291 lines
35 KiB
XML
291 lines
35 KiB
XML
<document id="B7CC4CF833C0BDBFF026EE32EFC01919" ID-CLB-Dataset="8717" ID-DOI="10.11646/zootaxa.4908.4.4" ID-GBIF-Dataset="8cb0c3e0-d9aa-4020-b0b5-fa55bdd5d4e0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4447189" ID-ZooBank="C7E21237-4524-460D-BD3F-DFAED905A60B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610978992805" checkinUser="plazi" docAuthor="Zakrzewska, Marta & Jankowska, Ariadna" docDate="2021" docId="575A879DF1306E5AC8818E0AFD40F83C" docLanguage="en" docName="zootaxa4908.4.4.pdf" docOrigin="Zootaxa 4908 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stempellinella gilkai Zakrzewska & Jankowska 2021, sp. nov." docType="treatment" docUuid="CE2F9673-970F-4DA7-80B9-0E67AF40DF44" docUuidSource="ZooBank" docVersion="9" lastPageNumber="507" masterDocId="AB63FFE5F1316E58C8168B3FFF93FFC5" masterDocTitle="A new fossil Stempellinella from Eocene Baltic amber, with systematic notes and amendment of the generic diagnosis (Diptera: Chironomidae)" masterLastPageNumber="514" masterPageNumber="505" pageNumber="506" updateTime="1698924581037" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="C0817C0C234D183A695B6A75EB21A180" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="068D9CE71686F02D44D39853AEAE6156">
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<mods:title id="B809ED0A6D79EB7F03D965C40B6002C9">A new fossil Stempellinella from Eocene Baltic amber, with systematic notes and amendment of the generic diagnosis (Diptera: Chironomidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C396AEB04ABEF4993A2F68088B7798B9" type="personal">
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<mods:role id="AFEA01D90E6CEAA213F3046E494050F8">
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<mods:roleTerm id="030C20713A722ADDF662459460278F2F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C4602CE12B73E94A2D21FE7CBACB75A5">Zakrzewska, Marta</mods:namePart>
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<mods:affiliation id="3BCAA8789A703ECA60EEE724DE119551">University of Gdańsk, Faculty of Biology, Department of Invertebrate Zoology and Parasitology, Laboratory of Systematic Zoology, Wita Stwosza 59, 80 - 308 Gdańsk, Poland</mods:affiliation>
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</mods:name>
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<mods:name id="4F6215824CEB8B6B87FA707696697AEE" type="personal">
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<mods:role id="C89B3EFEB2534B9F1B3627B819C112FC">
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<mods:roleTerm id="58D61E3E6ED3C77B9F5CA3DF45EF6AEF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6CA95CFAFCE41348FB852166206E29E7">Jankowska, Ariadna</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="BC7B007FA035B0FAAA8A238BA1C937BC">text</mods:typeOfResource>
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<mods:titleInfo id="DA882F068E642432193FDCE58133D4DF">
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<mods:title id="29F86C006994FA066B3221690B08F742">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E7F6EFFE990D2C3F76AE4F090C82EB55">
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<mods:date id="8C19444A7D11184449248777704765ED">2021</mods:date>
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<mods:detail id="33885FB34C193C61588208D27E8CD119" type="pubDate">
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<mods:number id="86821437A102F88541788262612B879A">2021-01-18</mods:number>
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<mods:number id="96C611137717A4909C10AEB765DAC0AF">4908</mods:number>
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<mods:detail id="78201FDD8466711D89CB41D99876F126" type="issue">
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<mods:number id="18AD5DF0AD4DAF68034B4506A132734E">4</mods:number>
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<mods:classification id="4BF8CEE0B7D0A902C939BD6959BFC692">journal article</mods:classification>
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<mods:identifier id="36B9A615902B1A289C1834D34FFF5588" type="DOI">10.11646/zootaxa.4908.4.4</mods:identifier>
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<mods:identifier id="1B399D7B5B7F44CBC863565524A46507" type="ZooBank">C7E21237-4524-460D-BD3F-DFAED905A60B</mods:identifier>
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</mods:mods>
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<treatment id="575A879DF1306E5AC8818E0AFD40F83C" ID-DOI="http://doi.org/10.5281/zenodo.4455437" ID-GBIF-Taxon="177013642" ID-Zenodo-Dep="4455437" LSID="urn:lsid:zoobank.org:act:CE2F9673-970F-4DA7-80B9-0E67AF40DF44" httpUri="http://treatment.plazi.org/id/575A879DF1306E5AC8818E0AFD40F83C" lastPageId="2" lastPageNumber="507" pageId="1" pageNumber="506">
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<subSubSection id="97E96500F1306E59C8818E0AFB83FAB1" pageId="1" pageNumber="506" type="nomenclature">
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<paragraph id="DF4C368BF1306E59C8818E0AFE66FA95" blockId="1.[151,1040,1333,1396]" box="[151,501,1333,1360]" pageId="1" pageNumber="506">
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<heading id="840481E7F1306E59C8818E0AFE66FA95" bold="true" box="[151,501,1333,1360]" fontSize="11" level="1" pageId="1" pageNumber="506" reason="1">
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<emphasis id="ED87EA99F1306E59C8818E0AFE66FA95" bold="true" box="[151,501,1333,1360]" pageId="1" pageNumber="506">
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<taxonomicName id="18F34D08F1306E59C8818E0AFE1CFA95" ID-CoL="8GTVN" authority="Zakrzewska & Jankowska, 2021" authorityName="Zakrzewska & Jankowska" authorityYear="2021" box="[151,399,1333,1360]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="species" species="gilkai" status="sp. nov.">
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<emphasis id="ED87EA99F1306E59C8818E0AFE1CFA95" bold="true" box="[151,399,1333,1360]" italics="true" pageId="1" pageNumber="506">Stempellinella gilkai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F6B457E2F1306E59C9818E09FE66FA95" box="[407,501,1334,1360]" pageId="1" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="DF4C368BF1306E59C8818E65FB83FAB1" blockId="1.[151,1040,1333,1396]" box="[151,1040,1370,1396]" pageId="1" pageNumber="506">
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LSID:
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<uri id="AB623A89F1306E59C8F28E65FB83FAB1" box="[228,1040,1370,1396]" pageId="1" pageNumber="506">
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urn:lsid:zoobank.org:act:
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<uuid id="AB550C5EF1306E59C9E38E65FB83FAB1" box="[501,1040,1370,1396]" pageId="1" pageNumber="506">CE2F9673-970F-4DA7-80B9-0E67AF40DF44</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="97E96500F1306E59C8818E9DFEF4F9ED" pageId="1" pageNumber="506" type="materials_examined">
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<paragraph id="DF4C368BF1306E59C8818E9DFEF4F9ED" blockId="1.[151,1437,1441,1937]" pageId="1" pageNumber="506">
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<emphasis id="ED87EA99F1306E59C8818E9DFED6FA7E" bold="true" box="[151,325,1441,1468]" pageId="1" pageNumber="506">Type material.</emphasis>
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<materialsCitation id="6F9B3CD6F1306E59C95D8E9DFEF4F9ED" ID-GBIF-Occurrence="3022776301" collectionCode="LSZ, DIZP, IR" pageId="1" pageNumber="506" specimenCount="2" specimenCount-adult="1" typeStatus="holotype">
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<typeStatus id="00488829F1306E59C95D8E9DFE24FA79" box="[331,439,1442,1468]" pageId="1" pageNumber="506" type="holotype">Holotype</typeStatus>
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. Adult male (tarsi of both fore, left mid and right hind leg missing) preserved in 30 x 24 x
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<quantity id="180B9B6EF1306E59C8818EF9FF4AFA25" box="[151,217,1478,1505]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="506" unit="mm" value="5.0">5 mm</quantity>
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piece of amber (
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<figureCitation id="47C82A0EF1306E59C9808EF9FD9DFA25" box="[406,526,1478,1504]" captionStart="FIGURE 2" captionStartId="3.[151,250,1508,1532]" captionTargetBox="[151,1435,181,1485]" captionTargetId="figure-19@3.[151,1435,181,1485]" captionTargetPageId="3" captionText="FIGURE 2. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A inclusion in amber, B habitus, C head and antenna (white arrows: borders between flagellomeres). Scale bars: 1 cm (A), 500 μm (B), 200 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.4447195" httpUri="https://zenodo.org/record/4447195/files/figure.png" pageId="1" pageNumber="506">Fig. 2A, B</figureCitation>
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) (Eocene Baltic amber, ~40–45 Ma, inventory no. F28a
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<collectionCode id="B9E2AE4EF1306E59CC9A8EF9FA94FA25" box="[1164,1287,1478,1504]" pageId="1" pageNumber="506">LSZ DIZP</collectionCode>
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;
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<collectionCode id="B9E2AE4EF1306E59CD048EF9FABDFA25" box="[1298,1326,1478,1504]" pageId="1" pageNumber="506">IR</collectionCode>
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spectrum no. 16218 IAA;
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<figureCitation id="47C82A0EF1306E59C95B8ED5FE1EF9C1" box="[333,397,1514,1540]" captionStart="FIGURE 1" captionStartId="2.[151,250,773,797]" captionTargetBox="[180,1415,183,749]" captionTargetId="figure-27@2.[170,1416,181,750]" captionTargetPageId="2" captionText="FIGURE 1. Fourier transform infrared spectroscopy (ATR) spectrum no. 16218 IAA obtained from the piece of Baltic amber (Eocene, ~40–45 Ma) with the examined inclusion." figureDoi="http://doi.org/10.5281/zenodo.4447191" httpUri="https://zenodo.org/record/4447191/files/figure.png" pageId="1" pageNumber="506">Fig. 1</figureCitation>
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). Syninclusions:
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<taxonomicName id="18F34D08F1306E59CA468ED5FD60F9C1" authorityName="Newman" authorityYear="1834" box="[592,755,1514,1540]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="subFamily" subFamily="Chironominae">Chironominae</taxonomicName>
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,
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<specimenCount id="C9F5FD02F1306E59CAE88ED5FCD9F9C1" box="[766,842,1514,1540]" pageId="1" pageNumber="506" type="adult">1 adult</specimenCount>
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male (F28b
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<collectionCode id="B9E2AE4EF1306E59CBC18ED5FBC1F9C1" box="[983,1106,1514,1540]" pageId="1" pageNumber="506">LSZ DIZP</collectionCode>
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), Sternorrhyncha,
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<specimenCount id="C9F5FD02F1306E59CD088ED5FA0FF9C1" box="[1310,1436,1514,1540]" pageId="1" pageNumber="506" type="generic">1 specimen</specimenCount>
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(F28c
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<collectionCode id="B9E2AE4EF1306E59C8F68D31FEC8F9ED" box="[224,347,1550,1576]" pageId="1" pageNumber="506">LSZ DIZP</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="97E96500F1306E59C8D18D0EFE94F9B5" pageId="1" pageNumber="506" type="etymology">
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<paragraph id="DF4C368BF1306E59C8D18D0EFE94F9B5" blockId="1.[151,1437,1441,1937]" pageId="1" pageNumber="506">
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<emphasis id="ED87EA99F1306E59C8D18D0EFE30F98E" bold="true" box="[199,419,1585,1611]" pageId="1" pageNumber="506">Derivatio nominis.</emphasis>
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Authors dedicate the specific epithet to their mentor, Prof. Wojciech Giłka, a
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<taxonomicName id="18F34D08F1306E59CCEA8D0DFA0FF989" authorityName="Newman" authorityYear="1834" box="[1276,1436,1586,1612]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="506" phylum="Arthropoda" rank="family">Chironomidae</taxonomicName>
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specialist.
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</paragraph>
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</subSubSection>
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<subSubSection id="97E96500F1306E59C8D18D46FEFAF88D" pageId="1" pageNumber="506" type="diagnosis">
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<paragraph id="DF4C368BF1306E59C8D18D46FEFAF88D" blockId="1.[151,1437,1441,1937]" pageId="1" pageNumber="506">
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<emphasis id="ED87EA99F1306E59C8D18D46FED1F956" bold="true" box="[199,322,1657,1683]" pageId="1" pageNumber="506">Diagnosis.</emphasis>
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Antennal flagellum composed of 13 flagellomeres. Sc ending far distal of RM; RM long, slightly oblique relative to M. Vein M
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<subScript id="437734CEF1306E59C9FA8D8FFD95F97B" attach="left" box="[492,518,1712,1726]" fontSize="6" pageId="1" pageNumber="506">3+4</subScript>
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ending opposite to R
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<subScript id="437734CEF1306E59CAEA8D8FFC85F97B" attach="left" box="[764,790,1712,1726]" fontSize="6" pageId="1" pageNumber="506">4+5</subScript>
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. Gonostylus slightly shorter than gonocoxite. Anal point stout, with strong crests framing numerous irregularly dispersed spinulae. Superior volsella elongate, peanut-shaped. Stem of median volsella long, bearing setiform and foliate lamellae, distal lamella with elongated filiform tip reaching apex of inferior volsella. Inferior volsella with spherical bulge bearing 4 medially directed distinct setae, apical part bent medially.
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</paragraph>
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</subSubSection>
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<subSubSection id="97E96500F1306E5AC8D18C6EFD40F83C" lastPageId="2" lastPageNumber="507" pageId="1" pageNumber="506" type="description">
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<paragraph id="DF4C368BF1306E59C8D18C6EFDA5F8A9" blockId="1.[151,1437,1441,1937]" box="[199,566,1873,1900]" pageId="1" pageNumber="506">
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<emphasis id="ED87EA99F1306E59C8D18C6EFECAF8AE" bold="true" box="[199,345,1873,1899]" pageId="1" pageNumber="506">Description.</emphasis>
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Adult male (n = 1).
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</paragraph>
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<paragraph id="DF4C368BF1306E59C8D18C49FD2CF855" blockId="1.[151,1437,1441,1937]" box="[199,703,1910,1937]" pageId="1" pageNumber="506">
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Total length ~
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<quantity id="180B9B6EF1306E59C9758C49FE59F855" box="[355,458,1910,1937]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="506" unit="mm" value="1.55">1.55 mm</quantity>
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; wing length 945 μm.
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</paragraph>
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<caption id="8B8C6603F1336E5AC881883AFD33FC87" ID-DOI="http://doi.org/10.5281/zenodo.4447191" ID-Zenodo-Dep="4447191" httpUri="https://zenodo.org/record/4447191/files/figure.png" pageId="2" pageNumber="507" startId="2.[151,250,773,797]" targetBox="[180,1415,183,749]" targetPageId="2">
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<paragraph id="DF4C368BF1336E5AC881883AFD33FC87" blockId="2.[151,1436,773,834]" pageId="2" pageNumber="507">
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||
<emphasis id="ED87EA99F1336E5AC881883AFE87FCDB" bold="true" box="[151,276,773,798]" pageId="2" pageNumber="507">FIGURE 1.</emphasis>
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Fourier transform infrared spectroscopy (ATR) spectrum no. 16218 IAA obtained from the piece of Baltic amber (Eocene, ~40–45 Ma) with the examined inclusion.
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||
</paragraph>
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||
</caption>
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||
<paragraph id="DF4C368BF1336E5AC8D1884FFE4EFC33" blockId="2.[151,1437,880,1338]" pageId="2" pageNumber="507">
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||
<emphasis id="ED87EA99F1336E5AC8D1884FFE97FC4F" box="[199,260,880,906]" italics="true" pageId="2" pageNumber="507">Head</emphasis>
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||
(
|
||
<figureCitation id="47C82A0EF1336E5AC903884FFE00FC4F" box="[277,403,880,906]" captionStart="FIGURE 2" captionStartId="3.[151,250,1508,1532]" captionTargetBox="[151,1435,181,1485]" captionTargetId="figure-19@3.[151,1435,181,1485]" captionTargetPageId="3" captionText="FIGURE 2. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A inclusion in amber, B habitus, C head and antenna (white arrows: borders between flagellomeres). Scale bars: 1 cm (A), 500 μm (B), 200 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.4447195" httpUri="https://zenodo.org/record/4447195/files/figure.png" pageId="2" pageNumber="507">Fig. 2B, C</figureCitation>
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). Eyes bare, without dorsomedian extension, broadly separated by frons. Frontal tubercles conical, length 8 μm, width 8 μm. Antenna with 13 flagellomeres (borders between flagellomeres 12 and 13 faintly visible but present) (
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<figureCitation id="47C82A0EF1336E5AC96B8887FE46FC17" box="[381,469,952,978]" captionStart="FIGURE 2" captionStartId="3.[151,250,1508,1532]" captionTargetBox="[151,1435,181,1485]" captionTargetId="figure-19@3.[151,1435,181,1485]" captionTargetPageId="3" captionText="FIGURE 2. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A inclusion in amber, B habitus, C head and antenna (white arrows: borders between flagellomeres). Scale bars: 1 cm (A), 500 μm (B), 200 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.4447195" httpUri="https://zenodo.org/record/4447195/files/figure.png" pageId="2" pageNumber="507">Fig. 2C</figureCitation>
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), AR 0.45, plume fully developed. Length of palpomeres 2–5 (in μm): 28, 60, 76, 117. Clypeus with at least 5 setae.
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</paragraph>
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<paragraph id="DF4C368BF1336E5AC8D18F3FFB2EFBDF" blockId="2.[151,1437,880,1338]" box="[199,1213,1024,1050]" pageId="2" pageNumber="507">
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||
<emphasis id="ED87EA99F1336E5AC8D18F3FFE8BFBDF" box="[199,280,1024,1050]" italics="true" pageId="2" pageNumber="507">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="47C82A0EF1336E5AC9318F3FFEECFBDF" box="[295,383,1024,1050]" captionStart="FIGURE 2" captionStartId="3.[151,250,1508,1532]" captionTargetBox="[151,1435,181,1485]" captionTargetId="figure-19@3.[151,1435,181,1485]" captionTargetPageId="3" captionText="FIGURE 2. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A inclusion in amber, B habitus, C head and antenna (white arrows: borders between flagellomeres). Scale bars: 1 cm (A), 500 μm (B), 200 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.4447195" httpUri="https://zenodo.org/record/4447195/files/figure.png" pageId="2" pageNumber="507">Fig. 2B</figureCitation>
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). Ac at least 7, Dc at least 7 on each side, Scts at least 3, Pa unobservable.
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</paragraph>
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<paragraph id="DF4C368BF1336E5AC8D18F1AFBA5FB6F" blockId="2.[151,1437,880,1338]" pageId="2" pageNumber="507">
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||
<emphasis id="ED87EA99F1336E5AC8D18F1AFE92FBFB" box="[199,257,1061,1086]" italics="true" pageId="2" pageNumber="507">Wing</emphasis>
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(
|
||
<figureCitation id="47C82A0EF1336E5AC9068F1BFEC7FBFB" box="[272,340,1060,1086]" captionStart="FIGURE 3" captionStartId="4.[151,250,1006,1030]" captionTargetBox="[170,1416,187,982]" captionTargetId="figure-27@4.[170,1416,181,982]" captionTargetPageId="4" captionText="FIGURE 3. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A, B wing. Scale bar: 100 μm. C—costa, Sc—subcosta, R, R1, R 4+5 —radial veins, RM—radial-median crossvein, M, M1+2, M3+4—median veins, Cu, Cu1—cubital veins, FCu—cubital crossvein, An—anal vein." figureDoi="http://doi.org/10.5281/zenodo.4447197" httpUri="https://zenodo.org/record/4447197/files/figure.png" pageId="2" pageNumber="507">Fig. 3</figureCitation>
|
||
). Slender, with anal lobe weak, broadest at 2/3 length; width: 285 μm, length/width ratio: 3.32. Sc ending well distal of RM and slightly distal of FCu, R
|
||
<subScript id="437734CEF1336E5ACAE28F65FC9EFBAD" attach="left" box="[756,781,1114,1128]" fontSize="6" pageId="2" pageNumber="507">2+3</subScript>
|
||
not visible. RM oblique relative to M. FCu placed distally of RM; VR
|
||
<subScript id="437734CEF1336E5AC90E8F41FEB8FB49" attach="left" box="[280,299,1150,1164]" fontSize="6" pageId="2" pageNumber="507">Cu</subScript>
|
||
1.36. Veins ending as follows (in order from base to tip): An, Sc, Cu
|
||
<subScript id="437734CEF1336E5ACC2D8F41FBD0FB49" attach="left" box="[1083,1091,1150,1164]" fontSize="6" pageId="2" pageNumber="507">1</subScript>
|
||
, R
|
||
<subScript id="437734CEF1336E5ACC728F41FBFFFB49" attach="left" box="[1124,1132,1150,1164]" fontSize="6" pageId="2" pageNumber="507">1</subScript>
|
||
, R
|
||
<subScript id="437734CEF1336E5ACC988F41FB3BFB49" attach="left" box="[1166,1192,1150,1164]" fontSize="6" pageId="2" pageNumber="507">4+5</subScript>
|
||
and M
|
||
<subScript id="437734CEF1336E5ACCE08F41FA83FB49" attach="left" box="[1270,1296,1150,1164]" fontSize="6" pageId="2" pageNumber="507">3+4</subScript>
|
||
, M
|
||
<subScript id="437734CEF1336E5ACD218F41FAC3FB49" attach="left" box="[1335,1360,1150,1164]" fontSize="6" pageId="2" pageNumber="507">1+2</subScript>
|
||
. Wing membrane and most veins (except subcosta and media) covered with macrotrichia.
|
||
</paragraph>
|
||
<paragraph id="DF4C368BF1336E5AC8D18F8AFD80FAFF" blockId="2.[151,1437,880,1338]" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC8D18F8AFF6FFB0B" box="[199,252,1205,1230]" italics="true" pageId="2" pageNumber="507">Legs</emphasis>
|
||
(
|
||
<figureCitation id="47C82A0EF1336E5AC91A8F8BFEF7FB0B" box="[268,356,1204,1230]" captionStart="FIGURE 2" captionStartId="3.[151,250,1508,1532]" captionTargetBox="[151,1435,181,1485]" captionTargetId="figure-19@3.[151,1435,181,1485]" captionTargetPageId="3" captionText="FIGURE 2. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A inclusion in amber, B habitus, C head and antenna (white arrows: borders between flagellomeres). Scale bars: 1 cm (A), 500 μm (B), 200 μm (C)." figureDoi="http://doi.org/10.5281/zenodo.4447195" httpUri="https://zenodo.org/record/4447195/files/figure.png" pageId="2" pageNumber="507">Fig. 2B</figureCitation>
|
||
). Tibial apices with their armature faintly visible. Foreleg tibia with fine spur, up to 10 μm long. Mid- and hindleg tibia with small combs ~10 μm long. One midleg comb bearing ~20 μm long spur, both hindleg combs with spurs (~15-25 μm long). Sensilla chaetica on ta
|
||
<subScript id="437734CEF1336E5ACB398E31FCA4FAD9" attach="left" box="[815,823,1294,1308]" fontSize="6" pageId="2" pageNumber="507">1</subScript>
|
||
of p
|
||
<subScript id="437734CEF1336E5ACB7D8E31FCE0FAD9" attach="left" box="[875,883,1294,1308]" fontSize="6" pageId="2" pageNumber="507">2</subScript>
|
||
not observed. Pulvilli absent. Lengths of leg segments and leg ratios as in
|
||
<tableCitation id="92710330F1336E5AC9AC8E1FFD9EFAFF" box="[442,525,1312,1338]" captionStart="TABLE 1" captionStartId="2.[151,239,1385,1409]" captionText="TABLE 1. Lengths (μm) of leg segments and leg ratios of male Stempellinella gilkai sp. nov., p 1 –p 3 = pair of legs 1–3, fe=femur, ti=tibia, ta 1 –ta 5 = tarsomeres 1–5, LR = leg ratio." pageId="2" pageNumber="507">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="8B8C6603F1336E5AC8818E56FC9FFA63" pageId="2" pageNumber="507">
|
||
<paragraph id="DF4C368BF1336E5AC8818E56FC9FFA63" blockId="2.[151,1436,1385,1452]" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC8818E56FE98FA47" bold="true" box="[151,267,1385,1410]" pageId="2" pageNumber="507">TABLE 1.</emphasis>
|
||
Lengths (μm) of leg segments and leg ratios of male
|
||
<taxonomicName id="18F34D08F1336E5ACB588E55FBBFFA47" authority="Zakrzewska & Jankowska, 2021" authorityName="Zakrzewska & Jankowska" authorityYear="2021" box="[846,1068,1386,1410]" class="Insecta" family="Chironomidae" genus="Stempellinella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="gilkai" status="sp. nov.">
|
||
<emphasis id="ED87EA99F1336E5ACB588E55FBBFFA47" box="[846,1068,1386,1410]" italics="true" pageId="2" pageNumber="507">Stempellinella gilkai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="ED87EA99F1336E5ACC258E55FB1CFA47" bold="true" box="[1075,1167,1386,1410]" pageId="2" pageNumber="507">
|
||
<taxonomicNameLabel id="F6B457E2F1336E5ACC258E55FB19FA47" box="[1075,1162,1386,1410]" pageId="2" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
p
|
||
<subScript id="437734CEF1336E5ACCB28E45FB3FFA4D" attach="left" box="[1188,1196,1402,1416]" fontSize="6" pageId="2" pageNumber="507">1</subScript>
|
||
–p
|
||
<subScript id="437734CEF1336E5ACCD18E45FB5CFA4D" attach="left" box="[1223,1231,1402,1416]" fontSize="6" pageId="2" pageNumber="507">3</subScript>
|
||
= pair of legs 1–3, fe=femur, ti=tibia, ta
|
||
<subScript id="437734CEF1336E5AC9638EA1FEEEFA69" attach="right" box="[373,381,1438,1452]" fontSize="6" pageId="2" pageNumber="507">1</subScript>
|
||
–ta
|
||
<subScript id="437734CEF1336E5AC98B8EA1FE36FA69" attach="left" box="[413,421,1438,1452]" fontSize="6" pageId="2" pageNumber="507">5</subScript>
|
||
= tarsomeres 1–5, LR = leg ratio.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DF4C368BF1336E5AC9228E89FADBF982" pageId="2" pageNumber="507">
|
||
<table id="ADF3C42BF13391A7C8B08E89FADBF988" box="[166,1352,1462,1613]" gridcols="9" gridrows="4" pageId="2" pageNumber="507">
|
||
<tr id="61C334C9F13391A7C8B08E89FADBFA11" box="[166,1352,1462,1492]" gridrow="0" pageId="2" pageNumber="507" rowspan-0="1">
|
||
<th id="22125DB5F13391A7C9228E89FEC9FA11" box="[308,346,1462,1492]" gridcol="1" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC9228E89FEDBFA0B" bold="true" box="[308,328,1462,1486]" pageId="2" pageNumber="507">fe</emphasis>
|
||
</th>
|
||
<th id="22125DB5F13391A7C9D58E89FE7AFA11" box="[451,489,1462,1492]" gridcol="2" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC9D58E89FE41FA0B" bold="true" box="[451,466,1462,1486]" pageId="2" pageNumber="507">ti</emphasis>
|
||
</th>
|
||
<th id="22125DB5F13391A7CA448E89FDEBFA11" box="[594,632,1462,1492]" gridcol="3" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5ACA448E89FDF4FA0B" bold="true" box="[594,615,1462,1486]" pageId="2" pageNumber="507">ta</emphasis>
|
||
<emphasis id="ED87EA99F1336E5ACA718EF9FDFDFA11" bold="true" box="[615,622,1478,1492]" pageId="2" pageNumber="507">1</emphasis>
|
||
</th>
|
||
<th id="22125DB5F13391A7CAF78E89FC94FA11" box="[737,775,1462,1492]" gridcol="4" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5ACAF78E89FD65FA0B" bold="true" box="[737,758,1462,1486]" pageId="2" pageNumber="507">ta</emphasis>
|
||
<emphasis id="ED87EA99F1336E5ACAE08EF9FD6EFA11" bold="true" box="[758,765,1478,1492]" pageId="2" pageNumber="507">2</emphasis>
|
||
</th>
|
||
<th id="22125DB5F13391A7CB798E89FC06FA11" box="[879,917,1462,1492]" gridcol="5" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5ACB798E89FC17FA0B" bold="true" box="[879,900,1462,1486]" pageId="2" pageNumber="507">ta</emphasis>
|
||
<emphasis id="ED87EA99F1336E5ACB938EF9FC1FFA11" bold="true" box="[901,908,1478,1492]" pageId="2" pageNumber="507">3</emphasis>
|
||
</th>
|
||
<th id="22125DB5F13391A7CBE88E89FB89FA11" box="[1022,1050,1462,1492]" gridcol="6" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5ACBE88E89FB80FA0B" bold="true" box="[1022,1043,1462,1486]" pageId="2" pageNumber="507">ta</emphasis>
|
||
<emphasis id="ED87EA99F1336E5ACC058EF9FB89FA11" bold="true" box="[1043,1050,1478,1492]" pageId="2" pageNumber="507">4</emphasis>
|
||
</th>
|
||
<th id="22125DB5F13391A7CC9B8E89FB3AFA11" box="[1165,1193,1462,1492]" gridcol="7" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5ACC9B8E89FB31FA0B" bold="true" box="[1165,1186,1462,1486]" pageId="2" pageNumber="507">ta</emphasis>
|
||
<emphasis id="ED87EA99F1336E5ACCB48EF9FB3AFA11" bold="true" box="[1186,1193,1478,1492]" pageId="2" pageNumber="507">5</emphasis>
|
||
</th>
|
||
<th id="22125DB5F13391A7CD0A8E89FADBFA11" box="[1308,1352,1462,1492]" gridcol="8" gridrow="0" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5ACD0A8E89FAACFA0B" bold="true" box="[1308,1343,1462,1486]" pageId="2" pageNumber="507">LR</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="61C334C9F13391A7C8B08EE1FADBFA39" box="[166,1352,1502,1532]" gridrow="1" pageId="2" pageNumber="507">
|
||
<th id="22125DB5F13391A7C8B08EE1FF28FA39" box="[166,187,1502,1532]" gridcol="0" gridrow="1" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC8B08EE0FF27FA32" bold="true" box="[166,180,1503,1527]" pageId="2" pageNumber="507">p</emphasis>
|
||
<emphasis id="ED87EA99F1336E5AC8A28ED1FF28FA39" bold="true" box="[180,187,1518,1532]" pageId="2" pageNumber="507">1</emphasis>
|
||
</th>
|
||
<td id="22125DB5F13391A7C9228EE1FEC9FA39" box="[308,346,1502,1532]" gridcol="1" gridrow="1" pageId="2" pageNumber="507">445</td>
|
||
<td id="22125DB5F13391A7C9D58EE1FE7AFA39" box="[451,489,1502,1532]" gridcol="2" gridrow="1" pageId="2" pageNumber="507">250</td>
|
||
<td id="22125DB5F13391A7CA448EE1FDEBFA39" box="[594,632,1502,1532]" gridcol="3" gridrow="1" pageId="2" pageNumber="507">-</td>
|
||
<td id="22125DB5F13391A7CAF78EE1FC94FA39" box="[737,775,1502,1532]" gridcol="4" gridrow="1" pageId="2" pageNumber="507">-</td>
|
||
<td id="22125DB5F13391A7CB798EE1FC06FA39" box="[879,917,1502,1532]" gridcol="5" gridrow="1" pageId="2" pageNumber="507">-</td>
|
||
<td id="22125DB5F13391A7CBE88EE1FB89FA39" box="[1022,1050,1502,1532]" gridcol="6" gridrow="1" pageId="2" pageNumber="507">-</td>
|
||
<td id="22125DB5F13391A7CC9B8EE1FB3AFA39" box="[1165,1193,1502,1532]" gridcol="7" gridrow="1" pageId="2" pageNumber="507">-</td>
|
||
<td id="22125DB5F13391A7CD0A8EE1FADBFA39" box="[1308,1352,1502,1532]" gridcol="8" gridrow="1" pageId="2" pageNumber="507">-</td>
|
||
</tr>
|
||
<tr id="61C334C9F13391A7C8B08D38FADBF9E1" box="[166,1352,1543,1572]" gridrow="2" pageId="2" pageNumber="507">
|
||
<th id="22125DB5F13391A7C8B08D38FF28F9E1" box="[166,187,1543,1572]" gridcol="0" gridrow="2" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC8B08D38FF27F9DA" bold="true" box="[166,180,1543,1567]" pageId="2" pageNumber="507">p</emphasis>
|
||
<emphasis id="ED87EA99F1336E5AC8A28D29FF28F9E1" bold="true" box="[180,187,1558,1572]" pageId="2" pageNumber="507">2</emphasis>
|
||
</th>
|
||
<td id="22125DB5F13391A7C9228D38FEC9F9E1" box="[308,346,1543,1572]" gridcol="1" gridrow="2" pageId="2" pageNumber="507">490</td>
|
||
<td id="22125DB5F13391A7C9D58D38FE7AF9E1" box="[451,489,1543,1572]" gridcol="2" gridrow="2" pageId="2" pageNumber="507">360</td>
|
||
<td id="22125DB5F13391A7CA448D38FDEBF9E1" box="[594,632,1543,1572]" gridcol="3" gridrow="2" pageId="2" pageNumber="507">230</td>
|
||
<td id="22125DB5F13391A7CAF78D38FC94F9E1" box="[737,775,1543,1572]" gridcol="4" gridrow="2" pageId="2" pageNumber="507">115</td>
|
||
<td id="22125DB5F13391A7CB798D38FC06F9E1" box="[879,917,1543,1572]" gridcol="5" gridrow="2" pageId="2" pageNumber="507">90</td>
|
||
<td id="22125DB5F13391A7CBE88D38FB89F9E1" box="[1022,1050,1543,1572]" gridcol="6" gridrow="2" pageId="2" pageNumber="507">60</td>
|
||
<td id="22125DB5F13391A7CC9B8D38FB3AF9E1" box="[1165,1193,1543,1572]" gridcol="7" gridrow="2" pageId="2" pageNumber="507">45</td>
|
||
<td id="22125DB5F13391A7CD0A8D38FADBF9E1" box="[1308,1352,1543,1572]" gridcol="8" gridrow="2" pageId="2" pageNumber="507">0.64</td>
|
||
</tr>
|
||
<tr id="61C334C9F13391A7C8B08D10FADBF988" box="[166,1352,1583,1613]" gridrow="3" pageId="2" pageNumber="507">
|
||
<th id="22125DB5F13391A7C8B08D10FF28F988" box="[166,187,1583,1613]" gridcol="0" gridrow="3" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC8B08D10FF27F982" bold="true" box="[166,180,1583,1607]" pageId="2" pageNumber="507">p</emphasis>
|
||
<emphasis id="ED87EA99F1336E5AC8A28D00FF28F988" bold="true" box="[180,187,1599,1613]" pageId="2" pageNumber="507">3</emphasis>
|
||
</th>
|
||
<td id="22125DB5F13391A7C9228D10FEC9F988" box="[308,346,1583,1613]" gridcol="1" gridrow="3" pageId="2" pageNumber="507">475</td>
|
||
<td id="22125DB5F13391A7C9D58D10FE7AF988" box="[451,489,1583,1613]" gridcol="2" gridrow="3" pageId="2" pageNumber="507">415</td>
|
||
<td id="22125DB5F13391A7CA448D10FDEBF988" box="[594,632,1583,1613]" gridcol="3" gridrow="3" pageId="2" pageNumber="507">295</td>
|
||
<td id="22125DB5F13391A7CAF78D10FC94F988" box="[737,775,1583,1613]" gridcol="4" gridrow="3" pageId="2" pageNumber="507">155</td>
|
||
<td id="22125DB5F13391A7CB798D10FC06F988" box="[879,917,1583,1613]" gridcol="5" gridrow="3" pageId="2" pageNumber="507">135</td>
|
||
<td id="22125DB5F13391A7CBE88D10FB89F988" box="[1022,1050,1583,1613]" gridcol="6" gridrow="3" pageId="2" pageNumber="507">90</td>
|
||
<td id="22125DB5F13391A7CC9B8D10FB3AF988" box="[1165,1193,1583,1613]" gridcol="7" gridrow="3" pageId="2" pageNumber="507">50</td>
|
||
<td id="22125DB5F13391A7CD0A8D10FADBF988" box="[1308,1352,1583,1613]" gridcol="8" gridrow="3" pageId="2" pageNumber="507">0.71</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="DF4C368BF1336E5AC8D18DA3FD40F83C" blockId="2.[151,1437,1691,2041]" pageId="2" pageNumber="507">
|
||
<emphasis id="ED87EA99F1336E5AC8D18DA3FEC5F970" box="[199,342,1692,1717]" italics="true" pageId="2" pageNumber="507">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="47C82A0EF1336E5AC9708DA4FE3BF970" box="[358,424,1691,1717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1912]" captionTargetBox="[158,1430,181,1858]" captionTargetId="figure-27@6.[158,1430,181,1864]" captionTargetPageId="6" captionText="FIGURE 4. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A–C hypopygium in dorsal aspect, stacked at different depth levels (A, B) and drawn (C), D, E median volsella (magnified c. 2 times relative to hypopygium photo). Scale bars: 50 μm. MS—median setae, AnP—anal point, SVo—superior volsella, IVo—inferior volsella, Gc—gonocoxite, Gs—gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4447199" httpUri="https://zenodo.org/record/4447199/files/figure.png" pageId="2" pageNumber="507">Fig. 4</figureCitation>
|
||
A–E). Gonostylus short, ~65 μm, slightly shorter than gonocoxite, gently tapering to blunt apex armed with both strong setae and microtrichia. Anal tergite subtriangular, with 5–6 posterolateral setae on each side of anal point, several irregularly dispersed fine median setae present. Anal point long and stout, reaching posterior margin of superior volsellae apices, tapering to blunt apex, with strong crests framing 21 irregularly dispersed spinulae (
|
||
<figureCitation id="47C82A0EF1336E5AC9608C14FE61F880" box="[374,498,1835,1861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1912]" captionTargetBox="[158,1430,181,1858]" captionTargetId="figure-27@6.[158,1430,181,1864]" captionTargetPageId="6" captionText="FIGURE 4. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A–C hypopygium in dorsal aspect, stacked at different depth levels (A, B) and drawn (C), D, E median volsella (magnified c. 2 times relative to hypopygium photo). Scale bars: 50 μm. MS—median setae, AnP—anal point, SVo—superior volsella, IVo—inferior volsella, Gc—gonocoxite, Gs—gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4447199" httpUri="https://zenodo.org/record/4447199/files/figure.png" pageId="2" pageNumber="507">Fig. 4A, C</figureCitation>
|
||
). Superior volsella elongate, peanut-shaped, apex with protuberances bearing up to 4 setae. Digitus not observed. Stem of median volsella long (~30 μm), more or less cylindrical with 2 subtle conical processes on apex, bearing two setiform lamellae on the proximal part and 5 to 6 foliate lamellae, distal lamella with elongated filiform tip reaching apex of inferior volsella (
|
||
<figureCitation id="47C82A0EF1336E5ACB558CA8FC48F874" box="[835,987,1943,1969]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1912]" captionTargetBox="[158,1430,181,1858]" captionTargetId="figure-27@6.[158,1430,181,1864]" captionTargetPageId="6" captionText="FIGURE 4. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A–C hypopygium in dorsal aspect, stacked at different depth levels (A, B) and drawn (C), D, E median volsella (magnified c. 2 times relative to hypopygium photo). Scale bars: 50 μm. MS—median setae, AnP—anal point, SVo—superior volsella, IVo—inferior volsella, Gc—gonocoxite, Gs—gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4447199" httpUri="https://zenodo.org/record/4447199/files/figure.png" pageId="2" pageNumber="507">Fig. 4B, D, E</figureCitation>
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). Inferior volsella reaching 2/3 length of gonostylus, with spherical bulge bearing 4 medially directed distinct setae at 1/3 of its length, with apical part bent medially, bearing several strong setae (
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<figureCitation id="47C82A0EF1336E5ACA5D8CE0FD1EF83C" box="[587,653,2015,2041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1888,1912]" captionTargetBox="[158,1430,181,1858]" captionTargetId="figure-27@6.[158,1430,181,1864]" captionTargetPageId="6" captionText="FIGURE 4. Stempellinella gilkai sp. nov., holotype, adult male (F28a LSZ DIZP, Eocene Baltic amber, ~40–45 Ma): A–C hypopygium in dorsal aspect, stacked at different depth levels (A, B) and drawn (C), D, E median volsella (magnified c. 2 times relative to hypopygium photo). Scale bars: 50 μm. MS—median setae, AnP—anal point, SVo—superior volsella, IVo—inferior volsella, Gc—gonocoxite, Gs—gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4447199" httpUri="https://zenodo.org/record/4447199/files/figure.png" pageId="2" pageNumber="507">Fig. 4</figureCitation>
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A–C).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |