375 lines
43 KiB
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375 lines
43 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1120.90086" ID-Pensoft-Pub="1313-2970-1120-177" ID-Pensoft-UUID="5875DFBEE22E54A687C53788A1E21180" ID-ZooBank="E576A469D3364D82AE487442B473D55C" ModsDocID="1313-2970-1120-177" checkinTime="1662649608510" checkinUser="pensoft" docAuthor="Luo, Jiu-Yang, Gong, Qiang-Bang & Xie, Qiang" docDate="2022" docId="E7E8D75FA85254EDBBC5E1B2D9EF2A6E" docLanguage="en" docName="ZooKeys 1120: 177-193" docOrigin="ZooKeys 1120" docPubDate="2022-09-07" docSource="http://dx.doi.org/10.3897/zookeys.1120.90086" docTitle="Cornonannus Luo & Gong & Xie 2022, gen. nov." docType="treatment" docUuid="1AEACAED-23C0-4690-AB8E-5287B3E9E5D9" docUuidSource="ZooBank" docVersion="1" id="5875DFBEE22E54A687C53788A1E21180" lastPageNumber="177" masterDocId="5875DFBEE22E54A687C53788A1E21180" masterDocTitle="A new genus and species of minute litter bugs family Schizopteridae Reuter, 1891 from China (Hemiptera, Heteroptera, Dipsocoromorpha)" masterLastPageNumber="193" masterPageNumber="177" pageNumber="177" updateTime="1662649608510" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus and species of minute litter bugs family Schizopteridae Reuter, 1891 from China (Hemiptera, Heteroptera, Dipsocoromorpha)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Luo, Jiu-Yang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2748-9534</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gong, Qiang-Bang</mods:namePart>
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<mods:affiliation>School of Life Sciences, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Xie, Qiang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6376-8808</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Biocontrol, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China</mods:affiliation>
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<mods:nameIdentifier type="email">xieq8@mail.sysu.edu.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1120</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>177</mods:start>
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<mods:end>193</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1120.90086</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1120.90086</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1120-177</mods:identifier>
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<mods:identifier type="ZooBank">E576A469D3364D82AE487442B473D55C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5875DFBEE22E54A687C53788A1E21180</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:1AEACAED-23C0-4690-AB8E-5287B3E9E5D9" httpUri="http://treatment.plazi.org/id/E7E8D75FA85254EDBBC5E1B2D9EF2A6E" lastPageNumber="177" pageId="0" pageNumber="177">
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<subSubSection pageId="0" pageNumber="177" type="nomenclature">
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<paragraph pageId="0" pageNumber="177">
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Genus
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<taxonomicName LSID="https://zoobank.org/1AEACAED-23C0-4690-AB8E-5287B3E9E5D9" authority="Luo & Gong & Xie, 2022" authorityName="Luo & Gong & Xie" authorityYear="2022" family="Schizopteridae" genus="Cornonannus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cornonannus" order="Hemiptera" pageId="0" pageNumber="177" rank="genus" status="gen. nov.">Cornonannus</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="177">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="177" type="description">
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<paragraph pageId="0" pageNumber="177">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cornonannus bui gen. et sp. nov. A-C male holotype A in dorsal view B in ventral view C in lateral view D male paratype in dorsal view (in dry condition)." figureDoi="10.3897/zookeys.1120.90086.figure1" httpUri="https://binary.pensoft.net/fig/739580" pageId="0" pageNumber="177">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Abdomen of Cornonannus bui gen. et sp. nov., male paratype. A in dorsal view B in lateral view C in ventral view D in caudal view." figureDoi="10.3897/zookeys.1120.90086.figure4" httpUri="https://binary.pensoft.net/fig/739583" pageId="0" pageNumber="177">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A metasternal spine B adhesive pad C abdomen D microtrichia on area near connexivum E middle region of basal sternite VII F middle region of sternite VII." figureDoi="10.3897/zookeys.1120.90086.figure6" httpUri="https://binary.pensoft.net/fig/739585" pageId="0" pageNumber="177">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in dorsal view A abdomen B groove and tubercles on tergite V C apex of abdomen D right region of tergite VIII E genitalia F apex of anophoric process." figureDoi="10.3897/zookeys.1120.90086.figure7" httpUri="https://binary.pensoft.net/fig/739586" pageId="0" pageNumber="177">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Drawings of Cornonannus bui gen. et sp. nov., male paratype A abdomen in dorsal view B tergite VIII in dorsal view C pygophore in dorsal view D, E two aspects of anophoric process F Aedeagus in dorsal view G-H two aspects of left paramere I, J two aspects of right paramere. Scale bars: 0.2 mm (A); 0.1 mm (B, C); 0.05 mm (D-J)." figureDoi="10.3897/zookeys.1120.90086.figure8" httpUri="https://binary.pensoft.net/fig/739587" pageId="0" pageNumber="177">, 8</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution map of Cornonannus bui gen. et sp. nov." figureDoi="10.3897/zookeys.1120.90086.figure9" httpUri="https://binary.pensoft.net/fig/739588" pageId="0" pageNumber="177">, 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="177" type="type species">
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<paragraph pageId="0" pageNumber="177">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="177">
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Type species by present designation:
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<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
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<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="177" type="diagnosis">
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<paragraph pageId="0" pageNumber="177">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="177">
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The genus
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<taxonomicName authorityName="Luo & Gong & Xie" authorityYear="2022" family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus" order="Hemiptera" pageId="0" pageNumber="177" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus</emphasis>
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</taxonomicName>
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gen. nov. can be distinguished from the other genera of
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<taxonomicName authorityName="Reuter" authorityYear="1891" lsidName="" pageId="0" pageNumber="177" rank="subFamily" subFamily="Schizopterinae">Schizopterinae</taxonomicName>
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by the following combined characteristics: 1) male with an upcurved horn-like frontal process on middle of head (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cornonannus bui gen. et sp. nov. A-C male holotype A in dorsal view B in ventral view C in lateral view D male paratype in dorsal view (in dry condition)." figureDoi="10.3897/zookeys.1120.90086.figure1" httpUri="https://binary.pensoft.net/fig/739580" pageId="0" pageNumber="177">1A-D</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A-C</figureCitation>
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); 2) pronotum without collar; 3) forewing without costal fracture (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">3A</figureCitation>
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); 4) forewing C+Sc, R1 distinctly thicker than other veins, R2 broad, about 2-3 times width of M, Cu or apical half of An (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">3A</figureCitation>
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); 5) tarsal formula 3-3-3 (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">3C-E</figureCitation>
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); 6) tergites I-VII of abdomen with transverse groove, tubercles on both sides of groove respectively or only on posterior side of groove (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Abdomen of Cornonannus bui gen. et sp. nov., male paratype. A in dorsal view B in lateral view C in ventral view D in caudal view." figureDoi="10.3897/zookeys.1120.90086.figure4" httpUri="https://binary.pensoft.net/fig/739583" pageId="0" pageNumber="177">4B, C</figureCitation>
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); 7) male anophoric process large and curved (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Abdomen of Cornonannus bui gen. et sp. nov., male paratype. A in dorsal view B in lateral view C in ventral view D in caudal view." figureDoi="10.3897/zookeys.1120.90086.figure4" httpUri="https://binary.pensoft.net/fig/739583" pageId="0" pageNumber="177">4B, C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="177" type="description">
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<paragraph pageId="0" pageNumber="177">Description.</paragraph>
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<paragraph pageId="0" pageNumber="177">
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Small (ca 1.5-1.6 mm), oblong and stout, forewing exceeding apex of abdomen (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cornonannus bui gen. et sp. nov. A-C male holotype A in dorsal view B in ventral view C in lateral view D male paratype in dorsal view (in dry condition)." figureDoi="10.3897/zookeys.1120.90086.figure1" httpUri="https://binary.pensoft.net/fig/739580" pageId="0" pageNumber="177">1A-D</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="177">Coloration</emphasis>
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: ground color brown to dark brown, compound eyes red to dark red, appendages yellowish brown to light brown, subapical of forewings with whitish area (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cornonannus bui gen. et sp. nov. A-C male holotype A in dorsal view B in ventral view C in lateral view D male paratype in dorsal view (in dry condition)." figureDoi="10.3897/zookeys.1120.90086.figure1" httpUri="https://binary.pensoft.net/fig/739580" pageId="0" pageNumber="177">1A, D</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1120.90086.figure1" httpUri="https://binary.pensoft.net/fig/739580" pageId="0" pageNumber="177" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="177">
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<emphasis bold="true" pageId="0" pageNumber="177">Figure 1.</emphasis>
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Habitus of
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<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
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<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
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</taxonomicName>
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gen. et sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="177">A-C</emphasis>
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male holotype
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<emphasis bold="true" pageId="0" pageNumber="177">A</emphasis>
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in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="177">B</emphasis>
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in ventral view
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<emphasis bold="true" pageId="0" pageNumber="177">C</emphasis>
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in lateral view
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<emphasis bold="true" pageId="0" pageNumber="177">D</emphasis>
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male paratype in dorsal view (in dry condition).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="177">
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<emphasis bold="true" pageId="0" pageNumber="177">
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Structure:
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<emphasis italics="true" pageId="0" pageNumber="177">head</emphasis>
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</emphasis>
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strongly declivent, short in lateral view, with dense punctures; frons slightly convex, with frontal process in the middle (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A, C</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">5A, C</figureCitation>
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), and several frontal cibarial muscle scars; vertex with cibarial scars consists of five contiguous coarse pits arranged in a row on both sides (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A</figureCitation>
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); areas near inner margin of eyes convex; maxillary plate and buccula with semi-erect, long setae, clypeus with three pairs of semi-erect setae, and one central long seta; labrum with one pair of long setae, labial segment IV with two pairs of long setae (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">5C, E</figureCitation>
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). Compound eye with about 30 ommatidia. Ocelli small, near inner margin of eyes, as large as about two ommatidia (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">5B</figureCitation>
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). Antennae four-segmented, antennal segments I and II stout, subequal in length, with several semi-erect setae (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A-C</figureCitation>
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); segments III and IV slender, segment IV longer than III, both segments with very long, semi-erect setae (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cornonannus bui gen. et sp. nov. A-C male holotype A in dorsal view B in ventral view C in lateral view D male paratype in dorsal view (in dry condition)." figureDoi="10.3897/zookeys.1120.90086.figure1" httpUri="https://binary.pensoft.net/fig/739580" pageId="0" pageNumber="177">1A-C</figureCitation>
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). Labium four-segmented, reaching to middle of mesosternum, segment I thickest, segment IV longest and tapering (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Cornonannus bui gen. et sp. nov. A-C male holotype A in dorsal view B in ventral view C in lateral view D male paratype in dorsal view (in dry condition)." figureDoi="10.3897/zookeys.1120.90086.figure1" httpUri="https://binary.pensoft.net/fig/739580" pageId="0" pageNumber="177">1A, B</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="177">Thorax</emphasis>
|
|
: prothorax with relatively dense punctures as in head (Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A, B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">5C-D</figureCitation>
|
|
). Pronotum near trapezoidal, declivent, without collar; callosite region with muscle scars consist by several pits (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2A</figureCitation>
|
|
); disk region convex; lateral margin sinuate, posterior margin slightly convex, middle near straight; proepisternal lobe inflated, almost reaching antennal insertions in lateral view (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2B</figureCitation>
|
|
). Proepimeral lobe wider than proepisternal lobe, subrectangular, posterior margin of proepimeron sinuate (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2B</figureCitation>
|
|
). Prosternum with wide longitudinal middle groove, apical portion of prosternal xiphus bilobate, sternal plate of prothorax trapezoidal, with straight posterior margin (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2C</figureCitation>
|
|
). Mesoscutum large; apex of mesoscutellum projecting posterodorsally in lateral view, base of mesoscutellum with a pair of pits (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2D-F</figureCitation>
|
|
); axillary cord sinuated; mesopostnotum with dense tubercles (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2E, F</figureCitation>
|
|
); mesoepisternal lobe large, mesepimeral lobe narrow, with tongue-shaped apex; upper area of mesopleura with a caudally directed triangular process (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2 D-F</figureCitation>
|
|
), mesopleura with groove along with middle coxal cleft, a deep concave located near the middle (Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2F</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">5F</figureCitation>
|
|
); mesosternum with longitudinal mesosternal ridge (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2F</figureCitation>
|
|
). Metascutum transverse, curved; metapostnotum transverse; a deep groove between mesopleura and metapleural (Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2F</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">5F</figureCitation>
|
|
). Metasternum with large keel-like metasternal spine, interlocking with notch between middle coxal cavities, basal portion of metasternal spine with round process, triangular apex (Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2F</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A metasternal spine B adhesive pad C abdomen D microtrichia on area near connexivum E middle region of basal sternite VII F middle region of sternite VII." figureDoi="10.3897/zookeys.1120.90086.figure6" httpUri="https://binary.pensoft.net/fig/739585" pageId="0" pageNumber="177">6A</figureCitation>
|
|
); apex of metendosternite bilobate (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Head and thorax of Cornonannus bui gen. et sp. nov., male paratype A-C male head and prothorax D-F male pterothorax A in frontal view B in lateral view C in ventral view D in ventral view E in dorsal view F in lateral view." figureDoi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177">2D-E</figureCitation>
|
|
). Forewings macropterous in males, female unknown; C+Sc and R thickened, with very narrow, slender subcostal cell; R2 distinctly wider than other veins; basal cell and trapezoidal discal cell slender; end of M subdivided into two branches (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">3A</figureCitation>
|
|
). Hind wings large and subdivided into four lobes, jugal lobe small; Sc+R+M and Cu subequal in length, 1An about three times as long as 2An, m-cu absent, (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">3B</figureCitation>
|
|
). Tarsal formula 3-3-3; apex of coxae, whole trochanters, femurs, tibiae, and tarsi with semi-erect setae (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">3C-E</figureCitation>
|
|
). Arolia absent; pulvilli fine and straight. Inner apex of foretibiae protuberant, with a long seta (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A frontal process B left eye and ocellus C head and thorax D microtrichia on head E mouth parts F pterothorax." figureDoi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177">5E</figureCitation>
|
|
); foretibiae bristle comb with about 10 setae; middle tibiae bristle comb with about 10 setae. Inner surface of hind coxae with adhesive pad (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Wings and legs of Cornonannus bui gen. et sp. nov., male paratype A forewing in dorsal view B hindwing in lateral view C foreleg D middle leg E hind leg." figureDoi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177">3E</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in ventro-lateral view A metasternal spine B adhesive pad C abdomen D microtrichia on area near connexivum E middle region of basal sternite VII F middle region of sternite VII." figureDoi="10.3897/zookeys.1120.90086.figure6" httpUri="https://binary.pensoft.net/fig/739585" pageId="0" pageNumber="177">6A, B</figureCitation>
|
|
); distal half of hind tibiae slightly curved, apex of hind tibiae with four thick setae.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="177">Pregenital abdomen</emphasis>
|
|
: abdomen strong sclerotized and rigid; terga interlocking, sterna also interlocking; tergites I-VI, sternites II-VI almost symmetric, tergite VII, sternite VII slightly asymmetric, tergite VIII strongly asymmetric (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Abdomen of Cornonannus bui gen. et sp. nov., male paratype. A in dorsal view B in lateral view C in ventral view D in caudal view." figureDoi="10.3897/zookeys.1120.90086.figure4" httpUri="https://binary.pensoft.net/fig/739583" pageId="0" pageNumber="177">4A-C</figureCitation>
|
|
). Tergites I and II fused, with transverse groove in the middle, tubercles on both sides of groove respectively or only on posterior side of groove (Figs
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Abdomen of Cornonannus bui gen. et sp. nov., male paratype. A in dorsal view B in lateral view C in ventral view D in caudal view." figureDoi="10.3897/zookeys.1120.90086.figure4" httpUri="https://binary.pensoft.net/fig/739583" pageId="0" pageNumber="177">4B, C</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scanning electron micrographs of Cornonannus bui gen. et sp. nov., male paratype in dorsal view A abdomen B groove and tubercles on tergite V C apex of abdomen D right region of tergite VIII E genitalia F apex of anophoric process." figureDoi="10.3897/zookeys.1120.90086.figure7" httpUri="https://binary.pensoft.net/fig/739586" pageId="0" pageNumber="177">7A, B</figureCitation>
|
|
). Middle area of sternites II and III less sclerotized, as sockets interlocking with metasternal plate; middle of sternite IV with longitudinal ridge; posterior margin of sternites VI and VII sinuated; sternite VII showing distinct sinistral asymmetry, sternites on both sides are approximately 90° from caudal view (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Abdomen of Cornonannus bui gen. et sp. nov., male paratype. A in dorsal view B in lateral view C in ventral view D in caudal view." figureDoi="10.3897/zookeys.1120.90086.figure4" httpUri="https://binary.pensoft.net/fig/739583" pageId="0" pageNumber="177">4D</figureCitation>
|
|
). Tergite VIII also showing sinistral asymmetry.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="177">Genitalia</emphasis>
|
|
: basal portion of pygophore near globular, overlapped by tergite VIII and sternite VII in repose; apical portion of pygophore flat, and exposed, with dense setae, asymmetrical. Anophore tubular, with anophoric process. Parameres strongly asymmetry; left paramere short, with a broad base, and a curved flattened distal projection; right paramere long, tapering, with a flat oval base provided with inner curved distal projection. Aedeagus complex, with a large, triangular basal plate; apical portion tubular, thin.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1120.90086.figure2" httpUri="https://binary.pensoft.net/fig/739581" pageId="0" pageNumber="177" start="Figure 2" startId="F2">
|
|
<paragraph pageId="0" pageNumber="177">
|
|
<emphasis bold="true" pageId="0" pageNumber="177">Figure 2.</emphasis>
|
|
Head and thorax of
|
|
<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
|
|
<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., male paratype
|
|
<emphasis bold="true" pageId="0" pageNumber="177">A-C</emphasis>
|
|
male head and prothorax
|
|
<emphasis bold="true" pageId="0" pageNumber="177">D-F</emphasis>
|
|
male pterothorax
|
|
<emphasis bold="true" pageId="0" pageNumber="177">A</emphasis>
|
|
in frontal view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">B</emphasis>
|
|
in lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">C</emphasis>
|
|
in ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">D</emphasis>
|
|
in ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">E</emphasis>
|
|
in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">F</emphasis>
|
|
in lateral view.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1120.90086.figure3" httpUri="https://binary.pensoft.net/fig/739582" pageId="0" pageNumber="177" start="Figure 3" startId="F3">
|
|
<paragraph pageId="0" pageNumber="177">
|
|
<emphasis bold="true" pageId="0" pageNumber="177">Figure 3.</emphasis>
|
|
Wings and legs of
|
|
<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
|
|
<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., male paratype
|
|
<emphasis bold="true" pageId="0" pageNumber="177">A</emphasis>
|
|
forewing in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">B</emphasis>
|
|
hindwing in lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">C</emphasis>
|
|
foreleg
|
|
<emphasis bold="true" pageId="0" pageNumber="177">D</emphasis>
|
|
middle leg
|
|
<emphasis bold="true" pageId="0" pageNumber="177">E</emphasis>
|
|
hind leg.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1120.90086.figure4" httpUri="https://binary.pensoft.net/fig/739583" pageId="0" pageNumber="177" start="Figure 4" startId="F4">
|
|
<paragraph pageId="0" pageNumber="177">
|
|
<emphasis bold="true" pageId="0" pageNumber="177">Figure 4.</emphasis>
|
|
Abdomen of
|
|
<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
|
|
<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., male paratype.
|
|
<emphasis bold="true" pageId="0" pageNumber="177">A</emphasis>
|
|
in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">B</emphasis>
|
|
in lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">C</emphasis>
|
|
in ventral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">D</emphasis>
|
|
in caudal view.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1120.90086.figure5" httpUri="https://binary.pensoft.net/fig/739584" pageId="0" pageNumber="177" start="Figure 5" startId="F5">
|
|
<paragraph pageId="0" pageNumber="177">
|
|
<emphasis bold="true" pageId="0" pageNumber="177">Figure 5.</emphasis>
|
|
Scanning electron micrographs of
|
|
<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
|
|
<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., male paratype in ventro-lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">A</emphasis>
|
|
frontal process
|
|
<emphasis bold="true" pageId="0" pageNumber="177">B</emphasis>
|
|
left eye and ocellus
|
|
<emphasis bold="true" pageId="0" pageNumber="177">C</emphasis>
|
|
head and thorax
|
|
<emphasis bold="true" pageId="0" pageNumber="177">D</emphasis>
|
|
microtrichia on head
|
|
<emphasis bold="true" pageId="0" pageNumber="177">E</emphasis>
|
|
mouth parts
|
|
<emphasis bold="true" pageId="0" pageNumber="177">F</emphasis>
|
|
pterothorax.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1120.90086.figure6" httpUri="https://binary.pensoft.net/fig/739585" pageId="0" pageNumber="177" start="Figure 6" startId="F6">
|
|
<paragraph pageId="0" pageNumber="177">
|
|
<emphasis bold="true" pageId="0" pageNumber="177">Figure 6.</emphasis>
|
|
Scanning electron micrographs of
|
|
<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
|
|
<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., male paratype in ventro-lateral view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">A</emphasis>
|
|
metasternal spine
|
|
<emphasis bold="true" pageId="0" pageNumber="177">B</emphasis>
|
|
adhesive pad
|
|
<emphasis bold="true" pageId="0" pageNumber="177">C</emphasis>
|
|
abdomen
|
|
<emphasis bold="true" pageId="0" pageNumber="177">D</emphasis>
|
|
microtrichia on area near connexivum
|
|
<emphasis bold="true" pageId="0" pageNumber="177">E</emphasis>
|
|
middle region of basal sternite VII
|
|
<emphasis bold="true" pageId="0" pageNumber="177">F</emphasis>
|
|
middle region of sternite VII.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1120.90086.figure7" httpUri="https://binary.pensoft.net/fig/739586" pageId="0" pageNumber="177" start="Figure 7" startId="F7">
|
|
<paragraph pageId="0" pageNumber="177">
|
|
<emphasis bold="true" pageId="0" pageNumber="177">Figure 7.</emphasis>
|
|
Scanning electron micrographs of
|
|
<taxonomicName family="Schizopteridae" genus="Cornonannus" kingdom="Animalia" lsidName="Cornonannus bui" order="Hemiptera" pageId="0" pageNumber="177" rank="species" species="bui">
|
|
<emphasis italics="true" pageId="0" pageNumber="177">Cornonannus bui</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., male paratype in dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="177">A</emphasis>
|
|
abdomen
|
|
<emphasis bold="true" pageId="0" pageNumber="177">B</emphasis>
|
|
groove and tubercles on tergite V
|
|
<emphasis bold="true" pageId="0" pageNumber="177">C</emphasis>
|
|
apex of abdomen
|
|
<emphasis bold="true" pageId="0" pageNumber="177">D</emphasis>
|
|
right region of tergite VIII
|
|
<emphasis bold="true" pageId="0" pageNumber="177">E</emphasis>
|
|
genitalia
|
|
<emphasis bold="true" pageId="0" pageNumber="177">F</emphasis>
|
|
apex of anophoric process.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="177" type="etymology">
|
|
<paragraph pageId="0" pageNumber="177">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="177">
|
|
The generic name is derived from the Greek prefix
|
|
<normalizedToken originalValue="“corn-”">"corn-"</normalizedToken>
|
|
(horned or having horns or horn-like appendages) and the Greek root
|
|
<normalizedToken originalValue="“nannus”">"nannus"</normalizedToken>
|
|
(a dwarf). The gender is masculine.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |