249 lines
35 KiB
XML
249 lines
35 KiB
XML
<document id="9174F6FE0D30ACB2A6C12152CD6D5651" ID-DOI="10.11646/zootaxa.4750.2.4" ID-GBIF-Dataset="2c856dad-dfc5-4983-a2e7-9494ba2b701f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3707352" ID-ZooBank="75FFA71A-AADD-4E2B-A154-29B2DF4F77BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584002027481" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="E34E3C29FFBD3E09268AF99FCD6EFC24" docLanguage="en" docName="zootaxa.4750.2.4.pdf" docOrigin="Zootaxa 4750 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protandroconnus akeratophorus Jałoszyński 2020, sp. n." docType="treatment" docVersion="5" lastPageNumber="209" masterDocId="1F774451FFBE3E0C261DFFCFC82CFFEF" masterDocTitle="First records of Trichocircus Jałoszyński and Protandroconnus Franz in Mexico, with descriptions of two new species (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="210" masterPageNumber="204" pageNumber="207" updateTime="1698727108048" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B4F8D445EEDD59FFFC111AE0B4D0C397">First records of Trichocircus Jałoszyński and Protandroconnus Franz in Mexico, with descriptions of two new species (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="28F9EE8FF7865B09C0DE587F2D336E88">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="FFF28AEAD32D9F779E52E40CB7EAABA9">Zootaxa</mods:title>
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<mods:date id="F5B21BCF4BF97BC85B3A6E5A516EC17E">2020</mods:date>
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<mods:number id="00E090406522392A6C7051126C82E8C1">2020-03-11</mods:number>
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<mods:number id="F3CD2BDFF31F758C4CD3B337450AB5B4">4750</mods:number>
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<treatment id="E34E3C29FFBD3E09268AF99FCD6EFC24" ID-DOI="http://doi.org/10.5281/zenodo.3717808" ID-GBIF-Taxon="162483873" ID-Zenodo-Dep="3717808" LSID="urn:lsid:plazi:treatment:E34E3C29FFBD3E09268AF99FCD6EFC24" httpUri="http://treatment.plazi.org/id/E34E3C29FFBD3E09268AF99FCD6EFC24" lastPageId="5" lastPageNumber="209" pageId="3" pageNumber="207">
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<subSubSection id="23FDDEB4FFBD3E0F268AF99FCA40F985" box="[151,620,1616,1643]" pageId="3" pageNumber="207" type="nomenclature">
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<paragraph id="6B588D3FFFBD3E0F268AF99FCA40F985" blockId="3.[151,620,1616,1678]" box="[151,620,1616,1643]" pageId="3" pageNumber="207">
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<heading id="30103A53FFBD3E0F268AF99FCA40F985" bold="true" box="[151,620,1616,1643]" fontSize="11" level="1" pageId="3" pageNumber="207" reason="1">
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<emphasis id="5993512DFFBD3E0F268AF99FCA40F985" bold="true" box="[151,620,1616,1643]" pageId="3" pageNumber="207">
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<taxonomicName id="ACE7F6BCFFBD3E0F268AF99FCA0EF984" ID-CoL="8Y8NP" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,546,1616,1643]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="207" phylum="Arthropoda" rank="species" species="akeratophorus" status="sp. nov.">
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<emphasis id="5993512DFFBD3E0F268AF99FCA0EF984" bold="true" box="[151,546,1616,1643]" italics="true" pageId="3" pageNumber="207">Protandroconnus akeratophorus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="42A0EC56FFBD3E0F2437F99FCA40F985" box="[554,620,1616,1642]" pageId="3" pageNumber="207" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="23FDDEB4FFBD3E0F268AF9BBC90AF961" box="[151,294,1652,1678]" pageId="3" pageNumber="207" type="description">
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<paragraph id="6B588D3FFFBD3E0F268AF9BBC90AF961" blockId="3.[151,620,1616,1678]" box="[151,294,1652,1678]" pageId="3" pageNumber="207">
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(
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<figureCitation id="F3DC91BAFFBD3E0F2682F9BBC932F961" box="[159,286,1652,1678]" captionStart-0="FIGURES 11–16" captionStart-1="FIGURES 17–23" captionStartId-0="4.[151,264,1897,1922]" captionStartId-1="5.[151,264,1785,1810]" captionTargetBox-0="[168,1428,705,1872]" captionTargetBox-1="[151,1430,993,1760]" captionTargetId-0="figure@4.[160,1428,705,1872]" captionTargetId-1="figure@5.[151,1436,993,1760]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 11–16. Protandroconnus akeratophorus sp. n., male. Dorsal habitus (11), head, prothorax and elytral base in dorsal view (12), head, prothorax and anterior portion of pterothorax in ventral view (13), head in dorsal (14), anterodorsal (15) and lateral (16) views. Abbreviations: bst, basisternal portion of prosternum; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; smn, submentum." captionText-1="FIGURES 17–23. Protandroconnus akeratophorus sp. n., male. Prothorax in ventral view (17), pterothorax in ventral view (18), pterothorax and abdomen in ventral view (19), aedeagus in ventral (20, 22) and lateral (21, 23) views. Abbreviations: bst, basisternal portion of prosternum; dlf, dorsolateral fovea; hyr, hypomeral ridge; lfa, lateral furcal arm; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi-0="http://doi.org/10.5281/zenodo.3707358" figureDoi-1="http://doi.org/10.5281/zenodo.3707360" httpUri-0="https://zenodo.org/record/3707358/files/figure.png" httpUri-1="https://zenodo.org/record/3707360/files/figure.png" pageId="3" pageNumber="207">Figs 11–23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="23FDDEB4FFBD3E0F268AF973CBA4F8AD" pageId="3" pageNumber="207" type="materials_examined">
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<paragraph id="6B588D3FFFBD3E0F268AF973CBA4F8AD" blockId="3.[151,1437,1723,2038]" pageId="3" pageNumber="207">
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<emphasis id="5993512DFFBD3E0F268AF973C968F939" bold="true" box="[151,324,1723,1750]" pageId="3" pageNumber="207">Type material.</emphasis>
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<materialsCitation id="DB8F8762FFBD3E0F2756F973CBD9F8F1" ID-GBIF-Occurrence="2575868655" collectionCode="MHNG" collectorName="Dr. A. Fenyes" country="Mexico" location="Veracruz State" pageId="3" pageNumber="207" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
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<emphasis id="5993512DFFBD3E0F2756F973CADDF939" bold="true" box="[331,753,1723,1750]" pageId="3" pageNumber="207">
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<typeStatus id="B45C339DFFBD3E0F2756F973C990F939" box="[331,444,1724,1750]" pageId="3" pageNumber="207" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="13F0CDAFFFBD3E0F27DAF974CA32F939" box="[455,542,1723,1750]" name="Mexico" pageId="3" pageNumber="207">Mexico</collectingCountry>
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(
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<collectingRegion id="A92343DDFFBD3E0F2431F974CACFF939" box="[556,739,1723,1750]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="207">Veracruz State</collectingRegion>
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):
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</emphasis>
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<specimenCount id="7DE146B6FFBD3E0F24EAF973CB22F939" box="[759,782,1724,1750]" pageId="3" pageNumber="207" type="male">♂</specimenCount>
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, four labels: “Cordoba / Mex. Ver. /
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<collectorName id="C612E8E9FFBD3E0F22BDF973CD1CF939" box="[1184,1328,1724,1750]" pageId="3" pageNumber="207">Dr.A.Fenyes</collectorName>
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” [creamy white, printed], “A. FENYES / Collection” [yellow, printed], small green star, “
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<taxonomicName id="ACE7F6BCFFBD3E0F2237F92FCD16F915" authorityName="Franz" authorityYear="1989" box="[1066,1338,1760,1786]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="207" phylum="Arthropoda" rank="genus">
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<emphasis id="5993512DFFBD3E0F2237F92FCD16F915" box="[1066,1338,1760,1786]" italics="true" pageId="3" pageNumber="207">PROTANDROCONNUS</emphasis>
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</taxonomicName>
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/
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<emphasis id="5993512DFFBD3E0F234EF92FC8D7F8F1" italics="true" pageId="3" pageNumber="207">akera- tophorus</emphasis>
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m. / P.
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<bibRefCitation id="0F76F0CEFFBD3E0F274FF8CBCA36F8F1" author="Jaloszynski, P." box="[338,538,1796,1822]" pageId="3" pageNumber="207" pagination="515 - 534" refId="ref2886" refString="Jaloszynski, P. (2019) Neotropical Glandulariini with male cephalic modifications: Protandroconnus Franz and Trichocircus gen. nov. (Coleoptera: Staphylinidae: Scydmaeninae). Annales Zoologici, 69 (3), 515 - 534. https: // doi. org / 10.3161 / 00034541 ANZ 2019.69.3.003" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
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/
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<typeStatus id="B45C339DFFBD3E0F242FF8CBCAF0F8F1" box="[562,732,1796,1822]" pageId="3" pageNumber="207" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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<collectionCode id="0DF615FAFFBD3E0F258CF8CBCBC1F8F1" box="[913,1005,1796,1822]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="207" type="Museum">MHNG</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="DB8F8762FFBD3E0F221DF8CBCBA9F8AD" ID-GBIF-Occurrence="2575868656" collectionCode="MHNG" collectorName="Dr. A. Fenyes" country="Mexico" location="Veracruz State" pageId="3" pageNumber="207" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
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<emphasis id="5993512DFFBD3E0F221DF8CBCC5AF8F1" bold="true" box="[1024,1142,1796,1822]" pageId="3" pageNumber="207">
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<typeStatus id="B45C339DFFBD3E0F221DF8CBCC5DF8F1" box="[1024,1137,1796,1822]" pageId="3" pageNumber="207" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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<specimenCount id="7DE146B6FFBD3E0F2263F8CBCCB9F8F1" box="[1150,1173,1796,1822]" pageId="3" pageNumber="207" type="male">♂</specimenCount>
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, same data as for holo- type, with standard yellow printed “
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<typeStatus id="B45C339DFFBD3E0F243AF8E7CAE7F8AD" box="[551,715,1832,1858]" pageId="3" pageNumber="207" type="paratype">PARATYPUS</typeStatus>
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” label (
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<collectionCode id="0DF615FAFFBD3E0F253DF8E7CB50F8AD" box="[800,892,1832,1858]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="207" type="Museum">MHNG</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="23FDDEB4FFBD3E0F26DAF884C94CF841" pageId="3" pageNumber="207" type="diagnosis">
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<paragraph id="6B588D3FFFBD3E0F26DAF884C94CF841" blockId="3.[151,1437,1723,2038]" pageId="3" pageNumber="207">
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<emphasis id="5993512DFFBD3E0F26DAF884C96EF889" bold="true" box="[199,322,1867,1894]" pageId="3" pageNumber="207">Diagnosis.</emphasis>
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Head in male lacking frontal ‘horn’, with transverse impression in the form of two lateral triangles connected at middle by narrow sulcus; apical, trapezoidal portion of aedagus in ventral view broader than long; parameres absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="23FDDEB4FFBD3E0926DAF878C9B2FE3C" lastPageId="5" lastPageNumber="209" pageId="3" pageNumber="207" type="description">
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<paragraph id="6B588D3FFFBD3E0F26DAF878CACAF819" blockId="3.[151,1437,1723,2038]" pageId="3" pageNumber="207">
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<emphasis id="5993512DFFBD3E0F26DAF878C975F83D" bold="true" box="[199,345,1975,2002]" pageId="3" pageNumber="207">Description.</emphasis>
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Body of male (
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<figureCitation id="F3DC91BAFFBD3E0F2410F877CA72F83D" box="[525,606,1976,2002]" captionStart="FIGURES 11–16" captionStartId="4.[151,264,1897,1922]" captionTargetBox="[168,1428,705,1872]" captionTargetId="figure@4.[160,1428,705,1872]" captionTargetPageId="4" captionText="FIGURES 11–16. Protandroconnus akeratophorus sp. n., male. Dorsal habitus (11), head, prothorax and elytral base in dorsal view (12), head, prothorax and anterior portion of pterothorax in ventral view (13), head in dorsal (14), anterodorsal (15) and lateral (16) views. Abbreviations: bst, basisternal portion of prosternum; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.3707358" httpUri="https://zenodo.org/record/3707358/files/figure.png" pageId="3" pageNumber="207">Fig. 11</figureCitation>
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) elongate, moderately slender, strongly convex, light brown and covered with yellowish setae and bristles; BL
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<quantity id="AC1F20DAFFBD3E0F2421F814CACEF819" box="[572,738,2011,2038]" metricMagnitude="-3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="3" pageNumber="207" unit="mm" value="1.065" valueMax="1.08" valueMin="1.05">1.05–1.08 mm</quantity>
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.
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</paragraph>
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<paragraph id="6B588D3FFFBA3E0826DAFF56CA5EFDF4" blockId="4.[151,1437,153,683]" pageId="4" pageNumber="208">
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Head (
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<figureCitation id="F3DC91BAFFBA3E08270EFF56C9B8FF5B" box="[275,404,153,180]" captionStart="FIGURES 11–16" captionStartId="4.[151,264,1897,1922]" captionTargetBox="[168,1428,705,1872]" captionTargetId="figure@4.[160,1428,705,1872]" captionTargetPageId="4" captionText="FIGURES 11–16. Protandroconnus akeratophorus sp. n., male. Dorsal habitus (11), head, prothorax and elytral base in dorsal view (12), head, prothorax and anterior portion of pterothorax in ventral view (13), head in dorsal (14), anterodorsal (15) and lateral (16) views. Abbreviations: bst, basisternal portion of prosternum; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.3707358" httpUri="https://zenodo.org/record/3707358/files/figure.png" pageId="4" pageNumber="208">Figs 12–16</figureCitation>
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) in dorsal view subpentagonal and slightly transverse, broadest at eyes, HL
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<quantity id="AC1F20DAFFBA3E0822E4FF56CD4DFF5C" box="[1273,1377,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="208" unit="mm" value="0.2">0.20 mm</quantity>
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, HW
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<quantity id="AC1F20DAFFBA3E08268AFF73C92DFF38" box="[151,257,188,215]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="208" unit="mm" value="0.25">0.25 mm</quantity>
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; vertex demarcated from frons by arcuate, anteriorly convex ridge that overhangs frons between eyes; frons between eyes with arcuate, posteriorly convex ridge, so that vertex and frons are separated by transverse impression composed of median sulcus between frontal and vertexal ridges and broad subtriangular lateral portions; supraantennal tubercles weakly elevated. Punctures on frons and vertex absent, setal insertions not impressed, setae sparse, short and suberect, sides of frons and lateral portions of transverse impression asetose. Antennae (
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<figureCitation id="F3DC91BAFFBA3E082372FE82C89EFE64" captionStart="FIGURES 11–16" captionStartId="4.[151,264,1897,1922]" captionTargetBox="[168,1428,705,1872]" captionTargetId="figure@4.[160,1428,705,1872]" captionTargetPageId="4" captionText="FIGURES 11–16. Protandroconnus akeratophorus sp. n., male. Dorsal habitus (11), head, prothorax and elytral base in dorsal view (12), head, prothorax and anterior portion of pterothorax in ventral view (13), head in dorsal (14), anterodorsal (15) and lateral (16) views. Abbreviations: bst, basisternal portion of prosternum; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.3707358" httpUri="https://zenodo.org/record/3707358/files/figure.png" pageId="4" pageNumber="208">Fig. 11</figureCitation>
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) slender; AnL
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<quantity id="AC1F20DAFFBA3E082740FEBFCA28FE64" box="[349,516,368,395]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="4" pageNumber="208" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
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; scape and pedicel elongate, antennomeres III–VI each about as long as broad or slightly transverse, VII enlarged, distinctly longer and broader than VI, about as long as broad, VIII slightly shorter and narrower than VII, weakly transverse, IX distinctly broader and longer than VIII, distinctly transverse, X distinctly broader but indistinctly longer than IX, distinctly transverse, XI as broad as X, slightly shorter than IX and X combined, about 1.6 × as long as broad.
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</paragraph>
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<paragraph id="6B588D3FFFBA3E0826DAFDEAC8FBFD44" blockId="4.[151,1437,153,683]" pageId="4" pageNumber="208">
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Pronotum (
|
||
<figureCitation id="F3DC91BAFFBA3E082755FDEAC9B0FDD0" box="[328,412,549,575]" captionStart="FIGURES 11–16" captionStartId="4.[151,264,1897,1922]" captionTargetBox="[168,1428,705,1872]" captionTargetId="figure@4.[160,1428,705,1872]" captionTargetPageId="4" captionText="FIGURES 11–16. Protandroconnus akeratophorus sp. n., male. Dorsal habitus (11), head, prothorax and elytral base in dorsal view (12), head, prothorax and anterior portion of pterothorax in ventral view (13), head in dorsal (14), anterodorsal (15) and lateral (16) views. Abbreviations: bst, basisternal portion of prosternum; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.3707358" httpUri="https://zenodo.org/record/3707358/files/figure.png" pageId="4" pageNumber="208">Fig. 12</figureCitation>
|
||
) broadest near middle; PL
|
||
<quantity id="AC1F20DAFFBA3E0824C6FDEACB68FDD0" box="[731,836,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="208" unit="mm" value="0.3">0.30 mm</quantity>
|
||
, PW
|
||
<quantity id="AC1F20DAFFBA3E08259BFDEACBDCFDD0" box="[902,1008,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="208" unit="mm" value="0.3">0.30 mm</quantity>
|
||
; sides strongly rounded, anterior and posterior margins indistinctly arcuate. Punctures on pronotal disc fine and inconspicuous, setae sparse, short and suberect, lateral bristles long and erect, in anterior 2/3 standing in various directions, in posterior third directed posterad.
|
||
</paragraph>
|
||
<caption id="3F98DDB7FFBA3E08268AF8A6CAB1F801" ID-DOI="http://doi.org/10.5281/zenodo.3707358" ID-Zenodo-Dep="3707358" httpUri="https://zenodo.org/record/3707358/files/figure.png" pageId="4" pageNumber="208" startId="4.[151,264,1897,1922]" targetBox="[168,1428,705,1872]" targetPageId="4">
|
||
<paragraph id="6B588D3FFFBA3E08268AF8A6CAB1F801" blockId="4.[151,1437,1897,2030]" pageId="4" pageNumber="208">
|
||
<emphasis id="5993512DFFBA3E08268AF8A6C97FF86D" bold="true" box="[151,339,1897,1922]" pageId="4" pageNumber="208">FIGURES 11–16.</emphasis>
|
||
<taxonomicName id="ACE7F6BCFFBA3E082745F8A5CA8EF86D" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[344,674,1898,1922]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="208" phylum="Arthropoda" rank="species" species="akeratophorus" status="sp. nov.">
|
||
<emphasis id="5993512DFFBA3E082745F8A5CA8EF86D" box="[344,674,1898,1922]" italics="true" pageId="4" pageNumber="208">Protandroconnus akeratophorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5993512DFFBA3E0824B5F8A5CACDF86D" bold="true" box="[680,737,1898,1922]" pageId="4" pageNumber="208">
|
||
<taxonomicNameLabel id="42A0EC56FFBA3E0824B5F8A5CACDF86D" box="[680,737,1898,1922]" pageId="4" pageNumber="208" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male. Dorsal habitus (11), head, prothorax and elytral base in dorsal view (12), head, prothorax and anterior portion of pterothorax in ventral view (13), head in dorsal (14), anterodorsal (15) and lateral (16) views. Abbreviations: bst, basisternal portion of prosternum; gp, gular plate; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; smn, submentum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6B588D3FFFBB3E0926DAFF56C920FF14" blockId="5.[151,1437,152,971]" pageId="5" pageNumber="209">
|
||
Elytra together oval, broadest slightly behind middle, EL
|
||
<quantity id="AC1F20DAFFBB3E092551FF57CBDDFF5C" box="[844,1009,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.8" metricValueMin="5.5" pageId="5" pageNumber="209" unit="mm" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 mm</quantity>
|
||
, EW
|
||
<quantity id="AC1F20DAFFBB3E092232FF56CCB9FF5C" box="[1071,1173,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="209" unit="mm" value="0.43">0.43 mm</quantity>
|
||
, EI 1.29–1.35; humeral calli weakly elevated, apices separately rounded. Punctures as on pronotal disc; setae slightly longer than those on pronotum.
|
||
</paragraph>
|
||
<paragraph id="6B588D3FFFBB3E0926DAFECACA2BFEF0" blockId="5.[151,1437,152,971]" box="[199,519,261,287]" pageId="5" pageNumber="209">Hind wings fully developed.</paragraph>
|
||
<paragraph id="6B588D3FFFBB3E0926DAFEE6CA7CFEAC" blockId="5.[151,1437,152,971]" box="[199,592,297,323]" pageId="5" pageNumber="209">Legs short and robust, unmodified.</paragraph>
|
||
<paragraph id="6B588D3FFFBB3E0926DAFE82C98AFE40" blockId="5.[151,1437,152,971]" pageId="5" pageNumber="209">
|
||
Aedeagus (
|
||
<figureCitation id="F3DC91BAFFBB3E092758FE82C9E9FE88" box="[325,453,333,359]" captionStart="FIGURES 17–23" captionStartId="5.[151,264,1785,1810]" captionTargetBox="[151,1430,993,1760]" captionTargetId="figure@5.[151,1436,993,1760]" captionTargetPageId="5" captionText="FIGURES 17–23. Protandroconnus akeratophorus sp. n., male. Prothorax in ventral view (17), pterothorax in ventral view (18), pterothorax and abdomen in ventral view (19), aedeagus in ventral (20, 22) and lateral (21, 23) views. Abbreviations: bst, basisternal portion of prosternum; dlf, dorsolateral fovea; hyr, hypomeral ridge; lfa, lateral furcal arm; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.3707360" httpUri="https://zenodo.org/record/3707360/files/figure.png" pageId="5" pageNumber="209">Figs 20–23</figureCitation>
|
||
) moderately slender, AeL
|
||
<quantity id="AC1F20DAFFBB3E0924F2FE82CB7BFE88" box="[751,855,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="209" unit="mm" value="0.13">0.13 mm</quantity>
|
||
; median lobe in ventral view broadest in basal half, apex subtrapezoidal, in lateral view apical portion strongly bent dorsad; endophallus with weakly sclerotized structures; parameres absent.
|
||
</paragraph>
|
||
<paragraph id="6B588D3FFFBB3E0926DAFE76C9B2FE3C" blockId="5.[151,1437,152,971]" box="[199,414,441,467]" pageId="5" pageNumber="209">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23FDDEB4FFBB3E0926DAFE13CA1DFE18" box="[199,561,476,503]" pageId="5" pageNumber="209" type="distribution">
|
||
<paragraph id="6B588D3FFFBB3E0926DAFE13CA1DFE18" blockId="5.[151,1437,152,971]" box="[199,561,476,503]" pageId="5" pageNumber="209">
|
||
<emphasis id="5993512DFFBB3E0926DAFE13C94DFE18" bold="true" box="[199,353,476,503]" pageId="5" pageNumber="209">Distribution.</emphasis>
|
||
Southern
|
||
<collectingCountry id="13F0CDAFFFBB3E0927C9FE12CA01FE18" box="[468,557,477,503]" name="Mexico" pageId="5" pageNumber="209">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23FDDEB4FFBB3E0926DAFDCECBA4FDD0" pageId="5" pageNumber="209" type="etymology">
|
||
<paragraph id="6B588D3FFFBB3E0926DAFDCECBA4FDD0" blockId="5.[151,1437,152,971]" pageId="5" pageNumber="209">
|
||
<emphasis id="5993512DFFBB3E0926DAFDCEC962FDF4" bold="true" box="[199,334,513,539]" pageId="5" pageNumber="209">Etymology.</emphasis>
|
||
The adjective
|
||
<taxonomicName id="ACE7F6BCFFBB3E0927EFFDCECAB9FDF4" authorityName="Jałoszyński" authorityYear="2020" box="[498,661,513,539]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="akeratophorus">
|
||
<emphasis id="5993512DFFBB3E0927EFFDCECAB9FDF4" box="[498,661,513,539]" italics="true" pageId="5" pageNumber="209">akeratophorus</emphasis>
|
||
</taxonomicName>
|
||
(= not horned) refers to lack of the frontal ‘horn’ in this species; after Greek
|
||
<emphasis id="5993512DFFBB3E0926FEFDE9C8DDFDD0" box="[227,241,550,575]" italics="true" pageId="5" pageNumber="209">a</emphasis>
|
||
(not) +
|
||
<emphasis id="5993512DFFBB3E092751FDEAC9A5FDD0" box="[332,393,549,575]" italics="true" pageId="5" pageNumber="209">keras</emphasis>
|
||
(horn) +
|
||
<emphasis id="5993512DFFBB3E0927EEFDEACA64FDD0" box="[499,584,549,575]" italics="true" pageId="5" pageNumber="209">pherein</emphasis>
|
||
(carry), in a Latinized form.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23FDDEB4FFBB3E0926DAFD86CD6EFC24" pageId="5" pageNumber="209" type="discussion">
|
||
<paragraph id="6B588D3FFFBB3E0926DAFD86CD6EFC24" blockId="5.[151,1437,152,971]" pageId="5" pageNumber="209">
|
||
<emphasis id="5993512DFFBB3E0926DAFD86C917FD8C" bold="true" box="[199,315,585,611]" pageId="5" pageNumber="209">Remarks.</emphasis>
|
||
<taxonomicName id="ACE7F6BCFFBB3E09275EFD86CA83FD8C" authorityName="Jałoszyński" authorityYear="2020" box="[323,687,585,611]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="akeratophorus">
|
||
<emphasis id="5993512DFFBB3E09275EFD86CA83FD8C" box="[323,687,585,611]" italics="true" pageId="5" pageNumber="209">Protandroconnus akeratophorus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="ACE7F6BCFFBB3E09255BFD85CBE1FD8C" authorityName="Franz" authorityYear="1989" box="[838,973,586,611]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="peruensis">
|
||
<emphasis id="5993512DFFBB3E09255BFD85CBE1FD8C" box="[838,973,586,611]" italics="true" pageId="5" pageNumber="209">P. peruensis</emphasis>
|
||
</taxonomicName>
|
||
not only in male cephalic modifications, but also in ventral prothoracic and male genital structures. The notosternal sutures in
|
||
<taxonomicName id="ACE7F6BCFFBB3E09225EFDA1CCE5FD68" authorityName="Franz" authorityYear="1989" box="[1091,1225,622,647]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="peruensis">
|
||
<emphasis id="5993512DFFBB3E09225EFDA1CCE5FD68" box="[1091,1225,622,647]" italics="true" pageId="5" pageNumber="209">P. peruensis</emphasis>
|
||
</taxonomicName>
|
||
are largely obliterated, but their short anterior and posterior portions are discernible, whereas in
|
||
<taxonomicName id="ACE7F6BCFFBB3E0925E4FD5DCC9BFD44" authorityName="Jałoszyński" authorityYear="2020" box="[1017,1207,657,683]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="akeratophorus">
|
||
<emphasis id="5993512DFFBB3E0925E4FD5DCC9BFD44" box="[1017,1207,657,683]" italics="true" pageId="5" pageNumber="209">P. akeratophorus</emphasis>
|
||
</taxonomicName>
|
||
the sutures (
|
||
<figureCitation id="F3DC91BAFFBB3E09235EFD5ECDB8FD44" box="[1347,1428,657,683]" captionStart="FIGURES 17–23" captionStartId="5.[151,264,1785,1810]" captionTargetBox="[151,1430,993,1760]" captionTargetId="figure@5.[151,1436,993,1760]" captionTargetPageId="5" captionText="FIGURES 17–23. Protandroconnus akeratophorus sp. n., male. Prothorax in ventral view (17), pterothorax in ventral view (18), pterothorax and abdomen in ventral view (19), aedeagus in ventral (20, 22) and lateral (21, 23) views. Abbreviations: bst, basisternal portion of prosternum; dlf, dorsolateral fovea; hyr, hypomeral ridge; lfa, lateral furcal arm; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.3707360" httpUri="https://zenodo.org/record/3707360/files/figure.png" pageId="5" pageNumber="209">Fig. 17</figureCitation>
|
||
) are entirely absent. In
|
||
<taxonomicName id="ACE7F6BCFFBB3E092790FD79CA3EFD20" authorityName="Franz" authorityYear="1989" box="[397,530,694,719]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="peruensis">
|
||
<emphasis id="5993512DFFBB3E092790FD79CA3EFD20" box="[397,530,694,719]" italics="true" pageId="5" pageNumber="209">P. peruensis</emphasis>
|
||
</taxonomicName>
|
||
the postcoxal lobe of hypomeron that protrudes mesad seems to be fused with the postcoxal portion of prosternum, whereas in
|
||
<taxonomicName id="ACE7F6BCFFBB3E09248CFD15CB7CFD1C" authorityName="Jałoszyński" authorityYear="2020" box="[657,848,729,755]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="akeratophorus">
|
||
<emphasis id="5993512DFFBB3E09248CFD15CB7CFD1C" box="[657,848,729,755]" italics="true" pageId="5" pageNumber="209">P. akeratophorus</emphasis>
|
||
</taxonomicName>
|
||
the hypomeral lobe (
|
||
<figureCitation id="F3DC91BAFFBB3E09225FFD16CCB8FD1C" box="[1090,1172,729,755]" captionStart="FIGURES 17–23" captionStartId="5.[151,264,1785,1810]" captionTargetBox="[151,1430,993,1760]" captionTargetId="figure@5.[151,1436,993,1760]" captionTargetPageId="5" captionText="FIGURES 17–23. Protandroconnus akeratophorus sp. n., male. Prothorax in ventral view (17), pterothorax in ventral view (18), pterothorax and abdomen in ventral view (19), aedeagus in ventral (20, 22) and lateral (21, 23) views. Abbreviations: bst, basisternal portion of prosternum; dlf, dorsolateral fovea; hyr, hypomeral ridge; lfa, lateral furcal arm; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.3707360" httpUri="https://zenodo.org/record/3707360/files/figure.png" pageId="5" pageNumber="209">Fig. 17</figureCitation>
|
||
) overlaps with the prosternum, but is not fused to it behind the procoxal cavity. The parameres in
|
||
<taxonomicName id="ACE7F6BCFFBB3E0925C6FD31CC4DFCF8" authorityName="Franz" authorityYear="1989" box="[987,1121,766,791]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="peruensis">
|
||
<emphasis id="5993512DFFBB3E0925C6FD31CC4DFCF8" box="[987,1121,766,791]" italics="true" pageId="5" pageNumber="209">P. peruensis</emphasis>
|
||
</taxonomicName>
|
||
are vestigial, conspicuously short, and in
|
||
<taxonomicName id="ACE7F6BCFFBB3E092737FCEDC9C5FCD4" authorityName="Jałoszyński" authorityYear="2020" box="[298,489,801,827]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="species" species="akeratophorus">
|
||
<emphasis id="5993512DFFBB3E092737FCEDC9C5FCD4" box="[298,489,801,827]" italics="true" pageId="5" pageNumber="209">P. akeratophorus</emphasis>
|
||
</taxonomicName>
|
||
the aedeagus (
|
||
<figureCitation id="F3DC91BAFFBB3E09248DFCEECB3CFCD4" box="[656,784,801,827]" captionStart="FIGURES 17–23" captionStartId="5.[151,264,1785,1810]" captionTargetBox="[151,1430,993,1760]" captionTargetId="figure@5.[151,1436,993,1760]" captionTargetPageId="5" captionText="FIGURES 17–23. Protandroconnus akeratophorus sp. n., male. Prothorax in ventral view (17), pterothorax in ventral view (18), pterothorax and abdomen in ventral view (19), aedeagus in ventral (20, 22) and lateral (21, 23) views. Abbreviations: bst, basisternal portion of prosternum; dlf, dorsolateral fovea; hyr, hypomeral ridge; lfa, lateral furcal arm; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest." figureDoi="http://doi.org/10.5281/zenodo.3707360" httpUri="https://zenodo.org/record/3707360/files/figure.png" pageId="5" pageNumber="209">Figs 20–23</figureCitation>
|
||
) is devoid of any traces of parameres. All other structures are shared by both species, and those different in the Mexican
|
||
<taxonomicName id="ACE7F6BCFFBB3E092558FC8ACC2BFCB0" authorityName="Franz" authorityYear="1989" box="[837,1031,837,863]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5993512DFFBB3E092558FC8ACC2BFCB0" box="[837,1031,837,863]" italics="true" pageId="5" pageNumber="209">Protandroconnus</emphasis>
|
||
</taxonomicName>
|
||
(except for the free hypomeral lobe) seem to be reductions. The differences are too minor to justify placing the new species in a separate genus or even subgenus, and the more reasonable approach seems to be modification of the generic diagnosis of
|
||
<taxonomicName id="ACE7F6BCFFBB3E0922CEFC42CDB9FC48" authorityName="Franz" authorityYear="1989" box="[1235,1429,909,935]" class="Insecta" family="Staphylinidae" genus="Protandroconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5993512DFFBB3E0922CEFC42CDB9FC48" box="[1235,1429,909,935]" italics="true" pageId="5" pageNumber="209">Protandroconnus</emphasis>
|
||
</taxonomicName>
|
||
, to include the aforementioned variability in notosternal sutures, postcoxal hypomeral lobes and parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |