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<document id="85F1304318B9FA22517BD7335EA1C4A8" ID="10.11646/zootaxa.4154.1.5" ID-DOI="10.11646/zootaxa.4154.1.5" ID-GBIF-Dataset="9c2d1a98-2453-43f6-9023-f4e2dd0c0123" ID-ISSN="1175-5326" ID-Zenodo-Dep="256492" ID-ZooBank="3CD75005-7EB2-4FC7-8569-7C9E80213FF1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1471503941698" checkinUser="plazi" docAuthor="Lyon, Robert Gary, Geiger, Matthias F. &amp; Freyhof, Jörg" docDate="2016" docId="B459F366442695176693F9BAFD4027C7" docLanguage="en" docName="zootaxa.4154.1.5.pdf" docOrigin="Zootaxa 4154 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra sindhi Lyon, Geiger &amp; Freyhof, 2016, new species" docType="treatment" docVersion="8" lastPageNumber="87" masterDocId="48608B1E4427951F6604FFE0FFD0266B" masterDocTitle="Garra sindhi, a new species from the Jebel Samhan Nature Reserve in Oman (Teleostei: Cyprinidae)" masterLastPageNumber="88" masterPageNumber="79" pageNumber="80" updateTime="1698382039167" updateUser="plazi">
<mods:mods id="C050D1AB24B6B20171D8A1AD1689383F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E7DA82DC613D04AA31B4DBEF749F5BA4">
<mods:title id="8757EBCA872D02CF93EB5A3F8C0E8AFA">Garra sindhi, a new species from the Jebel Samhan Nature Reserve in Oman (Teleostei: Cyprinidae)</mods:title>
</mods:titleInfo>
<mods:name id="74B06C3CEC8BEC2068A320A268A7CA5D" type="personal">
<mods:role id="FE91CB1DCB8ABD077230BCA69BDC0BC8">
<mods:roleTerm id="6B38D29C0CBF6665946B058F55AFA57A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7941346154E70AF1D0C6DF4B44824A8E">Lyon, Robert Gary</mods:namePart>
</mods:name>
<mods:name id="3059853E15ACCBDF02D114AC29EBA99B" type="personal">
<mods:role id="5516DC0D084764E269E69DC9EE06AF8D">
<mods:roleTerm id="4076A3B5C80515CC0E41AF8FFDDC4F00">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B0AE7C7E39F01E3FF8B1C1D8ECD796E0">Geiger, Matthias F.</mods:namePart>
</mods:name>
<mods:name id="66599D2C0E335E4205A6519AA465D511" type="personal">
<mods:role id="062C0099EEFC84AED5548500E8367C57">
<mods:roleTerm id="C4B5D0ADDFE292C5FC19FF7798C586D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6E1EBFFE82462621F7D2413E90D18BA">Freyhof, Jörg</mods:namePart>
</mods:name>
<mods:typeOfResource id="7F35177761414A921D5EBCCAAF953933">text</mods:typeOfResource>
<mods:relatedItem id="D50A69BC2AB7F493B4287D651645916A" type="host">
<mods:titleInfo id="102AC24D92F5F17D643A21312FA94447">
<mods:title id="38A4AA7DFCC8A7126BBDF2C863851B6E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="97C0D2CE27C6B87FCFA0658237F37860">
<mods:date id="2C4F7B49899326075D820855B768B2E0">2016</mods:date>
<mods:detail id="B1D579D554F4FCE473A882529750D00D" type="volume">
<mods:number id="7F483DBDA31096D4256178ED4E2CC7BA">4154</mods:number>
</mods:detail>
<mods:detail id="9F5723D1B24F171F036338BE7B52DFFE" type="issue">
<mods:number id="0979AAD355C109136907B9855DCF010F">1</mods:number>
</mods:detail>
<mods:extent id="D03D9C307EF5C239B07E1E759AF645CD" unit="page">
<mods:start id="6E97092343AC70C2194E243540A577A5">79</mods:start>
<mods:end id="1B1810C2DFCF8CF47DBE060AC25296A4">88</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2D5EBFAAFB92BA902D03440EAB5B08EC">journal article</mods:classification>
<mods:identifier id="2F2AFE26ECB9058261495C87E4368ED1" type="DOI">10.11646/zootaxa.4154.1.5</mods:identifier>
<mods:identifier id="27D46E20C151CB4AD1FAFAA8B375A366" type="GBIF-Dataset">9c2d1a98-2453-43f6-9023-f4e2dd0c0123</mods:identifier>
<mods:identifier id="0E224E39A16690EDD3CA554A16C28335" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="11ACF3C37C817B6F517C7885AB3EAD68" type="Zenodo-Dep">256492</mods:identifier>
<mods:identifier id="9A5D318B06F41F16A962FD1C1FF055A0" type="ZooBank">3CD75005-7EB2-4FC7-8569-7C9E80213FF1</mods:identifier>
</mods:mods>
<treatment id="B459F366442695176693F9BAFD4027C7" ID-DOI="http://doi.org/10.5281/zenodo.5662518" ID-GBIF-Taxon="124379756" ID-Zenodo-Dep="5662518" LSID="urn:lsid:plazi:treatment:B459F366442695176693F9BAFD4027C7" httpUri="http://treatment.plazi.org/id/B459F366442695176693F9BAFD4027C7" lastPageId="8" lastPageNumber="87" pageId="1" pageNumber="80">
<subSubSection id="74EA11FB4426951E6693F9BAFEC220FD" pageId="1" pageNumber="80" type="nomenclature">
<paragraph id="3C4F42704426951E6693F9BAFE01201E" blockId="1.[151,465,1626,1686]" box="[151,465,1626,1653]" pageId="1" pageNumber="80">
<heading id="6707F51C4426951E6693F9BAFE01201E" bold="true" box="[151,465,1626,1653]" fontSize="11" level="1" pageId="1" pageNumber="80" reason="1">
<emphasis id="0E849E624426951E6693F9BAFE01201E" bold="true" box="[151,465,1626,1653]" pageId="1" pageNumber="80">
<taxonomicName id="FBF039F34426951E6693F9BAFEE2201F" box="[151,306,1626,1652]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="80" phylum="Chordata" rank="species" species="sindhi" status="sp. nov.">
<emphasis id="0E849E624426951E6693F9BAFEE2201F" bold="true" box="[151,306,1626,1652]" italics="true" pageId="1" pageNumber="80">Garra sindhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="15B723194426951E6745F9BBFE01201E" box="[321,465,1627,1653]" pageId="1" pageNumber="80" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3C4F42704426951E6693F99DFEC220FD" blockId="1.[151,465,1626,1686]" box="[151,274,1661,1686]" pageId="1" pageNumber="80">
(
<figureCitation id="A4CB5EF54426951E669BF99DFEDA20FD" box="[159,266,1661,1686]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="3.[151,249,621,643]" captionStartId-1="5.[151,249,1134,1156]" captionStartId-2="5.[151,249,1950,1972]" captionStartId-3="7.[151,249,1429,1451]" captionTargetBox-0="[151,1436,193,599]" captionTargetBox-1="[151,1436,193,1112]" captionTargetBox-2="[151,1436,1206,1928]" captionTargetBox-3="[151,1435,445,1408]" captionTargetId-0="figure@3.[151,1437,193,599]" captionTargetId-1="figure@5.[151,1436,193,1113]" captionTargetId-2="figure@5.[151,1436,1206,1929]" captionTargetId-3="figure@7.[151,1436,445,1408]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="7" captionText-0="FIGURE 2. Garra sindhi, ONHM 4196, holotype, 82.5 mm SL; Oman: Wadi Andhur." captionText-1="FIGURE 4. Garra sindhi, ONHM 4219, paratype. 48.7 mm SL; Oman: Wadi Andhur." captionText-2="FIGURE 5. Garra sindhi, about 45 mm SL, not preserved; Oman: Wadi Andhur." captionText-3="FIGURE 6. Pool in Wadi Andhur." httpUri-0="https://zenodo.org/record/256493/files/figure.png" httpUri-1="https://zenodo.org/record/256494/files/figure.png" httpUri-2="https://zenodo.org/record/256495/files/figure.png" httpUri-3="https://zenodo.org/record/256496/files/figure.png" pageId="1" pageNumber="80">Figs. 2–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="74EA11FB4426951E6693F925FB012105" pageId="1" pageNumber="80" type="materials_examined">
<paragraph id="3C4F42704426951E6693F925FDB32169" blockId="1.[151,1436,1733,2010]" pageId="1" pageNumber="80">
<materialsCitation id="8C98482D4426951E6693F925FDB32169" ID-GBIF-Occurrence="1291640431" collectingDate="2015-08-03" collectionCode="ONHM" collectorName="R. G. Lyon." country="Oman" latitude="17.5686" location="Wadi Andhur" longLatPrecision="1" longitude="54.70487" pageId="1" pageNumber="80" specimenCode="ONHM 4196, 82" specimenCount="1" stateProvince="Dhofar" typeStatus="holotype">
<emphasis id="0E849E624426951E6693F925FEDB20B5" bold="true" box="[151,267,1733,1758]" pageId="1" pageNumber="80">
<typeStatus id="E34BFCD24426951E6693F925FED620B5" box="[151,262,1733,1758]" pageId="1" pageNumber="80" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="6C56EA0B4426951E6719F926FE2420B5" box="[285,500,1733,1758]" pageId="1" pageNumber="80">
<collectionCode id="5AE1DAB54426951E6719F926FEA220B5" box="[285,370,1734,1758]" pageId="1" pageNumber="80">ONHM</collectionCode>
4196, 82
</specimenCode>
mm SL;
<collectingCountry id="44E702E04426951E6484F925FCBF20B5" box="[640,879,1733,1758]" name="Oman" pageId="1" pageNumber="80">Sultanate of Oman</collectingCountry>
:
<collectingRegion id="FE348C924426951E6581F925FC0420B5" box="[901,980,1733,1758]" country="Oman" name="Zufar" pageId="1" pageNumber="80">Dhofar</collectingRegion>
prov.:
<location id="392F14AB4426951E623EF925FB0F20B5" LSID="urn:lsid:plazi:treatment:B459F366442695176693F9BAFD4027C7:392F14AB4426951E623EF925FB0F20B5" box="[1082,1247,1733,1758]" country="Oman" latitude="17.5686" longLatPrecision="1" longitude="54.70487" name="Wadi Andhur" pageId="1" pageNumber="80" stateProvince="Dhofar">Wadi Andhur</location>
,
<geoCoordinate id="59C424B74426951E62F1F925FA4C20B5" box="[1269,1436,1733,1758]" direction="north" orientation="latitude" pageId="1" pageNumber="80" precision="1" value="17.5686">17°34'6.956&quot;N</geoCoordinate>
<geoCoordinate id="59C424B74426951E6693F909FE9B2169" box="[151,331,1769,1794]" direction="east" orientation="longitude" pageId="1" pageNumber="80" precision="1" value="54.70487">54°42'17.518&quot;E</geoCoordinate>
;
<date id="484E64B04426951E6753F90AFE0F2169" box="[343,479,1769,1794]" pageId="1" pageNumber="80" value="2015-08-03">
<collectingDate id="580A9D584426951E6753F90AFE0F2169" box="[343,479,1769,1794]" pageId="1" pageNumber="80" value="2015-08-03">3 Aug 2015</collectingDate>
</date>
,
<collectorName id="910527A64426951E67EEF90AFDB32169" box="[490,611,1770,1794]" pageId="1" pageNumber="80">R.G. Lyon.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="3C4F42704426951E66C3F8EDFB012105" blockId="1.[151,1436,1733,2010]" pageId="1" pageNumber="80">
<materialsCitation id="8C98482D4426951E66C3F8EDFDE92121" ID-GBIF-Occurrence="1291640430" collectingDate="2015-08-03" collectionCode="ONHM" collectorName="R. G. Lyon." country="Oman" latitude="17.5686" location="Wadi Andhur" longLatPrecision="1" longitude="54.70487" pageId="1" pageNumber="80" specimenCode="ONHM 4197, 3" specimenCount="1" stateProvince="Dhofar" typeStatus="paratype">
<emphasis id="0E849E624426951E66C3F8EDFE96214D" bold="true" box="[199,326,1805,1830]" pageId="1" pageNumber="80">
<typeStatus id="E34BFCD24426951E66C3F8EDFE91214D" box="[199,321,1805,1830]" pageId="1" pageNumber="80" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="6C56EA0B4426951E674AF8EEFDD4214D" box="[334,516,1806,1830]" pageId="1" pageNumber="80">
<collectionCode id="5AE1DAB54426951E674AF8EEFE73214D" box="[334,419,1806,1830]" pageId="1" pageNumber="80">ONHM</collectionCode>
4197, 3
</specimenCode>
,
<quantity id="FB08EF954426951E6416F8EDFD5C214E" box="[530,652,1805,1830]" metricMagnitude="-2" metricUnit="m" metricValue="6.15" metricValueMax="8.5" metricValueMin="3.8" pageId="1" pageNumber="80" unit="mm" value="61.5" valueMax="85.0" valueMin="38.0">38–85 mm</quantity>
SL; same data as holotype; Genbank accession number:
<accessionNumber id="23A3DF934426951E631DF8EDFA48214D" box="[1305,1432,1805,1830]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244649" pageId="1" pageNumber="80">KX244649</accessionNumber>
,
<accessionNumber id="23A3DF934426951E6693F8D1FEC62121" box="[151,278,1841,1866]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244650" pageId="1" pageNumber="80">KX244650</accessionNumber>
,
<accessionNumber id="23A3DF934426951E6727F8D1FE712121" box="[291,417,1841,1866]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244651" pageId="1" pageNumber="80">KX244651</accessionNumber>
,
<accessionNumber id="23A3DF934426951E67AAF8D1FDFD2121" box="[430,557,1841,1866]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244652" pageId="1" pageNumber="80">KX244652</accessionNumber>
).
</materialsCitation>
—FSJF 3527, 5,
<quantity id="FB08EF954426951E64FAF8D1FCA82122" box="[766,888,1841,1866]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.2" metricValueMin="3.9" pageId="1" pageNumber="80" unit="mm" value="45.5" valueMax="52.0" valueMin="39.0">39–52 mm</quantity>
SL;
<materialsCitation id="8C98482D4426951E65B4F8D2FB012105" ID-GBIF-Occurrence="1291640429" collectingDate="2015-11-09" collectionCode="ONHM" collectorName="R. G. Lyon." country="Oman" latitude="17.570341" location="Wadi Andhur" longLatPrecision="1" longitude="54.703392" pageId="1" pageNumber="80" specimenCode="ONHM 4219, 30" specimenCount="1" stateProvince="Dhofar" typeStatus="paratype">
<specimenCode id="6C56EA0B4426951E65B4F8D2FBA42121" box="[944,1140,1842,1866]" pageId="1" pageNumber="80">
<collectionCode id="5AE1DAB54426951E65B4F8D2FBD52121" box="[944,1029,1842,1866]" pageId="1" pageNumber="80">ONHM</collectionCode>
4219, 30
</specimenCode>
,
<quantity id="FB08EF954426951E6285F8D1FB2B2122" box="[1153,1275,1841,1866]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="5.7" metricValueMin="3.0" pageId="1" pageNumber="80" unit="mm" value="43.5" valueMax="57.0" valueMin="30.0">30–57 mm</quantity>
SL;
<collectingCountry id="44E702E04426951E6330F8D1FF2E2105" name="Oman" pageId="1" pageNumber="80">Sultanate of Oman</collectingCountry>
:
<collectingRegion id="FE348C924426951E670CF8B5FE872105" box="[264,343,1877,1902]" country="Oman" name="Zufar" pageId="1" pageNumber="80">Dhofar</collectingRegion>
prov.:
<location id="392F14AB4426951E67A2F8B5FDEE2105" LSID="urn:lsid:plazi:treatment:B459F366442695176693F9BAFD4027C7:392F14AB4426951E67A2F8B5FDEE2105" box="[422,574,1877,1902]" country="Oman" latitude="17.570341" longLatPrecision="1" longitude="54.703392" name="Wadi Andhur" pageId="1" pageNumber="80" stateProvince="Dhofar">Wadi Andhur</location>
,
<geoCoordinate id="59C424B74426951E644EF8B6FD2F2105" box="[586,767,1878,1902]" direction="north" orientation="latitude" pageId="1" pageNumber="80" precision="1" value="17.570341">17°34'13.224&quot;N</geoCoordinate>
<geoCoordinate id="59C424B74426951E6502F8B5FC6A2105" box="[774,954,1877,1902]" direction="east" orientation="longitude" pageId="1" pageNumber="80" precision="1" value="54.703392">54°42'12.211&quot;E</geoCoordinate>
;
<date id="484E64B04426951E65C1F8B6FB9C2105" box="[965,1100,1877,1902]" pageId="1" pageNumber="80" value="2015-11-09">
<collectingDate id="580A9D584426951E65C1F8B6FB9C2105" box="[965,1100,1877,1902]" pageId="1" pageNumber="80" value="2015-11-09">9 Nov 2015</collectingDate>
</date>
,
<collectorName id="910527A64426951E625CF8B6FB012105" box="[1112,1233,1878,1902]" pageId="1" pageNumber="80">R.G. Lyon.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="74EA11FB4426951D66C3F899FB8B232A" lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80" type="diagnosis">
<paragraph id="3C4F42704426951D66C3F899FF39240B" blockId="1.[151,1436,1733,2010]" lastBlockId="2.[151,1437,151,608]" lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80">
<emphasis id="0E849E624426951E66C3F899FE9121F9" bold="true" box="[199,321,1913,1938]" pageId="1" pageNumber="80">Diagnosis.</emphasis>
<taxonomicName id="FBF039F34426951E6743F89AFE0721F9" box="[327,471,1913,1938]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="80" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624426951E6743F89AFE0721F9" box="[327,471,1913,1938]" italics="true" pageId="1" pageNumber="80">Garra sindhi</emphasis>
</taxonomicName>
is distinguished from the other two species of
<taxonomicName id="FBF039F34426951E65D9F89AFBF321FA" box="[989,1059,1914,1937]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="0E849E624426951E65D9F89AFBF321FA" box="[989,1059,1914,1937]" italics="true" pageId="1" pageNumber="80">Garra</emphasis>
</taxonomicName>
of the Omani
<collectingRegion id="FE348C924426951E62C6F899FAC121F9" box="[1218,1297,1913,1938]" country="Oman" name="Zufar" pageId="1" pageNumber="80">Dhofar</collectingRegion>
province by the following combination of characters. It differs from
<emphasis id="0E849E624426951E6537F87EFC4A21DE" box="[819,922,1950,1973]" italics="true" pageId="1" pageNumber="80">G. s m a r t i</emphasis>
by having the belly and breast behind the pectoral-fin origin fully covered by deeply embedded scales (vs. scales absent in front of the posterior pectoral-fin base), 8–12 gill rakers on lower limb of first gill arch (vs. 13–15), maximum body depth at about middle between nape and dorsal-fin origin (vs. maximum body depth at dorsal-fin origin), a prominent hump on back in some individuals (vs. no hump) and the oral disc being wider than long (vs. longer than wide).
<taxonomicName id="FBF039F34425951D62E0FF00FAAB2693" box="[1252,1403,223,248]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624425951D62E0FF00FAAB2693" box="[1252,1403,223,248]" italics="true" pageId="2" pageNumber="81">Garra sindhi</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="FBF039F34425951D677CFEE5FE252776" box="[376,501,260,285]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="dunsirei">
<emphasis id="0E849E624425951D677CFEE5FE252776" box="[376,501,260,285]" italics="true" pageId="2" pageNumber="81">G. dunsirei</emphasis>
</taxonomicName>
, a troglomorphic species, by having normally developed eyes, visible in ventral view (vs. very small, not visible in ventral view), 14–16 circumpeduncular scales (vs. 12), 8–12 gill rakers on lower limb of first gill arch (vs. 6–7) and breast and belly fully covered by deeply embedded scales (vs. scales reduced or absent).
<taxonomicName id="FBF039F34425951D6777FE90FDDB27E3" box="[371,523,367,392]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="lautior">
<emphasis id="0E849E624425951D6777FE90FDDB27E3" box="[371,523,367,392]" italics="true" pageId="2" pageNumber="81">Garra lautior</emphasis>
</taxonomicName>
and
<taxonomicName id="FBF039F34425951D6446FE90FD3727E3" box="[578,743,367,392]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="mamshuqua">
<emphasis id="0E849E624425951D6446FE90FD3727E3" box="[578,743,367,392]" italics="true" pageId="2" pageNumber="81">G. mamshuqua</emphasis>
</taxonomicName>
are two species from Wadi Hadramut, which is situated about
<quantity id="FB08EF954425951D6693FE74FF3E27C6" box="[151,238,404,429]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="81" unit="km" value="600.0">600 km</quantity>
west of Wadi Andhur.
