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<document id="DEB1B985A6A9B269E77D62624FF4545B" ID-DOI="10.3897/zookeys.1210.123986" ID-ZooBank="6972216E-2356-4DF7-BD24-2B6B68E639B4" ID-publisher-id="123986" URI-arpha="B6184626-6862-59C4-9047-EF9B7AC24E35" URI-zoobank="http://zoobank.org/6972216E-2356-4DF7-BD24-2B6B68E639B4" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1723830749198" checkinUser="pensoft" docAuthor="Xu, Xin, Zhan, Yi, Aung, Khin Pyae Pyae, Liu, Fengxiang, Yu, Long &amp; Li, Daiqin" docDate="2024" docId="1148FBC405EA553C96A0DBAFAAE5E4F3" docLanguage="en" docName="ZooKeys 1210: 99-115" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/123986/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Liphistius nawngau Zhan &amp; Xu 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="115" masterDocId="B6184626686259C49047EF9B7AC24E35" masterDocTitle="Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar" masterLastPageNumber="115" masterPageNumber="99" pageNumber="99" updateTime="1723832703504" updateUser="admin">
<mods:mods id="8F0F8C53C2BA037DBBE9EDE74426AA3A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="90E1ED2BB1661916B454F2A1B4BF7D06">
<mods:title id="BD0A73D762C5B6AD6D693CB10E62B2C0">Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar</mods:title>
</mods:titleInfo>
<mods:name id="A861F20157389300FCEDED7B7310B7DF" type="personal">
<mods:role id="55C8427D0BD3BF27B0BE10C0478B6A1B">
<mods:roleTerm id="B4C8E65269CE4FD1DD3139052593E545">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9BCA59A80E8FFDE7019C5C7179649B93">Xu, Xin</mods:namePart>
<mods:nameIdentifier id="F6467ADA8D53FE5EE6F25FA3CFE4113A" type="ORCID">0000-0001-5632-6622</mods:nameIdentifier>
<mods:affiliation id="D084964F3E7EE2F721C0381B4261094D">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
<mods:name id="C3CBA9DFCC21F0298DF154417BDDF022" type="personal">
<mods:role id="07113ADA6E42F688F02403268B33D5C5">
<mods:roleTerm id="118CAA0EB4E1684EE57DEA8514A35934">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4DBE4942D0DD15EFB04135A27C8C7B6B">Zhan, Yi</mods:namePart>
<mods:affiliation id="D4D0961F8825B6CE3C9E0C0D545F6F83">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
</mods:name>
<mods:name id="8B62E67464DE2582EE6E0B68AA162360" type="personal">
<mods:role id="4219823D3A6F013F65E0F3B1D9B36AB4">
<mods:roleTerm id="AD949FFCCE8A38586E4B71BA279C6C5B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="229DF82E4FB7D9E4B7A2F7C3B9D73050">Aung, Khin Pyae Pyae</mods:namePart>
<mods:affiliation id="04ACACF345F16EF9D040E0687D69F2B5">Department of Zoology, University of Yangon, Kamayut Township, Pyay Road, Yangon, 11041, Myanmar &amp; Department of Biology, Taungoo Education Degree College, Taungoo, 08101, Myanmar</mods:affiliation>
</mods:name>
<mods:name id="BE1A1341D55706221209AC5624782EC3" type="personal">
<mods:role id="EAA083297E000DDEDB190779B0DA2A8F">
<mods:roleTerm id="15F4B505FF781562777D668FEE412AF5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="11AB59A2A97D8CE996AB7A5CD6255D25">Liu, Fengxiang</mods:namePart>
<mods:affiliation id="6EAB8BBA2EEBA94E04BC69ED171DF190">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
<mods:name id="17D371480303A0B3A494C196E051DA43" type="personal">
<mods:role id="64AAE39064779078558348A5A1291CE3">
<mods:roleTerm id="3C3B3CEB7CE47EC5840B7BF8A469AEDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09647BA466A01462A2CB3FF04D5F606F">Yu, Long</mods:namePart>
<mods:nameIdentifier id="534EC1807E0DDDB2ADC9785B44D871C3" type="ORCID">0000-0001-5675-1864</mods:nameIdentifier>
<mods:affiliation id="C5C6D7BF1EDA1BEE2795996F12F00B12">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
<mods:name id="56B8C7A6D0D27218B4FEF0DA7A04F794" type="personal">
<mods:role id="FEFEC7E5F337032A99525A866655B0DC">
<mods:roleTerm id="2E3DB77028B7E981DA6550986D1AFEAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA971E9621864989C0D9CAC1A5172D84">Li, Daiqin</mods:namePart>
<mods:nameIdentifier id="8E56376FE427A42ABE602CFF1A04245F" type="ORCID">0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation id="BEE5CEE560E7777B80F19A3C3B3F7080">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China &amp; Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543</mods:affiliation>
</mods:name>
<mods:typeOfResource id="09B1E803EF22AEB21F305B926DA4B432">text</mods:typeOfResource>
