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<document id="989104578B02523283CB49598AF3FD2E" ID-DOI="10.11646/zootaxa.5027.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5448088" ID-ZooBank="D8FE51C2-C34F-47CE-87FC-80BAB16C54BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630464300981" checkinUser="plazi" docAuthor="Nazaré-Silva, Everton E. &amp; Silva, Fernando A. B." docDate="2021" docId="038A87A7FFF3FFA071F5FD2DB93E1F16" docLanguage="en" docName="zootaxa.5027.1.3.pdf" docOrigin="Zootaxa 5027 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocanthon vazdemelloi Nazare-Silva &amp; Silva 2021, new species" docType="treatment" docVersion="3" lastPageNumber="76" masterDocId="FFB3FFDFFFFDFFAF7162FFFBBD5A1E57" masterDocTitle="A taxonomic revision of the South American species of Pseudocanthon Bates, 1887 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="86" masterPageNumber="61" pageNumber="75" updateTime="1699280357525" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8E2BE176FEFF4642496DA85CD5779312">A taxonomic revision of the South American species of Pseudocanthon Bates, 1887 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:namePart id="6981B3496082D6E7F18568B7F84789CE">Nazaré-Silva, Everton E.</mods:namePart>
<mods:affiliation id="1D17764446CE2D3A398037128EFE3D6F">Universidade Federal do Pará, Instituto de Ciências Biológicas. Programa de Pós-graduação em Zoologia da Universidade Federal do Pará e Museu Paraense Emílio Goeldi. Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<mods:namePart id="1A70D052EA15D75ABC3A8F0D7BA86742">Silva, Fernando A. B.</mods:namePart>
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<subSubSection id="C339653AFFF3FFA171F5FD2DBECE1CA7" box="[151,916,726,753]" pageId="14" pageNumber="75" type="nomenclature">
<paragraph id="8B9C36B1FFF3FFA171F5FD2DBECE1CA7" blockId="14.[151,961,726,825]" box="[151,916,726,753]" pageId="14" pageNumber="75">
<heading id="D0D481DDFFF3FFA171F5FD2DBECE1CA7" bold="true" box="[151,916,726,753]" fontSize="11" level="1" pageId="14" pageNumber="75" reason="1">
<emphasis id="B957EAA3FFF3FFA171F5FD2DBECE1CA7" bold="true" box="[151,916,726,753]" pageId="14" pageNumber="75">
<taxonomicName id="4C234D32FFF3FFA171F5FD2DBFAD1CA7" ID-CoL="8NCCK" authority="Nazare-Silva &amp; Silva" authorityName="Nazare-Silva &amp; Silva" authorityYear="2021" box="[151,759,726,753]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="vazdemelloi" status="sp. nov.">
<emphasis id="B957EAA3FFF3FFA171F5FD2DBCB21CA6" bold="true" box="[151,488,726,753]" italics="true" pageId="14" pageNumber="75">Pseudocanthon vazdemelloi</emphasis>
Nazaré-Silva &amp; Silva
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,
<taxonomicNameLabel id="A26457D8FFF3FFA17266FD2CBECE1CA7" box="[772,916,726,753]" pageId="14" pageNumber="75" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C339653AFFF3FFA171F5FD00BE9B1D6E" pageId="14" pageNumber="75" type="description">
<paragraph id="8B9C36B1FFF3FFA171F5FD00BF7D1D42" blockId="14.[151,961,726,825]" box="[151,551,762,789]" pageId="14" pageNumber="75">
(
<figureCitation id="13182A34FFF3FFA171FDFD00BC681D42" box="[159,306,763,789]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2AD); 0.2 mm (2EH)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="14" pageNumber="75">Figs. 2A, 2C</figureCitation>
,
<figureCitation id="13182A34FFF3FFA1705CFD00BC381D42" box="[318,354,763,789]" captionStart="FIGURE 3" captionStartId="12.[151,250,1927,1951]" captionTargetBox="[292,1300,191,1893]" captionTargetId="figure-30@12.[280,1307,181,1903]" captionTargetPageId="12" captionText="FIGURE 3. South American species of Pseudocanthon Bates, 1887, dorsal view. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 1 mm (3AE); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448094" httpUri="https://zenodo.org/record/5448094/files/figure.png" pageId="14" pageNumber="75">3D</figureCitation>
,
<figureCitation id="13182A34FFF3FFA1700CFD00BCCB1D42" box="[366,401,763,789]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3AE); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="14" pageNumber="75">4C</figureCitation>
,
<figureCitation id="13182A34FFF3FFA170FFFD01BC9B1D42" box="[413,449,762,789]" captionStart="FIGURE 5" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[175,1410,894,1887]" captionTargetId="figure-346@16.[164,1422,885,1896]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagus in South American species of Pseudocanthon Bates, 1887, lateral and ventral views, respectively. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847) (arrows indicate variation at apex of parameres); D. P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5448098" httpUri="https://zenodo.org/record/5448098/files/figure.png" pageId="14" pageNumber="75">5D</figureCitation>
,
<figureCitation id="13182A34FFF3FFA170AFFD00BCAB1D42" box="[461,497,763,789]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">6D</figureCitation>
,
<figureCitation id="13182A34FFF3FFA1709FFD00BF441D42" box="[509,542,763,789]" captionStart="FIGURE 7" captionStartId="18.[151,250,1872,1896]" captionTargetBox="[156,1423,190,1839]" captionTargetId="figure-30@18.[155,1432,181,1847]" captionTargetPageId="18" captionText="FIGURE 7. Known distribution in South America of species of Pseudocanthon Bates, 1887. A, P. perplexus (LeConte, 1847) and P. xanthurus (Blanchard, 1847); B, P. vazdemelloi new species, P. pantanensis new species, and P. chaquensis new species." figureDoi="http://doi.org/10.5281/zenodo.5448102" httpUri="https://zenodo.org/record/5448102/files/figure.png" pageId="14" pageNumber="75">7B</figureCitation>
)
</paragraph>
<paragraph id="8B9C36B1FFF3FFA171F5FCE5BE9B1D6E" blockId="14.[151,961,726,825]" box="[151,961,798,825]" pageId="14" pageNumber="75">
<uri id="FFB23AB3FFF3FFA171F5FCE5BE9B1D6E" box="[151,961,798,825]" pageId="14" pageNumber="75">
urn:lsid:zoobank.org:act:
<uuid id="FF850C64FFF3FFA170C8FCE5BE9B1D6E" box="[426,961,798,825]" pageId="14" pageNumber="75">FF662C35-03A0-4A96-A93C-8567FC58F15C</uuid>
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</subSubSection>
<subSubSection id="C339653AFFF3FFA171F5FC9CBC9B1D9E" pageId="14" pageNumber="75" type="etymology">
<paragraph id="8B9C36B1FFF3FFA171F5FC9CBC9B1D9E" blockId="14.[151,1437,871,2013]" pageId="14" pageNumber="75">
<emphasis id="B957EAA3FFF3FFA171F5FC9CBC421DD6" bold="true" box="[151,280,871,897]" pageId="14" pageNumber="75">Etymology</emphasis>
. The species is named after the Brazilian scarabaeidologist Fernando Zagury Vaz-de-Mello as recognition for so many years of fruitful collaboration with the second author, and for his valuable contributions towards the taxonomy of dung beetles.
</paragraph>
</subSubSection>
<subSubSection id="C339653AFFF3FFA171A5FC29BC751B96" pageId="14" pageNumber="75" type="diagnosis">
<paragraph id="8B9C36B1FFF3FFA171A5FC29BC751B96" blockId="14.[151,1437,871,2013]" pageId="14" pageNumber="75">
<emphasis id="B957EAA3FFF3FFA171A5FC29BC601DBB" bold="true" box="[199,314,978,1004]" pageId="14" pageNumber="75">Diagnosis</emphasis>
.
<taxonomicName id="4C234D32FFF3FFA1702BFC28BFD91DBA" authorityName="Nazare-Silva &amp; Silva" authorityYear="2021" box="[329,643,979,1005]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B957EAA3FFF3FFA1702BFC28BFD91DBA" box="[329,643,979,1005]" italics="true" pageId="14" pageNumber="75">Pseudocanthon vazdemelloi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C234D32FFF3FFA17274FC28BEE11DBB" authorityName="Nazaré-Silva &amp; Silva" authorityYear="2021" box="[790,955,979,1005]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="pantanensis">
<emphasis id="B957EAA3FFF3FFA17274FC28BE7D1DBB" box="[790,807,979,1004]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA17257FC28BEE11DBB" box="[821,955,979,1004]" italics="true" pageId="14" pageNumber="75">pantanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D32FFF3FFA17290FC28B9D71DBA" authorityName="Nazaré-Silva &amp; Silva" authorityYear="2021" box="[1010,1165,979,1005]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="chaquensis">
<emphasis id="B957EAA3FFF3FFA17290FC28B9591DBB" box="[1010,1027,979,1004]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA17573FC28B9D71DBA" box="[1041,1165,979,1005]" italics="true" pageId="14" pageNumber="75">chaquensis</emphasis>
</taxonomicName>
in having the pronotum with dense, well-impressed granulate microsculpture centrally (
<figureCitation id="13182A34FFF3FFA1720CFC0CBE901A46" box="[878,970,1015,1041]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2AD); 0.2 mm (2EH)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="14" pageNumber="75">Fig. 2A</figureCitation>
). However,
<taxonomicName id="4C234D32FFF3FFA17534FC0CB9A41A46" authorityName="Nazare-Silva &amp; Silva" authorityYear="2021" box="[1110,1278,1015,1041]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B957EAA3FFF3FFA17534FC0CB93D1A47" box="[1110,1127,1015,1040]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA1751AFC0CB9A41A46" box="[1144,1278,1015,1041]" italics="true" pageId="14" pageNumber="75">vazdemelloi</emphasis>
</taxonomicName>
can be easily distinguished by the eighth elytral stria absent or reduced to basal one-fourth of the elytra (
<figureCitation id="13182A34FFF3FFA175F2FBE0B9B21A62" box="[1168,1256,1051,1077]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2AD); 0.2 mm (2EH)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="14" pageNumber="75">Fig. 2C</figureCitation>
); posterior edge of metafemur regularly curved, not projected towards the abdominal ventrites (
<figureCitation id="13182A34FFF3FFA1756CFBC4B93D1A0E" box="[1038,1127,1087,1113]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3AE); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="14" pageNumber="75">Fig. 4C</figureCitation>
); apex of parameres curved inward, forming an acute spiniform projection at ventral edge (
<figureCitation id="13182A34FFF3FFA17230FB98BEF11A2A" box="[850,939,1122,1149]" captionStart="FIGURE 5" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[175,1410,894,1887]" captionTargetId="figure-346@16.[164,1422,885,1896]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagus in South American species of Pseudocanthon Bates, 1887, lateral and ventral views, respectively. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847) (arrows indicate variation at apex of parameres); D. P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5448098" httpUri="https://zenodo.org/record/5448098/files/figure.png" pageId="14" pageNumber="75">Fig. 5D</figureCitation>
); frontolateral peripheral (FLP) endophallite elongate, bifurcate at inferior portion (
<figureCitation id="13182A34FFF3FFA17322FB7CBFC31AF5" box="[576,665,1159,1186]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">Fig. 6D</figureCitation>
); and accessory endophallite (AE) small and elongate (
<figureCitation id="13182A34FFF3FFA17599FB7CB80E1AF5" box="[1275,1364,1159,1186]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">Fig. 6D</figureCitation>
). In
<taxonomicName id="4C234D32FFF3FFA174E6FB7CBC471A93" authorityName="Nazaré-Silva &amp; Silva" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="pantanensis">
<emphasis id="B957EAA3FFF3FFA174E6FB7CB8CF1AF7" box="[1412,1429,1159,1184]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA171F5FB50BC471A93" box="[151,285,1195,1220]" italics="true" pageId="14" pageNumber="75">pantanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D32FFF3FFA17036FB50BCAA1A92" authorityName="Nazaré-Silva &amp; Silva" authorityYear="2021" box="[340,496,1195,1221]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="chaquensis">
<emphasis id="B957EAA3FFF3FFA17036FB50BC3F1A93" box="[340,357,1195,1220]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA17016FB50BCAA1A92" box="[372,496,1195,1221]" italics="true" pageId="14" pageNumber="75">chaquensis</emphasis>
</taxonomicName>
, in turn, the eighth stria extends to across the basal half of the elytra (
<figureCitation id="13182A34FFF3FFA17466FB50B8051A92" box="[1284,1375,1195,1221]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2AD); 0.2 mm (2EH)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="14" pageNumber="75">Fig. 2D</figureCitation>
); and the posterior edge of metafemur is lightly angulate at basal third and expanded towards the abdominal ventrites (
<figureCitation id="13182A34FFF3FFA1740DFB34BD871B5A" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3AE); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="14" pageNumber="75">Fig. 4DE</figureCitation>
). In
<taxonomicName id="4C234D32FFF3FFA17073FB08BCED1B5B" authorityName="Nazaré-Silva &amp; Silva" authorityYear="2021" box="[273,439,1267,1293]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="pantanensis">
<emphasis id="B957EAA3FFF3FFA17073FB08BC781B5B" box="[273,290,1267,1292]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA17053FB08BCED1B5B" box="[305,439,1267,1292]" italics="true" pageId="14" pageNumber="75">pantanensis</emphasis>
</taxonomicName>
, in particular, the parameres are curved and pointed apically (
<figureCitation id="13182A34FFF3FFA17518FB08B98B1B5A" box="[1146,1233,1266,1293]" captionStart="FIGURE 5" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[175,1410,894,1887]" captionTargetId="figure-346@16.[164,1422,885,1896]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagus in South American species of Pseudocanthon Bates, 1887, lateral and ventral views, respectively. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847) (arrows indicate variation at apex of parameres); D. P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5448098" httpUri="https://zenodo.org/record/5448098/files/figure.png" pageId="14" pageNumber="75">Fig. 5E</figureCitation>
), the frontolateral peripheral endophallite is sinuous, with inferior portion rounded (
<figureCitation id="13182A34FFF3FFA1721AFAECBE971B65" box="[888,973,1303,1330]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">Fig. 6E</figureCitation>
) and the AE absent (
<figureCitation id="13182A34FFF3FFA175DAFAECB8551B65" box="[1208,1295,1303,1330]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">Fig. 6E</figureCitation>
), whereas in
<taxonomicName id="4C234D32FFF3FFA171F5FAC0BC681B02" authorityName="Nazaré-Silva &amp; Silva" authorityYear="2021" box="[151,306,1339,1365]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="chaquensis">
<emphasis id="B957EAA3FFF3FFA171F5FAC0BDF21B03" box="[151,168,1339,1364]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA171D4FAC0BC681B02" box="[182,306,1339,1365]" italics="true" pageId="14" pageNumber="75">chaquensis</emphasis>
</taxonomicName>
the parameres are truncate apically (
<figureCitation id="13182A34FFF3FFA173AFFAC0BE781B02" box="[717,802,1338,1365]" captionStart="FIGURE 5" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[175,1410,894,1887]" captionTargetId="figure-346@16.[164,1422,885,1896]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagus in South American species of Pseudocanthon Bates, 1887, lateral and ventral views, respectively. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847) (arrows indicate variation at apex of parameres); D. P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5448098" httpUri="https://zenodo.org/record/5448098/files/figure.png" pageId="14" pageNumber="75">Fig. 5F</figureCitation>
). We did not find any significant differences in the shape of frontolateral peripheral endophallite and accessory endophallite between
<taxonomicName id="4C234D32FFF3FFA17286FAA4B9D31B2E" authorityName="Nazare-Silva &amp; Silva" authorityYear="2021" box="[996,1161,1375,1401]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B957EAA3FFF3FFA17286FAA4BEAF1B2F" box="[996,1013,1375,1400]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA17561FAA4B9D31B2E" box="[1027,1161,1375,1401]" italics="true" pageId="14" pageNumber="75">vazdemelloi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D32FFF3FFA175DCFAA4B8021B2E" authorityName="Nazaré-Silva &amp; Silva" authorityYear="2021" box="[1214,1368,1375,1401]" class="Insecta" family="Scarabaeidae" genus="Pseudocanthon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="chaquensis">
<emphasis id="B957EAA3FFF3FFA175DCFAA4B9951B2F" box="[1214,1231,1375,1400]" italics="true" pageId="14" pageNumber="75">P</emphasis>
.
<emphasis id="B957EAA3FFF3FFA175BEFAA4B8021B2E" box="[1244,1368,1375,1401]" italics="true" pageId="14" pageNumber="75">chaquensis</emphasis>
</taxonomicName>
. Both species have frontolateral peripheral endophallite elongate, bifurcate at inferior portion, and AE small, elongate (
<figureCitation id="13182A34FFF3FFA171FDFA5CBC781B96" box="[159,290,1447,1473]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">Figs. 6D, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C339653AFFF3FFA171A5FA31BEBC1919" pageId="14" pageNumber="75" type="description">
<paragraph id="8B9C36B1FFF3FFA171A5FA31BEBC1919" blockId="14.[151,1437,871,2013]" pageId="14" pageNumber="75">
<emphasis id="B957EAA3FFF3FFA171A5FA31BC0B1BB3" bold="true" box="[199,337,1482,1508]" pageId="14" pageNumber="75">Description</emphasis>
.
<emphasis id="B957EAA3FFF3FFA17002FA30BF6B1BB2" bold="true" box="[352,561,1483,1509]" pageId="14" pageNumber="75">
<typeStatus id="54988813FFF3FFA17002FA30BC951BB2" box="[352,463,1483,1509]" pageId="14" pageNumber="75" type="holotype">Holotype</typeStatus>
. Length
</emphasis>
. 5.0 mm.
<emphasis id="B957EAA3FFF3FFA173C6FA31BFAA1BB3" bold="true" box="[676,752,1482,1508]" pageId="14" pageNumber="75">Width</emphasis>
.
<quantity id="4CDB9B54FFF3FFA1739CFA30BE021BB2" box="[766,856,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="14" pageNumber="75" unit="mm" value="3.3">3.3 mm</quantity>
.
<emphasis id="B957EAA3FFF3FFA17201FA31BEED1BB3" bold="true" box="[867,951,1482,1508]" pageId="14" pageNumber="75">Colour</emphasis>
. Surface of body dark brown with metallic green reflections; legs orange; orange spot on clypeus and on last abdominal ventrite.
<emphasis id="B957EAA3FFF3FFA1751DFA15BC7A187B" bold="true" pageId="14" pageNumber="75">Surface punctation and sculpturing</emphasis>
. In general, surface punctation consists of foveolate punctures. Surface of head, mesoventrite, metaventrite, elytra, abdomen, tibiae, and femora with dense and well-impressed granulate microsculpture; lateral edges and anterior angle of pronotum, hypomera, lateral edges, and anterior portion of metaventrite, mesepimeron, metanepisternum, epipleuron, tibiae, and femora with imbricate microsculpture. Pronotum with dense and wellimpressed granulate microsculpture centrally (
<figureCitation id="13182A34FFF3FFA173D2F958BE5418EA" box="[688,782,1699,1725]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2AD); 0.2 mm (2EH)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="14" pageNumber="75">Fig. 2A</figureCitation>
).
<emphasis id="B957EAA3FFF3FFA17244F958BE2818EA" bold="true" box="[806,882,1699,1725]" pageId="14" pageNumber="75">Elytra</emphasis>
. Eighth elytral stria absent or reduced to basal one-fourth of the elytra (
<figureCitation id="13182A34FFF3FFA170D3F93CBF5118B6" box="[433,523,1735,1761]" captionStart="FIGURE 2" captionStartId="11.[151,250,1807,1831]" captionTargetBox="[265,1315,191,1773]" captionTargetId="figure-21@11.[261,1326,181,1783]" captionTargetPageId="11" captionText="FIGURE 2. Details of the external morphology in species of Pseudocanthon Bates, 1887. A, P. vazdemelloi new species pronotal disc; B, P. xanthurus (Blanchard, 1847) pronotal disc; C, P. vazdemelloi new species detail of left lateral of elytron; D, P. pantanensis new species detail of left lateral of elytron (arrow indicates eighth elytral stria present); E, P. perplexus (LeConte, 1847) protibia in dorsal view (arrow indicates median emargination); F, P. xanthurus (Blanchard, 1847) protibia in dorsal view; G. P. pantanensis new species pygidium; H, P. chaquensis new species pygidium. Scale bar: 0.5 mm (2AD); 0.2 mm (2EH)." figureDoi="http://doi.org/10.5281/zenodo.5448092" httpUri="https://zenodo.org/record/5448092/files/figure.png" pageId="14" pageNumber="75">Fig. 2C</figureCitation>
).
<emphasis id="B957EAA3FFF3FFA17342F93CBF0218B6" bold="true" box="[544,600,1735,1761]" pageId="14" pageNumber="75">Legs</emphasis>
. Posterior edge of metafemur regularly curved, not projected towards the abdominal ventrites (
<figureCitation id="13182A34FFF3FFA170E4F910BCBA1952" box="[390,480,1771,1797]" captionStart="FIGURE 4" captionStartId="13.[151,250,1868,1892]" captionTargetBox="[171,1413,608,1832]" captionTargetId="figure-218@13.[162,1424,597,1845]" captionTargetPageId="13" captionText="FIGURE 4. South American species of Pseudocanthon Bates, 1887, ventral view. A, Pseudocanthon perplexus (LeConte, 1847); B, P. xanthurus (Blanchard, 1847); C, P. vazdemelloi new species; D, P. pantanensis new species (arrows indicate angulation at posterior edge of metafemur); E, P. chaquensis new species (arrows indicate angulation at posterior edge of metafemur). Scale bar: 1 mm (3AE); 2 mm (3F)." figureDoi="http://doi.org/10.5281/zenodo.5448096" httpUri="https://zenodo.org/record/5448096/files/figure.png" pageId="14" pageNumber="75">Fig. 4C</figureCitation>
).
<emphasis id="B957EAA3FFF3FFA17096F910BF081952" bold="true" box="[500,594,1771,1797]" pageId="14" pageNumber="75">Tegmen</emphasis>
. Apex of parameres curved inward, forming an acute spiniform projection at ventral edge (
<figureCitation id="13182A34FFF3FFA1702CF8F4BCF3197E" box="[334,425,1806,1833]" captionStart="FIGURE 5" captionStartId="16.[151,250,1919,1943]" captionTargetBox="[175,1410,894,1887]" captionTargetId="figure-346@16.[164,1422,885,1896]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagus in South American species of Pseudocanthon Bates, 1887, lateral and ventral views, respectively. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847) (arrows indicate variation at apex of parameres); D. P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5448098" httpUri="https://zenodo.org/record/5448098/files/figure.png" pageId="14" pageNumber="75">Fig. 5D</figureCitation>
).
<emphasis id="B957EAA3FFF3FFA170DCF8F4BF0F197E" bold="true" box="[446,597,1807,1833]" pageId="14" pageNumber="75">Endophallus</emphasis>
. Frontolateral peripheral (FLP) endophallite elongate, bifurcate at inferior portion (
<figureCitation id="13182A34FFF3FFA17195F8C8BC081919" box="[247,338,1843,1870]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">Fig. 6D</figureCitation>
); acessory endophallite (AE) small and elongate (
<figureCitation id="13182A34FFF3FFA1721DF8C8BE831919" box="[895,985,1843,1870]" captionStart="FIGURE 6" captionStartId="17.[151,250,1414,1438]" captionTargetBox="[155,1422,192,1385]" captionTargetId="figure-21@17.[152,1436,181,1390]" captionTargetPageId="17" captionText="FIGURE 6. Endophallites in South American species of Pseudocanthon Bates, 1887. A, Pseudocanthon perplexus (LeConte, 1847); BC, P. xanthurus (Blanchard, 1847); D, P. vazdemelloi new species; E, P. pantanensis new species; F, P. chaquensis new species. Accessory endophallite (AE). Complex of axial and subaxial (A+SA) endophallite. Frontolateral peripheral (FLP) endophallite. Superior right peripheral (SRP) endophallite. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5448100" httpUri="https://zenodo.org/record/5448100/files/figure.png" pageId="14" pageNumber="75">Fig. 6D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C339653AFFF3FFA071A5F8ACB8101E82" lastPageId="15" lastPageNumber="76" pageId="14" pageNumber="75" type="materials_examined">
<paragraph id="8B9C36B1FFF3FFA171A5F8ACB8C7198A" blockId="14.[151,1437,871,2013]" pageId="14" pageNumber="75">
<emphasis id="B957EAA3FFF3FFA171A5F8ACBC2E1927" bold="true" box="[199,372,1878,1905]" pageId="14" pageNumber="75">Type material.</emphasis>
<materialsCitation id="3B4B3CECFFF3FFA17018F8ACBCA919C2" collectingDate="1998-12-03" country="Brazil" latitude="-16.512053" location="Pousada Araras" longLatPrecision="1" longitude="-56.71156" municipality="Pocone" pageId="14" pageNumber="75" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Mato Grosso" typeStatus="holotype">
<specimenCount id="9D25FD38FFF3FFA17018F8ACBCC61926" box="[378,412,1879,1905]" pageId="14" pageNumber="75" type="female">6♀</specimenCount>
and
<specimenCount id="9D25FD38FFF3FFA170B3F8ACBCAD1926" box="[465,503,1879,1905]" pageId="14" pageNumber="75" type="male">6♂</specimenCount>
.
<emphasis id="B957EAA3FFF3FFA17363F8ACBFAA1927" bold="true" box="[513,752,1878,1905]" pageId="14" pageNumber="75">
<typeStatus id="54988813FFF3FFA17363F8ACBF2A1926" box="[513,624,1879,1905]" pageId="14" pageNumber="75" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F3347621FFF3FFA17319F8ADBFB11927" box="[635,747,1878,1904]" name="Brazil" pageId="14" pageNumber="75">BRAZIL</collectingCountry>
:
</emphasis>
<collectingRegion id="49E7F853FFF3FFA17394F8ACBEE71926" box="[758,957,1879,1905]" country="Brazil" name="Mato Grosso" pageId="14" pageNumber="75">MATO GROSSO</collectingRegion>
:
<collectingMunicipality id="6BF8ACCBFFF3FFA172A5F8ACB9461926" box="[967,1052,1879,1905]" pageId="14" pageNumber="75">Poconé</collectingMunicipality>
,
<location id="8EFC606AFFF3FFA17544F8ACB98F1926" LSID="urn:lsid:plazi:treatment:038A87A7FFF3FFA071F5FD2DB93E1F16:8EFC606AFFF3FFA17544F8ACB98F1926" box="[1062,1237,1879,1905]" country="Brazil" latitude="-16.512053" longLatPrecision="1" longitude="-56.71156" municipality="Pocone" name="Pousada Araras" pageId="14" pageNumber="75" stateProvince="Mato Grosso">Pousada Araras</location>
, [
<geoCoordinate id="EE175076FFF3FFA17585F8ACB8CD1926" box="[1255,1431,1879,1905]" degrees="16" direction="south" minutes="30" orientation="latitude" pageId="14" pageNumber="75" precision="1" seconds="43.39" value="-16.512053">16°3043.39S</geoCoordinate>
;
<geoCoordinate id="EE175076FFF3FFA171F5F881BC0819C2" box="[151,338,1914,1941]" degrees="56" direction="west" minutes="42" orientation="longitude" pageId="14" pageNumber="75" precision="1" seconds="41.61" value="-56.71156">56°4241.61W</geoCoordinate>
],
<date id="FF9D1071FFF3FFA1700AF880BCA919C2" box="[360,499,1915,1941]" pageId="14" pageNumber="75" value="1998-12-03">
<collectingDate id="EFD9E999FFF3FFA1700AF880BCA919C2" box="[360,499,1915,1941]" pageId="14" pageNumber="75" value="1998-12-03">03.XII.1998</collectingDate>
</date>
</materialsCitation>
, J. Moreno &amp; T. Mestre—
<materialsCitation id="3B4B3CECFFF3FFA17252F880BE9C19EE" collectingDate="1998-12-03" collectionCode="CEMT" country="Brazil" latitude="-16.512053" location="Pousada Araras" longLatPrecision="1" longitude="-56.71156" municipality="Pocone" pageId="14" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCount id="9D25FD38FFF3FFA17252F880BE1119C2" box="[816,843,1915,1941]" pageId="14" pageNumber="75" type="male">1♂</specimenCount>
(
<collectionCode id="ED32AE74FFF3FFA1723FF880BEEB19C2" box="[861,945,1915,1941]" pageId="14" pageNumber="75">CEMT</collectionCode>
).
<emphasis id="B957EAA3FFF3FFA172A4F880B99919C3" bold="true" box="[966,1219,1914,1941]" pageId="14" pageNumber="75">
<typeStatus id="54988813FFF3FFA172A4F880B91A19C2" box="[966,1088,1915,1941]" pageId="14" pageNumber="75" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F3347621FFF3FFA1752DF881B9E519C3" box="[1103,1215,1914,1940]" name="Brazil" pageId="14" pageNumber="75">BRAZIL</collectingCountry>
:
</emphasis>
<collectingRegion id="49E7F853FFF3FFA175AFF880B8C319C2" box="[1229,1433,1915,1941]" country="Brazil" name="Mato Grosso" pageId="14" pageNumber="75">MATO GROSSO</collectingRegion>
:
<collectingMunicipality id="6BF8ACCBFFF3FFA171F5F864BDB619EE" box="[151,236,1951,1977]" pageId="14" pageNumber="75">Poconé</collectingMunicipality>
,
<location id="8EFC606AFFF3FFA17194F864BCFE19EE" LSID="urn:lsid:plazi:treatment:038A87A7FFF3FFA071F5FD2DB93E1F16:8EFC606AFFF3FFA17194F864BCFE19EE" box="[246,420,1951,1977]" country="Brazil" latitude="-16.512053" longLatPrecision="1" longitude="-56.71156" municipality="Pocone" name="Pousada Araras" pageId="14" pageNumber="75" stateProvince="Mato Grosso">Pousada Araras</location>
, [
<geoCoordinate id="EE175076FFF3FFA170D7F864BF3F19EE" box="[437,613,1951,1977]" degrees="16" direction="south" minutes="30" orientation="latitude" pageId="14" pageNumber="75" precision="1" seconds="43.39" value="-16.512053">16°3043.39S</geoCoordinate>
;
<geoCoordinate id="EE175076FFF3FFA1730DF865BE7019EE" box="[623,810,1950,1977]" degrees="56" direction="west" minutes="42" orientation="longitude" pageId="14" pageNumber="75" precision="1" seconds="41.61" value="-56.71156">56°4241.61W</geoCoordinate>
],
<date id="FF9D1071FFF3FFA17259F864BE9C19EE" box="[827,966,1951,1977]" pageId="14" pageNumber="75" value="1998-12-03">
<collectingDate id="EFD9E999FFF3FFA17259F864BE9C19EE" box="[827,966,1951,1977]" pageId="14" pageNumber="75" value="1998-12-03">03.XII.1998</collectingDate>
</date>
</materialsCitation>
, J. Moreno &amp; T. Mestre—
<materialsCitation id="3B4B3CECFFF3FFA1758EF864B8C2198A" collectingDate="1990-12-17" collectionCode="CEMT" country="Argentina" elevation="76" latitude="-25.145308" location="Parque Nacional Pilcomayo" longLatPrecision="1" longitude="-58.186607" pageId="14" pageNumber="75" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D25FD38FFF3FFA1758EF864B85D19EE" box="[1260,1287,1951,1977]" pageId="14" pageNumber="75" type="female">4♀</specimenCount>
<specimenCount id="9D25FD38FFF3FFA1746FF864B86A19EE" box="[1293,1328,1951,1977]" pageId="14" pageNumber="75" type="male">3♂</specimenCount>
(
<collectionCode id="ED32AE74FFF3FFA1745FF864B8CB19EE" box="[1341,1425,1951,1977]" pageId="14" pageNumber="75">CEMT</collectionCode>
).
<emphasis id="B957EAA3FFF3FFA171F5F839BC16198B" bold="true" box="[151,332,1986,2012]" pageId="14" pageNumber="75">
<collectingCountry id="F3347621FFF3FFA171F5F839BC1D198B" box="[151,327,1986,2012]" name="Argentina" pageId="14" pageNumber="75">ARGENTINA</collectingCountry>
:
</emphasis>
FORMOSA:
<location id="8EFC606AFFF3FFA17088F838BE72198A" LSID="urn:lsid:plazi:treatment:038A87A7FFF3FFA071F5FD2DB93E1F16:8EFC606AFFF3FFA17088F838BE72198A" box="[490,808,1987,2013]" country="Argentina" latitude="-25.145308" longLatPrecision="1" longitude="-58.186607" name="Parque Nacional Pilcomayo" pageId="14" pageNumber="75">Parque Nacional Pilcomayo</location>
, [
<geoCoordinate id="EE175076FFF3FFA17259F839BEB0198A" box="[827,1002,1986,2013]" degrees="25" direction="south" minutes="08" orientation="latitude" pageId="14" pageNumber="75" precision="1" seconds="43.11" value="-25.145308">25°0843.11S</geoCoordinate>
;
<geoCoordinate id="EE175076FFF3FFA17295F839B9EA198A" box="[1015,1200,1986,2013]" degrees="58" direction="west" minutes="11" orientation="longitude" pageId="14" pageNumber="75" precision="1" seconds="11.78" value="-58.186607">58°1111.78W</geoCoordinate>
],
<quantity id="4CDB9B54FFF3FFA175A6F838B85A198A" box="[1220,1280,1987,2013]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="75" unit="m" value="76.0">
<elevation id="000ED182FFF3FFA175A6F838B85A198A" box="[1220,1280,1987,2013]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="75" unit="m" value="76.0">76 m</elevation>
</quantity>
,
<date id="FF9D1071FFF3FFA1746EF838B8C2198A" box="[1292,1432,1987,2013]" pageId="14" pageNumber="75" value="1990-12-17">
<collectingDate id="EFD9E999FFF3FFA1746EF838B8C2198A" box="[1292,1432,1987,2013]" pageId="14" pageNumber="75" value="1990-12-17">17.XII.1990</collectingDate>
</date>
</materialsCitation>
,
</paragraph>
<paragraph id="8B9C36B1FFF2FFA071F5FF6CB8101E82" blockId="15.[151,1437,150,321]" pageId="15" pageNumber="76">
<materialsCitation id="3B4B3CECFFF2FFA071F5FF6CBCC31EE6" box="[151,409,151,177]" collectionCode="CMNC" collectorName="J. Peck" pageId="15" pageNumber="76" specimenCount="1" specimenCount-male="1">
<collectorName id="26D65367FFF2FFA071F5FF6CBDB41EE6" box="[151,238,151,177]" pageId="15" pageNumber="76">J. Peck</collectorName>
<specimenCount id="9D25FD38FFF2FFA0706EFF6CBC7D1EE6" box="[268,295,151,177]" pageId="15" pageNumber="76" type="male">1♂</specimenCount>
(
<collectionCode id="ED32AE74FFF2FFA0705AFF6CBCCB1EE6" box="[312,401,151,177]" pageId="15" pageNumber="76">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3CECFFF2FFA070C7FF6CBC701E82" collectingDate="1939-01" collectionCode="CMNC" elevation="32" latitude="-32.958702" location="Rosario" longLatPrecision="1" longitude="-60.693043" pageId="15" pageNumber="76" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Santa Fe">
<collectingRegion id="49E7F853FFF2FFA070C7FF6CBF771EE4" box="[421,557,151,179]" country="Argentina" name="Santa Fe" pageId="15" pageNumber="76">SANTA FÉ</collectingRegion>
:
<location id="8EFC606AFFF2FFA0735BFF6CBFC91EE6" LSID="urn:lsid:plazi:treatment:038A87A7FFF3FFA071F5FD2DB93E1F16:8EFC606AFFF2FFA0735BFF6CBFC91EE6" box="[569,659,151,177]" latitude="-32.958702" longLatPrecision="1" longitude="-60.693043" name="Rosario" pageId="15" pageNumber="76" stateProvince="Santa Fe">Rosário</location>
, [
<geoCoordinate id="EE175076FFF2FFA073C5FF6DBE0D1EE6" box="[679,855,150,177]" degrees="32" direction="south" minutes="57" orientation="latitude" pageId="15" pageNumber="76" precision="1" seconds="31.33" value="-32.958702">32°5731.33S</geoCoordinate>
;
<geoCoordinate id="EE175076FFF2FFA07207FF6DB97B1EE6" box="[869,1057,150,177]" degrees="60" direction="west" minutes="41" orientation="longitude" pageId="15" pageNumber="76" precision="1" seconds="34.95" value="-60.693043">60°4134.95W</geoCoordinate>
],
<quantity id="4CDB9B54FFF2FFA07556FF6CB92B1EE6" box="[1076,1137,151,177]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="76" unit="m" value="32.0">
<elevation id="000ED182FFF2FFA07556FF6CB92B1EE6" box="[1076,1137,151,177]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="15" pageNumber="76" unit="m" value="32.0">32 m</elevation>
</quantity>
,
<date id="FF9D1071FFF2FFA0751DFF6CB9931EE6" box="[1151,1225,151,177]" pageId="15" pageNumber="76" value="1939-01">
<collectingDate id="EFD9E999FFF2FFA0751DFF6CB9931EE6" box="[1151,1225,151,177]" pageId="15" pageNumber="76" value="1939-01">I.1939</collectingDate>
</date>
, no collector—
<specimenCount id="9D25FD38FFF2FFA074E4FF6CB8C61EE6" box="[1414,1436,151,177]" pageId="15" pageNumber="76" type="female">1♀</specimenCount>
<specimenCount id="9D25FD38FFF2FFA071F5FF40BDE01E82" box="[151,186,187,213]" pageId="15" pageNumber="76" type="male">1♂</specimenCount>
(
<collectionCode id="ED32AE74FFF2FFA071ABFF40BC781E82" box="[201,290,187,213]" pageId="15" pageNumber="76">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B4B3CECFFF2FFA07057FF40B81C1E82" box="[309,1350,186,213]" collectingDate="1976-03" collectionCode="CMNC" collectorName="Martinez" elevation="63" latitude="-27.469213" location="CORRIENTES" longLatPrecision="1" longitude="-58.830635" pageId="15" pageNumber="76" specimenCount="1" specimenCount-female="1" stateProvince="Corrientes">
<collectingRegion id="49E7F853FFF2FFA07057FF40BCBC1E82" box="[309,486,187,213]" country="Argentina" name="Corrientes" pageId="15" pageNumber="76">CORRIENTES</collectingRegion>
: [
<geoCoordinate id="EE175076FFF2FFA0709AFF40BFF21E82" box="[504,680,187,213]" degrees="27" direction="south" minutes="28" orientation="latitude" pageId="15" pageNumber="76" precision="1" seconds="09.17" value="-27.469213">27°2809.17S</geoCoordinate>
;
<geoCoordinate id="EE175076FFF2FFA073D6FF41BE351E82" box="[692,879,186,213]" degrees="58" direction="west" minutes="49" orientation="longitude" pageId="15" pageNumber="76" precision="1" seconds="50.29" value="-58.830635">58°4950.29W</geoCoordinate>
],
<quantity id="4CDB9B54FFF2FFA072E0FF40BEE71E82" box="[898,957,187,213]" metricMagnitude="1" metricUnit="m" metricValue="6.3" pageId="15" pageNumber="76" unit="m" value="63.0">
<elevation id="000ED182FFF2FFA072E0FF40BEE71E82" box="[898,957,187,213]" metricMagnitude="1" metricUnit="m" metricValue="6.3" pageId="15" pageNumber="76" unit="m" value="63.0">63 m</elevation>
</quantity>
,
<date id="FF9D1071FFF2FFA072ABFF40B97C1E82" box="[969,1062,187,213]" pageId="15" pageNumber="76" value="1976-03">
<collectingDate id="EFD9E999FFF2FFA072ABFF40B97C1E82" box="[969,1062,187,213]" pageId="15" pageNumber="76" value="1976-03">III.1976</collectingDate>
</date>
,
<collectorName id="26D65367FFF2FFA07550FF40B9FB1E82" box="[1074,1185,187,213]" pageId="15" pageNumber="76">Martínez</collectorName>
<specimenCount id="9D25FD38FFF2FFA075A2FF40B98C1E82" box="[1216,1238,187,213]" pageId="15" pageNumber="76" type="female">1♀</specimenCount>
(
<collectionCode id="ED32AE74FFF2FFA07587FF40B8641E82" box="[1253,1342,187,213]" pageId="15" pageNumber="76">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C339653AFFF2FFA071A5FF25B93E1F16" pageId="15" pageNumber="76" type="distribution">
<paragraph id="8B9C36B1FFF2FFA071A5FF25B93E1F16" blockId="15.[151,1437,150,321]" pageId="15" pageNumber="76">
<emphasis id="B957EAA3FFF2FFA071A5FF25BC031EAF" bold="true" box="[199,345,222,248]" pageId="15" pageNumber="76">Distribution</emphasis>
. Known from
<collectingCountry id="F3347621FFF2FFA0736AFF24BF2D1EAE" box="[520,631,223,249]" name="Argentina" pageId="15" pageNumber="76">Argentina</collectingCountry>
(
<collectingRegion id="49E7F853FFF2FFA073E8FF24BE591EAE" box="[650,771,223,249]" country="Argentina" name="Corrientes" pageId="15" pageNumber="76">Corrientes</collectingRegion>
,
<collectingRegion id="49E7F853FFF2FFA07270FF24BE2D1EAE" box="[786,887,223,249]" country="Argentina" name="Formosa" pageId="15" pageNumber="76">Formosa</collectingRegion>
, and
<collectingRegion id="49E7F853FFF2FFA072DBFF24B97A1EAE" box="[953,1056,223,249]" country="Argentina" name="Santa Fe" pageId="15" pageNumber="76">Santa Fé</collectingRegion>
) and
<collectingCountry id="F3347621FFF2FFA07507FF24B9F31EAE" box="[1125,1193,223,249]" name="Brazil" pageId="15" pageNumber="76">Brazil</collectingCountry>
(
<collectingRegion id="49E7F853FFF2FFA075DEFF24B80E1EAE" box="[1212,1364,223,249]" country="Brazil" name="Mato Grosso" pageId="15" pageNumber="76">Mato Grosso</collectingRegion>
) (
<figureCitation id="13182A34FFF2FFA0740DFF24BDE61F4A" captionStart="FIGURE 7" captionStartId="18.[151,250,1872,1896]" captionTargetBox="[156,1423,190,1839]" captionTargetId="figure-30@18.[155,1432,181,1847]" captionTargetPageId="18" captionText="FIGURE 7. Known distribution in South America of species of Pseudocanthon Bates, 1887. A, P. perplexus (LeConte, 1847) and P. xanthurus (Blanchard, 1847); B, P. vazdemelloi new species, P. pantanensis new species, and P. chaquensis new species." figureDoi="http://doi.org/10.5281/zenodo.5448102" httpUri="https://zenodo.org/record/5448102/files/figure.png" pageId="15" pageNumber="76">Fig. 7B</figureCitation>
).
<emphasis id="B957EAA3FFF2FFA071B2FEF9BE241F4B" bold="true" box="[208,894,258,284]" pageId="15" pageNumber="76">Areas of endemism [South America]: Brazilian subregion</emphasis>
: South Brazilian dominion:
<collectingRegion id="49E7F853FFF2FFA075DDFEF8B8C31F4A" box="[1215,1433,259,285]" country="Brazil" name="Rondonia" pageId="15" pageNumber="76">Rondônia province</collectingRegion>
.
<emphasis id="B957EAA3FFF2FFA071F5FEDDBC241F17" bold="true" box="[151,382,294,320]" pageId="15" pageNumber="76">Chacoan subregion</emphasis>
: Chacoan dominion: Chaco province (see
<bibRefCitation id="EFB24B40FFF2FFA07239FEDCB95A1F16" author="Morrone, J." box="[859,1024,295,321]" pageId="15" pageNumber="76" pagination="1 - 110" refId="ref21576" refString="Morrone, J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782, 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1." type="journal article" year="2014">Morrone 2014</bibRefCitation>
: fig. 12).
</paragraph>
</subSubSection>
</treatment>
</document>