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<document id="7BCA649C866074988A15A8B28534218F" ID-DOI="10.5281/zenodo.5170505" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170505" ID-ZooBank="B39DE284-15C2-4ADE-B1B9-9146D3DFE764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628285093964" checkinUser="felipe" docAuthor="Ferro, Michael L." docDate="2016" docId="CB384340917CFFAD6CA2BB662C185323" docLanguage="en" docName="InsectaMundi.0472.1-57.pdf" docOrigin="Insecta Mundi 2016 (472)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Sonoma twaini Ferro 2016, new species" docType="treatment" docVersion="4" lastPageNumber="17" masterDocId="37013B38916EFFBF6C62B9072D725479" masterDocTitle="Fourteen new species of Sonoma Casey (Coleoptera: Staphylinidae: Pselaphinae) with a key to species from western North America" masterLastPageNumber="57" masterPageNumber="1" pageNumber="17" updateTime="1699040133542" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="05E824C827817C2632D8E9835CC59C02" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4831F4906CB9D62E9D807B8F88220F43">
<mods:title id="BD8CCCC94CBC222091B50753E39B2768">Fourteen new species of Sonoma Casey (Coleoptera: Staphylinidae: Pselaphinae) with a key to species from western North America</mods:title>
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<mods:namePart id="0071B4C355CFA77C95E875DCFD54F6C1">Ferro, Michael L.</mods:namePart>
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<mods:title id="C0755955552FD49D50D2282EFE6A95A2">Insecta Mundi</mods:title>
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<mods:part id="AC458F58F12E07629325B1006CE5210A">
<mods:date id="3301384E6657B7AC4734EE64B6D9366C">2016</mods:date>
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<mods:number id="82ABBBD27029665978739C6E6596495F">2016-03-11</mods:number>
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<mods:number id="C08E6985BEA2B650CDFBF95A9AC10921">472</mods:number>
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<mods:identifier id="BA5DFD30AA0CFE5D246CAEA1D50F594E" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="1D9EED66B43B81BC82C9F223817CFF03" type="Zenodo-Dep">5170505</mods:identifier>
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<treatment id="CB384340917CFFAD6CA2BB662C185323" ID-DOI="http://doi.org/10.5281/zenodo.5185131" ID-Zenodo-Dep="5185131" LSID="urn:lsid:plazi:treatment:CB384340917CFFAD6CA2BB662C185323" httpUri="http://treatment.plazi.org/id/CB384340917CFFAD6CA2BB662C185323" lastPageNumber="17" pageId="18" pageNumber="17">
<subSubSection id="0B8BA1DD917CFFAD6CA2BB662F185600" box="[192,618,609,634]" pageId="18" pageNumber="17" type="nomenclature">
<paragraph id="432EF256917CFFAD6CA2BB662F185600" blockId="18.[192,618,609,666]" box="[192,618,609,634]" pageId="18" pageNumber="17">
<heading id="1866453A917CFFAD6CA2BB662F185600" bold="true" box="[192,618,609,634]" fontSize="10" level="2" pageId="18" pageNumber="17" reason="3">
<emphasis id="71E52E44917CFFAD6CA2BB662F185600" bold="true" box="[192,618,609,634]" pageId="18" pageNumber="17">
13.
<taxonomicName id="849189D5917CFFAD6C8CBB662CCF5600" ID-CoL="4Y9W8" authority="Ferro, 2016" authorityName="Ferro" authorityYear="2016" box="[238,445,609,633]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="twaini" status="sp. nov.">
<emphasis id="71E52E44917CFFAD6C8CBB662CCF5600" bold="true" box="[238,445,609,633]" italics="true" pageId="18" pageNumber="17">Sonoma twaini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AD6933F917CFFAD6DA7BB652F185600" box="[453,618,609,634]" pageId="18" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD917CFFAD6CA2BB852E625000" pageId="18" pageNumber="17" type="description">
<paragraph id="432EF256917CFFAD6CA2BB852CC656E3" blockId="18.[192,618,609,666]" box="[192,436,642,666]" pageId="18" pageNumber="17">
<figureCitation id="DBAAEED3917CFFAD6CA2BB852C6556E3" box="[192,279,642,666]" captionStart="Figures 12–14" captionStartId="24.[192,288,1688,1710]" captionTargetBox="[426,1267,217,1634]" captionTargetId="figure-17@24.[328,1321,192,1690]" captionTargetPageId="24" captionText="Figures 12–14. Sonoma spp. holotype habiti. 12) S. stewarti. 13) S. twaini. 14) S. virgo." figureDoi="http://doi.org/10.5281/zenodo.5170511" httpUri="https://zenodo.org/record/5170511/files/figure.png" pageId="18" pageNumber="17">Fig. 13</figureCitation>
,
<figureCitation id="DBAAEED3917CFFAD6D46BB852C3656E3" box="[292,324,642,666]" captionStart="Figures 15–18" captionStartId="25.[192,288,1821,1843]" captionTargetBox="[217,1410,194,1796]" captionTargetId="figure-13@25.[192,1421,192,1801]" captionTargetPageId="25" captionText="Figures 15–18. Aedeagi of Sonoma spp. 15) S. stewarti new species. 16) S. wintuorum Chandler. Modified from Chandler (2003). 17) S. twaini new species. 18) S. colberti new species. Right side of figures is anatomical left." figureDoi="http://doi.org/10.5281/zenodo.5170513" httpUri="https://zenodo.org/record/5170513/files/figure.png" pageId="18" pageNumber="17">17</figureCitation>
; Map 39.
</paragraph>
<paragraph id="432EF256917CFFAD6CA2BBC628745763" blockId="18.[192,1440,705,1146]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6CA2BBC62C1856A0" bold="true" box="[192,362,705,729]" pageId="18" pageNumber="17">Description.</emphasis>
<typeStatus id="9C2A4CF4917CFFAD6D0DBBC52C9256A3" box="[367,480,706,730]" pageId="18" pageNumber="17" type="holotype">Holotype</typeStatus>
, male. Measurements: head 0.25 long, 0.31 wide; pronotum 0.30 long, 0.34 wide; elytra 0.59 long, 0.28 wide; antennomeres 1–11 total 0.57; total length 1.92. Body brown, maxillary palpi and legs paler. Body with moderate length setae (approximately 1/2 or less width of eye).
</paragraph>
<paragraph id="432EF256917CFFAD6C92BA2529BF5723" blockId="18.[192,1440,705,1146]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6C92BA252C435743" box="[240,305,802,826]" italics="true" pageId="18" pageNumber="17">Head</emphasis>
. Eyes prominent, maximum length in dorsal view 0.47x length of first antennal segment, with approximately 30 facets. Antennomere 2 approximately 0.90x width of 1; 3 smallest.
</paragraph>
<paragraph id="432EF256917CFFAD6C92BA652E4257E3" blockId="18.[192,1440,705,1146]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6C92BA652C3A5703" box="[240,328,866,890]" italics="true" pageId="18" pageNumber="17">Thorax</emphasis>
. Elytra with indistinct sutural foveae; 2 foveae lateral to sutural fovea; central row of 3 foveae in basal 1/3. Winged. Metatibia unmodified.
</paragraph>
<paragraph id="432EF256917CFFAD6C92BAA529B357A3" blockId="18.[192,1440,705,1146]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6C92BAA52C1157C3" box="[240,355,930,954]" italics="true" pageId="18" pageNumber="17">Abdomen</emphasis>
. Tergite IV with transverse patch of microtrichia narrowly interrupted at midline. Basal lateral foveae on ventrites V–VII. Basal pubescence present on all visible ventrites.
</paragraph>
<paragraph id="432EF256917CFFAD6C92BAE52E625000" blockId="18.[192,1440,705,1146]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6C92BAE52C165783" box="[240,356,994,1018]" italics="true" pageId="18" pageNumber="17">Aedeagus</emphasis>
. Elongate. Left paramere: equal in length to endophallus; subequal in width from base to apex; with 4–5 stout apical setae and 3 ventral subapical setae. Endophallus: widest at basal 1/5; with two ventral processes arising 2/3 from base projecting perpendicular to aedeagus; apex bent 90° to the left. Right paramere: 2/3 length left paramere; subequal in width from base to apex; with 4–5 stout apical setae and 3 ventral subapical setae.
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD917CFFAD6CA2BDA52F3C51E3" pageId="18" pageNumber="17" type="materials_examined">
<paragraph id="432EF256917CFFAD6CA2BDA52C4150A3" blockId="18.[192,1440,1185,1338]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6CA2BDA52CFF50C0" bold="true" box="[192,397,1185,1210]" pageId="18" pageNumber="17">Type Material.</emphasis>
<materialsCitation id="F3F9F80B917CFFAD6DF4BDA52C4150A3" collectionCode="T, R, CSCA" collectorName="Haig Whiskeytown Shasta Co. Cal." county="Haig Whiskeytown Shasta Co." location="Haig Whiskeytown Shasta Co." pageId="18" pageNumber="17" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9C2A4CF4917CFFAD6DF4BDA52F7A50C3" box="[406,520,1186,1210]" pageId="18" pageNumber="17" type="holotype">Holotype</typeStatus>
, male: *
<collectionCode id="25806A93917CFFAD6E0DBDA52FF150C3" box="[623,643,1186,1210]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="17">T</collectionCode>
.
<collectionCode id="25806A93917CFFAD6EE5BDA52FEF50C3" box="[647,669,1186,1210]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="17">R</collectionCode>
.
<collectorName id="EE649780917CFFAD6EC2BDA5292650C3" box="[672,1108,1186,1210]" pageId="18" pageNumber="17">
<collectingCounty id="AA4F8ADA917CFFAD6EC2BDA5295150C3" box="[672,1059,1186,1210]" pageId="18" pageNumber="17">Haig Whiskeytown Shasta Co.</collectingCounty>
Cal.
</collectorName>
III.9.1973 (
<specimenCount id="559739DF917CFFAD6889BDA5287E50C3" box="[1259,1292,1185,1210]" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
). Deposited in
<collectionCode id="25806A93917CFFAD6C82BDC52C5D50A3" box="[224,303,1218,1242]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="18" pageNumber="17" type="Museum">CSCA</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="432EF256917CFFAD6C92BDE529E35143" blockId="18.[192,1440,1185,1338]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6C92BDE52C085083" bold="true" box="[240,378,1250,1274]" pageId="18" pageNumber="17">
<typeStatus id="9C2A4CF4917CFFAD6C92BDE52C085083" box="[240,378,1250,1274]" pageId="18" pageNumber="17" type="paratype">Paratypes</typeStatus>
</emphasis>
(n=4).
<emphasis id="71E52E44917CFFAD6DAEBDE5296C5080" bold="true" box="[460,1054,1249,1274]" pageId="18" pageNumber="17">
<collectingCountry id="3B86B2C6917CFFAD6DAEBDE52FCA5080" box="[460,696,1249,1274]" name="United States of America" pageId="18" pageNumber="17">UNITED STATES</collectingCountry>
:
<collectingRegion id="81553CB4917CFFAD6EA1BDE62EF25080" box="[707,896,1249,1273]" country="United States of America" name="California" pageId="18" pageNumber="17">CALIFORNIA</collectingRegion>
:
<collectingCounty id="AA4F8ADA917CFFAD6FE8BDE6296A5080" box="[906,1048,1249,1273]" pageId="18" pageNumber="17">Shasta Co.</collectingCounty>
:
</emphasis>
*
<collectionCode id="25806A93917CFFAD684DBDE529305083" box="[1071,1090,1250,1274]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="17">T</collectionCode>
.
<collectionCode id="25806A93917CFFAD6824BDE529295083" box="[1094,1115,1250,1274]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="17">R</collectionCode>
.Haig Whiskeytown Shasta Co.Cal. III.9.1973 (
<collectionCode id="25806A93917CFFAD6DC6BC052C825163" box="[420,496,1282,1306]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="18" pageNumber="17" type="Museum">CSCA</collectionCode>
) (
<specimenCount id="559739DF917CFFAD6E64BC052F5B5163" box="[518,553,1281,1306]" pageId="18" pageNumber="17" type="male">2♂</specimenCount>
). *
<collectionCode id="25806A93917CFFAD6E29BC052F2C5163" box="[587,606,1282,1306]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="17">T</collectionCode>
.
<collectionCode id="25806A93917CFFAD6E00BC052F0A5163" box="[610,632,1282,1306]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="17">R</collectionCode>
.Haig Whiskeytown Shasta Co.Cal. I.31.1973 (
<collectionCode id="25806A93917CFFAD68C6BC0529825163" box="[1188,1264,1282,1306]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="18" pageNumber="17" type="Museum">CSCA</collectionCode>
) (
<specimenCount id="559739DF917CFFAD696ABC05285A5163" box="[1288,1320,1281,1306]" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
).
<emphasis id="71E52E44917CFFAD695CBC062D865140" bold="true" pageId="18" pageNumber="17">
<collectingCounty id="AA4F8ADA917CFFAD695CBC062D9C5140" pageId="18" pageNumber="17">Trinity Co.</collectingCounty>
:
</emphasis>
*
<collectionCode id="25806A93917CFFAD6D65BC252C105143" box="[263,354,1314,1338]" pageId="18" pageNumber="17">CALIF</collectionCode>
:Trinity Co. Junction City
<date id="372FD496917CFFAD6EC0BC252E5F5143" box="[674,813,1314,1338]" pageId="18" pageNumber="17" value="1981-03-12">III-12-1981</date>
<collectionCode id="25806A93917CFFAD6F57BC252E3A5143" box="[821,840,1314,1338]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="17">T</collectionCode>
.
<collectionCode id="25806A93917CFFAD6F2EBC252E105143" box="[844,866,1314,1338]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="17">R</collectionCode>
.Haig coll. (
<collectionCode id="25806A93917CFFAD6F8FBC2529345143" box="[1005,1094,1314,1338]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="18" pageNumber="17" type="Museum">FMNH</collectionCode>
) (
<specimenCount id="559739DF917CFFAD6802BC2529F25143" box="[1120,1152,1313,1338]" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
).
</paragraph>
<paragraph id="432EF256917CFFAD6CA2BC662F3C51E3" blockId="18.[192,1440,1377,1434]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6CA2BC662F4A5100" bold="true" box="[192,568,1377,1401]" pageId="18" pageNumber="17">Geographical Distribution.</emphasis>
<taxonomicName id="849189D5917CFFAD6E23BC652F8F5100" authorityName="Ferro" authorityYear="2016" box="[577,765,1378,1401]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E44917CFFAD6E23BC652F8F5100" box="[577,765,1378,1401]" italics="true" pageId="18" pageNumber="17">Sonoma twaini</emphasis>
</taxonomicName>
is only known from two locations, one each in Shasta and Trinity counties,
<collectingRegion id="81553CB4917CFFAD6DA8BC852F3851E3" box="[458,586,1410,1434]" country="United States of America" name="California" pageId="18" pageNumber="17">California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD917CFFAD6CA2BCC6285752C0" pageId="18" pageNumber="17" type="discussion">
<paragraph id="432EF256917CFFAD6CA2BCC6281451A3" blockId="18.[192,1440,1473,1722]" box="[192,1382,1473,1498]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6CA2BCC62C2951A0" bold="true" box="[192,347,1473,1497]" pageId="18" pageNumber="17">Comments.</emphasis>
<taxonomicName id="849189D5917CFFAD6D00BCC52F6E51A0" authorityName="Ferro" authorityYear="2016" box="[354,540,1474,1497]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E44917CFFAD6D00BCC52F6E51A0" box="[354,540,1474,1497]" italics="true" pageId="18" pageNumber="17">Sonoma twaini</emphasis>
</taxonomicName>
was collected in January and March, collection method is unknown.
</paragraph>
<paragraph id="432EF256917CFFAD6C92BCE5285752C0" blockId="18.[192,1440,1473,1722]" pageId="18" pageNumber="17">
Aedeagal characters of
<taxonomicName id="849189D5917CFFAD6E78BCE52FFF5180" authorityName="Ferro" authorityYear="2016" box="[538,653,1506,1529]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E44917CFFAD6E78BCE52FFF5180" box="[538,653,1506,1529]" italics="true" pageId="18" pageNumber="17">S. twaini</emphasis>
</taxonomicName>
are superficially similar to
<taxonomicName id="849189D5917CFFAD6F96BCE529D35180" authorityName="Chandler. Modified" authorityYear="2003" box="[1012,1185,1506,1529]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="wintuorum">
<emphasis id="71E52E44917CFFAD6F96BCE529D35180" box="[1012,1185,1506,1529]" italics="true" pageId="18" pageNumber="17">S. wintuorum</emphasis>
</taxonomicName>
,
<taxonomicName id="849189D5917CFFAD68D6BCE528495180" authorityName="Ferro" authorityYear="2016" box="[1204,1339,1506,1529]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="stewarti">
<emphasis id="71E52E44917CFFAD68D6BCE528495180" box="[1204,1339,1506,1529]" italics="true" pageId="18" pageNumber="17">S. stewarti</emphasis>
</taxonomicName>
, and
<taxonomicName id="849189D5917CFFAD69E5BCE52C695263" authorityName="Ferro" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E44917CFFAD69E5BCE52C695263" italics="true" pageId="18" pageNumber="17">S. colberti</emphasis>
</taxonomicName>
all of which share a similar gestalt comprised of narrow, elongate parameres and an elongate endophallus with elaborate apical hooks or processes. The flowing combination of characters will separate
<taxonomicName id="849189D5917CFFAD6C99BF452C195220" authorityName="Ferro" authorityYear="2016" box="[251,363,1602,1625]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E44917CFFAD6C99BF452C195220" box="[251,363,1602,1625]" italics="true" pageId="18" pageNumber="17">S. twaini</emphasis>
</taxonomicName>
from the others: endophallus with two ventral processes arising 2/3 from base projecting perpendicular to aedeagus [absent in
<taxonomicName id="849189D5917CFFAD6EDFBF652E155200" authorityName="Chandler. Modified" authorityYear="2003" box="[701,871,1634,1657]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="wintuorum">
<emphasis id="71E52E44917CFFAD6EDFBF652E155200" box="[701,871,1634,1657]" italics="true" pageId="18" pageNumber="17">S. wintuorum</emphasis>
</taxonomicName>
,
<taxonomicName id="849189D5917CFFAD6F17BF652E8B5200" authorityName="Ferro" authorityYear="2016" box="[885,1017,1634,1657]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="stewarti">
<emphasis id="71E52E44917CFFAD6F17BF652E8B5200" box="[885,1017,1634,1657]" italics="true" pageId="18" pageNumber="17">S. stewarti</emphasis>
</taxonomicName>
, and
<taxonomicName id="849189D5917CFFAD685CBF6529CB5203" authorityName="Ferro" authorityYear="2016" box="[1086,1209,1634,1658]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E44917CFFAD685CBF6529CB5203" box="[1086,1209,1634,1658]" italics="true" pageId="18" pageNumber="17">S. colberti</emphasis>
</taxonomicName>
]; without elongate thin posteriorly directed process arising 2/3 from base of endophallus [present in
<taxonomicName id="849189D5917CFFAD68C3BF85283952E0" authorityName="Chandler. Modified" authorityYear="2003" box="[1185,1355,1666,1689]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="wintuorum">
<emphasis id="71E52E44917CFFAD68C3BF85283952E0" box="[1185,1355,1666,1689]" italics="true" pageId="18" pageNumber="17">S. wintuorum</emphasis>
</taxonomicName>
and
<taxonomicName id="849189D5917CFFAD69E5BF852C5152C0" authorityName="Ferro" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="stewarti">
<emphasis id="71E52E44917CFFAD69E5BF852C5152C0" italics="true" pageId="18" pageNumber="17">S. stewarti</emphasis>
</taxonomicName>
]; endophallus with single apical process [two in
<taxonomicName id="849189D5917CFFAD6F1ABFA5295352C0" authorityName="Chandler. Modified" authorityYear="2003" box="[888,1057,1698,1721]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="wintuorum">
<emphasis id="71E52E44917CFFAD6F1ABFA5295352C0" box="[888,1057,1698,1721]" italics="true" pageId="18" pageNumber="17">S. wintuorum</emphasis>
</taxonomicName>
, three in
<taxonomicName id="849189D5917CFFAD68FBBFA5286652C3" authorityName="Ferro" authorityYear="2016" box="[1177,1300,1698,1722]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="colberti">
<emphasis id="71E52E44917CFFAD68FBBFA5286652C3" box="[1177,1300,1698,1722]" italics="true" pageId="18" pageNumber="17">S. colberti</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="0B8BA1DD917CFFAD6CA2BFE52C185323" pageId="18" pageNumber="17" type="etymology">
<paragraph id="432EF256917CFFAD6CA2BFE52C185323" blockId="18.[192,1440,1762,1882]" pageId="18" pageNumber="17">
<emphasis id="71E52E44917CFFAD6CA2BFE52C2E5283" bold="true" box="[192,348,1762,1786]" pageId="18" pageNumber="17">Etymology.</emphasis>
<taxonomicName id="849189D5917CFFAD6D00BFE52F6E5280" authorityName="Ferro" authorityYear="2016" box="[354,540,1762,1785]" class="Insecta" family="Staphylinidae" genus="Sonoma" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="twaini">
<emphasis id="71E52E44917CFFAD6D00BFE52F6E5280" box="[354,540,1762,1785]" italics="true" pageId="18" pageNumber="17">Sonoma twaini</emphasis>
</taxonomicName>
is named for the character Mark Twain, developed by Samuel Langhorne Clemens, an author, lecturer, philosopher, humanitarian, champion of science, and humorist. Clemens lived in
<collectingRegion id="81553CB4917CFFAD6D42BE252CEE5343" box="[288,412,1826,1850]" country="United States of America" name="California" pageId="18" pageNumber="17">California</collectingRegion>
for awhile, but traveled nowhere near where this species occurs—the author forgives the oversight.
</paragraph>
</subSubSection>
</treatment>
</document>
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