138 lines
10 KiB
XML
138 lines
10 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.72.744" ID-GBIF-Dataset="9b029a3f-f31b-421f-a62e-fb4dd8ff6731" ID-PMC="PMC3083004" ID-Pensoft-Pub="1313-2970-72-23" ID-PubMed="21594103" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-72-23" ModsDocOrigin="ZooKeys 72" ModsDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451250861919" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak" docDate="2010" docId="F028A40B624B60FDCB7FB733A6BAC641" docLanguage="en" docName="ZooKeys 72: 23-68" docOrigin="ZooKeys 72" docSource="http://dx.doi.org/10.3897/zookeys.72.744" docTitle="Tylopus grandis Likhitrakarn, Golovatch, Prateepasen & Panha, 2010, sp. n." docType="treatment" docVersion="5" lastPageNumber="28" masterDocId="D7146C3215665021FFDFFFACFF8CFFD0" masterDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="68" masterPageNumber="23" pageNumber="27" updateTime="1668165214761" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prateepasen, Rujiporn</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>72</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>68</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.72.744</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.72.744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-72-23</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159360925" LSID="urn:lsid:zoobank.org:act:148C0F96-F560-437A-92FC-6FA159B96699" httpUri="http://treatment.plazi.org/id/F028A40B624B60FDCB7FB733A6BAC641" lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27">
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<subSubSection pageId="4" pageNumber="27" type="nomenclature">
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<paragraph pageId="4" pageNumber="27">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:148C0F96-F560-437A-92FC-6FA159B96699" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus grandis" order="Polydesmida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="grandis">Tylopus grandis</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="27">sp. n.</taxonomicNameLabel>
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Figs 46
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="27" type="holotype">
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<paragraph pageId="4" pageNumber="27">Holotype</paragraph>
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<paragraph pageId="4" pageNumber="27">♂ (CUMZ), Thailand, Mae Hong Son Province, Pangmapha District, near Cave Pha Mon, 19.07.2008, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="27" type="paratypes">
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<paragraph pageId="4" pageNumber="27">Paratypes:</paragraph>
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<paragraph pageId="4" pageNumber="27">1 ♂, 2 ♀ (CUMZ), same lcality, together with holotype. 1 ♂ (CUMZ), same District, Mae Lana crossroads, 19.07.2008, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="27" type="name">
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<paragraph pageId="4" pageNumber="27">Name:</paragraph>
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<paragraph pageId="4" pageNumber="27">To emphasize the large size of this species</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="27" type="diagnosis">
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<paragraph pageId="4" pageNumber="27">Diagnosis:</paragraph>
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<paragraph pageId="4" pageNumber="27">Differs from congeners in the large size, coupled with a short spiniform process h, a basally only poorly delimited lobe l, and a small lobiform process z of the gonopod.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27" type="description">
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<paragraph pageId="4" pageNumber="27">Description:</paragraph>
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<paragraph lastPageId="5" lastPageNumber="28" pageId="4" pageNumber="27">
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Length 41 mm (holotype), 40-42 mm (♂), 38-39 mm (♀), width of midbody pro- and metazona 3.0 and 4.5 mm (holotype), 2.8-3.0 and 4.3-4.5 mm (♂), 3.6-3.8 and 4.7-5.0 mm (♀), respectively. Coloration in alcohol dark
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<pageBreakToken pageId="5" pageNumber="28" start="start">brown</pageBreakToken>
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to black-brown (Fig. 4
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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): calluses, venter and antennomeres 1-5 slightly to considerably lighter, brown to light yellow-brown (Fig. 1
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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), antennomeres 6 and 7 dark brown.
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</paragraph>
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<paragraph pageId="5" pageNumber="28">
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All characters as in
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus bispinosus" order="Polydesmida" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="bispinosus">Tylopus bispinosus</taxonomicName>
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sp. n., except as follows.
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</paragraph>
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<paragraph pageId="5" pageNumber="28">
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Antennae short and slender (Fig. 4B), reaching behind segment 3 (♂) or 2 (♀) dorsally. In width, head <collum = segments 3-4 <2 = 5-16 (♂), or head <segments 3-4 <collum <segment 2 <5-16 (♀); thereafter body gradually and gently tapering towards telson (Fig. 4
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="5" pageNumber="28">
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Tegument generally rather smooth and either dull (only in places modestly shining) or shining (Fig. 4
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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). Paraterga strongly developed (Fig. 4
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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), lying high (at 1/4-1/5 midbody height), subhorizontal to slightly upturned laterally (Fig. 4
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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). Transverse sulcus either absent or poorly developed, then not reaching bases of paraterga 4, always evident and reaching bases of paraterga 5-18, rather faintly rugulose at bottom. Stricture between pro- and metazona rather faintly beaded to striolate (Fig. 4
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Epiproct tip evidently emarginate, pre-apical papillae very distinct (Fig. 4F, G). Hypoproct semi-circular, caudal setae strongly separated (Fig. 4G). Pleurosternal carinae visible on segments 2-15(16) (♂) or segments 2-6 (♀), mostly as low bulges anteriorly and a more or less distinct denticle posteriorly (Fig. 4B, D).
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</paragraph>
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<paragraph pageId="5" pageNumber="28">Sterna moderately setose, without modifications; a slightly notched sternal lobe between ♂ coxae 4 (Fig. 4H, I). Legs long and slender (Fig. 1B, C, H), especially so in ♂ compared to ♀ (1.7-1.8 versus 1.5-1.6 times as long as midbody height); ♂ prefemora distinctly bulged laterally (Fig. 4J), acropodites with particularly dense, nearly adpressed setae, but tarsal brushes missing.</paragraph>
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<paragraph pageId="5" pageNumber="28">Gonopods (Figs 5, 6) with lobe l poorly demarcated at base; spine h very small; process z not spiniform, but like a short lobe.</paragraph>
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<caption pageId="5" pageNumber="28">
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<paragraph pageId="5" pageNumber="28">
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Figure 4.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus grandis" order="Polydesmida" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="grandis">Tylopus grandis</taxonomicName>
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sp. n., ♂ paratype from Mae Lana (
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<normalizedToken originalValue="A–J">A-J</normalizedToken>
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). A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E, F, G posterior part of body, lateral, dorsal and ventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively J midbody leg.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="28">
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<paragraph pageId="5" pageNumber="28">
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Figure 5.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus grandis" order="Polydesmida" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="grandis">Tylopus grandis</taxonomicName>
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sp. n., ♂ paratype from Mae Lana. A, B right gonopod, mesal and lateral views, respectively
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<normalizedToken originalValue="C–E">C-E</normalizedToken>
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distal part of right gonopod, mesal, lateral and suboral views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="28">
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<paragraph pageId="5" pageNumber="28">
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Figure 6.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus grandis" order="Polydesmida" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="grandis">Tylopus grandis</taxonomicName>
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sp. n., ♂ paratype from Mae Lana. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |