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<document id="AB52A6CBB1C95A39862F064E096C51FB" ID-DOI="10.11646/zootaxa.3077.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5243536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629750005925" checkinUser="felipe" docAuthor="Peck, Stewart B. &amp; Cook, Joyce" docDate="2011" docId="03B375533D58A15F9DC0F8A1FDBA0A96" docLanguage="en" docName="zt03077p118.pdf" docOrigin="Zootaxa 3077 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3077.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Pinodytes haidagwaii Peck &amp; Cook 2011, new species" docType="treatment" docVersion="3" lastPageNumber="46" masterDocId="FF8A0D2B3D74A1729D57FF87FFB30F0A" masterDocTitle="Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077" masterLastPageNumber="118" masterPageNumber="1" pageNumber="45" updateTime="1699105988326" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="97BAD11F04B81B4ACFA5FC0878224E3A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BAC43A1166DE0DD104696AFFFDC2C66F">
<mods:title id="6EA7232C33DEB197B5D649516ABAA69D">Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077</mods:title>
</mods:titleInfo>
<mods:name id="4D85F5D483A5685DA26CAB7FE51ECA50" type="personal">
<mods:role id="89C6A5EA617CCD604047E54A089D3057">
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</mods:role>
<mods:namePart id="397DD6A1D261130D929A5DCF75EBD029">Peck, Stewart B.</mods:namePart>
</mods:name>
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<mods:namePart id="89E419231E2F1D8303C6F42E39F02974">Cook, Joyce</mods:namePart>
</mods:name>
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<mods:title id="45246AB5ADA2C93D7EECA0273F521EA6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5C2C566B8DB364E0DF293EF8381B1532">
<mods:date id="47D57EE7491D3803C7FC59836404314C">2011</mods:date>
<mods:detail id="CC065A873FD3C9B6EB614384241495B8" type="pubDate">
<mods:number id="96C766A9FBA6C1F28D887E30FEE9D66C">2011-10-28</mods:number>
</mods:detail>
<mods:detail id="55D7B0D9F2913A19693E33F257F570D9" type="volume">
<mods:number id="75C1A02C987E087D28AC82766CD5818A">3077</mods:number>
</mods:detail>
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<mods:number id="1CA27FDDDBD831DD6FAF31D13D9F5C2C">1</mods:number>
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<mods:url id="994813CA588F06480AB9A24B62E6FF14">https://biotaxa.org/Zootaxa/article/view/zootaxa.3077.1.1</mods:url>
</mods:location>
<mods:classification id="1BD4770A6FCFD1410066E14DF3F82D05">journal article</mods:classification>
<mods:identifier id="4EA02610FD58FFC6B1DF762CF4338EA2" type="DOI">10.11646/zootaxa.3077.1.1</mods:identifier>
<mods:identifier id="9B0EF3DB28417237DAE1B0DA319AFAF5" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="B6B9EC439DB4C05F2E1BB95CA33727A7" type="Zenodo-Dep">5243536</mods:identifier>
</mods:mods>
<treatment id="03B375533D58A15F9DC0F8A1FDBA0A96" LSID="urn:lsid:plazi:treatment:03B375533D58A15F9DC0F8A1FDBA0A96" httpUri="http://treatment.plazi.org/id/03B375533D58A15F9DC0F8A1FDBA0A96" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<subSubSection id="C30097CE3D58A15E9DC0F8A1FD56084A" box="[151,741,1830,1856]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BA5C4453D58A15E9DC0F8A1FD56084A" blockId="44.[151,741,1830,1889]" box="[151,741,1830,1856]" pageId="44" pageNumber="45">
<heading id="D0ED73293D58A15E9DC0F8A1FD56084A" bold="true" box="[151,741,1830,1856]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<emphasis id="B96E18573D58A15E9DC0F8A1FD56084A" bold="true" box="[151,741,1830,1856]" pageId="44" pageNumber="45">
<taxonomicName id="4C1ABFC63D58A15E9DC0F8A1FDF4084A" ID-CoL="4HZCK" authority="Peck &amp; Cook" authorityName="Peck &amp; Cook" authorityYear="2011" box="[151,583,1830,1856]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="haidagwaii" status="sp. nov.">
<emphasis id="B96E18573D58A15E9DC0F8A1FE2A084A" bold="true" box="[151,409,1830,1856]" italics="true" pageId="44" pageNumber="45">Pinodytes haidagwaii</emphasis>
Peck &amp; Cook
</taxonomicName>
,
<taxonomicNameLabel id="A25DA52C3D58A15E9F02F8A1FD56084A" box="[597,741,1830,1856]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D58A15E9DC0F8CEFE0C086B" box="[151,447,1865,1889]" pageId="44" pageNumber="45" type="description">
<paragraph id="8BA5C4453D58A15E9DC0F8CEFE0C086B" blockId="44.[151,741,1830,1889]" box="[151,447,1865,1889]" pageId="44" pageNumber="45">
(
<figureCitation id="1321D8C03D58A15E9DC8F8CEFF4E086B" box="[159,253,1865,1889]" captionStart="FIGURES 37–43" captionStartId="57.[151,264,1419,1442]" captionTargetBox="[185,1402,183,1398]" captionTargetId="figure-77@57.[185,1402,183,1398]" captionTargetPageId="57" captionText="FIGURES 37–43. Pinodytes habitus photos. 37. P.garibaldi, total length 2.80 mm. 38. P. haidagwaii, total length 3.04 mm. 39. P. isabella, total length 2.18 mm. 40. P.orca, total length 4.20 mm. 41. P.setosus, total length 1.96 mm. 42. P. shoshone, total length 2.18 mm. 43. P. sinuatus, total length 2.28 mm." figureDoi="http://doi.org/10.5281/zenodo.5243555" httpUri="https://zenodo.org/record/5243555/files/figure.png" pageId="44" pageNumber="45">Figs. 38</figureCitation>
,
<figureCitation id="1321D8C03D58A15E9C5FF8CEFEF4086B" box="[264,327,1865,1889]" captionStart="FIGURE265" captionStartId="90.[151,251,1103,1126]" captionTargetBox="[212,1385,193,1082]" captionTargetId="figure-110@90.[202,1385,193,1082]" captionTargetPageId="90" captionText="FIGURE265. Distributions of species in the Pinodytes cryptophagoides species group in northwestern North America. Map A. P. cryptophagoides, dots, extending from southeastern Washington State, through coastal mainland and insular British Columbia, to Sitka, Alaska. The distribution may have been achieved by coastal currents and dispersal on slide rafts. Map B. P. haidagwaii, dots, restricted to the Haida Gwaii (Formerly Queen Charlotte) Islands. This distribution argues for Pleistocene survival on the islands, which served as a Pleistocene refugium, and possibly for a pre-Pleistocene speciation event on the islands." figureDoi="http://doi.org/10.5281/zenodo.5243667" httpUri="https://zenodo.org/record/5243667/files/figure.png" pageId="44" pageNumber="45">265B</figureCitation>
,
<figureCitation id="1321D8C03D58A15E9C04F8CEFE04086B" box="[339,439,1865,1889]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="44" pageNumber="45">343–350</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D58A15F9DC0F811FC900E16" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="8BA5C4453D58A15E9DC0F811FA2808FC" blockId="44.[151,1436,1941,2038]" pageId="44" pageNumber="45">
Type material.
<materialsCitation id="3B72CE183D58A15E9C1BF811FCF308DB" collectingDate="1983-08-17" collectionCode="CNCI" collectorName="Queen Charlotte Is. &amp; Graham Is., J. M" country="Canada" location="British Columbia" pageId="44" pageNumber="45" specimenCount="1" stateProvince="British Columbia" typeStatus="holotype">
<emphasis id="B96E18573D58A15E9C1BF811FE7008A4" bold="true" box="[332,451,1942,1966]" pageId="44" pageNumber="45">
<typeStatus id="54A17AE73D58A15E9C1BF811FE0E08A4" box="[332,445,1942,1966]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (
<collectionCode id="ED0B5C803D58A15E9F47F811FDEA08A4" box="[528,601,1942,1966]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="44" pageNumber="45">CNCI</collectionCode>
).