<mods:relatedItem id="ECCDEB424ECFDC6AD775B10D578AC3C7" type="host">
<mods:titleInfo id="B278D3F3EBE0BCA2D0414D349537225D">
<mods:title id="CFD63E1A6C87061AA604F0B443487E2E">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="E8786F42066324E7578417FA33ED4256">
<mods:date id="A1987529B21A126F4F4B7090B2EB5274">2024</mods:date>
<mods:detail id="31103D4A18022124DDE4D88773BD8A88" type="pubDate">
<mods:number id="3D78189DFBB93C9975DFFE55A457ABE7">2024-08-16</mods:number>
</mods:detail>
<mods:detail id="35FB7722FFE9D0F05FCB53DC2660F635" type="volume">
<mods:number id="C1BEC6FAD690469C69AFEB889FD8C9B6">1210</mods:number>
</mods:detail>
<mods:extent id="47F24B3E5584C1F64AB802BC6B8E566F" unit="page">
<mods:start id="DAF76EBD7213A041110770FF70D689DF">99</mods:start>
<mods:end id="02148F5BD9EBFDF4F4F4EF8E91642344">115</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F6B57EC7BABE6C735A273164AB3814C8">journal article</mods:classification>
<mods:identifier id="EB34FDA84D38DFA49C4E0FBE24051DFD" type="DOI">10.3897/zookeys.1210.123986</mods:identifier>
<mods:identifier id="DA9B468EE348C5EC826DA3BD927DDA79" type="ZooBank">6972216E-2356-4DF7-BD24-2B6B68E639B4</mods:identifier>
</mods:mods>
<treatment id="1148FBC405EA553C96A0DBAFAAE5E4F3" ID-DOI="http://doi.org/10.5281/zenodo.13333482" ID-Zenodo-Dep="13333482" ID-arpha="1148FBC4-05EA-553C-96A0-DBAFAAE5E4F3" ID-zoobank="https://zoobank.org/1ADFCE8F-66EE-43F9-B61A-927F6B88189F" LSID="urn:lsid:plazi:treatment:1148FBC405EA553C96A0DBAFAAE5E4F3" LSID-ZBK="urn:lsid:zoobank.org:act:1ADFCE8F-66EE-43F9-B61A-927F6B88189F" httpUri="http://treatment.plazi.org/id/1148FBC405EA553C96A0DBAFAAE5E4F3">
<subSubSection id="91EC8776201CB8D09910ABB853F27E75" type="nomenclature">
<paragraph id="15085F835D034EA16CF6B316CD414D3B">
<taxonomicName id="351F29596982D3F6CCCA2BAA2C8513BA" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="984167406BB6BA6CBBAF12D4335FF7EF" italics="true">Liphistius nawngau</emphasis>
Zhan &amp; Xu
</taxonomicName>
<taxonomicNameLabel id="8C1F53FB7EC2EA139188E081E7B6A350" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="65E8133EBF945D84B4A72FC1051B2879" type="description">
<paragraph id="B7BDADFCD483D9FBD47B9DC65CCC03D0">
<figureCitation id="5B9D23331179DB5F3A2EAA46ECA6A39A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">Figs 7</figureCitation>
,
<figureCitation id="397A66BF49634B4074C959AB83F5FC88" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Vulva of Liphistius nawngau sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure8" httpUri="https://binary.pensoft.net/fig/1115134">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBXAG" type="type material">
<paragraph id="B2427413558002C8C651BAD42C6317D6">
<heading id="48808AE7FF11DA07348FEBBDF736175D" reason="title">Type material.</heading>
</paragraph>
<paragraph id="C27AC64F39B6B2D8552C857AEF9C94DC">
<materialsCitation id="98B02842AC81616341CF770C7D8F188B" collectingDate="2019-07-26" collectionCode="XUX" collectorName="D. Li" country="Myanmar" county="Kyaukme Dist." elevation="1096" latitude="22.26" location="Nawng Au Vill." longLatPrecision="756" longitude="96.83" municipality="Nawnghkio Township" specimenCode="XUX-2019-054A" specimenCount="1" specimenCount-male="1" stateProvince="Shan State" typeStatus="holotype">
<emphasis id="3967BE0C03BDAFD15E2F720F8C5BD23C" bold="true" italics="true">
<typeStatus id="8331C2623EA9CA57598E737FA09963D6" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="82C41F59AAFBB83F819C51ABA6B3976B" name="Myanmar">Myanmar</collectingCountry>
<specimenCount id="27F1C66CB74841F134A89DB2F205F2DB" count="1" type="male">♂</specimenCount>
,
<collectingRegion id="7FC831130EB4E3B05E9C83A881787D77" country="Myanmar" name="Shan">Shan State</collectingRegion>
,
<collectingCounty id="3DC684B598ACBFB6F43210EEA13F97DF">Kyaukme Dist.</collectingCounty>
,
<collectingMunicipality id="25DE4FCE47AD34C6DA14F4C7F2A3D788">Nawnghkio Township</collectingMunicipality>
,
<location id="9C115DF75981A5C3823413ACB7C6F46F" LSID="urn:lsid:plazi:treatment:1148FBC405EA553C96A0DBAFAAE5E4F3:9C115DF75981A5C3823413ACB7C6F46F" country="Myanmar" county="Kyaukme Dist." latitude="22.26" longLatPrecision="756" longitude="96.83" municipality="Nawnghkio Township" name="Nawng Au Vill." stateProvince="Shan State">Nawng Au Vill.</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPXAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.830000,22.260000]}">
<geoCoordinate id="227AE6416CDD76731B1A80B6F22A2689" degrees="22.26" direction="north" orientation="latitude" precision="555" value="22.26">22.26 ° N</geoCoordinate>
,
<geoCoordinate id="CB3FAF6ED323CD62361EFC8C78E458D9" degrees="96.83" direction="east" orientation="longitude" precision="555" value="96.83">96.83 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="B79E0E489DE4EE3B6902F0109307F486" metricMagnitude="3" metricUnit="m" metricValue="1.0959999999999999" unit="m" value="1096.0">
<elevation id="300F25DAF517E7FE4B5477CCD530D8B4" metricMagnitude="3" metricUnit="m" metricValue="1.0959999999999999" unit="m" value="1096.0">1096 m</elevation>
</quantity>
;
<date id="D22521C200B8D6B04DDC3D16955576EE" value="2019-07-26">
<collectingDate id="890F7379B699B3BD75B5198C9D009A52" value="2019-07-26">26. 07. 2019</collectingDate>
</date>
;
<collectorName id="4C09A07E9F5C76BD1EDBB0DA053C28B6">D. Li</collectorName>
et al. leg.;
<specimenCode id="BF8B0035A30FB5323E1714092EA06F77">XUX-2019-054A</specimenCode>
</materialsCitation>
.
<materialsCitation id="291A6CA20E91EA2A6741433BCC693983" collectingDate="2019-07-26" collectionCode="XUX" collectorName="D. Li" country="Myanmar" county="Kyaukme Dist." elevation="1096" latitude="22.26" location="Nawng Au Vill." longLatPrecision="756" longitude="96.83" municipality="Nawnghkio Township" specimenCode="XUX-2019-054" specimenCount="12" specimenCount-female="3" specimenCount-male="9" stateProvince="Shan State" typeStatus="paratype">
<emphasis id="87D2C1ECEE3C6D2433D9B2F5630AAB87" bold="true" italics="true">
<typeStatus id="5175CA05864765D837E6BEB4200C6377" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="12782989DF2711F719CDA6182C0E9C6A" count="9" type="male">9 ♂♂</specimenCount>
<specimenCount id="BE40E20D3F6262F735F43D209513ADC8" count="3" type="female">3 ♀♀</specimenCount>
, same data as for holotype;
<specimenCode id="400C9A824AA10B4C5286CE53637695F8">XUX-2019-054</specimenCode>
/ 054 A / 055 / 055 A / 055 B / 055 D / 056 / 056 A / 057 / 058 / 059 / 060
</materialsCitation>
;
<materialsCitation id="6EC957B7BDC930E46B93A5A650B5A196" collectingDate="2018-07-14" collectingDateMax="2019-07-26" collectingDateMin="2018-07-14" collectionCode="XUX" collectorName="D. Li" country="Myanmar" county="Kyaukme Dist." elevation="845" latitude="22.3" location="Nawng Au Vill." longLatPrecision="756" longitude="96.73" municipality="Nawnghkio Township" specimenCode="XUX-2019-049, XUX-2018-116" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Shan State">
<specimenCount id="959E3C847DB5BEA621CC4D9D5064B946" count="5" type="male">5 ♂♂</specimenCount>
<specimenCount id="786D548C5965EFE48F6501A7C8826BCB" count="2" type="female">2 ♀♀</specimenCount>
, same township as for holotype;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1XAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.730000,22.300000]}">
<geoCoordinate id="C079624369EC4049B17118E4C2D0C485" degrees="22.30" direction="north" orientation="latitude" precision="555" value="22.3">22.30 ° N</geoCoordinate>
,
<geoCoordinate id="1D2DAD04DEC1FB2727F417733318C787" degrees="96.73" direction="east" orientation="longitude" precision="555" value="96.73">96.73 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="900D0F6B1816747D688313A62529FD47" metricMagnitude="2" metricUnit="m" metricValue="8.45" unit="m" value="845.0">
<elevation id="8B6176691117F4B726E0CA060443133A" metricMagnitude="2" metricUnit="m" metricValue="8.45" unit="m" value="845.0">845 m</elevation>
</quantity>
;
<date id="60360393FD5A2E3A868C1276994DE804" value="2019-07-26">
<collectingDate id="D20563F0558F2BF27524F30C432DCE1A" value="2019-07-26">26. 07. 2019</collectingDate>
</date>
and
<date id="8E9A96EF38BE6748B2DDE64053394C26" value="2018-07-14">
<collectingDate id="B23504197B4CC73D98ADA038ACD505FC" value="2018-07-14">14. 07. 2018</collectingDate>
</date>
;
<specimenCode id="4F96082BA20986567356BA0DA47837E5">XUX-2019-049</specimenCode>
/ 051 / 052 / 053,
<specimenCode id="AED5DC82D96615C3AA6B07E971689E16">XUX-2018-116</specimenCode>
/ 119 / 123
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6XAG" type="diagnosis">
<paragraph id="E622171C10E234893C37A94C1A290FDE">
<heading id="677442942DABB5F6313A398673E134AB" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="CB6CF9E6CA03FADAA9C74D7940C7D77F">
The male of
<taxonomicName id="E48D430159D7DD8EF7C07AF787EB8872" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="AED3D523C2716F33A17BCBA4DC36B3E1" italics="true">L. nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3247E53B2A878AEBC50484E3555579B1" rank="species">sp. nov.</taxonomicNameLabel>
resembles those of
<taxonomicName id="95E18D322FE590564A4D40AB6528D48D" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lordae">
<emphasis id="1BA94450FE7C24C73739B5334C2146AC" italics="true">L. lordae</emphasis>
</taxonomicName>
and
<taxonomicName id="38A60B1481E877F408DD8CA58114A90E" authorityName="Xu, Yu, Aung, Yu, Liu, Lwin, Sang &amp; Li" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="pyinoolwin">
<emphasis id="6C1D4AF212F839CAA5510679372CB83D" italics="true">L. pyinoolwin</emphasis>
</taxonomicName>
in having an adpressed proximal tegular margin (Fig.
<figureCitation id="36C500FC34254F0DC9C68E74987F9DFC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 C, F</figureCitation>
), but it can be distinguished from them by the tegulum (
<collectionCode id="7601C6A74046EE1B7E71F994AAE9318B" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
), which bears a distinct transversal ridge in retrolateral view (Fig.
<figureCitation id="E2F08F526CE359A53490D70DB6621A00" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 C, F</figureCitation>
vs lacking transversal ridge, figs 5,
<quantity id="A69941A57D68D5155096EE39CF535A3A" metricMagnitude="-1" metricUnit="m" metricValue="1.524" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="D0C44E1564AECCBB7B6D52C0FE3099A1" DOI="10.1111/j.1463-6409.1990.tb00262.x" author="Schwendinger" firstAuthor="Schwendinger" issue="3" journalOrPublisher="Zoologica Scripta" pagination="331-351" refId="B2" refString="Schwendinger PJ (1990) On the spider genus Liphistius (Araneae: Mesothelae) in Thailand and Burma. Zoologica Scripta 19 (3): 331–351. https://doi.org/10.1111/j.1463-6409.1990.tb00262.x" title="On the spider genus Liphistius (Araneae: Mesothelae) in Thailand and Burma." volume="19" year="1990">Schwendinger 1990</bibRefCitation>
; fig. 4 F in
<bibRefCitation id="3C7DEF065A7EA75F5F1CACC4B1C030AB" DOI="10.3897/zookeys.1031.59102" author="Xu" etAl="et al." firstAuthor="Xu" journalOrPublisher="ZooKeys" pagination="41-58" refId="B9" refString="Xu X, Yu L, Aung KPP, Yu L, Liu FX, Lwin WW, Sang MZ, Li DQ (2021) A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae). ZooKeys 1031: 41–58. https://doi.org/10.3897/zookeys.1031.59102" title="A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae)." volume="1031" year="2021">Xu et al. 2021</bibRefCitation>
), and by the embolus (E), which has smooth longitudinal ridges reaching tip (Fig.
<figureCitation id="5E06C243BD665C5FFFADA78E7D9D432C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 E</figureCitation>
vs having several denticles along longitudinal ridges, fig. 4 F in
<bibRefCitation id="24DDC3B8F403C959EF46219AE197A54D" DOI="10.3897/zookeys.1031.59102" author="Xu" etAl="et al." firstAuthor="Xu" journalOrPublisher="ZooKeys" pagination="41-58" refId="B9" refString="Xu X, Yu L, Aung KPP, Yu L, Liu FX, Lwin WW, Sang MZ, Li DQ (2021) A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae). ZooKeys 1031: 41–58. https://doi.org/10.3897/zookeys.1031.59102" title="A new species of Liphistius from Myanmar and description of the actual male of L. birmanicus Thorell, 1897 (Araneae, Mesothelae, Liphistiidae)." volume="1031" year="2021">Xu et al. 2021</bibRefCitation>
). The female of
<taxonomicName id="AC42B1FF463303D449FAAA9E14DF69F6" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="A2EEF69F4593FF8C909CCDE112AFAA60" italics="true">L. nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="220D031B3544CD812EEBCB3788818077" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from that of
<taxonomicName id="C70E2A240CFB13E5123CC40649A301DC" authorityName="Aung, Xu, Lwin, Sang, Yu, Liu, Liu &amp; Li" authorityYear="2019" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="hpruso">
<emphasis id="92D66A463D100963F8DF9C88A7999499" italics="true">L. hpruso</emphasis>
</taxonomicName>
by having the posterior stalk triangular (
<collectionCode id="30DC087311331A2E87FBFE68C86B0ACC">PS</collectionCode>
) (Fig.
