treatments-xml/data/03/A3/5D/03A35D13FFB7ED78FF3AFD76EC996AF4.xml
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<document id="36F7D9DBDE1F4725BD00B90CD0745C73" ID-CLB-Dataset="59775" ID-DOI="10.11646/zootaxa.5323.1.7" ID-GBIF-Dataset="184392fa-4901-4720-891c-e4e7847d6382" ID-ISSN="1175-5326" ID-Zenodo-Dep="8204051" ID-ZooBank="F06E94A9-1D19-471F-853B-5856E2362A36" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690886483627" checkinUser="plazi" docAuthor="Holloway, Graham J. &amp; Herrmann, Andreas" docDate="2023" docId="03A35D13FFB7ED78FF3AFD76EC996AF4" docLanguage="en" docName="zootaxa.5323.1.7.pdf" docOrigin="Zootaxa 5323 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5323.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthrenus (Anthrenus) semipallens Holloway &amp; Herrmann 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="131" masterDocId="FF9A256BFFB6ED7DFFADFFC4EC7B6A26" masterDocTitle="Anthrenus (s. str.) semipallens sp. nov., a new species from Spain (Coleoptera: Dermestidae: Anthreninae)" masterLastPageNumber="132" masterPageNumber="126" pageNumber="127" updateTime="1691067701831" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8E41A931BAE6458B077F2778BA3F22E9">Anthrenus (s. str.) semipallens sp. nov., a new species from Spain (Coleoptera: Dermestidae: Anthreninae)</mods:title>
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<mods:namePart id="2F842ED2C38B487EBF5B661F93EF8AF2">Holloway, Graham J.</mods:namePart>
<mods:affiliation id="E1AD3915AAC3BF0AC302AA29D1CB8414">Cole Museum of Zoology, Biological Sciences, HLS Building, University of Reading, Whiteknights, Reading RG 6 6 EX, UK.</mods:affiliation>
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<mods:namePart id="E0B5DB3A3BD839A9688DADB632FEEC2C">Herrmann, Andreas</mods:namePart>
<mods:affiliation id="78C067C1FD9760978CC600251FFB3398">Bremervörder Strasse 123, 21682 Stade, Germany.</mods:affiliation>
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<treatment id="03A35D13FFB7ED78FF3AFD76EC996AF4" ID-DOI="http://doi.org/10.5281/zenodo.8211635" ID-Zenodo-Dep="8211635" LSID="urn:lsid:plazi:treatment:03A35D13FFB7ED78FF3AFD76EC996AF4" httpUri="http://treatment.plazi.org/id/03A35D13FFB7ED78FF3AFD76EC996AF4" lastPageId="5" lastPageNumber="131" pageId="1" pageNumber="127">
<subSubSection id="C310BF8EFFB7ED7CFF3AFD76EF2168EA" box="[151,858,690,717]" pageId="1" pageNumber="127" type="nomenclature">
<paragraph id="8BB5EC05FFB7ED7CFF3AFD76EF2168EA" blockId="1.[151,858,690,717]" box="[151,858,690,717]" pageId="1" pageNumber="127">
<heading id="D0FD5B69FFB7ED7CFF3AFD76EF2168EA" bold="true" box="[151,858,690,717]" fontSize="11" level="1" pageId="1" pageNumber="127" reason="1">
<emphasis id="B97E3017FFB7ED7CFF3AFD76EF2168EA" bold="true" box="[151,858,690,717]" pageId="1" pageNumber="127">
<taxonomicName id="4C0A9786FFB7ED7CFF3AFD76EE3268EB" authority="Holloway &amp; Herrmann, 2023" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[151,585,690,717]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="semipallens" status="sp. nov." subGenus="Anthrenus">
<emphasis id="B97E3017FFB7ED7CFF3AFD76ED6368EB" bold="true" box="[151,280,690,717]" italics="true" pageId="1" pageNumber="127">Anthrenus</emphasis>
(
<emphasis id="B97E3017FFB7ED7CFE84FD76EDD168EB" bold="true" box="[297,426,690,717]" italics="true" pageId="1" pageNumber="127">Anthrenus</emphasis>
)
<emphasis id="B97E3017FFB7ED7CFE16FD76EE3268EB" bold="true" box="[443,585,690,717]" italics="true" pageId="1" pageNumber="127">semipallens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D8D6CFFB7ED7CFDFDFD77EED568EB" box="[592,686,691,717]" pageId="1" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1331F080FFB7ED7CFD10FD76EF2968EA" box="[701,850,690,716]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[152,255,776,800]" captionStartId-1="2.[152,255,1919,1943]" captionTargetBox-0="[151,1428,181,752]" captionTargetBox-1="[151,1436,934,1896]" captionTargetId-0="figure-17@2.[151,1436,181,752]" captionTargetId-1="figure-45@2.[151,1436,933,1896]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURE 1. Anthrenus semipallens sp. nov. Holotype. A: habitus dorsal aspect, B: elytral scales, C: ventrites. Scale bars = 1 mm." captionText-1="FIGURE 2. Anthrenus semipallens sp. nov. Holotype. A: Antenna, B: aedeagus ventral aspect, C: sternite IX. Scale bars = 100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.8204232" figureDoi-1="http://doi.org/10.5281/zenodo.8204057" httpUri-0="https://zenodo.org/record/8204232/files/figure.png" httpUri-1="https://zenodo.org/record/8204057/files/figure.png" pageId="1" pageNumber="127">Figures 12</figureCitation>
)
</emphasis>
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</paragraph>
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<subSubSection id="C310BF8EFFB7ED78FF3AFD3FEC996AF4" lastPageId="5" lastPageNumber="131" pageId="1" pageNumber="127" type="description">
<paragraph id="8BB5EC05FFB7ED7CFF3AFD3FE89C6983" blockId="1.[151,1437,763,1797]" pageId="1" pageNumber="127">
<emphasis id="B97E3017FFB7ED7CFF3AFD3FED076933" box="[151,380,763,789]" italics="true" pageId="1" pageNumber="127">Specimen examined.</emphasis>
<materialsCitation id="3B62E658FFB7ED7CFE2FFD3FE830691F" collectingDate="2017-07-04" collectionCode="AHEC" collectorName="J. Marmaneu &amp; E. Mico" country="Spain" county="Villarejo" latitude="40.472" location="New" municipality="Salamanca" pageId="1" pageNumber="127" specimenCount="2" typeStatus="holotype">
<location id="8ED5BADEFFB7ED7CFE2FFD3FEDCC6933" LSID="urn:lsid:plazi:treatment:03A35D13FFB7ED78FF3AFD76EC996AF4:8ED5BADEFFB7ED7CFE2FFD3FEDCC6933" box="[386,439,763,789]" country="Spain" county="Villarejo" latitude="40.472" municipality="Salamanca" name="New" pageId="1" pageNumber="127">New</location>
record (holotype) for
<taxonomicName id="4C0A9786FFB7ED7CFD02FD3FEF846933" authority="Holloway &amp; Herrmann, 2023" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[687,1023,763,789]" class="Insecta" family="Dermestidae" genus="Anthrenus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="semipallens" status="sp. nov.">
<emphasis id="B97E3017FFB7ED7CFD02FD3FEF5A6933" box="[687,801,763,789]" italics="true" pageId="1" pageNumber="127">Anthrenus</emphasis>
(s. str.)
<emphasis id="B97E3017FFB7ED7CFCD6FD3FEF846933" box="[891,1023,763,789]" italics="true" pageId="1" pageNumber="127">semipallens</emphasis>
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<taxonomicNameLabel id="A24D8D6CFFB7ED7CFBABFD3FE81B6933" box="[1030,1120,763,789]" pageId="1" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<collectingCountry id="F31DAC95FFB7ED7CFBCBFD3FE8D26933" box="[1126,1193,763,789]" name="Spain" pageId="1" pageNumber="127">Spain</collectingCountry>
,
<collectingMunicipality id="6BD1767FFFB7ED7CFB19FD3FE94B6933" box="[1204,1328,763,789]" pageId="1" pageNumber="127">Salamanca</collectingMunicipality>
,
<collectingCounty id="62D49489FFB7ED7CFA97FD3FE9E76933" box="[1338,1436,763,789]" pageId="1" pageNumber="127">Villarejo</collectingCounty>
(N 40.472, E -6.457),
<date id="FFB4CAC5FFB7ED7CFE26FCDBEE63691F" box="[395,536,798,825]" pageId="1" pageNumber="127" value="2017-07-04">
<collectingDate id="EFF0332DFFB7ED7CFE26FCDBEE63691F" box="[395,536,798,825]" pageId="1" pageNumber="127" value="2017-07-04">
4
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July 2017
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<collectorName id="26FF89D3FFB7ED7CFDB2FCDBEECB691F" box="[543,688,799,825]" pageId="1" pageNumber="127">J. Marmaneu</collectorName>
and
<collectorName id="26FF89D3FFB7ED7CFD49FCDBEF45691F" box="[740,830,799,825]" pageId="1" pageNumber="127">E. Micó</collectorName>
leg.
<typeStatus id="54B152A7FFB7ED7CFCDEFCDBEFA0691F" box="[883,987,799,825]" pageId="1" pageNumber="127">Holotype</typeStatus>
<collectionCode id="ED1B74C0FFB7ED7CFC56FCDBE830691F" box="[1019,1099,799,825]" pageId="1" pageNumber="127">AHEC</collectionCode>
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.
<materialsCitation id="3B62E658FFB7ED7CFBF9FCDBE8986983" collectingDate="2017-07-04" collectingDateMax="2017-07-05" collectingDateMin="2017-07-04" collectionCode="NHML, AHEC" country="Spain" latitude="40.97" location="Salamanca" municipality="Escurial de la Sierra" pageId="1" pageNumber="127" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54B152A7FFB7ED7CFBF9FCDBE8BE691F" box="[1108,1221,799,825]" pageId="1" pageNumber="127">Paratypes</typeStatus>
:
<collectingCountry id="F31DAC95FFB7ED7CFB62FCDBE969691F" box="[1231,1298,799,825]" name="Spain" pageId="1" pageNumber="127">Spain</collectingCountry>
,
<location id="8ED5BADEFFB7ED7CFAB1FCDBE9E3691F" LSID="urn:lsid:plazi:treatment:03A35D13FFB7ED78FF3AFD76EC996AF4:8ED5BADEFFB7ED7CFAB1FCDBE9E3691F" box="[1308,1432,799,825]" country="Spain" latitude="40.97" municipality="Escurial de la Sierra" name="Salamanca" pageId="1" pageNumber="127">Salamanca</location>
,
<collectorName id="26FF89D3FFB7ED7CFF3AFC87ED04697B" box="[151,383,835,861]" pageId="1" pageNumber="127">
<collectingMunicipality id="6BD1767FFFB7ED7CFF3AFC87ED04697B" box="[151,383,835,861]" pageId="1" pageNumber="127">Escurial de la Sierra</collectingMunicipality>
</collectorName>
(N 40.618, E -5.95)
<date id="FFB4CAC5FFB7ED7CFDDDFC87EF78697B" box="[624,771,834,861]" pageId="1" pageNumber="127" value="2017-07-04">
<collectingDate id="EFF0332DFFB7ED7CFDDDFC87EF78697B" box="[624,771,834,861]" pageId="1" pageNumber="127" value="2017-07-04">
4
<superScript id="7C7F414DFFB7ED7CFDD3FC86EEF06976" attach="left" box="[638,651,834,848]" fontSize="6" pageId="1" pageNumber="127">th</superScript>
July 2017
</collectingDate>
</date>
<collectorName id="26FF89D3FFB7ED7CFCA1FC87EFE4697B" box="[780,927,835,861]" pageId="1" pageNumber="127">J. Marmaneu</collectorName>
and
<collectorName id="26FF89D3FFB7ED7CFC74FC87E84E697B" box="[985,1077,835,861]" pageId="1" pageNumber="127">E. Micó</collectorName>
leg. (
<specimenCount id="9D0C278CFFB7ED7CFBDAFC87E89B697B" box="[1143,1248,835,861]" pageId="1" pageNumber="127" type="male">one male</specimenCount>
<collectionCode id="ED1B74C0FFB7ED7CFB44FC87E93A697B" box="[1257,1345,835,861]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="1" pageNumber="127">NHML</collectionCode>
); Spain,
<collectorName id="26FF89D3FFB7ED7CFF3AFCA3ED6869A7" box="[151,275,871,897]" pageId="1" pageNumber="127">Salamanca</collectorName>
,
<collectorName id="26FF89D3FFB7ED7CFE8DFCA3ED0D69A7" box="[288,374,871,897]" pageId="1" pageNumber="127">Morillo</collectorName>
(N 40.972, E -5.663)
<date id="FFB4CAC5FFB7ED7CFDDBFCA3EF7269A7" box="[630,777,870,897]" pageId="1" pageNumber="127" value="2017-07-04">
<collectingDate id="EFF0332DFFB7ED7CFDDBFCA3EF7269A7" box="[630,777,870,897]" pageId="1" pageNumber="127" value="2017-07-04">
4
<superScript id="7C7F414DFFB7ED7CFD29FCA2EEEA6952" attach="left" box="[644,657,870,884]" fontSize="6" pageId="1" pageNumber="127">th</superScript>
July 2017
</collectingDate>
</date>
<collectorName id="26FF89D3FFB7ED7CFCBFFCA3EFDD69A7" box="[786,934,871,897]" pageId="1" pageNumber="127">J. Marmaneu</collectorName>
and
<collectorName id="26FF89D3FFB7ED7CFC4DFCA3E84769A7" box="[992,1084,871,897]" pageId="1" pageNumber="127">E. Micó</collectorName>
leg. (
<specimenCount id="9D0C278CFFB7ED7CFBD2FCA3E89369A7" box="[1151,1256,871,897]" pageId="1" pageNumber="127" type="male">one male</specimenCount>
<collectionCode id="ED1B74C0FFB7ED7CFB42FCA3E93B69A7" box="[1263,1344,871,897]" pageId="1" pageNumber="127">AHEC</collectionCode>
); Spain,
<collectorName id="26FF89D3FFB7ED7CFF3AFC4FED686983" box="[151,275,907,933]" pageId="1" pageNumber="127">Salamanca</collectorName>
, (N 40.970, E -5.668)
<date id="FFB4CAC5FFB7ED7CFDA2FC4EEEE56983" box="[527,670,906,933]" pageId="1" pageNumber="127" value="2017-07-05">
<collectingDate id="EFF0332DFFB7ED7CFDA2FC4EEEE56983" box="[527,670,906,933]" pageId="1" pageNumber="127" value="2017-07-05">
5
<superScript id="7C7F414DFFB7ED7CFDB0FC4EEE5169BE" attach="left" box="[541,554,906,920]" fontSize="6" pageId="1" pageNumber="127">th</superScript>
July 2017
</collectingDate>
</date>
<collectorName id="26FF89D3FFB7ED7CFD08FC4FEF4C6983" box="[677,823,907,933]" pageId="1" pageNumber="127">J. Marmaneu</collectorName>
and
<collectorName id="26FF89D3FFB7ED7CFCC3FC4FEFB36983" box="[878,968,907,933]" pageId="1" pageNumber="127">E. Micó</collectorName>
leg. (
<specimenCount id="9D0C278CFFB7ED7CFBAAFC4FE8FF6983" box="[1031,1156,907,933]" pageId="1" pageNumber="127" type="female">one female</specimenCount>
<collectionCode id="ED1B74C0FFB7ED7CFB27FC4FE8A06983" box="[1162,1243,907,933]" pageId="1" pageNumber="127">AHEC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB5EC05FFB7ED7CFF6AFC6BEFFD6F17" blockId="1.[151,1437,763,1797]" pageId="1" pageNumber="127">
<emphasis id="B97E3017FFB7ED7CFF6AFC6BEDAF69EF" box="[199,468,943,969]" italics="true" pageId="1" pageNumber="127">External characteristics</emphasis>
.
<typeStatus id="54B152A7FFB7ED7CFE4CFC6BEE3269EF" box="[481,585,943,969]" pageId="1" pageNumber="127">Holotype</typeStatus>
habitus dorsal aspect
<figureCitation id="1331F080FFB7ED7CFC91FC6BEFD769EF" box="[828,940,943,969]" captionStart="FIGURE 1" captionStartId="2.[152,255,776,800]" captionTargetBox="[151,1428,181,752]" captionTargetId="figure-17@2.[151,1436,181,752]" captionTargetPageId="2" captionText="FIGURE 1. Anthrenus semipallens sp. nov. Holotype. A: habitus dorsal aspect, B: elytral scales, C: ventrites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8204232" httpUri="https://zenodo.org/record/8204232/files/figure.png" pageId="1" pageNumber="127">Figure 1A</figureCitation>
(BL =
<quantity id="4CF241E0FFB7ED7CFC57FC6AE82A69EF" box="[1018,1105,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="127" unit="mm" value="2.5">2.5 mm</quantity>
, BW =
<quantity id="4CF241E0FFB7ED7CFB09FC6AE97069EF" box="[1188,1291,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="127" unit="mm" value="1.75">1.75 mm</quantity>
). Integument head and pronotum black, margin of pronotum tinged red, basal 1/6
<superScript id="7C7F414DFFB7ED7CFC1FFC16EFC469C6" attach="left" box="[946,959,978,992]" fontSize="6" pageId="1" pageNumber="127">th</superScript>
of elytra black, rest of elytra red. Sharp demarcation between black and red integument of elytra. Elytra loosely covered with white and yellow to pale orange scales, many of which appear to be rectangular and square-ended (
<figureCitation id="1331F080FFB7ED7CFC4FFBDFE8236E13" box="[994,1112,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[152,255,776,800]" captionTargetBox="[151,1428,181,752]" captionTargetId="figure-17@2.[151,1436,181,752]" captionTargetPageId="2" captionText="FIGURE 1. Anthrenus semipallens sp. nov. Holotype. A: habitus dorsal aspect, B: elytral scales, C: ventrites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8204232" httpUri="https://zenodo.org/record/8204232/files/figure.png" pageId="1" pageNumber="127">Figure 1B</figureCitation>
). Scales not arranged in any particular patterns with no obvious spots or fasciae. Scutellum small and black. Ventrites (
<figureCitation id="1331F080FFB7ED7CFBD5FBFBE8926E7F" box="[1144,1257,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[152,255,776,800]" captionTargetBox="[151,1428,181,752]" captionTargetId="figure-17@2.[151,1436,181,752]" captionTargetPageId="2" captionText="FIGURE 1. Anthrenus semipallens sp. nov. Holotype. A: habitus dorsal aspect, B: elytral scales, C: ventrites. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8204232" httpUri="https://zenodo.org/record/8204232/files/figure.png" pageId="1" pageNumber="127">Figure 1C</figureCitation>
) covered in offwhite, slightly translucent, loosely packed scales. Scales along outer margins of sternites III, IV, and V more densely packed, have a hint of yellow in them, and increase in density towards the tip of sternite V. Single brown ocellus between eyes. Head entirely covered in black scales. Eyes emarginated on inner margin. Femora, tibiae, and tarsi entirely red with a few whitish scales on the femora. 11-segmented antenna (
<figureCitation id="1331F080FFB7ED7CFC59FB0BE8116ECF" box="[1012,1130,1231,1257]" captionStart="FIGURE 2" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1436,934,1896]" captionTargetId="figure-45@2.[151,1436,933,1896]" captionTargetPageId="2" captionText="FIGURE 2. Anthrenus semipallens sp. nov. Holotype. A: Antenna, B: aedeagus ventral aspect, C: sternite IX. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.8204057" httpUri="https://zenodo.org/record/8204057/files/figure.png" pageId="1" pageNumber="127">Figure 2A</figureCitation>
) red throughout. Last three antennomeres forming a well-defined, slightly asymmetric, tear-drop shaped club that expands from antennomere 9 to antennomere 11. Broad sutures separating antennomeres 9 to 11.
</paragraph>
<paragraph id="8BB5EC05FFB7ED7CFF6AFAFFE9F86FC3" blockId="1.[151,1437,763,1797]" pageId="1" pageNumber="127">
<emphasis id="B97E3017FFB7ED7CFF6AFAFFEDA96F73" box="[199,466,1339,1365]" italics="true" pageId="1" pageNumber="127">Internal characteristics</emphasis>
. Aedeagus (
<figureCitation id="1331F080FFB7ED7CFDCEFAFFEEA16F73" box="[611,730,1339,1365]" captionStart="FIGURE 2" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1436,934,1896]" captionTargetId="figure-45@2.[151,1436,933,1896]" captionTargetPageId="2" captionText="FIGURE 2. Anthrenus semipallens sp. nov. Holotype. A: Antenna, B: aedeagus ventral aspect, C: sternite IX. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.8204057" httpUri="https://zenodo.org/record/8204057/files/figure.png" pageId="1" pageNumber="127">Figure 2B</figureCitation>
) has two broad, symmetrical parameres diverging from base before curving evenly inwards ending in blunt tips. Dorsal (concave) surface of each paramere carry inward pointing hairs. Base of median lobe broad, margins converge but are slightly sinuous, ending in a very thin finger-like projection. Sternite IX (
<figureCitation id="1331F080FFB7ED7CFE0BFA63EE676FE7" box="[422,540,1447,1473]" captionStart="FIGURE 2" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1436,934,1896]" captionTargetId="figure-45@2.[151,1436,933,1896]" captionTargetPageId="2" captionText="FIGURE 2. Anthrenus semipallens sp. nov. Holotype. A: Antenna, B: aedeagus ventral aspect, C: sternite IX. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.8204057" httpUri="https://zenodo.org/record/8204057/files/figure.png" pageId="1" pageNumber="127">Figure 2C</figureCitation>
) has a broad, slightly convex posterior tip carrying stout setae concentrated at outer corners. Strong setae continue down the outer margins beyond the narrowest part of the well-defined neck.
