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<document id="3AD16347FAD8552B8A311027FBC2DCBC" ID-CLB-Dataset="27764" ID-DOI="10.11646/zootaxa.4532.1.1" ID-GBIF-Dataset="b3fb460f-20b4-4c62-98ea-a07d901bbfc0" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614967" ID-ZooBank="4E9D0908-0933-48AF-A6ED-F3B8D39E8994" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553787614041" checkinUser="plazi" docAuthor="Cairns, Stephen D." docDate="2018" docId="03B0147FFFFCFFD676CC6B52426FF9E5" docLanguage="en" docName="zootaxa.4532.1.1.pdf" docOrigin="Zootaxa 4532 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macroprimnoa ornata Cairns 2018, n. sp." docType="treatment" docVersion="13" lastPageNumber="19" masterDocId="FF896C07FFEEFFC4765B6A1F4732FF92" masterDocTitle="Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific" masterLastPageNumber="43" masterPageNumber="1" pageNumber="19" updateTime="1698714965598" updateUser="ExternalLinkService">
<mods:mods id="B0F3049A7A4CB47A3B82E6AB4F21F2FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8D2429432A3080DE546E65A9D84CB42C">
<mods:title id="4E7FFB3233EE1BE221338463554A0375">Primnoidae (Cnidaria: Octocorallia: Calcaxonia) of the Okeanos Explorer expeditions (CAPSTONE) to the central Pacific</mods:title>
</mods:titleInfo>
<mods:name id="59E4664B4BE105F733E8EE13E8EF8A7E" type="personal">
<mods:role id="C363CB8A4C6FD39E8564D40A76A48A57">
<mods:roleTerm id="60BF91DBE3AC3E7C8329577E8EF522E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9EB91E08DE427F8634A7086B29319F03">Cairns, Stephen D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5E04C570778FD3C0E1546563D7B90C3A">text</mods:typeOfResource>
<mods:relatedItem id="102161485C44093888763E407DE3E45B" type="host">
<mods:titleInfo id="676D42AEB9EB36360A9259FECF79D89E">
<mods:title id="312E69D2CB1748CD1BE1C6460B3BDD38">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5B670CE7CD42F146F640C55B72FE6D1A">
<mods:date id="BBBEADB8327F86124873657DFD399DB2">2018</mods:date>
<mods:detail id="E7C437082E89E42F0FC66B009D1E02AF" type="pubDate">
<mods:number id="5253DD1573E3FB45E0465C8747A39225">2018-12-17</mods:number>
</mods:detail>
<mods:detail id="ECEDC6EC1306A8DC9F2F2F448D05B1A1" type="volume">
<mods:number id="BF4DD86BF2D5C25ADA4E472737A554FC">4532</mods:number>
</mods:detail>
<mods:detail id="427C4F0F46CDB1D3B6643D42E261A002" type="issue">
<mods:number id="D03045F2BCB8D5BCE5D624F1E1B8CC3E">1</mods:number>
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<mods:start id="B1C746BD1C730D981A371C1128EC07E2">1</mods:start>
<mods:end id="BF173024215CD20ACDD57CF06852AE7A">43</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F24C0CCAC73D42AD37B7C73D5C7A99E5">journal article</mods:classification>
<mods:identifier id="CB47220EF0968C77E092694C0DA0A45F" type="CLB-Dataset">27764</mods:identifier>
<mods:identifier id="8E7952F70F7BFF9B2EA91FC8A8B06125" type="DOI">10.11646/zootaxa.4532.1.1</mods:identifier>
<mods:identifier id="F58DFE4AF0F61E6AEB967D2382EC6940" type="GBIF-Dataset">b3fb460f-20b4-4c62-98ea-a07d901bbfc0</mods:identifier>
<mods:identifier id="AE95F7DACAAE69B8E1BB54D9D832FFFF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9FB48DC68BCFF135BBD3233D1A82CABA" type="Zenodo-Dep">2614967</mods:identifier>
<mods:identifier id="F86C3C77D77D4A6D587420CC228A77B2" type="ZooBank">4E9D0908-0933-48AF-A6ED-F3B8D39E8994</mods:identifier>
</mods:mods>
<treatment id="03B0147FFFFCFFD676CC6B52426FF9E5" ID-DOI="http://doi.org/10.5281/zenodo.5951519" ID-GBIF-Taxon="154521079" ID-Zenodo-Dep="5951519" LSID="urn:lsid:plazi:treatment:03B0147FFFFCFFD676CC6B52426FF9E5" httpUri="http://treatment.plazi.org/id/03B0147FFFFCFFD676CC6B52426FF9E5" lastPageNumber="19" pageId="18" pageNumber="19">
<subSubSection id="C303F6E2FFFCFFD676CC6B5246C4FEF5" box="[151,502,333,359]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BA6A569FFFCFFD676CC6B5246C4FEF5" blockId="18.[151,502,333,395]" box="[151,502,333,359]" pageId="18" pageNumber="19">
<heading id="D0EE1205FFFCFFD676CC6B5246C4FEF5" bold="true" box="[151,502,333,359]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B96D797BFFFCFFD676CC6B5246C4FEF5" bold="true" box="[151,502,333,359]" pageId="18" pageNumber="19">
<taxonomicName id="4C19DEEAFFFCFFD676CC6B524697FEF5" ID-CoL="3X8PC" authority="Cairns, 2018" authorityName="Cairns" authorityYear="2018" box="[151,421,333,359]" class="Anthozoa" family="Primnoidae" genus="Macroprimnoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B96D797BFFFCFFD676CC6B524697FEF5" bold="true" box="[151,421,333,359]" italics="true" pageId="18" pageNumber="19">Macroprimnoa ornata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25EC400FFFCFFD677EF6B5246C4FEF5" box="[436,502,333,359]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFCFFD676CC6B6E460DFE19" box="[151,319,369,395]" pageId="18" pageNumber="19" type="description">
<paragraph id="8BA6A569FFFCFFD676CC6B6E460DFE19" blockId="18.[151,502,333,395]" box="[151,319,369,395]" pageId="18" pageNumber="19">
<figureCitation id="1322B9ECFFFCFFD676CC6B6E47DCFE19" box="[151,238,369,395]" captionStart="FIGURE 1" captionStartId="3.[151,250,1774,1799]" captionTargetBox="[174,1414,166,1753]" captionTargetId="figure@3.[174,1414,166,1753]" captionTargetPageId="3" captionText="FIGURE 1. In situ images of various species: A, Callogorgia cracentis, holotype. B, Calyptrophora lyra, holotype (not in situ). C, C. diaphana, EX1606-9-1. D, C. carinata, holotype. E, C. clarki, EX1605-L3-6. F, C. pourtalesi, holotype. G, C. distolos, holotype. H, Candidella gigantea, EX1606-6. I, Macroprimnoa ornata, holotype. J, Narella virgosa, holotype. K-L, N. aurantiaca, holotype. M, N. merga, holotype. N, N. horrida, EX1703-2-3. O, N. fordi, holotype. P, N. calamus, holotype. Q, N. virgosa, paratype, EX1504-L4-9-4. R, N. macrocalyx, EX1606-7. S, N. hawaiiensis, EX1504-9-2. T, Paracalyotrophora hawaiiensis, EX1703-2-1. U, P. echinata, EX1703-10-1. V, P. spiralis, holotype. W, Plumarella circumoperculum, EX1603-2-3. X, Thouarella hilgendorfi, EX1605-L1-17." httpUri="https://zenodo.org/record/2614969/files/figure.png" pageId="18" pageNumber="19">
Figs.
<date id="FFA783A9FFFCFFD6768E6B6E47DCFE19" box="[213,238,369,395]" pageId="18" pageNumber="19">1I</date>
</figureCitation>
,
<figureCitation id="1322B9ECFFFCFFD676A06B6E463AFE19" box="[251,264,369,395]" captionStart="FIGURE 9" captionStartId="19.[151,250,1758,1783]" captionTargetBox="[174,1414,151,1738]" captionTargetId="figure@19.[174,1414,151,1738]" captionTargetPageId="19" captionText="FIGURE 9. Macroprimnoa ornata, paratype from USNM 142423: A–C, stereo views of lateral and opercular sides of a polyp. D, stereo view of coenenchymal scales, in situ. E, body wall scales, in situ. F, highly ridged body wall scales. G, opercular scales. H, ridged coenenchymal scales. Scale bars in mm." httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="18" pageNumber="19">9</figureCitation>
A–H
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFCFFD676CC6BA647C1FD5D" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BA6A569FFFCFFD676CC6BA6462BFE65" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD676CC6BA6462AFE41" bold="true" box="[151,280,441,467]" pageId="18" pageNumber="19">Etymology</emphasis>
. Named
<taxonomicName id="4C19DEEAFFFCFFD677276BA546F5FE41" authorityName="Cairns" authorityYear="2018" box="[380,455,442,467]" class="Anthozoa" family="Primnoidae" genus="Macroprimnoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="19" phylum="Cnidaria" rank="species" species="ornata">
<emphasis id="B96D797BFFFCFFD677276BA546F5FE41" box="[380,455,442,467]" italics="true" pageId="18" pageNumber="19">ornata</emphasis>
</taxonomicName>
(Latin for decorated or adorned), an allusion to the somewhat ornate ridging on the body wall scales.
</paragraph>
<paragraph id="8BA6A569FFFCFFD6769C681E47C1FD5D" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD6769C681E46DFFD89" bold="true" box="[199,493,512,539]" pageId="18" pageNumber="19">Types and Type Locality</emphasis>
.
<materialsCitation id="3B71AF34FFFCFFD677A0681E433EFD89" ID-GBIF-Occurrence="2117750060" box="[507,1036,512,539]" collectionCode="USNM" pageId="18" pageNumber="19" specimenCode="USNM 1463530" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A21BCBFFFCFFD677A0681E455AFD89" box="[507,616,513,539]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
: a complete colony,
<specimenCode id="DBBF0D12FFFCFFD67515681E433EFD89" box="[846,1036,512,539]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 1463530</specimenCode>
</materialsCitation>
.
<typeStatus id="54A21BCBFFFCFFD6724C681E43BAFD89" box="[1047,1160,513,539]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B96D797BFFFCFFD672C8681D4387FD89" box="[1171,1205,514,539]" italics="true" pageId="18" pageNumber="19">EX</emphasis>
<date id="FFA783A9FFFCFFD672EE681E422DFD89" box="[1205,1311,513,539]" pageId="18" pageNumber="19" value="1708-01-02">1708-2-1</date>
, 25.553˚N, 158.214˚W (topotypic),
<quantity id="4CE1088CFFFCFFD677FD683A46CFFDAD" box="[422,509,549,575]" metricMagnitude="3" metricUnit="m" metricValue="3.282" pageId="18" pageNumber="19" unit="m" value="3282.0">3282 m</quantity>
,
<materialsCitation id="3B71AF34FFFCFFD67452683A4464FDF1" ID-GBIF-Occurrence="2117771105" collectingDate="2005-04-01" collectionCode="USNM" country="Guam" elevation="3643" location="Enigma Seamount" pageId="18" pageNumber="19" specimenCode="USNM 1467528" specimenCount="1" typeStatus="paratype">
colony fragment,
<specimenCode id="DBBF0D12FFFCFFD67494683A44BFFDAD" box="[719,909,548,575]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 1467528</specimenCode>
;
<emphasis id="B96D797BFFFCFFD675CC6839448BFDAD" box="[919,953,550,575]" italics="true" pageId="18" pageNumber="19">EX</emphasis>
1605-L
<collectingDate id="EFE37A41FFFCFFD6724A683B436BFDAD" box="[1041,1113,548,575]" pageId="18" pageNumber="19" value="2005-04-01">
<date id="FFA783A9FFFCFFD6724A683B436BFDAD" box="[1041,1113,548,575]" pageId="18" pageNumber="19" value="2005-04-01">1-4-05</date>
</collectingDate>
, 11.41˚N, 144.78˚E (
<location id="8EC6F3B2FFFCFFD67318683A4638FDF1" LSID="urn:lsid:plazi:treatment:03B0147FFFFCFFD676CC6B52426FF9E5:8EC6F3B2FFFCFFD67318683A4638FDF1" country="Guam" name="Enigma Seamount" pageId="18" pageNumber="19">Enigma Seamount</location>
, south of
<collectingCountry id="F30EE5F9FFFCFFD6772C6856468CFDF1" box="[375,446,585,611]" name="Guam" pageId="18" pageNumber="19">Guam</collectingCountry>
,
<collectingCountry id="F30EE5F9FFFCFFD67792685645DEFDF6" box="[457,748,585,612]" name="Northern Mariana Islands" pageId="18" pageNumber="19">Northern Mariana Islands</collectingCountry>
),
<elevation id="0034425AFFFCFFD6755B68564464FDF1" box="[768,854,585,611]" metricMagnitude="3" metricUnit="m" metricValue="3.643" pageId="18" pageNumber="19" unit="m" value="3643.0">
<quantity id="4CE1088CFFFCFFD6755B68564464FDF1" box="[768,854,585,611]" metricMagnitude="3" metricUnit="m" metricValue="3.643" pageId="18" pageNumber="19" unit="m" value="3643.0">3643 m</quantity>
</elevation>
</materialsCitation>
,
<materialsCitation id="3B71AF34FFFCFFD67539685644FAFD15" ID-GBIF-Occurrence="2117750024" collectingDate="1606-03-13" collectionCode="USNM" country="Wake Island" elevation="3066" location="Batfish Seamount" pageId="18" pageNumber="19" specimenCode="USNM 1424086" specimenCount="1" typeStatus="paratype">
colony fragment,
<specimenCode id="DBBF0D12FFFCFFD67273685643D4FDF1" box="[1064,1254,585,611]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 1424086</specimenCode>
;
<emphasis id="B96D797BFFFCFFD672AB68554220FDF1" box="[1264,1298,586,611]" italics="true" pageId="18" pageNumber="19">EX</emphasis>
<collectingDate id="EFE37A41FFFCFFD67348685642AAFDF1" box="[1299,1432,585,611]" pageId="18" pageNumber="19" value="1606-03-13">
<date id="FFA783A9FFFCFFD67348685642AAFDF1" box="[1299,1432,585,611]" pageId="18" pageNumber="19" value="1606-03-13">1606-13-03</date>
</collectingDate>
, 16.581˚N, 166.472˚E (
<location id="8EC6F3B2FFFCFFD677CF68724553FD15" LSID="urn:lsid:plazi:treatment:03B0147FFFFCFFD676CC6B52426FF9E5:8EC6F3B2FFFCFFD677CF68724553FD15" box="[404,609,621,647]" country="Wake Island" name="Batfish Seamount" pageId="18" pageNumber="19">Batfish Seamount</location>
, south of
<collectingCountry id="F30EE5F9FFFCFFD674956872446FFD1A" box="[718,861,621,648]" name="Wake Island" pageId="18" pageNumber="19">Wake Island</collectingCountry>
),
<elevation id="0034425AFFFCFFD6752B687244FAFD15" box="[880,968,621,647]" metricMagnitude="3" metricUnit="m" metricValue="3.066" pageId="18" pageNumber="19" unit="m" value="3066.0">
<quantity id="4CE1088CFFFCFFD6752B687244FAFD15" box="[880,968,621,647]" metricMagnitude="3" metricUnit="m" metricValue="3.066" pageId="18" pageNumber="19" unit="m" value="3066.0">3066 m</quantity>
</elevation>
</materialsCitation>
,
<materialsCitation id="3B71AF34FFFCFFD6758F687247DDFD5D" ID-GBIF-Occurrence="2117750047" collectingDate="2386" collectingDateMax="2388" collectingDateMin="2386" collectionCode="SEM" country="Wake Island" elevation="3207" location="Beach Ridge" municipality="Musicians Seamounts" pageId="18" pageNumber="19" specimenCode="USNM 1424231" specimenCount="1" typeStatus="paratype">
colony fragment,
<collectionCode id="ED083DACFFFCFFD672C1687243E6FD15" box="[1178,1236,621,647]" pageId="18" pageNumber="19">SEM</collectionCode>
stubs
<collectingDate id="EFE37A41FFFCFFD67347687242AAFD15" box="[1308,1432,621,647]" pageId="18" pageNumber="19" value="2386" valueMax="2388">
<date id="FFA783A9FFFCFFD67347687242AAFD15" box="[1308,1432,621,647]" pageId="18" pageNumber="19" value="2386" valueMax="2388">2386-2388</date>
</collectingDate>
,
<specimenCode id="DBBF0D12FFFCFFD676CC688E4661FD39" box="[151,339,657,683]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 1424231</specimenCode>
. Type Locality:
<emphasis id="B96D797BFFFCFFD67459688D4516FD39" box="[514,548,658,683]" italics="true" pageId="18" pageNumber="19">EX</emphasis>
1708-2-GEO2-A2, 25.553˚N, 158.215˚W (
<location id="8EC6F3B2FFFCFFD675A6688E43A2FD39" LSID="urn:lsid:plazi:treatment:03B0147FFFFCFFD676CC6B52426FF9E5:8EC6F3B2FFFCFFD675A6688E43A2FD39" box="[1021,1168,657,683]" country="Wake Island" municipality="Musicians Seamounts" name="Beach Ridge" pageId="18" pageNumber="19">Beach Ridge</location>
,
<collectingMunicipality id="6BC23F13FFFCFFD672C2688E42A2FD39" box="[1177,1424,657,683]" pageId="18" pageNumber="19">Musicians Seamounts</collectingMunicipality>
),
<elevation id="0034425AFFFCFFD676CC68AA47DDFD5D" box="[151,239,693,719]" metricMagnitude="3" metricUnit="m" metricValue="3.207" pageId="18" pageNumber="19" unit="m" value="3207.0">
<quantity id="4CE1088CFFFCFFD676CC68AA47DDFD5D" box="[151,239,693,719]" metricMagnitude="3" metricUnit="m" metricValue="3.207" pageId="18" pageNumber="19" unit="m" value="3207.0">3207 m</quantity>
</elevation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFCFFD6769C68C7460CFC85" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BA6A569FFFCFFD6769C68C7460CFC85" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD6769C68C7469CFD61" bold="true" box="[199,430,728,755]" pageId="18" pageNumber="19">Material Examined</emphasis>
.
<materialsCitation id="3B71AF34FFFCFFD677E168C64608FC85" ID-GBIF-Occurrence="2117750071" country="Northern Mariana Islands" elevation="3676" location="Saipan" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Saipan" typeStatus="holotype">
<typeStatus id="54A21BCBFFFCFFD677E168C64532FD61" box="[442,512,729,755]" pageId="18" pageNumber="19">Types</typeStatus>
;
<emphasis id="B96D797BFFFCFFD6745168C5451EFD61" box="[522,556,730,755]" italics="true" pageId="18" pageNumber="19">EX</emphasis>
1605-L1-13, 16.027˚N, 147.113˚E (off
<collectingRegion id="49DD6B8BFFFCFFD6758468C6431CFD61" box="[991,1070,729,755]" country="Northern Mariana Islands" name="Saipan" pageId="18" pageNumber="19">Saipan</collectingRegion>
,
<collectingCountry id="F30EE5F9FFFCFFD6726368C64260FD66" box="[1080,1362,729,756]" name="Northern Mariana Islands" pageId="18" pageNumber="19">northern Mariana Islands</collectingCountry>
),
<elevation id="0034425AFFFCFFD6733F68C6479DFC85" metricMagnitude="3" metricUnit="m" metricValue="3.676" pageId="18" pageNumber="19" unit="m" value="3676.0">
<quantity id="4CE1088CFFFCFFD6733F68C6479DFC85" metricMagnitude="3" metricUnit="m" metricValue="3.676" pageId="18" pageNumber="19" unit="m" value="3676.0">3676 m</quantity>
</elevation>
, image only
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFCFFD6769C693F46C4FA75" pageId="18" pageNumber="19" type="description">
<paragraph id="8BA6A569FFFCFFD6769C693F4552FC7D" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD6769C693F4663FCA9" bold="true" box="[199,337,800,827]" pageId="18" pageNumber="19">Description</emphasis>
. Colonies are unbranched (flagelliform), the
<typeStatus id="54A21BCBFFFCFFD67506693E448DFCA9" box="[861,959,801,827]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
(
<figureCitation id="1322B9ECFFFCFFD6758A693E4312FCA9" box="[977,1056,801,827]" captionStart="FIGURE 1" captionStartId="3.[151,250,1774,1799]" captionTargetBox="[174,1414,166,1753]" captionTargetId="figure@3.[174,1414,166,1753]" captionTargetPageId="3" captionText="FIGURE 1. In situ images of various species: A, Callogorgia cracentis, holotype. B, Calyptrophora lyra, holotype (not in situ). C, C. diaphana, EX1606-9-1. D, C. carinata, holotype. E, C. clarki, EX1605-L3-6. F, C. pourtalesi, holotype. G, C. distolos, holotype. H, Candidella gigantea, EX1606-6. I, Macroprimnoa ornata, holotype. J, Narella virgosa, holotype. K-L, N. aurantiaca, holotype. M, N. merga, holotype. N, N. horrida, EX1703-2-3. O, N. fordi, holotype. P, N. calamus, holotype. Q, N. virgosa, paratype, EX1504-L4-9-4. R, N. macrocalyx, EX1606-7. S, N. hawaiiensis, EX1504-9-2. T, Paracalyotrophora hawaiiensis, EX1703-2-1. U, P. echinata, EX1703-10-1. V, P. spiralis, holotype. W, Plumarella circumoperculum, EX1603-2-3. X, Thouarella hilgendorfi, EX1605-L1-17." httpUri="https://zenodo.org/record/2614969/files/figure.png" pageId="18" pageNumber="19">
Fig.
<date id="FFA783A9FFFCFFD67252693E4312FCA9" box="[1033,1056,801,827]" pageId="18" pageNumber="19">1I</date>
</figureCitation>
) an intact stem
<quantity id="4CE1088CFFFCFFD67285693E421FFCA9" box="[1246,1325,801,827]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" pageId="18" pageNumber="19" unit="cm" value="8.7">8.7 cm</quantity>
in height supporting 12 polyps. A larger colony (USNM 1424086,
<typeStatus id="54A21BCBFFFCFFD67570695A44BFFCCD" box="[811,909,837,863]" pageId="18" pageNumber="19" type="paratype">paratype</typeStatus>
) was estimated to have been
<quantity id="4CE1088CFFFCFFD672BE695A421FFCCD" box="[1253,1325,837,863]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="19" unit="cm" value="27.0">27 cm</quantity>
in height at the time of collection). The polyps are uniserially arranged on one side of the stem and very widely spaced (1 polyp/cm), about
<quantity id="4CE1088CFFFCFFD6770669924688FC35" box="[349,442,909,935]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="18" pageNumber="19" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
of branch coenenchyme between each polyp (
<figureCitation id="1322B9ECFFFCFFD675E569924338FC35" box="[958,1034,909,935]" captionStart="FIGURE 1" captionStartId="3.[151,250,1774,1799]" captionTargetBox="[174,1414,166,1753]" captionTargetId="figure@3.[174,1414,166,1753]" captionTargetPageId="3" captionText="FIGURE 1. In situ images of various species: A, Callogorgia cracentis, holotype. B, Calyptrophora lyra, holotype (not in situ). C, C. diaphana, EX1606-9-1. D, C. carinata, holotype. E, C. clarki, EX1605-L3-6. F, C. pourtalesi, holotype. G, C. distolos, holotype. H, Candidella gigantea, EX1606-6. I, Macroprimnoa ornata, holotype. J, Narella virgosa, holotype. K-L, N. aurantiaca, holotype. M, N. merga, holotype. N, N. horrida, EX1703-2-3. O, N. fordi, holotype. P, N. calamus, holotype. Q, N. virgosa, paratype, EX1504-L4-9-4. R, N. macrocalyx, EX1606-7. S, N. hawaiiensis, EX1504-9-2. T, Paracalyotrophora hawaiiensis, EX1703-2-1. U, P. echinata, EX1703-10-1. V, P. spiralis, holotype. W, Plumarella circumoperculum, EX1603-2-3. X, Thouarella hilgendorfi, EX1605-L1-17." httpUri="https://zenodo.org/record/2614969/files/figure.png" pageId="18" pageNumber="19">
Fig.
<date id="FFA783A9FFFCFFD675A969924338FC35" box="[1010,1034,909,935]" pageId="18" pageNumber="19">1I</date>
</figureCitation>
). The polyps stand perpendicular to the branch, are
<quantity id="4CE1088CFFFCFFD6771E69AF46FFFC59" box="[325,461,944,971]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="18" pageNumber="19" unit="mm" value="3.6" valueMax="3.7" valueMin="3.5">3.5–3.7 mm</quantity>
in height, and are shaped as a squat cylinders topped with a conical operculum, which accounts for well over half of the height.
</paragraph>
<paragraph id="8BA6A569FFFCFFD6769C69E646B1FB31" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
The body wall scales consist of eight longitudinal rows or two or three roughly rectangular (
<quantity id="4CE1088CFFFCFFD672A869E7424CFB81" box="[1267,1406,1016,1043]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="18" pageNumber="19" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.8–2.5 mm</quantity>
in width) scales, resulting in only 16–24 body wall scales (
<figureCitation id="1322B9ECFFFCFFD675456E024450FBA5" box="[798,866,1053,1079]" captionStart="FIGURE 9" captionStartId="19.[151,250,1758,1783]" captionTargetBox="[174,1414,151,1738]" captionTargetId="figure@19.[174,1414,151,1738]" captionTargetPageId="19" captionText="FIGURE 9. Macroprimnoa ornata, paratype from USNM 142423: A–C, stereo views of lateral and opercular sides of a polyp. D, stereo view of coenenchymal scales, in situ. E, body wall scales, in situ. F, highly ridged body wall scales. G, opercular scales. H, ridged coenenchymal scales. Scale bars in mm." httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
A–B). The body wall scales have a sinuous upper margin, sometimes similar to a
<emphasis id="B96D797BFFFCFFD674586E5E45A0FBC9" box="[515,658,1089,1115]" italics="true" pageId="18" pageNumber="19">Parastenella</emphasis>
-like flute (
<figureCitation id="1322B9ECFFFCFFD6754F6E5E4459FBC9" box="[788,875,1089,1115]" captionStart="FIGURE 9" captionStartId="19.[151,250,1758,1783]" captionTargetBox="[174,1414,151,1738]" captionTargetId="figure@19.[174,1414,151,1738]" captionTargetPageId="19" captionText="FIGURE 9. Macroprimnoa ornata, paratype from USNM 142423: A–C, stereo views of lateral and opercular sides of a polyp. D, stereo view of coenenchymal scales, in situ. E, body wall scales, in situ. F, highly ridged body wall scales. G, opercular scales. H, ridged coenenchymal scales. Scale bars in mm." httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="18" pageNumber="19">Fig. 9F</figureCitation>
), and those nearest the branch often bear several vertical ridges on their outer surface (
<figureCitation id="1322B9ECFFFCFFD6746F6E7A45BFFBED" box="[564,653,1125,1151]" captionStart="FIGURE 9" captionStartId="19.[151,250,1758,1783]" captionTargetBox="[174,1414,151,1738]" captionTargetId="figure@19.[174,1414,151,1738]" captionTargetPageId="19" captionText="FIGURE 9. Macroprimnoa ornata, paratype from USNM 142423: A–C, stereo views of lateral and opercular sides of a polyp. D, stereo view of coenenchymal scales, in situ. E, body wall scales, in situ. F, highly ridged body wall scales. G, opercular scales. H, ridged coenenchymal scales. Scale bars in mm." httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="18" pageNumber="19">Fig. 9H</figureCitation>
). The proximal margin of the body wall scales bear a series or irregular digitiform processes.
</paragraph>
<paragraph id="8BA6A569FFFCFFD6769C6EB24250FAC5" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
The operculum is prominent (
<figureCitation id="1322B9ECFFFCFFD6744B6EB24563FB55" box="[528,593,1197,1223]" captionStart="FIGURE 9" captionStartId="19.[151,250,1758,1783]" captionTargetBox="[174,1414,151,1738]" captionTargetId="figure@19.[174,1414,151,1738]" captionTargetPageId="19" captionText="FIGURE 9. Macroprimnoa ornata, paratype from USNM 142423: A–C, stereo views of lateral and opercular sides of a polyp. D, stereo view of coenenchymal scales, in situ. E, body wall scales, in situ. F, highly ridged body wall scales. G, opercular scales. H, ridged coenenchymal scales. Scale bars in mm." httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
A–B), composing over half the height of the polyp. The symmetrical abaxial operculars are up to
<quantity id="4CE1088CFFFCFFD677246ECE46E4FB79" box="[383,470,1233,1259]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="18" pageNumber="19" unit="mm" value="3.1">3.1 mm</quantity>
in length, have an L:W of 2.6–3.1, and have a pointed tip and irregular lateral edges. The asymmetrical lateral operculars are only slightly shorter (
<quantity id="4CE1088CFFFCFFD675396EEB44DDFA9D" box="[866,1007,1268,1295]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="18" pageNumber="19" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
) and have an L:W of about 1.6. The symmetrical adaxials are about
<quantity id="4CE1088CFFFCFFD677A16F074562FAA1" box="[506,592,1304,1331]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="2.5">2.5 mm</quantity>
in length, with an L:W of about 1.9. All operculars (
<figureCitation id="1322B9ECFFFCFFD672C56F0643C4FAA1" box="[1182,1270,1305,1331]" captionStart="FIGURE 9" captionStartId="19.[151,250,1758,1783]" captionTargetBox="[174,1414,151,1738]" captionTargetId="figure@19.[174,1414,151,1738]" captionTargetPageId="19" captionText="FIGURE 9. Macroprimnoa ornata, paratype from USNM 142423: A–C, stereo views of lateral and opercular sides of a polyp. D, stereo view of coenenchymal scales, in situ. E, body wall scales, in situ. F, highly ridged body wall scales. G, opercular scales. H, ridged coenenchymal scales. Scale bars in mm." httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="18" pageNumber="19">Fig. 9G</figureCitation>
) have a highly concave outer surface, a keeled inner surface, and a digitiform base, like the body wall scales, and are fragile.
</paragraph>
<paragraph id="8BA6A569FFFCFFD6769C6F7E451FFA51" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
The coenenchymal scales (
<figureCitation id="1322B9ECFFFCFFD677AD6F7E457DFAE9" box="[502,591,1377,1403]" captionStart="FIGURE 9" captionStartId="19.[151,250,1758,1783]" captionTargetBox="[174,1414,151,1738]" captionTargetId="figure@19.[174,1414,151,1738]" captionTargetPageId="19" captionText="FIGURE 9. Macroprimnoa ornata, paratype from USNM 142423: A–C, stereo views of lateral and opercular sides of a polyp. D, stereo view of coenenchymal scales, in situ. E, body wall scales, in situ. F, highly ridged body wall scales. G, opercular scales. H, ridged coenenchymal scales. Scale bars in mm." httpUri="https://zenodo.org/record/2614983/files/figure.png" pageId="18" pageNumber="19">Fig. 9H</figureCitation>
) are elongate (up to
<quantity id="4CE1088CFFFCFFD675626F7E44A2FAE9" box="[825,912,1377,1403]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="19" unit="mm" value="3.3">3.3 mm</quantity>
in length with an L:W = 2.6–5.6), and usually pointed on both ends. Each coenenchymal bears tall (up to
<quantity id="4CE1088CFFFCFFD675776F9B44B6FA0D" box="[812,900,1412,1439]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
, sail scales) ridges that sometimes coalesce into a network of interconnecting ridges.
</paragraph>
<paragraph id="8BA6A569FFFCFFD6769C6FD346C4FA75" blockId="18.[151,1437,441,1655]" box="[199,502,1484,1511]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD6769C6FD34654FA75" bold="true" box="[199,358,1484,1511]" pageId="18" pageNumber="19">Comparisons</emphasis>
. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFCFFD6769C6FEE455FF9C1" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BA6A569FFFCFFD6769C6FEE455FF9C1" blockId="18.[151,1437,441,1655]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD6769C6FEE4606F999" bold="true" box="[199,308,1521,1547]" pageId="18" pageNumber="19">Remarks</emphasis>
. The length of each pinnate tentacle is approximately
<quantity id="4CE1088CFFFCFFD675CD6FEF44DCF999" box="[918,1006,1520,1547]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="19" unit="mm" value="6.5">6.5 mm</quantity>
, with 16–18 pinnules on either side of each tentacle. This might explain why the polyps are so widely spaced and the opercular scales so long, but even so, tentacles from contiguous polyps overlap.
</paragraph>
</subSubSection>
<subSubSection id="C303F6E2FFFCFFD6769C6C43426FF9E5" box="[199,1373,1628,1655]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BA6A569FFFCFFD6769C6C43426FF9E5" blockId="18.[151,1437,441,1655]" box="[199,1373,1628,1655]" pageId="18" pageNumber="19">
<emphasis id="B96D797BFFFCFFD6769C6C43466BF9E5" bold="true" box="[199,345,1628,1655]" pageId="18" pageNumber="19">Distribution</emphasis>
. Musicians Seamounts,
<collectingCountry id="F30EE5F9FFFCFFD674316C4244B5F9E5" box="[618,903,1629,1655]" name="Northern Mariana Islands" pageId="18" pageNumber="19">northern Mariana Islands</collectingCountry>
, and south of
<collectingCountry id="F30EE5F9FFFCFFD672786C424382F9E5" box="[1059,1200,1629,1655]" name="Wake Island" pageId="18" pageNumber="19">Wake Island</collectingCountry>
,
<quantity id="4CE1088CFFFCFFD672E06C42426BF9E5" box="[1211,1369,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="3.371" metricValueMax="3.676" metricValueMin="3.066" pageId="18" pageNumber="19" unit="m" value="3371.0" valueMax="3676.0" valueMin="3066.0">3066–3676 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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