<figureCitation id="1CB69AACBC82AEFCFE7D9CD1E0653C07" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Vulva of Liphistius nawngau sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure8" httpUri="https://binary.pensoft.net/fig/1115134">8</figureCitation>
vs nearly oval, fig. 3 B – E in
<bibRefCitation id="7572601299EA0288AC6D5DE38590CBD8" DOI="10.3897/zookeys.882.38811" author="Aung" etAl="et al." firstAuthor="Aung" journalOrPublisher="ZooKeys" pagination="29-39" refId="B1" refString="Aung KPP, Xu X, Lwin WW, Sang MZ, Yu L, Liu H, Liu FX, Li DQ (2019) Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar. ZooKeys 882: 29–39. https://doi.org/10.3897/zookeys.882.38811" title="Two new species of the primitively segmented spider genus Liphistius Schiödte, 1849 (Mesothelae, Liphistiidae) from Myanmar." volume="882" year="2019">Aung et al. 2019</bibRefCitation>
).
</paragraph>
<caption id="55EC15F6C8177BCCB377C0D6AD86E5A4" ID-DOI="10.3897/zookeys.1210.123986.figure7" ID-arpha="167E0A7B-A8FC-5F8B-93C9-EF4CCFD53C17" httpUri="https://binary.pensoft.net/fig/1115133" startId="F7">
<paragraph id="B5EF217E0AC9A38DC1987D435AF81FAB">
<label id="13B70BADA264291994BC80504B5A733D">Figure 7.</label>
</paragraph>
<paragraph id="BBE9FC7E8BEA3D8AA3724CEAC8E7846B">
Male palp of
<taxonomicName id="02DA1C254D20CFA7D986BA823CE081DA" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="F37E9C261CF90F98C7ACD5CFB9DE5733" italics="true">Liphistius nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CBF0B573733A03D14C8DFB7FB01BDD0B" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="E2E9A079D9465B7B1D81DDE4A5BFCC25" bold="true">A, E</emphasis>
prolateral view
<emphasis id="FAC31AE70C43CA98ECFFF512372EAFF3" bold="true">B</emphasis>
ventral view
<emphasis id="8B867D17AD762AC8515C2344A478E31D" bold="true">C, F</emphasis>
retrolateral view
<emphasis id="F73C6918063F1E5C5F4BB1165B449D04" bold="true">D</emphasis>
dorsal view, red arrows show arched projection and triangular process of
<abbrev id="ABBRID0EX2AG" xlink_title="contrategulum">CT</abbrev>
<emphasis id="96B6E710A9D594DBC9C5E9FE33D7ABFC" bold="true">G</emphasis>
distal view. Abbreviations:
<abbrev id="ABBRID0E42AG" xlink_title="contrategulum">CT</abbrev>
= contrategulum;
<abbrev id="ABBRID0EB3AG" xlink_title="contrategular process">cp</abbrev>
= contrategular process;
<abbrev id="ABBRID0EF3AG" xlink_title="cumulus">Cu</abbrev>
= cumulus; E = embolus;
<abbrev id="ABBRID0EJ3AG" xlink_title="paracymbium">PC</abbrev>
= paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum;
<abbrev id="ABBRID0EN3AG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
= tibial apophysis. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="F9500FD4FA1C399A329CE9FB808ADE7B" ID-DOI="10.3897/zookeys.1210.123986.figure8" ID-arpha="B49B9831-E68E-5884-9F01-491C46F8CA97" httpUri="https://binary.pensoft.net/fig/1115134" startId="F8">
<paragraph id="FCBA55ACD07F3BE5B4DF5500AFCE3303">
<label id="D545C0EAE4F0ABE0728345C99AB16C26">Figure 8.</label>
</paragraph>
<paragraph id="7681864D046949A8BE6D09EFE17A4666">
Vulva of
<taxonomicName id="DFAB499436899E303DBA0091F40B57F1" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau" status="sp. nov.">
<emphasis id="204F60E36FF289A6F6F5C7D826D7BDB6" italics="true">Liphistius nawngau</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54FCABF6559B170A5177DBB1F12BD840" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="9D24A29FE825A83C0A3F7C78D1242781" bold="true">A, C</emphasis>
dorsal view
<emphasis id="F7C1B150387C93F12F7F37A7E9058B1C" bold="true">B, D</emphasis>
ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk;
<abbrev id="ABBRID0EN4AG" xlink_title="sclerotized poreplate">PPl</abbrev>
= poreplate;
<abbrev id="ABBRID0ER4AG" xlink_title="receptacular cluster">RC</abbrev>
= receptacular cluster. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E14AG" type="description">
<paragraph id="EC0B07C0424E6B7621A12D815D456854">
<heading id="F0AA6AE3F9BD974655C66A4EC08B06DE" reason="title">Description.</heading>
</paragraph>
<paragraph id="A43F56A996BB9D44E3015B5CD6D01863">
<emphasis id="5071D814A174CD7580E569BC16C6113D" bold="true">
Male
<typeStatus id="536D7EDCE0B7ECE5835A92691638218B" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 2–6
<superScript id="C26C15B5BD5E388F4C454A6E67953ADE">th</superScript>
larger than others, 5
<superScript id="00ED626ECA47D52B3D332A0429DB72EB">th</superScript>
largest; chelicerae robust, promargin of cheliceral groove with 11 denticles of variable size; labium and sternum yellowish brown, sternum with few short setae on anterior tip and many long setae on elongated posterior tip; legs yellowish brown, with strong setae and spines. Measurements:
<abbrev id="ABBRID0EG5AG" xlink_title="body length">BL</abbrev>
15.47,
<abbrev id="ABBRID0EK5AG" xlink_title="carapace length">CL</abbrev>
6.63,
<abbrev id="ABBRID0EO5AG" xlink_title="carapace width">CW</abbrev>
6.57,
<abbrev id="ABBRID0ES5AG" xlink_title="opisthosoma length">OL</abbrev>
7.73,
<abbrev id="ABBRID0EW5AG" xlink_title="opisthosoma width">OW</abbrev>
5.56; eye sizes and interdistances:
<abbrev id="ABBRID0E15AG" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0E55AG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.72,
<abbrev id="ABBRID0EC6AG" xlink_title="posterior median eyes">PME</abbrev>
0.35,
<abbrev id="ABBRID0EG6AG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.49,
<abbrev id="ABBRID0EK6AG" xlink_title="anterior median eyes">AME</abbrev>
–
<abbrev id="ABBRID0EO6AG" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0ES6AG" xlink_title="anterior median eyes">AME</abbrev>
–
<abbrev id="ABBRID0EW6AG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.20,
<abbrev id="ABBRID0E16AG" xlink_title="posterior median eyes">PME</abbrev>
–
<abbrev id="ABBRID0E56AG" xlink_title="posterior median eyes">PME</abbrev>
0.08,
<abbrev id="ABBRID0ECABG" xlink_title="posterior median eyes">PME</abbrev>
–
<abbrev id="ABBRID0EGABG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.07,
<abbrev id="ABBRID0EKABG" xlink_title="anterior lateral eyes">ALE</abbrev>
–
<abbrev id="ABBRID0EOABG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.81,
<abbrev id="ABBRID0ESABG" xlink_title="anterior lateral eyes">ALE</abbrev>
–
<abbrev id="ABBRID0EWABG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.11,
<abbrev id="ABBRID0E1ABG" xlink_title="posterior lateral eyes">PLE</abbrev>
–
<abbrev id="ABBRID0E5ABG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.16,
<abbrev id="ABBRID0ECBBG" xlink_title="anterior median eyes">AME</abbrev>
–
<abbrev id="ABBRID0EGBBG" xlink_title="posterior median eyes">PME</abbrev>
0.12; labium 0.81 long and 0.58 wide; sternum 2.59 long and 1.01 wide; legs: I 19.58 (5.67, 2.78, 4.03, 4.68, 2.42), II 20.42 (5.66, 2.78, 4.13, 4.96, 2.89), III 22.62 (5.96, 2.63, 4.54, 6.29, 3.20), IV 28.26 (7.16, 2.75, 5.82, 8.55, 3.98).
</paragraph>
<paragraph id="13DEF346FCB55E63AA0F505965C689DE">
Palp: tibial apophysis (
<abbrev id="ABBRID0EMBBG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
) with four setae of similar length (Fig.
<figureCitation id="18F4A67B9B7504651D213AD9E8020364" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A – C</figureCitation>
); paracymbium (
<abbrev id="ABBRID0EUBBG" xlink_title="paracymbium">PC</abbrev>
) with several setae situated on tip (Fig.
<figureCitation id="8D5B01052486702AA3B803E844EE00F1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A – C</figureCitation>
); cumulus (
<abbrev id="ABBRID0E3BBG" xlink_title="cumulus">Cu</abbrev>
) slightly elevated, with 8 tapering spines (Fig.
<figureCitation id="DAB077CD2E370910E8412BD3C22A9874" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A – C</figureCitation>
); subtegulum (ST) with weakly developed apophysis (Fig.
<figureCitation id="AB1EF823FC0F717952355E33E5B94DDF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 B, F</figureCitation>
); contrategulum (
<abbrev id="ABBRID0EICBG" xlink_title="contrategulum">CT</abbrev>
) with an arched projection distally, and triangular process with wide base (Fig.
<figureCitation id="F746B0CE3585B6394BF8A74A91C2ADD5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 D</figureCitation>
); tegulum (
<collectionCode id="D62A7A93BB629BC587F8E76B70679944" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
) with distinct transversal ridge in retrolateral view (Fig.
<figureCitation id="07C3CCC9B4432F82F6C5FFAF95037B28" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 C, F</figureCitation>
); paraembolic plate (PeP) narrow, sclerotized (Fig.
<figureCitation id="17A39763E185FE2B72C2430FA3F76076" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 A, F</figureCitation>
); embolus (E) with several longitudinal ridges reaching tip (Fig.
<figureCitation id="D637E3432CB30856623963395F26BD1F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male palp of Liphistius nawngau sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view, red arrows show arched projection and triangular process of CT G distal view. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure7" httpUri="https://binary.pensoft.net/fig/1115133">7 E, F</figureCitation>
).
</paragraph>
<paragraph id="F6BF0DB69BC88FEB5B9D6A6430F9FF46">
<emphasis id="4F9B2F36A76E9DE6AE1B7F8DAF7B2521" bold="true">
Female
<typeStatus id="27DCE62D2D9F0711C1B19C8B34FC31ED" type="paratype">paratype</typeStatus>
(
<specimenCode id="F5AD8278BA49F4E4F378753738839793">XUX- 2019-053</specimenCode>
).
</emphasis>
Carapace yellowish brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, with gray patches, 2–6
<superScript id="BC76F4729343AF25F0626CE05492E997">th</superScript>
larger than others, 5
<superScript id="A806F6080CD91B54E609DD60312A03CF">th</superScript>
largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 12 denticles of variable size; labium and sternum yellowish brown; legs with strong setae and spines. Measurements:
<abbrev id="ABBRID0EEDBG" xlink_title="body length">BL</abbrev>
14.24,
<abbrev id="ABBRID0EIDBG" xlink_title="carapace length">CL</abbrev>
6.10,
<abbrev id="ABBRID0EMDBG" xlink_title="carapace width">CW</abbrev>
5.30,
<abbrev id="ABBRID0EQDBG" xlink_title="opisthosoma length">OL</abbrev>
7.12,
<abbrev id="ABBRID0EUDBG" xlink_title="opisthosoma width">OW</abbrev>
4.96; eye sizes and interdistances:
<abbrev id="ABBRID0EYDBG" xlink_title="anterior median eyes">AME</abbrev>
0.08,
<abbrev id="ABBRID0E3DBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.57,
<abbrev id="ABBRID0EAEBG" xlink_title="posterior median eyes">PME</abbrev>
0.31,
<abbrev id="ABBRID0EEEBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.45,
<abbrev id="ABBRID0EIEBG" xlink_title="anterior median eyes">AME</abbrev>
–
<abbrev id="ABBRID0EMEBG" xlink_title="anterior median eyes">AME</abbrev>
0.10,
<abbrev id="ABBRID0EQEBG" xlink_title="anterior median eyes">AME</abbrev>
–
<abbrev id="ABBRID0EUEBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.17,
<abbrev id="ABBRID0EYEBG" xlink_title="posterior median eyes">PME</abbrev>
–
<abbrev id="ABBRID0E3EBG" xlink_title="posterior median eyes">PME</abbrev>
0.09,
<abbrev id="ABBRID0EAFBG" xlink_title="posterior median eyes">PME</abbrev>
–
<abbrev id="ABBRID0EEFBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.08,
<abbrev id="ABBRID0EIFBG" xlink_title="anterior lateral eyes">ALE</abbrev>
–
<abbrev id="ABBRID0EMFBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.09,
<abbrev id="ABBRID0EQFBG" xlink_title="anterior lateral eyes">ALE</abbrev>
–
<abbrev id="ABBRID0EUFBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.10,
<abbrev id="ABBRID0EYFBG" xlink_title="posterior lateral eyes">PLE</abbrev>
–
<abbrev id="ABBRID0E3FBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.46,
<abbrev id="ABBRID0EAGBG" xlink_title="anterior median eyes">AME</abbrev>
–
<abbrev id="ABBRID0EEGBG" xlink_title="posterior median eyes">PME</abbrev>
0.06; labium 1.23 long and 0.77 wide; sternum 2.68 long and 1.13 wide; palp 10.42 (3.63, 1.84, 2.77, 2.18), leg I 13.33 (4.16, 2.15, 2.90, 2.58, 1.54), II 16.85 (4.35, 2.26, 2.84, 2.84, 1.72), III 15.04 (4.37, 2.35, 2.90, 3.31, 2.11), IV 20.39 (5.92, 1.92, 4.48, 5.33, 2.74).
</paragraph>
<paragraph id="6FD506E4523891E20ABF022291F4FF64">
Vulva: poreplate (
<abbrev id="ABBRID0EKGBG" xlink_title="sclerotized poreplate">PPl</abbrev>
) with pair of large anterior lobes and pair of relatively small anterolateral lobes; central dorsal opening (
<collectionCode id="E46B961EB81896055B2684E7E78BFDEE">CDO</collectionCode>
) located at center of poreplate (
<abbrev id="ABBRID0EOGBG" xlink_title="sclerotized poreplate">PPl</abbrev>
); receptacular cluster (
<abbrev id="ABBRID0ESGBG" xlink_title="receptacular cluster">
<collectionCode id="87CBA80D545C8628F27374CFF471848D">RC</collectionCode>
</abbrev>
) racemose, protrudes upper edge of poreplate (
<abbrev id="ABBRID0EWGBG" xlink_title="sclerotized poreplate">PPl</abbrev>
); posterior margin of triangular posterior stalk (
<collectionCode id="3A30862C8C429FB335C0B1580DB7DC29">PS</collectionCode>
) almost straight; lateral margins of genital atrium (GA) slightly sclerotized (Fig.
<figureCitation id="C3FC2A0EE0676BD640A01CF5E8CB357C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Vulva of Liphistius nawngau sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure8" httpUri="https://binary.pensoft.net/fig/1115134">8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5GBG" type="variation">
<paragraph id="2B90B85B60376980224C958908C5A36A">
<heading id="068D6CC6F97FE8644086276C31A6E78E" reason="title">Variation.</heading>
</paragraph>
<paragraph id="84ACCB9E393456A9DA61EB3D0812AE94">
Males (
<emphasis id="BE05782BC2912D2287482EB475FA6084" italics="true">N</emphasis>
= 14):
<abbrev id="ABBRID0EGHBG" xlink_title="body length">BL</abbrev>
12.64–16.98,
<abbrev id="ABBRID0EKHBG" xlink_title="carapace length">CL</abbrev>
6.27–7.56,
<abbrev id="ABBRID0EOHBG" xlink_title="carapace width">CW</abbrev>
6.02–7.56,
<abbrev id="ABBRID0ESHBG" xlink_title="opisthosoma length">OL</abbrev>
6.95–8.50,
<abbrev id="ABBRID0EWHBG" xlink_title="opisthosoma width">OW</abbrev>
4.60–6.40; females (
<emphasis id="C6B7BEA17496CFB7EFDBDCC5EFA7F2D2" italics="true">N</emphasis>
= 5):
<abbrev id="ABBRID0E3HBG" xlink_title="body length">BL</abbrev>
10.55–14.24,
<abbrev id="ABBRID0EAIBG" xlink_title="carapace length">CL</abbrev>
5.26–6.37,
<abbrev id="ABBRID0EEIBG" xlink_title="carapace width">CW</abbrev>
4.47–5.63,
<abbrev id="ABBRID0EIIBG" xlink_title="opisthosoma length">OL</abbrev>
5.24–7.12,
<abbrev id="ABBRID0EMIBG" xlink_title="opisthosoma width">OW</abbrev>
4.33–4.96. The number of denticles on the promargin of male cheliceral groove varies from 10 to 13 (
<emphasis id="DA3EF92C33B33DB300E9323658E1FB7A" italics="true">N</emphasis>
= 12); in females, the number of denticles on the promargin of cheliceral groove varies from 11 to 12 (
<emphasis id="1D833BD5A78B28FC37ED503C44B73688" italics="true">N</emphasis>
= 5). The number of setae on tibial apophysis varies from 4 to 6.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUIBG" type="etymology">
<paragraph id="9547AE76BAB6288E82691D4CB49D6211">
<heading id="92856A1013A630F9BAF55F41E45B28F7" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="8B7EA0EAC35CA90B5E231292F4587935">
The species epithet
<taxonomicName id="1B0F9E4BB7AC45DAEC85C08312D6821A" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nawngau">
“
<emphasis id="8AEB66DE2593A547BBABD28E96214083" italics="true">nawngau</emphasis>
</taxonomicName>
” refers to the type locality, Nawng Au Village; it is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDJBG" type="distribution">
<paragraph id="D5267B716836F9C3F677E8EB9BD2C3BD">
<heading id="891C354375E2ED1C6222187E3106511B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="1D8EC46997F88337EE944622C3BE4074">
<collectingRegion id="CAAAE54240867612AB45B5794A10D990" country="Myanmar" name="Shan">Shan State</collectingRegion>
, (Kyaukme District),
<collectingCountry id="FCD4D0FA5A5AEE5063C962A6CB820501" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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