</paragraph>
<paragraph id="8BB5EC05FFB7ED7CFF6AFA2BED5E6C0B" blockId="1.[151,1437,763,1797]" pageId="1" pageNumber="127">
<emphasis id="B97E3017FFB7ED7CFF6AFA2BED3E6C2F" box="[199,325,1519,1545]" italics="true" pageId="1" pageNumber="127">Etymology.</emphasis>
The name
<taxonomicName id="4C0A9786FFB7ED7CFE6CFA2BEE3E6C2F" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[449,581,1519,1545]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB7ED7CFE6CFA2BEE3E6C2F" box="[449,581,1519,1545]" italics="true" pageId="1" pageNumber="127">semipallens</emphasis>
</taxonomicName>
translates from Latin as almost pale referring to the pale appearance of the new species.
</paragraph>
<paragraph id="8BB5EC05FFB7ED7CFF6AF9F3EFEF6D23" blockId="1.[151,1437,763,1797]" pageId="1" pageNumber="127">
<emphasis id="B97E3017FFB7ED7CFF6AF9F3EDC66C77" box="[199,445,1591,1617]" italics="true" pageId="1" pageNumber="127">Differential diagnosis</emphasis>
. The only
<taxonomicName id="4C0A9786FFB7ED7CFD90F9F3EED46C77" authorityName="Geoffroy" authorityYear="1762" box="[573,687,1591,1617]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B97E3017FFB7ED7CFD90F9F3EED46C77" box="[573,687,1591,1617]" italics="true" pageId="1" pageNumber="127">Anthrenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D8D6CFFB7ED7CFD15F9F3EE8E6C77" box="[696,757,1591,1617]" pageId="1" pageNumber="127" sensu="stricto">s. str.</taxonomicNameLabel>
species from
<collectingCountry id="F31DAC95FFB7ED7CFC35F9F3EFA36C77" box="[920,984,1591,1617]" name="Spain" pageId="1" pageNumber="127">Spain</collectingCountry>
(
<bibRefCitation id="EF9B91F4FFB7ED7CFC44F9F3E89B6C77" author="Holloway, G. J. &amp; Canada Luna, I. &amp; Kadej, M." box="[1001,1248,1591,1617]" pageId="1" pageNumber="127" pagination="1 - 8" refId="ref3428" refString="Holloway, G. J., Canada Luna, I. &amp; Kadej, M. (2019) A review of the literature and a checklist of the Spanish Dermestidae (Coleoptera). The Coleopterists Bulletin, 73, 1 - 8. https: // doi. org / 10.1649 / 0010 - 065 X- 73.3.513" type="journal article" year="2019">
Holloway
<emphasis id="B97E3017FFB7ED7CFBCFF9F3E8ED6C77" box="[1122,1174,1591,1617]" italics="true" pageId="1" pageNumber="127">et al</emphasis>
. 2019
</bibRefCitation>
) that marginally resembles
<taxonomicName id="4C0A9786FFB7ED7CFEA2F99FEDC86C53" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[271,435,1627,1653]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB7ED7CFEA2F99FEDC86C53" box="[271,435,1627,1653]" italics="true" pageId="1" pageNumber="127">A. semipallens</emphasis>
</taxonomicName>
externally are examples of very pale
<taxonomicName id="4C0A9786FFB7ED7CFCF2F99FE8EA6C53" authority="Kuster, 1848" authorityName="Kuster" authorityYear="1848" box="[863,1169,1627,1653]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="isabellinus">
<emphasis id="B97E3017FFB7ED7CFCF2F99FEF836C53" box="[863,1016,1627,1653]" italics="true" pageId="1" pageNumber="127">A. isabellinus</emphasis>
Küster, 1848
</taxonomicName>
(
<figureCitation id="1331F080FFB7ED7CFB0CF99FE9786C53" box="[1185,1283,1627,1653]" captionStart="FIGURE 3" captionStartId="3.[152,255,1320,1344]" captionTargetId="figure-28@3.[366,1222,181,1296]" captionTargetPageId="3" captionText="FIGURE 3. Anthrenus isabellinus pale example. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8211637" httpUri="https://zenodo.org/record/8211637/files/figure.png" pageId="1" pageNumber="127">Figure 3</figureCitation>
), although
<taxonomicName id="4C0A9786FFB7ED7CFA29F99FED6B6CBF" authorityName="Kuster" authorityYear="1848" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="isabellinus">
<emphasis id="B97E3017FFB7ED7CFA29F99FED6B6CBF" italics="true" pageId="1" pageNumber="127">A. isabellinus</emphasis>
</taxonomicName>
is generally much larger and rounder than
<taxonomicName id="4C0A9786FFB7ED7CFD5AF9BBEFE06CBF" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[759,923,1663,1689]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB7ED7CFD5AF9BBEFE06CBF" box="[759,923,1663,1689]" italics="true" pageId="1" pageNumber="127">A. semipallens</emphasis>
</taxonomicName>
. There are species in different subgenera that generally resemble
<taxonomicName id="4C0A9786FFB7ED7CFED8F967EE626C9B" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[373,537,1699,1725]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB7ED7CFED8F967EE626C9B" box="[373,537,1699,1725]" italics="true" pageId="1" pageNumber="127">A. semipallens</emphasis>
</taxonomicName>
, including
<taxonomicName id="4C0A9786FFB7ED7CFD36F967E8276C9B" authority="Kocher, 1955" authorityName="Kocher" authorityYear="1955" box="[667,1116,1698,1725]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="bellulus" subGenus="Anthrenodes">
<emphasis id="B97E3017FFB7ED7CFD36F967EEC86C9B" box="[667,691,1699,1725]" italics="true" pageId="1" pageNumber="127">A.</emphasis>
(
<emphasis id="B97E3017FFB7ED7CFD68F967EF2A6C9B" box="[709,849,1699,1725]" italics="true" pageId="1" pageNumber="127">Anthrenodes</emphasis>
)
<emphasis id="B97E3017FFB7ED7CFCCEF967EFC76C9B" box="[867,956,1699,1725]" italics="true" pageId="1" pageNumber="127">bellulus</emphasis>
Kocher, 1955
</taxonomicName>
and
<taxonomicName id="4C0A9786FFB7ED7CFB38F967ED3D6CC7" authority="Erichson, 1846" authorityName="Erichson" authorityYear="1846" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="127" phylum="Arthropoda" rank="species" species="minutus" subGenus="Helocerus">
<emphasis id="B97E3017FFB7ED7CFB38F967E8D56C9B" box="[1173,1198,1699,1725]" italics="true" pageId="1" pageNumber="127">A.</emphasis>
(
<emphasis id="B97E3017FFB7ED7CFB12F967E9496C9B" box="[1215,1330,1699,1725]" italics="true" pageId="1" pageNumber="127">Helocerus</emphasis>
)
<emphasis id="B97E3017FFB7ED7CFAE9F967E9E66C9A" box="[1348,1437,1699,1724]" italics="true" pageId="1" pageNumber="127">minutus</emphasis>
Erichson, 1846
</taxonomicName>
. Careful inspection of the number of antennal segments and the absence of a notch on the inner margin of the eye will quickly eliminate these species as candidates.
</paragraph>
<caption id="DF75BC8DFFB4ED7FFF35FCCCECBD6962" ID-DOI="http://doi.org/10.5281/zenodo.8204232" ID-Zenodo-Dep="8204232" httpUri="https://zenodo.org/record/8204232/files/figure.png" pageId="2" pageNumber="128" startId="2.[152,255,776,800]" targetBox="[151,1428,181,752]" targetPageId="2" targetType="figure">
<paragraph id="8BB5EC05FFB4ED7FFF35FCCCECBD6962" blockId="2.[151,1437,776,836]" pageId="2" pageNumber="128">
<emphasis id="B97E3017FFB4ED7FFF35FCCCED626906" bold="true" box="[152,281,776,800]" pageId="2" pageNumber="128">FIGURE 1.</emphasis>
<taxonomicName id="4C0A9786FFB4ED7FFE8CFCCCEE7C6906" authority="Holloway &amp; Herrmann, 2023" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[289,519,776,800]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="128" phylum="Arthropoda" rank="species" species="semipallens" status="sp. nov.">
<emphasis id="B97E3017FFB4ED7FFE8CFCCCEE7C6906" box="[289,519,776,800]" italics="true" pageId="2" pageNumber="128">Anthrenus semipallens</emphasis>
</taxonomicName>
<emphasis id="B97E3017FFB4ED7FFDA3FCCDEE246907" bold="true" box="[526,607,777,801]" pageId="2" pageNumber="128">
<taxonomicNameLabel id="A24D8D6CFFB4ED7FFDA3FCCDEE246907" box="[526,607,777,801]" pageId="2" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype. A: habitus dorsal aspect, B: elytral scales, C: ventrites. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF75BC8DFFB4ED7FFF35F8BBECBB6D9A" ID-DOI="http://doi.org/10.5281/zenodo.8204057" ID-Zenodo-Dep="8204057" httpUri="https://zenodo.org/record/8204057/files/figure.png" pageId="2" pageNumber="128" startId="2.[152,255,1919,1943]" targetBox="[151,1436,934,1896]" targetPageId="2" targetType="figure">
<paragraph id="8BB5EC05FFB4ED7FFF35F8BBECBB6D9A" blockId="2.[151,1436,1919,1980]" pageId="2" pageNumber="128">
<emphasis id="B97E3017FFB4ED7FFF35F8BBED636DB1" bold="true" box="[152,280,1919,1943]" pageId="2" pageNumber="128">FIGURE 2.</emphasis>
<taxonomicName id="4C0A9786FFB4ED7FFEB3F8BBEE786DB1" authority="Holloway &amp; Herrmann, 2023" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[286,515,1919,1943]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="128" phylum="Arthropoda" rank="species" species="semipallens" status="sp. nov.">
<emphasis id="B97E3017FFB4ED7FFEB3F8BBEE786DB1" box="[286,515,1919,1943]" italics="true" pageId="2" pageNumber="128">Anthrenus semipallens</emphasis>
</taxonomicName>
<emphasis id="B97E3017FFB4ED7FFDA5F8BBEE236DB1" bold="true" box="[520,600,1919,1943]" pageId="2" pageNumber="128">
<taxonomicNameLabel id="A24D8D6CFFB4ED7FFDA5F8BBEE236DB1" box="[520,600,1919,1943]" pageId="2" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype. A: Antenna, B: aedeagus ventral aspect, C: sternite IX. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="DF75BC8DFFB5ED7EFF35FAECEF356F19" ID-DOI="http://doi.org/10.5281/zenodo.8211637" ID-Zenodo-Dep="8211637" box="[152,846,1320,1344]" httpUri="https://zenodo.org/record/8211637/files/figure.png" pageId="3" pageNumber="129" startId="3.[152,255,1320,1344]" targetBox="[366,1190,181,1296]" targetPageId="3" targetType="figure">
<paragraph id="8BB5EC05FFB5ED7EFF35FAECEF356F19" blockId="3.[152,846,1320,1344]" box="[152,846,1320,1344]" pageId="3" pageNumber="129">
<emphasis id="B97E3017FFB5ED7EFF35FAECED636F66" bold="true" box="[152,280,1320,1344]" pageId="3" pageNumber="129">FIGURE 3.</emphasis>
<taxonomicName id="4C0A9786FFB5ED7EFEB2FAECED816F66" authorityName="Kuster" authorityYear="1848" box="[287,506,1320,1344]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="isabellinus">
<emphasis id="B97E3017FFB5ED7EFEB2FAECED816F66" box="[287,506,1320,1344]" italics="true" pageId="3" pageNumber="129">Anthrenus isabellinus</emphasis>
</taxonomicName>
pale example. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BB5EC05FFB5ED7EFF6AFA56ED9C6DEE" blockId="3.[151,1437,1426,2029]" pageId="3" pageNumber="129">
There are
<taxonomicName id="4C0A9786FFB5ED7EFEECFA56EE6B6F8A" box="[321,528,1426,1452]" pageId="3" pageNumber="129">
<emphasis id="B97E3017FFB5ED7EFEECFA56EDC86F8A" box="[321,435,1426,1452]" italics="true" pageId="3" pageNumber="129">Anthrenus</emphasis>
species
</taxonomicName>
that display colour variation, either fairly discrete variants as in
<taxonomicName id="4C0A9786FFB5ED7EFAB9FA56ED0A6FF7" authority="Kiesenwetter, 1851" authorityName="Kiesenwetter" authorityYear="1851" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="B97E3017FFB5ED7EFAB9FA56E9E76F8A" box="[1300,1436,1426,1452]" italics="true" pageId="3" pageNumber="129">A. delicatus</emphasis>
Kiesenwetter, 1851
</taxonomicName>
(
<bibRefCitation id="EF9B91F4FFB5ED7EFE2CFA72EE436FF6" author="Herrmann, A." box="[385,568,1462,1488]" pageId="3" pageNumber="129" refId="ref2896" refString="Herrmann, A. (2023) Dermestidae (Coleoptera) of the World. Dermestidae (Coleoptera) - Homepage of Andreas Herrmann. Available from: http: // www. dermestidae. com / (accessed 12 April 2023)" type="url" year="2023">Herrmann 2023</bibRefCitation>
) or continuous colour pattern plasticity as in
<taxonomicName id="4C0A9786FFB5ED7EFB92FA72ECA76FD2" authority="(Holloway et al. 2022)" baseAuthorityName="Holloway" baseAuthorityYear="2022" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="isabellinus">
<emphasis id="B97E3017FFB5ED7EFB92FA72E8A26FF6" box="[1087,1241,1462,1488]" italics="true" pageId="3" pageNumber="129">A. isabellinus</emphasis>
(Holloway
<emphasis id="B97E3017FFB5ED7EFACEFA73E9EE6FF6" box="[1379,1429,1462,1488]" italics="true" pageId="3" pageNumber="129">et al</emphasis>
. 2022)
</taxonomicName>
. Is it possible that
<taxonomicName id="4C0A9786FFB5ED7EFE00FA1EEE2A6FD2" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[429,593,1498,1524]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB5ED7EFE00FA1EEE2A6FD2" box="[429,593,1498,1524]" italics="true" pageId="3" pageNumber="129">A. semipallens</emphasis>
</taxonomicName>
is a colour variant of another species?
<taxonomicName id="4C0A9786FFB5ED7EFBA5FA1EE97E6FD2" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[1032,1285,1498,1524]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB5ED7EFBA5FA1EE97E6FD2" box="[1032,1285,1498,1524]" italics="true" pageId="3" pageNumber="129">Anthrenus semipallens</emphasis>
</taxonomicName>
is small. The male
<typeStatus id="54B152A7FFB5ED7EFF7BFA3BED3A6C3F" box="[214,321,1535,1561]" pageId="3" pageNumber="129">paratypes</typeStatus>
measure BL
<quantity id="4CF241E0FFB5ED7EFE76FA3AEE3B6C3F" box="[475,576,1534,1561]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="129" unit="mm" value="2.75">2.75 mm</quantity>
x BW
<quantity id="4CF241E0FFB5ED7EFD39FA3AEE906C3F" box="[660,747,1534,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="129" unit="mm" value="1.9">1.9 mm</quantity>
and BL
<quantity id="4CF241E0FFB5ED7EFCE2FA3AEFDC6C3F" box="[847,935,1534,1561]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="129" unit="mm" value="2.5">2.5 mm</quantity>
x BW
<quantity id="4CF241E0FFB5ED7EFC57FA3AE82F6C3F" box="[1018,1108,1534,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="129" unit="mm" value="1.8">1.8 mm</quantity>
, and the female
<typeStatus id="54B152A7FFB5ED7EFABCFA3BE90A6C3F" box="[1297,1393,1535,1561]" pageId="3" pageNumber="129">paratype</typeStatus>
BL
<quantity id="4CF241E0FFB5ED7EFF3AF9E6EC946C1B" box="[151,239,1570,1597]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="129" unit="mm" value="2.6">2.6 mm</quantity>
x BW
<quantity id="4CF241E0FFB5ED7EFEE9F9E6EDD76C1B" box="[324,428,1570,1597]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="3" pageNumber="129" unit="mm" value="1.85">1.85 mm</quantity>
, average BL
<quantity id="4CF241E0FFB5ED7EFDEBF9E6EED76C1B" box="[582,684,1570,1597]" metricMagnitude="-3" metricUnit="m" metricValue="2.59" pageId="3" pageNumber="129" unit="mm" value="2.59">2.59 mm</quantity>
x BW
<quantity id="4CF241E0FFB5ED7EFCACF9E6EF126C1B" box="[769,873,1570,1597]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="3" pageNumber="129" unit="mm" value="1.82">1.82 mm</quantity>
, BW/BL = 0.703. These dimensions eliminate
<taxonomicName id="4C0A9786FFB5ED7EFA29F9E6ED996C46" authority="Mulsant &amp; Rey, 1868" authorityName="Mulsant &amp; Rey" authorityYear="1868" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="goliath">
<emphasis id="B97E3017FFB5ED7EFA29F9E6EC9D6C46" italics="true" pageId="3" pageNumber="129">A. goliath</emphasis>
Mulsant &amp; Rey, 1868
</taxonomicName>
(
<bibRefCitation id="EF9B91F4FFB5ED7EFE5FF982EEDC6C46" author="Herrmann, A." box="[498,679,1606,1632]" pageId="3" pageNumber="129" refId="ref2896" refString="Herrmann, A. (2023) Dermestidae (Coleoptera) of the World. Dermestidae (Coleoptera) - Homepage of Andreas Herrmann. Available from: http: // www. dermestidae. com / (accessed 12 April 2023)" type="url" year="2023">Herrmann 2023</bibRefCitation>
) and
<taxonomicName id="4C0A9786FFB5ED7EFD4AF982E8756C47" authority="LeConte, 1854" authorityName="LeConte" authorityYear="1854" box="[743,1038,1606,1633]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="B97E3017FFB5ED7EFD4AF982EF256C46" box="[743,862,1606,1632]" italics="true" pageId="3" pageNumber="129">A. flavipes</emphasis>
LeConte, 1854
</taxonomicName>
(
<bibRefCitation id="EF9B91F4FFB5ED7EFBB0F982E9606C46" author="Armstrong, E. &amp; Bakaloudis, D. E. &amp; Holloway, G. J." box="[1053,1307,1606,1632]" pageId="3" pageNumber="129" refId="ref2636" refString="Armstrong, E., Bakaloudis, D. E. &amp; Holloway, G. J. (2023) A morphometric examination of Anthrenus flavipes flavipes Le Conte 1854 (Coleoptera: Dermestidae: Anthrenini). Bulletin of Insectology. [in press]" type="book" year="2023">
Armstrong
<emphasis id="B97E3017FFB5ED7EFB0DF983E8A26C46" box="[1184,1241,1606,1632]" italics="true" pageId="3" pageNumber="129">et al.</emphasis>
2023
</bibRefCitation>
) since they tend to be larger.
<taxonomicName id="4C0A9786FFB5ED7EFEF4F9AEEF646CA3" authority="(Fabricius, 1775)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[345,799,1642,1669]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="pimpinellae">
<emphasis id="B97E3017FFB5ED7EFEF4F9AEEE2D6CA2" box="[345,598,1642,1668]" italics="true" pageId="3" pageNumber="129">Anthrenus pimpinellae</emphasis>
(Fabricius, 1775)
</taxonomicName>
,
<taxonomicName id="4C0A9786FFB5ED7EFC86F9AEE8116CA2" authority="Holloway, 2019" authorityName="Holloway" authorityYear="2019" box="[811,1130,1642,1668]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="amandae">
<emphasis id="B97E3017FFB5ED7EFC86F9AEEFCA6CA2" box="[811,945,1642,1668]" italics="true" pageId="3" pageNumber="129">A. amandae</emphasis>
Holloway, 2019
</taxonomicName>
(Holloway and Bakaloudis 2020) and
<taxonomicName id="4C0A9786FFB5ED7EFEA3F94AEEF36C8F" authority="(Linnaeus 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[270,648,1678,1705]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="scrophulariae">
<emphasis id="B97E3017FFB5ED7EFEA3F94AEDB26C8E" box="[270,457,1678,1704]" italics="true" pageId="3" pageNumber="129">A. scrophulariae</emphasis>
(Linnaeus 1758)
</taxonomicName>
(Holloway unpublished data) have BW/BL less than 0.703.
<taxonomicName id="4C0A9786FFB5ED7EFA87F94AEE656CEA" authority="Ganglbauer, 1904" authorityName="Ganglbauer" authorityYear="1904" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="angustefasciatus">
<emphasis id="B97E3017FFB5ED7EFA87F94AED296CEA" italics="true" pageId="3" pageNumber="129">Anthrenus angustefasciatus</emphasis>
Ganglbauer, 1904
</taxonomicName>
has BW/BL greater than 0.703 (
<bibRefCitation id="EF9B91F4FFB5ED7EFC2FF976E8A06CEA" author="Holloway, G. J. &amp; Herrmann, A." box="[898,1243,1714,1741]" pageId="3" pageNumber="129" refId="ref3488" refString="Holloway, G. J. &amp; Herrmann, A. (2023) Anthrenus canadaluni sp. nov. new to Turkey and a comparison with Anthrenus angustefasciatus and Anthrenus mroczkowskii. Annales Zoologici. [in press]" type="book" year="2023">Holloway and Herrmann 2023</bibRefCitation>
). The dimensions of
<taxonomicName id="4C0A9786FFB5ED7EFF19F912EE766CD6" authority="Holloway, 2020" authorityName="Holloway" authorityYear="2020" box="[180,525,1750,1776]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="chikatunovi">
<emphasis id="B97E3017FFB5ED7EFF19F912ED2D6CD6" box="[180,342,1750,1776]" italics="true" pageId="3" pageNumber="129">A. chikatunovi</emphasis>
Holloway, 2020
</taxonomicName>
(Holloway 2020),
<taxonomicName id="4C0A9786FFB5ED7EFD72F912EF196CD6" authorityName="Kiesenwetter" authorityYear="1851" box="[735,866,1750,1776]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="B97E3017FFB5ED7EFD72F912EF196CD6" box="[735,866,1750,1776]" italics="true" pageId="3" pageNumber="129">A. delicatus</emphasis>
</taxonomicName>
(Holloway unpublished data),
<taxonomicName id="4C0A9786FFB5ED7EFB17F912ECB46D32" authority="Erichson, 1846" authorityName="Erichson" authorityYear="1846" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="B97E3017FFB5ED7EFB17F912E9576CD6" box="[1210,1324,1750,1776]" italics="true" pageId="3" pageNumber="129">A. festivus</emphasis>
Erichson, 1846
</taxonomicName>
(Holloway 2023), and
<taxonomicName id="4C0A9786FFB5ED7EFE61F93EEEAB6D32" authority="Hinton, 1943" authorityName="Hinton" authorityYear="1943" box="[460,720,1786,1812]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="munroi">
<emphasis id="B97E3017FFB5ED7EFE61F93EEE416D32" box="[460,570,1786,1812]" italics="true" pageId="3" pageNumber="129">A. munroi</emphasis>
Hinton, 1943
</taxonomicName>
(
<bibRefCitation id="EF9B91F4FFB5ED7EFD70F93EE8296D32" author="Holloway, G. J. &amp; Canada Luna, I." box="[733,1106,1786,1813]" pageId="3" pageNumber="129" pagination="289 - 298" refId="ref3356" refString="Holloway, G. J. &amp; Canada Luna, I. (2022) A morphometric analysis of Anthrenus munroi Hinton, 1943, and a key for citizen scientists to the Western European species in the Anthrenus pimpinellae complex (Coleoptera: Dermestidae). The Entomologists Monthly Magazine, 158 (2), 289 - 298. https: // doi. org / 10.31184 / M 00138908.1584.4147" type="journal article" year="2022">
Holloway and
<collectingCountry id="F31DAC95FFB5ED7EFC2DF93EEFAF6D32" box="[896,980,1786,1812]" name="Canada" pageId="3" pageNumber="129">Cañada</collectingCountry>
Luna 2022
</bibRefCitation>
) correspond to
<taxonomicName id="4C0A9786FFB5ED7EFB56F93EE9E76D32" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[1275,1436,1786,1812]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB5ED7EFB56F93EE9E76D32" box="[1275,1436,1786,1812]" italics="true" pageId="3" pageNumber="129">A. semipallens</emphasis>
</taxonomicName>
both in terms of BL and BW/BL. Finally,
<taxonomicName id="4C0A9786FFB5ED7EFDC9F8DAEFEA6D1E" authority="Bedel, 1884" authorityName="Bedel" authorityYear="1884" box="[612,913,1822,1848]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="miniopictus">
<emphasis id="B97E3017FFB5ED7EFDC9F8DAEF7E6D1E" box="[612,773,1822,1848]" italics="true" pageId="3" pageNumber="129">A. miniopictus</emphasis>
Bedel, 1884
</taxonomicName>
is recorded as having BL =
<quantity id="4CF241E0FFB5ED7EFB6AF8DAE9676D1F" box="[1223,1308,1822,1849]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="129" unit="mm" value="2.5">2.5 mm</quantity>
(
<bibRefCitation id="EF9B91F4FFB5ED7EFA84F8DAECAF6D7A" author="Herrmann, A." pageId="3" pageNumber="129" refId="ref2896" refString="Herrmann, A. (2023) Dermestidae (Coleoptera) of the World. Dermestidae (Coleoptera) - Homepage of Andreas Herrmann. Available from: http: // www. dermestidae. com / (accessed 12 April 2023)" type="url" year="2023">Herrmann 2023</bibRefCitation>
).
<taxonomicName id="4C0A9786FFB5ED7EFF4AF886ED9F6D7A" authorityName="Holloway" authorityYear="2020" box="[231,484,1858,1884]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="chikatunovi">
<emphasis id="B97E3017FFB5ED7EFF4AF886ED9F6D7A" box="[231,484,1858,1884]" italics="true" pageId="3" pageNumber="129">Anthrenus chikatunovi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0A9786FFB5ED7EFE5EF886EE036D7A" authorityName="Kiesenwetter" authorityYear="1851" box="[499,632,1858,1884]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="B97E3017FFB5ED7EFE5EF886EE036D7A" box="[499,632,1858,1884]" italics="true" pageId="3" pageNumber="129">A. delicatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0A9786FFB5ED7EFD2AF886EE806D7A" authorityName="Erichson" authorityYear="1846" box="[647,763,1858,1884]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="B97E3017FFB5ED7EFD2AF886EE806D7A" box="[647,763,1858,1884]" italics="true" pageId="3" pageNumber="129">A. festivus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0A9786FFB5ED7EFCA7F886EFD66D7A" authorityName="Bedel" authorityYear="1884" box="[778,941,1858,1884]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="miniopictus">
<emphasis id="B97E3017FFB5ED7EFCA7F886EFD66D7A" box="[778,941,1858,1884]" italics="true" pageId="3" pageNumber="129">A. miniopictus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C0A9786FFB5ED7EFC46F886E8206D7A" authorityName="Hinton" authorityYear="1943" box="[1003,1115,1858,1884]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="munroi">
<emphasis id="B97E3017FFB5ED7EFC46F886E8206D7A" box="[1003,1115,1858,1884]" italics="true" pageId="3" pageNumber="129">A. munroi</emphasis>
</taxonomicName>
all need to be considered as potential confusion species. In addition,
<taxonomicName id="4C0A9786FFB5ED7EFDC0F8A2EF736DA6" box="[621,776,1894,1920]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="isabellinus">
<emphasis id="B97E3017FFB5ED7EFDC0F8A2EF736DA6" box="[621,776,1894,1920]" italics="true" pageId="3" pageNumber="129">A. isabellinus</emphasis>
</taxonomicName>
is included courtesy of superficial external resemblance.
<figureCitation id="1331F080FFB5ED7EFF3AF84EEC826D82" box="[151,249,1930,1956]" captionStart="FIGURE 4" captionStartId="4.[152,255,1934,1958]" captionTargetBox="[162,1435,181,1910]" captionTargetId="figure-17@4.[151,1436,181,1910]" captionTargetPageId="4" captionText="FIGURE 4. Aedeagi. A: Anthrenus chikatunovi, B: Anthrenus delicatus, C: Anthrenus festivus, D: Anthrenus isabellinus. E: Anthrenus miniopictus, F: Anthrenus munroi. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.8204059" httpUri="https://zenodo.org/record/8204059/files/figure.png" pageId="3" pageNumber="129">Figure 4</figureCitation>
shows the aedeagi of these six species. None of them resemble the aedeagus shown in
<figureCitation id="1331F080FFB5ED7EFAA8F84EE9E36D82" box="[1285,1432,1930,1956]" captionStart="FIGURE 2" captionStartId="2.[152,255,1919,1943]" captionTargetBox="[151,1436,934,1896]" captionTargetId="figure-45@2.[151,1436,933,1896]" captionTargetPageId="2" captionText="FIGURE 2. Anthrenus semipallens sp. nov. Holotype. A: Antenna, B: aedeagus ventral aspect, C: sternite IX. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.8204057" httpUri="https://zenodo.org/record/8204057/files/figure.png" pageId="3" pageNumber="129">
Figure 2B
<taxonomicName id="4C0A9786FFB5ED7EFA29F84EE9E36D82" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[1412,1432,1930,1956]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB5ED7EFA29F84EE9E36D82" box="[1412,1432,1930,1956]" italics="true" pageId="3" pageNumber="129">A</emphasis>
</taxonomicName>
</figureCitation>
. semipallens is a valid species.
</paragraph>
<paragraph id="8BB5EC05FFB5ED7EFF6AF816E9F26DCA" blockId="3.[151,1437,1426,2029]" box="[199,1417,2002,2029]" pageId="3" pageNumber="129">
Given these results,
<tableCitation id="C688D9BEFFB5ED7EFE07F816ED866DCA" box="[426,509,2002,2028]" captionStart="TABLE 1" captionStartId="5.[152,240,150,174]" captionTargetBox="[163,1426,226,1034]" captionText="TABLE 1. Brief key to separate A. semipallens from other Anthrenus spp. occurring in Spain according to Holloway et al." httpUri="http://table.plazi.org/id/DF75BC8DFFB3ED78FF35FF52E9E76A88" pageId="3" pageNumber="129" tableUuid="DF75BC8DFFB3ED78FF35FF52E9E76A88">Table 1</tableCitation>
shows a key to the
<taxonomicName id="4C0A9786FFB5ED7EFD74F816EF306DCA" authorityName="Geoffroy" authorityYear="1762" box="[729,843,2002,2028]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B97E3017FFB5ED7EFD74F816EF306DCA" box="[729,843,2002,2028]" italics="true" pageId="3" pageNumber="129">Anthrenus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D8D6CFFB5ED7EFCFFF817EFF66DCB" box="[850,909,2003,2029]" pageId="3" pageNumber="129" sensu="stricto">s. str.</taxonomicNameLabel>
in
<collectingCountry id="F31DAC95FFB5ED7EFC1CF816EF8A6DCA" box="[945,1009,2002,2028]" name="Spain" pageId="3" pageNumber="129">Spain</collectingCountry>
according to
<bibRefCitation id="EF9B91F4FFB5ED7EFB27F816E9F26DCA" author="Holloway, G. J. &amp; Canada Luna, I. &amp; Kadej, M." box="[1162,1417,2002,2029]" pageId="3" pageNumber="129" pagination="1 - 8" refId="ref3428" refString="Holloway, G. J., Canada Luna, I. &amp; Kadej, M. (2019) A review of the literature and a checklist of the Spanish Dermestidae (Coleoptera). The Coleopterists Bulletin, 73, 1 - 8. https: // doi. org / 10.1649 / 0010 - 065 X- 73.3.513" type="journal article" year="2019">
Holloway
<emphasis id="B97E3017FFB5ED7EFB52F817E94B6DCA" box="[1279,1328,2002,2028]" italics="true" pageId="3" pageNumber="129">et al</emphasis>
. (2019)
</bibRefCitation>
</paragraph>
<caption id="DF75BC8DFFB2ED79FF35F84AEF466DED" ID-DOI="http://doi.org/10.5281/zenodo.8204059" ID-Zenodo-Dep="8204059" httpUri="https://zenodo.org/record/8204059/files/figure.png" pageId="4" pageNumber="130" startId="4.[152,255,1934,1958]" targetBox="[162,1435,181,1910]" targetPageId="4" targetType="figure">
<paragraph id="8BB5EC05FFB2ED79FF35F84AEF466DED" blockId="4.[151,1437,1934,1995]" pageId="4" pageNumber="130">
<emphasis id="B97E3017FFB2ED79FF35F84AED616D80" bold="true" box="[152,282,1934,1958]" pageId="4" pageNumber="130">FIGURE 4.</emphasis>
Aedeagi. A:
<taxonomicName id="4C0A9786FFB2ED79FE09F84AEEF26D80" authorityName="Holloway" authorityYear="2020" box="[420,649,1934,1958]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="chikatunovi">
<emphasis id="B97E3017FFB2ED79FE09F84AEEF26D80" box="[420,649,1934,1958]" italics="true" pageId="4" pageNumber="130">Anthrenus chikatunovi</emphasis>
</taxonomicName>
, B:
<taxonomicName id="4C0A9786FFB2ED79FD15F84AEFF96D80" authorityName="Kiesenwetter" authorityYear="1851" box="[696,898,1934,1958]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="B97E3017FFB2ED79FD15F84AEFF96D80" box="[696,898,1934,1958]" italics="true" pageId="4" pageNumber="130">Anthrenus delicatus</emphasis>
</taxonomicName>
, C:
<taxonomicName id="4C0A9786FFB2ED79FC1DF84AE8106D80" authorityName="Erichson" authorityYear="1846" box="[944,1131,1934,1958]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="B97E3017FFB2ED79FC1DF84AE8106D80" box="[944,1131,1934,1958]" italics="true" pageId="4" pageNumber="130">Anthrenus festivus</emphasis>
</taxonomicName>
, D:
<taxonomicName id="4C0A9786FFB2ED79FB36F84AE9036D80" box="[1179,1400,1934,1958]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="isabellinus">
<emphasis id="B97E3017FFB2ED79FB36F84AE9036D80" box="[1179,1400,1934,1958]" italics="true" pageId="4" pageNumber="130">Anthrenus isabellinus</emphasis>
</taxonomicName>
. E:
<taxonomicName id="4C0A9786FFB2ED79FF3AF876ED006DEC" authorityName="Bedel" authorityYear="1884" box="[151,379,1970,1994]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="miniopictus">
<emphasis id="B97E3017FFB2ED79FF3AF876ED006DEC" box="[151,379,1970,1994]" italics="true" pageId="4" pageNumber="130">Anthrenus miniopictus</emphasis>
</taxonomicName>
, F:
<taxonomicName id="4C0A9786FFB2ED79FE0EF876EE226DEC" authorityName="Hinton" authorityYear="1943" box="[419,601,1970,1994]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="munroi">
<emphasis id="B97E3017FFB2ED79FE0EF876EE226DEC" box="[419,601,1970,1994]" italics="true" pageId="4" pageNumber="130">Anthrenus munroi</emphasis>
</taxonomicName>
. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="DF75BC8DFFB3ED78FF35FF52E9E76A88" ID-Table-UUID="DF75BC8DFFB3ED78FF35FF52E9E76A88" box="[152,1436,150,174]" httpUri="http://table.plazi.org/id/DF75BC8DFFB3ED78FF35FF52E9E76A88" pageId="5" pageNumber="131" startId="5.[152,240,150,174]" targetBox="[163,1426,226,1034]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="8BB5EC05FFB3ED78FF35FF52E9E76A88" blockId="5.[151,1436,150,210]" box="[152,1436,150,174]" pageId="5" pageNumber="131">
<emphasis id="B97E3017FFB3ED78FF35FF52ED706A88" bold="true" box="[152,267,150,174]" pageId="5" pageNumber="131">TABLE 1.</emphasis>
Brief key to separate
<taxonomicName id="4C0A9786FFB3ED78FE58FF53EEF06A88" authorityName="Holloway &amp; Herrmann" authorityYear="2023" box="[501,651,150,174]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="131" phylum="Arthropoda" rank="species" species="semipallens">
<emphasis id="B97E3017FFB3ED78FE58FF53EEF06A88" box="[501,651,150,174]" italics="true" pageId="5" pageNumber="131">A. semipallens</emphasis>
</taxonomicName>
from other
<taxonomicName id="4C0A9786FFB3ED78FCA4FF52EFD86A88" box="[777,931,150,174]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="131" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97E3017FFB3ED78FCA4FF52EF0B6A88" box="[777,880,150,174]" italics="true" pageId="5" pageNumber="131">Anthrenus</emphasis>
spp.
</taxonomicName>
occurring in Spain according to Holloway
<emphasis id="B97E3017FFB3ED78FACAFF53E9ED6A88" box="[1383,1430,150,174]" italics="true" pageId="5" pageNumber="131">et al</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BB5EC05FFB3ED78FF3AFF7EEC996AF4" blockId="5.[151,1436,150,210]" box="[151,226,186,210]" pageId="5" pageNumber="131">(2019).</paragraph>
</subSubSection>
</treatment>
</document>