. org / 0000 - 0002 - 1190 - 4019</mods:affiliation>
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<mods:namePart id="654C984B3CBC2AB96C79F7D9E17CE187">Mudge, Alan D.</mods:namePart>
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<mods:namePart id="6F02DEC1A93377C8D30F45564255ABB1">Orozco, Jesus</mods:namePart>
<mods:affiliation id="2A4C53C5B440CAE82D0588AFEEA90644">Insect Collection, Agricultural Science and Production Department, Zamorano University (Escuela Agrícola Panamericana), Zamorano, Honduras.</mods:affiliation>
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<mods:namePart id="A6ABA0309965A4B445030964E1262F88">Neidlinger, Robert</mods:namePart>
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<mods:affiliation id="F5C8E671FEEE7D14B69FC1A01374037C">Systematics and Evolution Laboratory, Department of Biology, Western Kentucky University, Bowling Green, KY 42101 - 3576 USA. Keith. Philips @ wku. edu; https: // orcid. org / 0000 - 0002 - 1190 - 4019</mods:affiliation>
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<mods:namePart id="5AC0C25306269ECD1BE694962A224082">Beinhundner, Gerhard</mods:namePart>
<mods:affiliation id="82C0E600007827C8A651C66CBE5E5F95">Am Steigberg 25, D- 97502 Eurbach, Germany.</mods:affiliation>
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<mods:namePart id="4D9C81AB533492247B631F935DA5CDD1">Joly, Claude</mods:namePart>
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<paragraph id="8B9C36A0FF9FF814FF13FA7DDD4692A3" blockId="26.[151,542,1503,1530]" box="[151,542,1503,1530]" pageId="26" pageNumber="177">
<heading id="D0D481CCFF9FF814FF13FA7DDD4692A3" bold="true" box="[151,542,1503,1530]" fontSize="11" level="1" pageId="26" pageNumber="177" reason="1">
<taxonomicName id="4C234D23FF9FF814FF13FA7DDD4692A3" authority="Burmeister, 1842" authorityName="Burmeister" authorityYear="1842" box="[151,542,1503,1530]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAB2FF9FF814FF13FA7DDD4692A3" bold="true" box="[151,542,1503,1530]" pageId="26" pageNumber="177">
<emphasis id="B957EAB2FF9FF814FF13FA7DDE1A92A3" bold="true" box="[151,322,1503,1530]" italics="true" pageId="26" pageNumber="177">Charadronota</emphasis>
Burmeister, 1842
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<subSubSection id="C339652BFF9FF814FF13F98ADB5891F7" pageId="26" pageNumber="177" type="multiple">
<paragraph id="8B9C36A0FF9FF814FF13F98ADB5891F7" blockId="26.[151,1437,1576,2034]" pageId="26" pageNumber="177">
The genus
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<emphasis id="B957EAB2FF9FF814FE96F98ADEE9911B" box="[274,433,1576,1602]" italics="true" pageId="26" pageNumber="177">Charadronota</emphasis>
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includes six or seven species of medium-sized cetoniines (
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) widely distributed in tropical Africa (
<bibRefCitation id="EFB24B51FF9FF814FEF6F9EEDD32913F" author="Schoolmeesters" box="[370,618,1612,1638]" firstAuthor="Schoolmeesters" pageId="26" pageNumber="177" refId="ref18501" refString="Schoolmeesters, P. (2021) SCARABS: World Scarabaeidae Database. In: Banki, O., Roskov, Y., Vandepitte, L., DeWalt, R. E., Remsen, D., Schalk, P., Orrell, T., Keping, M., Miller, J., Aalbu, R., Adlard, R., Adriaenssens, E., Aedo, C., Aescht, E., Akkari, N., Alonso-Zarazaga, M. A., Alvarez, B., Alvarez, F., Anderson, G. et al, Catalogue of Life Checklis. Version 2021 - 09 - 01. Catalogue of Life. Leiden. https: // doi. org / 10.48580 / d 4 sl- 38 g" type="book" year="2021">Schoolmeesters 2021</bibRefCitation>
,
<bibRefCitation id="EFB24B51FF9FF814FDFDF9EEDC0A913F" author="Beinhundner" box="[633,850,1612,1638]" firstAuthor="Beinhundner" pageId="26" pageNumber="177" refId="ref16250" refString="Beinhundner, G. (2017) The Cetoniinae of Africa. Privately published, Euerbach, 1174 + XXIV pp." type="book" year="2017">Beinhundner 2017</bibRefCitation>
). Several of these, e.g.
<taxonomicName id="4C234D23FF9FF814FBEFF9EEDE0491D3" authority="(Gory &amp; Percheron, 1883)" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1883" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B957EAB2FF9FF814FBEFF9EEDA75913F" box="[1131,1325,1612,1638]" italics="true" pageId="26" pageNumber="177">C. quadrisignata</emphasis>
(Gory &amp; Percheron, 1883)
</taxonomicName>
,
<taxonomicName id="4C234D23FF9FF814FEEEF9D2DD2791D3" authority="Janson, 1888" authorityName="Janson" authorityYear="1888" box="[362,639,1648,1674]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="curvata">
<emphasis id="B957EAB2FF9FF814FEEEF9D2DEBA91D3" box="[362,482,1648,1674]" italics="true" pageId="26" pageNumber="177">C. curvata</emphasis>
Janson, 1888
</taxonomicName>
and
<taxonomicName id="4C234D23FF9FF814FD3DF9D2DCEB91D3" authority="Kraatz, 1883" authorityName="Kraatz" authorityYear="1883" box="[697,947,1648,1674]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="soror">
<emphasis id="B957EAB2FF9FF814FD3DF9D2DC4091D3" box="[697,792,1648,1674]" italics="true" pageId="26" pageNumber="177">C. soror</emphasis>
Kraatz, 1883
</taxonomicName>
are highly variable in color and markings, which has led to a number of forms, varieties, and synonyms being described.
</paragraph>
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<paragraph id="8B9C36A0FF9FF814FF43F91ADB7290F3" blockId="26.[151,1437,1576,2034]" pageId="26" pageNumber="177">
<taxonomicName id="4C234D23FF9FF814FF43F91ADD53918B" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1883" box="[199,523,1720,1746]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B957EAB2FF9FF814FF43F91ADD53918B" box="[199,523,1720,1746]" italics="true" pageId="26" pageNumber="177">Charadronota quadrisignata</emphasis>
</taxonomicName>
has been listed from several countries including
<collectingCountry id="F3347630FF9FF814FBB5F91ADB22918B" box="[1073,1146,1720,1746]" name="Ghana" pageId="26" pageNumber="177">Ghana</collectingCountry>
(
<bibRefCitation id="EFB24B51FF9FF814FB0DF91ADBA3918B" author="Joly" box="[1161,1275,1720,1746]" firstAuthor="Joly" pageId="26" pageNumber="177" pagination="1 - 10" refId="ref17099" refString="Joly, C. (2001) Contribution to the knowledge of Cetoniinae of Ghana (Coleoptera, Scarabaeidae, Cetoniinae). Lambillonea, CI, 1 - 10." type="book chapter" year="2001">Joly 2001</bibRefCitation>
,
<bibRefCitation id="EFB24B51FF9FF814FA82F91ADA19918B" author="Joly" box="[1286,1345,1720,1746]" firstAuthor="Joly" pageId="26" pageNumber="177" pagination="28 - 89" refId="ref17157" refString="Joly, C. (2018) Deuxieme Contribution a la connaissance des Cetoniidae du Ghana, Sous - familles des Cetoniinae et des Trichinae (Coleoptera, Scarabaeidae). Lambillionea, 118 (1), 28 - 89." type="journal article" year="2018">2018</bibRefCitation>
;
<bibRefCitation id="EFB24B51FF9FF814FAC9F91ADE4E91AF" author="Mudge" etAl="et al." firstAuthor="Mudge" pageId="26" pageNumber="177" pagination="113 - 174" refId="ref17706" refString="Mudge, A. D., Orozco, J., Philips, T. K. &amp; Antoine, P. (2012) The cetoniine fauna of the Upper Guinean forests and savannas of Ghana (Coleoptera: Scarabaeidae: Cetoniinae). Terrestrial Arthropod Reviews, 5, 113 - 174. https: // doi. org / 10.1163 / 187498312 X 635319" type="journal article" year="2012">
Mudge
<emphasis id="B957EAB2FF9FF814FF13F97FDF9291AF" box="[151,202,1756,1782]" italics="true" pageId="26" pageNumber="177">et al</emphasis>
. 2012
</bibRefCitation>
);
<collectingCountry id="F3347630FF9FF814FEAEF97EDE3591AF" box="[298,365,1756,1782]" name="Benin" pageId="26" pageNumber="177">Benin</collectingCountry>
(
<bibRefCitation id="EFB24B51FF9FF814FEF9F97EDDC791AF" author="Touroult &amp; Le Gall" box="[381,671,1756,1782]" firstAuthor="Touroult" pageId="26" pageNumber="177" pagination="29 - 39" refId="ref18903" refString="Touroult, J. &amp; Le Gall, P. (2001) Les Cetoines du Sud-Benin. Etude comparative du peuplement de differents biotopes. Cetoniimania, 1, 29 - 39." type="journal article" year="2001">Touroult &amp; Le Gall 2001</bibRefCitation>
,
<bibRefCitation id="EFB24B51FF9FF814FD2FF97EDDBF91AF" author="Touroult &amp; Le Gall" box="[683,743,1756,1782]" firstAuthor="Touroult" pageId="26" pageNumber="177" pagination="23 - 34" refId="ref18937" refString="Touroult, J. &amp; Le Gall, P. (2012) Fruit feeding Cetoniinae community structure in an anthropogenic landscape in West Africa. Journal of Insect Conservation, 17, 23 - 34. https: // doi. org / 10.1007 / s 10841 - 012 - 9483 - 2" type="journal article" year="2012">2012</bibRefCitation>
;
<bibRefCitation id="EFB24B51FF9FF814FD77F97EDCD691AE" author="Le Gall" box="[755,910,1756,1783]" firstAuthor="Le Gall" pageId="26" pageNumber="177" pagination="25 - 53" refId="ref17355" refString="Le Gall, P. (2015) La faune entomologique du Benin: Les Cetoniinae (Coleoptera, Cetoniidae: Cetoniinae). Cetoniimania NS, 8, 25 - 53." type="journal article" year="2015">Le Gall 2015</bibRefCitation>
),
<collectingCountry id="F3347630FF9FF814FC26F97EDB7F91AF" box="[930,1063,1756,1782]" name="Ivory Coast" pageId="26" pageNumber="177">Ivory Coast</collectingCountry>
(
<bibRefCitation id="EFB24B51FF9FF814FBBDF97EDA4B91AF" author="Fediere" box="[1081,1299,1756,1782]" etAl="et al." firstAuthor="Fediere" pageId="26" pageNumber="177" refId="ref16910" refString="Fediere, G., Lery, X., Herder, S. &amp; Kouassi, N. K. (1989) Cetoines de Cote d'Ivoire (Coleoptera, Cetoniidae, Cetoniinae). Institut International De Resherche Por Le Development a Adiopodoume (IIRSDA), Abidjan, Cote d'Ivoire, number of pages" type="book" year="1989">
Fédière
<emphasis id="B957EAB2FF9FF814FB11F97FDB9091AF" box="[1173,1224,1756,1782]" italics="true" pageId="26" pageNumber="177">et al</emphasis>
. 1989
</bibRefCitation>
),
<collectingCountry id="F3347630FF9FF814FAACF97EDAC591AF" box="[1320,1437,1756,1782]" name="Cameroon" pageId="26" pageNumber="177">Cameroon</collectingCountry>
(
<bibRefCitation id="EFB24B51FF9FF814FF1AF8A2DE6B9043" author="Krajcik" box="[158,307,1792,1818]" firstAuthor="Krajcik" pageId="26" pageNumber="177" refId="ref17199" refString="Krajcik, M. (1998) Cetoniidae of the world catalogue. Part 1. Typos Studio, Most, 96 + xxxvi pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
) and
<collectingCountry id="F3347630FF9FF814FEEBF8A2DDDE9043" box="[367,646,1792,1818]" name="Central African Republic" pageId="26" pageNumber="177">Central African Republic</collectingCountry>
(
<bibRefCitation id="EFB24B51FF9FF814FD17F8A2DC219043" author="Sakai &amp; Nagai" box="[659,889,1792,1818]" firstAuthor="Sakai" pageId="26" pageNumber="177" refId="ref18420" refString="Sakai, K. &amp; Nagai, S. (1998) The cetoniine beetles of the world. Mushi-Sha, Tokyo, 421 pp." type="book" year="1998">Sakai &amp; Nagai 1998</bibRefCitation>
).
<bibRefCitation id="EFB24B51FF9FF814FC0EF8A2DB339043" author="Beinhundner" box="[906,1131,1792,1818]" firstAuthor="Beinhundner" pageId="26" pageNumber="177" refId="ref16250" refString="Beinhundner, G. (2017) The Cetoniinae of Africa. Privately published, Euerbach, 1174 + XXIV pp." type="book" year="2017">Beinhundner (2017)</bibRefCitation>
lists
<taxonomicName id="4C234D23FF9FF814FB20F8A2DA389043" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1883" box="[1188,1376,1792,1818]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B957EAB2FF9FF814FB20F8A2DA389043" box="[1188,1376,1792,1818]" italics="true" pageId="26" pageNumber="177">C. quadrisignata</emphasis>
</taxonomicName>
from
<collectingCountry id="F3347630FF9FF814FF13F886DFA89067" box="[151,240,1828,1854]" name="Senegal" pageId="26" pageNumber="177">Senegal</collectingCountry>
(the
<typeStatus id="54988802FF9FF814FEA9F886DE059067" box="[301,349,1828,1854]" pageId="26" pageNumber="177">type</typeStatus>
locality) and
<collectingCountry id="F3347630FF9FF814FE7AF886DD0F9067" box="[510,599,1828,1854]" name="Gambia" pageId="26" pageNumber="177">Gambia</collectingCountry>
only, and
<taxonomicName id="4C234D23FF9FF814FD55F886DC129067" authorityName="Janson" authorityYear="1888" box="[721,842,1828,1854]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="curvata">
<emphasis id="B957EAB2FF9FF814FD55F886DC129067" box="[721,842,1828,1854]" italics="true" pageId="26" pageNumber="177">C. curvata</emphasis>
</taxonomicName>
as the widespread and highly variable species from
<collectingCountry id="F3347630FF9FF814FF13F8EADE47903B" box="[151,287,1864,1890]" name="Ivory Coast" pageId="26" pageNumber="177">Ivory Coast</collectingCountry>
,
<collectingCountry id="F3347630FF9FF814FEAFF8EADE33903B" box="[299,363,1864,1890]" name="Niger" pageId="26" pageNumber="177">Niger</collectingCountry>
(
<typeStatus id="54988802FF9FF814FEFFF8EADEF5903B" box="[379,429,1864,1890]" pageId="26" pageNumber="177">type</typeStatus>
locality
<collectingCountry id="F3347630FF9FF814FD95F8EADD09903B" box="[529,593,1864,1890]" name="Niger" pageId="26" pageNumber="177">Niger</collectingCountry>
District),
<collectingCountry id="F3347630FF9FF814FD42F8EADC4A903B" box="[710,786,1864,1890]" name="Ghana" pageId="26" pageNumber="177">Ghana</collectingCountry>
,
<collectingCountry id="F3347630FF9FF814FC9BF8EADB66903B" box="[799,1086,1864,1890]" name="Central African Republic" pageId="26" pageNumber="177">Central African Republic</collectingCountry>
,
<collectingCountry id="F3347630FF9FF814FBCDF8EADBDD903B" box="[1097,1157,1864,1890]" name="Togo" pageId="26" pageNumber="177">Togo</collectingCountry>
,
<collectingCountry id="F3347630FF9FF814FB16F8EADB8D903B" box="[1170,1237,1864,1890]" name="Benin" pageId="26" pageNumber="177">Benin</collectingCountry>
and
<collectingCountry id="F3347630FF9FF814FA8AF8EADA3E903B" box="[1294,1382,1864,1890]" name="Nigeria" pageId="26" pageNumber="177">Nigeria</collectingCountry>
. The two species are most reliably separated by differences in the apices of the male parameres. The
<taxonomicName id="4C234D23FF9FF814FB46F8CEDA2790DF" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1883" box="[1218,1407,1900,1926]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B957EAB2FF9FF814FB46F8CEDA2790DF" box="[1218,1407,1900,1926]" italics="true" pageId="26" pageNumber="177">C. quadrisignata</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFB24B51FF9FF814FF13F832DE3C90F3" author="Mudge" box="[151,356,1936,1962]" etAl="et al." firstAuthor="Mudge" pageId="26" pageNumber="177" pagination="113 - 174" refId="ref17706" refString="Mudge, A. D., Orozco, J., Philips, T. K. &amp; Antoine, P. (2012) The cetoniine fauna of the Upper Guinean forests and savannas of Ghana (Coleoptera: Scarabaeidae: Cetoniinae). Terrestrial Arthropod Reviews, 5, 113 - 174. https: // doi. org / 10.1163 / 187498312 X 635319" type="journal article" year="2012">
Mudge
<emphasis id="B957EAB2FF9FF814FF6AF833DE4790F3" box="[238,287,1936,1962]" italics="true" pageId="26" pageNumber="177">et al</emphasis>
. 2012
</bibRefCitation>
(and likely by other authors as well) may in fact be
<taxonomicName id="4C234D23FF9FF814FC29F832DB7C90F3" authorityName="Janson" authorityYear="1888" box="[941,1060,1936,1962]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="curvata">
<emphasis id="B957EAB2FF9FF814FC29F832DB7C90F3" box="[941,1060,1936,1962]" italics="true" pageId="26" pageNumber="177">C. curvata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36A0FF9FF815FF43F816DA6A97A0" blockId="26.[151,1437,1576,2034]" lastBlockId="27.[151,1437,151,249]" lastPageId="27" lastPageNumber="178" pageId="26" pageNumber="177">
<bibRefCitation id="EFB24B51FF9FF814FF43F816DE189097" author="Joly" box="[199,320,1972,1998]" firstAuthor="Joly" pageId="26" pageNumber="177" pagination="1 - 10" refId="ref17099" refString="Joly, C. (2001) Contribution to the knowledge of Cetoniinae of Ghana (Coleoptera, Scarabaeidae, Cetoniinae). Lambillonea, CI, 1 - 10." type="book chapter" year="2001">Joly (2001</bibRefCitation>
,
<bibRefCitation id="EFB24B51FF9FF814FECEF816DEDE9097" author="Joly" box="[330,390,1972,1998]" firstAuthor="Joly" pageId="26" pageNumber="177" pagination="28 - 89" refId="ref17157" refString="Joly, C. (2018) Deuxieme Contribution a la connaissance des Cetoniidae du Ghana, Sous - familles des Cetoniinae et des Trichinae (Coleoptera, Scarabaeidae). Lambillionea, 118 (1), 28 - 89." type="journal article" year="2018">2018</bibRefCitation>
),
<bibRefCitation id="EFB24B51FF9FF814FE1CF816DDD39097" author="Sakai &amp; Nagai" box="[408,651,1972,1998]" firstAuthor="Sakai" pageId="26" pageNumber="177" refId="ref18420" refString="Sakai, K. &amp; Nagai, S. (1998) The cetoniine beetles of the world. Mushi-Sha, Tokyo, 421 pp." type="book" year="1998">Sakai &amp; Nagai (1998)</bibRefCitation>
and
<bibRefCitation id="EFB24B51FF9FF814FD3AF816DCBE9097" author="Touroult &amp; Le Gall" box="[702,998,1972,1998]" firstAuthor="Touroult" pageId="26" pageNumber="177" pagination="23 - 34" refId="ref18937" refString="Touroult, J. &amp; Le Gall, P. (2012) Fruit feeding Cetoniinae community structure in an anthropogenic landscape in West Africa. Journal of Insect Conservation, 17, 23 - 34. https: // doi. org / 10.1007 / s 10841 - 012 - 9483 - 2" type="journal article" year="2012">Touroult &amp; Le Gall (2012)</bibRefCitation>
, list
<taxonomicName id="4C234D23FF9FF814FB9DF816DA389097" authority="Burmeister, 1842" authorityName="Burmeister" authorityYear="1842" box="[1049,1376,1972,1998]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="leprieuri">
<emphasis id="B957EAB2FF9FF814FB9DF816DBC29097" box="[1049,1178,1972,1998]" italics="true" pageId="26" pageNumber="177">C. leprieuri</emphasis>
Burmeister, 1842
</taxonomicName>
from
<collectingCountry id="F3347630FF9FF814FF13F87ADFBB90AB" box="[151,227,2008,2034]" name="Ghana" pageId="26" pageNumber="177">Ghana</collectingCountry>
,
<collectingCountry id="F3347630FF9FF814FF6BF87ADE7390AB" box="[239,299,2008,2034]" name="Togo" pageId="26" pageNumber="177">Togo</collectingCountry>
, and
<collectingCountry id="F3347630FF9FF814FEE3F87ADEF290AB" box="[359,426,2008,2034]" name="Benin" pageId="26" pageNumber="177">Benin</collectingCountry>
respectively and we list it again here.
<taxonomicName id="4C234D23FF9FF814FCDBF87ADB3090AB" authorityName="Burmeister" authorityYear="1842" box="[863,1128,2008,2034]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="leprieuri">
<emphasis id="B957EAB2FF9FF814FCDBF87ADB3090AB" box="[863,1128,2008,2034]" italics="true" pageId="26" pageNumber="177">Charadronota leprieuri</emphasis>
</taxonomicName>
, however, is considered by others to be a synonym of
<taxonomicName id="4C234D23FF9EF815FE31FF35DD2997E8" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1883" box="[437,625,151,177]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B957EAB2FF9EF815FE31FF35DD2997E8" box="[437,625,151,177]" italics="true" pageId="27" pageNumber="178">C. quadrisignata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB24B51FF9EF815FDFAFF35DC2897E8" author="Schoolmeesters" box="[638,880,151,177]" firstAuthor="Schoolmeesters" pageId="27" pageNumber="178" refId="ref18501" refString="Schoolmeesters, P. (2021) SCARABS: World Scarabaeidae Database. In: Banki, O., Roskov, Y., Vandepitte, L., DeWalt, R. E., Remsen, D., Schalk, P., Orrell, T., Keping, M., Miller, J., Aalbu, R., Adlard, R., Adriaenssens, E., Aedo, C., Aescht, E., Akkari, N., Alonso-Zarazaga, M. A., Alvarez, B., Alvarez, F., Anderson, G. et al, Catalogue of Life Checklis. Version 2021 - 09 - 01. Catalogue of Life. Leiden. https: // doi. org / 10.48580 / d 4 sl- 38 g" type="book" year="2021">Schoolmeesters 2021</bibRefCitation>
,
<bibRefCitation id="EFB24B51FF9EF815FCFDFF35DB5597E8" author="Krajcik" box="[889,1037,151,177]" firstAuthor="Krajcik" pageId="27" pageNumber="178" refId="ref17199" refString="Krajcik, M. (1998) Cetoniidae of the world catalogue. Part 1. Typos Studio, Most, 96 + xxxvi pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
,
<bibRefCitation id="EFB24B51FF9EF815FB92FF35DBB197E8" author="Beinhundner" box="[1046,1257,151,177]" firstAuthor="Beinhundner" pageId="27" pageNumber="178" refId="ref16250" refString="Beinhundner, G. (2017) The Cetoniinae of Africa. Privately published, Euerbach, 1174 + XXIV pp." type="book" year="2017">Beinhundner 2017</bibRefCitation>
). Until a generic revision is done, we believe the true number of species of
<taxonomicName id="4C234D23FF9EF815FC91FF19DCEC978C" authorityName="Burmeister" authorityYear="1842" box="[789,948,187,213]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAB2FF9EF815FC91FF19DCEC978C" box="[789,948,187,213]" italics="true" pageId="27" pageNumber="178">Charadronota</emphasis>
</taxonomicName>
and their distribution will remain uncertain. At this moment, we list six species from
<collectingCountry id="F3347630FF9EF815FDE5FF7DDDF597A0" box="[609,685,223,249]" name="Ghana" pageId="27" pageNumber="178">Ghana</collectingCountry>
, including
<taxonomicName id="4C234D23FF9EF815FCADFF7DDCF897A1" authorityName="Janson" authorityYear="1888" box="[809,928,223,249]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="curvata">
<emphasis id="B957EAB2FF9EF815FCADFF7DDCF897A1" box="[809,928,223,249]" italics="true" pageId="27" pageNumber="178">C. curvata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C234D23FF9EF815FC29FF7DDB3397A0" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1883" box="[941,1131,223,249]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="quadrisignata">
<emphasis id="B957EAB2FF9EF815FC29FF7DDC9897A0" box="[941,960,223,249]" italics="true" pageId="27" pageNumber="178">C</emphasis>
.
<emphasis id="B957EAB2FF9EF815FC4AFF7DDB3397A0" box="[974,1131,223,249]" italics="true" pageId="27" pageNumber="178">quadrisignata</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C234D23FF9EF815FB2DFF7DDA7397A0" authorityName="Burmeister" authorityYear="1842" box="[1193,1323,223,249]" class="Insecta" family="Cetoniidae" genus="Charadronota" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="leprieuri">
<emphasis id="B957EAB2FF9EF815FB2DFF7DDA7397A0" box="[1193,1323,223,249]" italics="true" pageId="27" pageNumber="178">C. leprieuri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>