<emphasis id="B96E18573D58A15E9F3BF812FC7D08A4" bold="true" box="[620,974,1941,1966]" pageId="44" pageNumber="45">
<collectingCountry id="F30D84D53D58A15E9F3BF812FD5408A4" box="[620,743,1941,1966]" name="Canada" pageId="44" pageNumber="45">CANADA</collectingCountry>
.
<collectingRegion id="49DE0AA73D58A15E9FA3F812FC7A08A4" box="[756,969,1941,1966]" country="Canada" name="British Columbia" pageId="44" pageNumber="45">British Columbia</collectingRegion>
:
</emphasis>
<collectorName id="26EFA1933D58A15E9E82F811FB0708A7" box="[981,1204,1942,1966]" pageId="44" pageNumber="45">Queen Charlotte Is.</collectorName>
: Kiusta,
<collectorName id="26EFA1933D58A15E984FF811FF7608DB" pageId="44" pageNumber="45">Graham Is., J.M</collectorName>
. Campbell,
<date id="FFA4E2853D58A15E9C1CF83EFE5308DB" box="[331,480,1977,2001]" pageId="44" pageNumber="45" value="1983-08-17">
<collectingDate id="EFE01B6D3D58A15E9C1CF83EFE5308DB" box="[331,480,1977,2001]" pageId="44" pageNumber="45" value="1983-08-17">17.VIII.1983</collectingDate>
</date>
, 83-90, sifting squirrel midden
</materialsCitation>
.
<materialsCitation id="3B72CE183D58A15E9E1CF83DFA2808FC" collectingDate="1983-08-17" collectionCode="CNCI, MCZC" collectorName="Graham Is. &amp; Queen Charlotte" country="Canada" location="British Columbia" pageId="44" pageNumber="45" specimenCount="1" stateProvince="British Columbia" typeStatus="paratype">
<emphasis id="B96E18573D58A15E9E1CF83DFC7008D8" bold="true" box="[843,963,1978,2002]" pageId="44" pageNumber="45">
<typeStatus id="54A17AE73D58A15E9E1CF83DFC7008D8" box="[843,963,1978,2002]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
</emphasis>
(5).
<emphasis id="B96E18573D58A15E9EAEF83EFAE808D8" bold="true" box="[1017,1371,1977,2002]" pageId="44" pageNumber="45">
<collectingCountry id="F30D84D53D58A15E9EAEF83EFBC708D8" box="[1017,1140,1977,2002]" name="Canada" pageId="44" pageNumber="45">CANADA</collectingCountry>
.
<collectingRegion id="49DE0AA73D58A15E99D6F83EFAE508D8" box="[1153,1366,1977,2002]" country="Canada" name="British Columbia" pageId="44" pageNumber="45">British Columbia</collectingRegion>
:
</emphasis>
same data as holotype, 1 (
<collectionCode id="ED0B5C803D58A15E9C21F859FE0C08FC" box="[374,447,2014,2038]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="44" pageNumber="45">CNCI</collectionCode>
); Massett,
<collectorName id="26EFA1933D58A15E9F60F859FD0608FF" box="[567,693,2014,2038]" pageId="44" pageNumber="45">Graham Is.</collectorName>
, III.46, Mrs. Clark, sifting moss, etc., 1 (
<collectionCode id="ED0B5C803D58A15E992CF859FB6208FC" box="[1147,1233,2014,2038]" pageId="44" pageNumber="45">MCZC</collectionCode>
);
<collectorName id="26EFA1933D58A15E99B3F859FA2808FC" box="[1252,1435,2014,2038]" pageId="44" pageNumber="45">Queen Charlotte</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BA5C4453D59A15F9DC0FF1FFA2F0FFD" blockId="45.[151,1440,151,1436]" pageId="45" pageNumber="46">
<materialsCitation id="3B72CE183D59A15F9DC0FF1FFB660FA5" box="[151,1237,151,175]" collectionCode="SBPC" collectorName="Graham Is. &amp; R. S. Anderson" pageId="45" pageNumber="46" specimenCount="1">
Is.,
<collectorName id="26EFA1933D59A15F9D95FF10FEF20FA5" box="[194,321,151,175]" pageId="45" pageNumber="46">Graham Is.</collectorName>
, 1miNW Tlell, 9.VII.84,
<collectorName id="26EFA1933D59A15F9F33FF10FCBE0FA5" box="[612,781,151,175]" pageId="45" pageNumber="46">R.S. Anderson</collectorName>
, Sitka spruce/hemlock forest, 1 (
<collectionCode id="ED0B5C803D59A15F99D4FF10FB7E0FA5" box="[1155,1229,151,175]" pageId="45" pageNumber="46">SBPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72CE183D59A15F99B5FF10FB150FDE" collectingDate="1983-08-10" collectionCode="CNCI" collectorName="Lyall Is. &amp; Gate Ck. &amp; J. M. Campbell" location="Queen" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Charlotte">
<location id="8EC5929E3D59A15F99B5FF10FA980FA5" LSID="urn:lsid:plazi:treatment:03B375533D58A15F9DC0F8A1FDBA0A96:8EC5929E3D59A15F99B5FF10FA980FA5" box="[1250,1323,151,175]" name="Queen" pageId="45" pageNumber="46" stateProvince="Charlotte">Queen</location>
<collectingRegion id="49DE0AA73D59A15F9864FF10FA280FA5" box="[1331,1435,151,175]" country="Saint Vincent and the Grenadines" name="Charlotte" pageId="45" pageNumber="46">Charlotte</collectingRegion>
Is.,
<collectorName id="26EFA1933D59A15F9D94FF3BFE970FD9" box="[195,292,188,212]" pageId="45" pageNumber="46">Lyall Is.</collectorName>
,
<collectorName id="26EFA1933D59A15F9C64FF3BFE2E0FDE" box="[307,413,188,212]" pageId="45" pageNumber="46">Gate Ck.</collectorName>
,
<date id="FFA4E2853D59A15F9CFCFF3BFD8C0FDE" box="[427,575,188,212]" pageId="45" pageNumber="46" value="1983-08-10">
<collectingDate id="EFE01B6D3D59A15F9CFCFF3BFD8C0FDE" box="[427,575,188,212]" pageId="45" pageNumber="46" value="1983-08-10">10.VIII.1983</collectingDate>
</date>
,
<collectorName id="26EFA1933D59A15F9F19FF3BFD490FDE" box="[590,762,188,212]" pageId="45" pageNumber="46">J.M. Campbell</collectorName>
, 83-71, sifting alder litter, 1 (
<collectionCode id="ED0B5C803D59A15F9903FF3BFB2E0FDE" box="[1108,1181,188,212]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="45" pageNumber="46">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72CE183D59A15F99E3FF3BFA2B0FFD" collectingDate="1983-08-14" collectionCode="CNCI" collectorName="Moresby Is. &amp; J. M. Campbell" location="Cumshewa" municipality="Queen" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Charlotte">
<collectingMunicipality id="6BC15E3F3D59A15F99E3FF3BFB4E0FDE" box="[1204,1277,188,212]" pageId="45" pageNumber="46">Queen</collectingMunicipality>
<collectingRegion id="49DE0AA73D59A15F9850FF3BFADC0FDE" box="[1287,1391,188,212]" country="Saint Vincent and the Grenadines" name="Charlotte" pageId="45" pageNumber="46">Charlotte</collectingRegion>
Is.,
<collectorName id="26EFA1933D59A15F9DC0FF58FEAE0FFD" box="[151,285,223,247]" pageId="45" pageNumber="46">Moresby Is.</collectorName>
,
<location id="8EC5929E3D59A15F9C7EFF58FE1B0FFD" LSID="urn:lsid:plazi:treatment:03B375533D58A15F9DC0F8A1FDBA0A96:8EC5929E3D59A15F9C7EFF58FE1B0FFD" box="[297,424,223,247]" municipality="Queen" name="Cumshewa" pageId="45" pageNumber="46" stateProvince="Charlotte">Cumshewa</location>
,
<date id="FFA4E2853D59A15F9CE4FF58FDFB0FFD" box="[435,584,223,247]" pageId="45" pageNumber="46" value="1983-08-14">
<collectingDate id="EFE01B6D3D59A15F9CE4FF58FDFB0FFD" box="[435,584,223,247]" pageId="45" pageNumber="46" value="1983-08-14">14.VIII.1983</collectingDate>
</date>
,
<collectorName id="26EFA1933D59A15F9F04FF67FD480FFD" box="[595,763,223,247]" pageId="45" pageNumber="46">J.M. Campbell</collectorName>
, 83-82, sifting squirrel midden beside totem pole, 1 (
<collectionCode id="ED0B5C803D59A15F9810FF58FA230FFD" box="[1351,1424,223,247]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="45" pageNumber="46">CNCI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5C4453D59A15F9D90FE84FC900E16" blockId="45.[151,1440,151,1436]" box="[199,803,259,284]" pageId="45" pageNumber="46">
<emphasis id="B96E18573D59A15F9D90FE84FE030E11" box="[199,432,259,284]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">
<emphasis id="B96E18573D59A15F9D90FE84FE1A0E16" bold="true" box="[199,425,259,284]" pageId="45" pageNumber="46">Material examined</emphasis>
.
</emphasis>
We have examined
<specimenCount id="9D1C0FCC3D59A15F9FC5FE83FCAC0E16" box="[658,799,260,284]" pageId="45" pageNumber="46" type="generic">6 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D59A15F9D90FEA0FEEF0A5E" pageId="45" pageNumber="46" type="distribution">
<paragraph id="8BA5C4453D59A15F9D90FEA0FB9E0EFE" blockId="45.[151,1440,151,1436]" pageId="45" pageNumber="46">
<emphasis id="B96E18573D59A15F9D90FEA0FED30E4A" bold="true" box="[199,352,295,320]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">Distribution.</emphasis>
Specimens (
<figureCitation id="1321D8C03D59A15F9CA5FEA0FDD60E35" box="[498,613,295,319]" captionStart="FIGURE265" captionStartId="90.[151,251,1103,1126]" captionTargetBox="[212,1385,193,1082]" captionTargetId="figure-110@90.[202,1385,193,1082]" captionTargetPageId="90" captionText="FIGURE265. Distributions of species in the Pinodytes cryptophagoides species group in northwestern North America. Map A. P. cryptophagoides, dots, extending from southeastern Washington State, through coastal mainland and insular British Columbia, to Sitka, Alaska. The distribution may have been achieved by coastal currents and dispersal on slide rafts. Map B. P. haidagwaii, dots, restricted to the Haida Gwaii (Formerly Queen Charlotte) Islands. This distribution argues for Pleistocene survival on the islands, which served as a Pleistocene refugium, and possibly for a pre-Pleistocene speciation event on the islands." figureDoi="http://doi.org/10.5281/zenodo.5243667" httpUri="https://zenodo.org/record/5243667/files/figure.png" pageId="45" pageNumber="46">Fig. 265B</figureCitation>
) are known only from Graham, Lyall, and Moresby Islands of the Haida Gwaii (formerly Queen
<collectingRegion id="49DE0AA73D59A15F9CE5FECBFDAF0E6E" box="[434,540,332,356]" country="Saint Vincent and the Grenadines" name="Charlotte" pageId="45" pageNumber="46">Charlotte</collectingRegion>
) Islands,
<collectingRegion id="49DE0AA73D59A15F9FC5FECBFCEF0E6E" box="[658,860,332,356]" country="Canada" name="British Columbia" pageId="45" pageNumber="46">British Columbia</collectingRegion>
. The species is endemic to these islands, and its restriction to this island group argues that it survived Pleistocene glaciation on the islands. Therefore the islands (and their enlarged areas with lower sea levels) were not entirely ice covered, and they served as a glacial refugium. Other plants and animals are also known to occur only on these islands, and they are additional evidence for their being at least partly ice free and acting as a refugium (
<bibRefCitation id="EF8BB9B43D59A15F9FA2FE5BFB920EFE" author="Scudder, G. G. E. &amp; Gessler, N." box="[757,1057,476,500]" pageId="45" pageNumber="46" refId="ref57161" refString="Scudder, G. G. E. and Gessler, N. (Eds.). 1989. The outer shores, based on the proceedings of the Queen Charlotte Islands First International Scientific Symposium, University of British Columbia, August 1984. Queen Charlotte Islands Museum Press, Skidgate, BC." type="proceedings" year="1989">Scudder and Gessler 1989</bibRefCitation>
).
</paragraph>
<paragraph id="8BA5C4453D59A15F9D90FE78FD540D8E" blockId="45.[151,1440,151,1436]" pageId="45" pageNumber="46">
This is the only species that has been found in sympatry (co-occurrence) with another species of
<taxonomicName id="4C1ABFC63D59A15F985FFD87FAC50D12" authorityName="Horn" authorityYear="1880" box="[1288,1398,512,536]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B96E18573D59A15F985FFD87FAC50D12" box="[1288,1398,512,536]" italics="true" pageId="45" pageNumber="46">Pinodytes</emphasis>
</taxonomicName>
(
<taxonomicName id="4C1ABFC63D59A15F98D0FD86FEE10D36" authorityName=": Hatch" authorityYear="1928" class="Insecta" family="Leiodidae" genus="Catopocerus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cryptophagoides">
<emphasis id="B96E18573D59A15F98D0FD86FEE10D36" italics="true" pageId="45" pageNumber="46">P. cryptophagoides</emphasis>
</taxonomicName>
). We interpret the case to be one in which
<taxonomicName id="4C1ABFC63D59A15F9E17FDA2FC680D36" authorityName="Peck &amp; Cook" authorityYear="2011" box="[832,987,548,572]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="haidagwaii">
<emphasis id="B96E18573D59A15F9E17FDA2FCFD0D36" box="[832,846,549,572]" italics="true" pageId="45" pageNumber="46">P</emphasis>
.
<emphasis id="B96E18573D59A15F9E0AFDA3FC680D36" box="[861,987,548,572]" italics="true" pageId="45" pageNumber="46">haidagwaii</emphasis>
</taxonomicName>
originated on the Haida Gwaii Islands, and
<taxonomicName id="4C1ABFC63D59A15F9D9EFDCEFE110D6A" authorityName=": Hatch" authorityYear="1928" box="[201,418,584,608]" class="Insecta" family="Leiodidae" genus="Catopocerus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="cryptophagoides">
<emphasis id="B96E18573D59A15F9D9EFDCEFE110D6A" box="[201,418,584,608]" italics="true" pageId="45" pageNumber="46">P. cryptophagoides</emphasis>
</taxonomicName>
dispersed to the islands in a later, possibly postglacial, time. There are three collection events in which the two species were found together.
</paragraph>
<paragraph id="8BA5C4453D59A15F9D90FD08FEEF0A5E" blockId="45.[151,1440,151,1436]" pageId="45" pageNumber="46">
<emphasis id="B96E18573D59A15F9D90FD08FE690DAD" box="[199,474,655,680]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">
<emphasis id="B96E18573D59A15F9D90FD08FE600DA2" bold="true" box="[199,467,655,680]" pageId="45" pageNumber="46">Diagnostic description</emphasis>
.
</emphasis>
Total length
<quantity id="4CE269A03D59A15F9F25FD08FCAA0DAD" box="[626,793,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="3.04" metricValueMin="2.3" pageId="45" pageNumber="46" unit="mm" value="2.67" valueMax="3.04" valueMin="2.3">2.30–3.04 mm</quantity>
; greatest width
<quantity id="4CE269A03D59A15F9E99FD08FBC60DAD" box="[974,1141,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.145" metricValueMax="1.24" metricValueMin="1.05" pageId="45" pageNumber="46" unit="mm" value="1.145" valueMax="1.24" valueMin="1.05">1.05–1.24 mm</quantity>
. Reddish brown; elongate in shape (
<figureCitation id="1321D8C03D59A15F9C52FD33FEE90DC6" box="[261,346,692,716]" captionStart="FIGURES 37–43" captionStartId="57.[151,264,1419,1442]" captionTargetBox="[185,1402,183,1398]" captionTargetId="figure-77@57.[185,1402,183,1398]" captionTargetPageId="57" captionText="FIGURES 37–43. Pinodytes habitus photos. 37. P.garibaldi, total length 2.80 mm. 38. P. haidagwaii, total length 3.04 mm. 39. P. isabella, total length 2.18 mm. 40. P.orca, total length 4.20 mm. 41. P.setosus, total length 1.96 mm. 42. P. shoshone, total length 2.18 mm. 43. P. sinuatus, total length 2.28 mm." figureDoi="http://doi.org/10.5281/zenodo.5243555" httpUri="https://zenodo.org/record/5243555/files/figure.png" pageId="45" pageNumber="46">Fig. 38</figureCitation>
).
<emphasis id="B96E18573D59A15F9C38FD33FE1C0DC6" bold="true" box="[367,431,692,716]" pageId="45" pageNumber="46">Head</emphasis>
. Finely punctate, punctures separated by 2–5 diameters; with a mixture of reticulate and substriate microsculpture. Eyes absent. Antenna (
<figureCitation id="1321D8C03D59A15F9FE8FD50FCAE0DE5" box="[703,797,727,751]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 343</figureCitation>
) with antennomeres 2 and 3 subequal in length; antennomere 5 larger than 4 and 6; antennomere 7 clearly larger than 8; antennomeres 9 and 10 lack visible sensory vesicles.
<emphasis id="B96E18573D59A15F9D82FCA7FEFD0C32" bold="true" box="[213,334,800,824]" pageId="45" pageNumber="46">Pronotum</emphasis>
. Finely punctate, punctures separated by 2–4 diameters, with a few larger punctures, usually paired; with reticulate microsculpture. Sides weakly rounded, widest near middle; apical margin weakly emarginate, basal margin straight; apical angles rounded, basal angles weakly obtuse.
<emphasis id="B96E18573D59A15F995DFCEFFBE50C8A" bold="true" box="[1034,1110,872,896]" pageId="45" pageNumber="46">Elytra</emphasis>
. Strial punctures moderately coarse; striae irregularly impressed; interstrial punctures fine; imbricate microsculpture near base. Joined elytra slightly wider than pronotum; sides subparallel in basal one-half, narrowing apically.
<emphasis id="B96E18573D59A15F992BFC37FB070CC2" bold="true" box="[1148,1204,944,968]" pageId="45" pageNumber="46">Legs</emphasis>
. Protibia (
<figureCitation id="1321D8C03D59A15F9867FC28FA270CCD" box="[1328,1428,943,967]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 344</figureCitation>
) broad at apex in male; two curved spines at apex of outer margin; apical one-half of inner margin with fine, dense spines. Mesotibia (
<figureCitation id="1321D8C03D59A15F9C27FC70FE610B05" box="[368,466,1015,1039]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 345</figureCitation>
) weakly curved in male, with strong spines on outer margin and apically. Metatibia (
<figureCitation id="1321D8C03D59A15F9DC8FB9BFEB50B3E" box="[159,262,1052,1076]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 346</figureCitation>
) slender, widened apically. Metafemur (
<figureCitation id="1321D8C03D59A15F9FB9FB9BFCE60B3E" box="[750,853,1052,1076]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 346</figureCitation>
) slender. Male protarsomeres (
<figureCitation id="1321D8C03D59A15F999AFB9BFA870B3E" box="[1229,1332,1052,1076]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 344</figureCitation>
) weakly expanded; protarsomere 1 as long as 2–4 combined; bearing elongate setae laterally and thin, broad, colorless phanerae ventrally. Mesotarsomeres without phanerae.
<emphasis id="B96E18573D59A15F9E77FBE3FCDD0B76" bold="true" box="[800,878,1124,1148]" pageId="45" pageNumber="46">Venter</emphasis>
. Mesoventrite (
<figureCitation id="1321D8C03D59A15F997BFBE3FB210B76" box="[1068,1170,1124,1148]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 350</figureCitation>
) carinate; longitudinal carina with a median tooth; depressed anterior to median tooth; excavation behind transverse carina.
<emphasis id="B96E18573D59A15F9852FB0FFF070BCE" bold="true" pageId="45" pageNumber="46">Male genitalia</emphasis>
. Median lobe of aedeagus (
<figureCitation id="1321D8C03D59A15F9CA6FB2BFD250BCE" box="[497,662,1196,1220]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Figs. 347, 348</figureCitation>
) elongate, broad, evenly curved at basal one-third; sharply angulate before elongate, flattened apex. Inverted internal sac (
<figureCitation id="1321D8C03D59A15F9FA2FB48FCE70BED" box="[757,852,1231,1255]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 348</figureCitation>
) with a cluster of short, broad spines apically, elongate spines medially, and large spines basally. Parameres (
<figureCitation id="1321D8C03D59A15F9E63FB73FC680A06" box="[820,987,1268,1292]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Figs. 347, 348</figureCitation>
) slender, less than two-thirds length of median lobe; each paramere bearing one apical and one subapical seta.
<emphasis id="B96E18573D59A15F9EE3FA90FBE20A3A" bold="true" box="[948,1105,1303,1328]" pageId="45" pageNumber="46">Spermatheca</emphasis>
. Elongate (
<figureCitation id="1321D8C03D59A15F9986FA90FA800A25" box="[1233,1331,1303,1327]" captionStart="FIGURES 343–350" captionStartId="102.[151,263,1604,1627]" captionTargetBox="[183,1408,200,1577]" captionTargetId="figure-70@102.[172,1414,193,1583]" captionTargetPageId="102" captionText="FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350)." figureDoi="http://doi.org/10.5281/zenodo.5243709" httpUri="https://zenodo.org/record/5243709/files/figure.png" pageId="45" pageNumber="46">Fig. 349</figureCitation>
), tubular, sigmoid in shape.
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D59A15F9D90FAE7FDBA0A96" pageId="45" pageNumber="46" type="etymology">
<paragraph id="8BA5C4453D59A15F9D90FAE7FDBA0A96" blockId="45.[151,1440,151,1436]" pageId="45" pageNumber="46">
<emphasis id="B96E18573D59A15F9D90FAE7FEFD0A72" bold="true" box="[199,334,1376,1400]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">Etymology.</emphasis>
The name
<taxonomicName id="4C1ABFC63D59A15F9C9BFAE7FDF90A72" authorityName="Peck &amp; Cook" authorityYear="2011" box="[460,586,1376,1400]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="haidagwaii">
<emphasis id="B96E18573D59A15F9C9BFAE7FDF90A72" box="[460,586,1376,1400]" italics="true" pageId="45" pageNumber="46">haidagwaii</emphasis>
</taxonomicName>
, a noun in apposition, refers to the distribution of this species on the Haida Gwaii Islands,
<collectingRegion id="49DE0AA73D59A15F9C68FA03FDB60A96" box="[319,517,1412,1436]" country="Canada" name="British Columbia" pageId="45" pageNumber="46">British Columbia</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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