<taxonomicName id="FBF039F34425951D67F8FE75FD5F27C6" box="[508,655,404,429]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624425951D67F8FE75FD5F27C6" box="[508,655,404,429]" italics="true" pageId="2" pageNumber="81">Garra sindhi</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="FBF039F34425951D6597FE75FBD027C6" box="[915,1024,404,429]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="lautior">
<emphasis id="0E849E624425951D6597FE75FBD027C6" box="[915,1024,404,429]" italics="true" pageId="2" pageNumber="81">G. lautior</emphasis>
</taxonomicName>
by having a fully-scaled breast and belly (vs. naked in front of anal fin) and from
<taxonomicName id="FBF039F34425951D64ACFE58FC9F27BB" box="[680,847,439,464]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="mamshuqua">
<emphasis id="0E849E624425951D64ACFE58FC9F27BB" box="[680,847,439,464]" italics="true" pageId="2" pageNumber="81">G. mamshuqua</emphasis>
</taxonomicName>
by lacking tubercles on snout (vs. present), and the central callous pad being much longer than the anteromedian fold (vs. about equal). Our molecular data show that
<taxonomicName id="FBF039F34425951D6693FDE0FF292473" box="[151,249,511,536]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624425951D6693FDE0FF292473" box="[151,249,511,536]" italics="true" pageId="2" pageNumber="81">G. sindhi</emphasis>
</taxonomicName>
is only distantly related to the
<taxonomicName id="FBF039F34425951D6450FDE0FD4A247C" box="[596,666,512,535]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis id="0E849E624425951D6450FDE0FD4A247C" box="[596,666,512,535]" italics="true" pageId="2" pageNumber="81">Garra</emphasis>
</taxonomicName>
species found in the north of
<collectingCountry id="44E702E04425951D65E2FDE0FBFB2473" box="[998,1067,512,536]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
(
<taxonomicName id="FBF039F34425951D623FFDE0FB052473" box="[1083,1237,511,536]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E624425951D623FFDE0FB052473" box="[1083,1237,511,536]" italics="true" pageId="2" pageNumber="81">G. barreimiae</emphasis>
</taxonomicName>
,
<emphasis id="0E849E624425951D62E7FDE0FEC52456" italics="true" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D62E7FDE0FAAA2473" box="[1251,1402,511,536]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="gallagheri">G. gallagheri</taxonomicName>
,
<taxonomicName id="FBF039F34425951D6380FDE0FEC52456" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="longipinnis">G. longipinnis</taxonomicName>
</emphasis>
) and it is distinguished from these by lacking tubercles on the head (vs. present at least in mature males).
</paragraph>
<paragraph id="3C4F42704425951D64C6FD53FA5D253B" blockId="2.[360,1421,687,1704]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D64C6FD53FB1324AD" authority="Wadi Bani Khalid Oman 2578" authorityName="Wadi Bani Khalid Oman" authorityYear="2578" box="[706,1219,687,710]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="0E849E624425951D64C6FD53FCA524AE" box="[706,885,690,709]" italics="true" pageId="2" pageNumber="81">Garra gallagheri</emphasis>
Wadi Bani Khalid
<collectingCountry id="44E702E04425951D6244FD4FFB5424AD" box="[1088,1156,687,710]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
2578
</taxonomicName>
<accessionNumber id="23A3DF934425951D62CDFD4FFAEE24AD" box="[1225,1342,687,710]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244647" pageId="2" pageNumber="81">KX244647</accessionNumber>
93-82-97
<taxonomicName id="FBF039F34425951D64C6FD2FFB132489" authority="Wadi Bani Khalid Oman 2578" authorityName="Wadi Bani Khalid Oman" authorityYear="2578" box="[706,1219,715,738]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="0E849E624425951D64C6FD2FFCA5248B" box="[706,885,717,737]" italics="true" pageId="2" pageNumber="81">Garra gallagheri</emphasis>
Wadi Bani Khalid
<collectingCountry id="44E702E04425951D6244FD2BFB542489" box="[1088,1156,715,738]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
2578
</taxonomicName>
<accessionNumber id="23A3DF934425951D62CDFD2BFAEE2489" box="[1225,1342,715,738]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244648" pageId="2" pageNumber="81">KX244648</accessionNumber>
<taxonomicName id="FBF039F34425951D64CCFD0AFB192496" authority="Wadi Bani Khalid Oman 2578" authorityName="Wadi Bani Khalid Oman" authorityYear="2578" box="[712,1225,742,765]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="0E849E624425951D64CCFD0AFCAB2497" box="[712,891,745,764]" italics="true" pageId="2" pageNumber="81">Garra gallagheri</emphasis>
Wadi Bani Khalid
<collectingCountry id="44E702E04425951D6242FD06FB5A2496" box="[1094,1162,742,765]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
2578
</taxonomicName>
<accessionNumber id="23A3DF934425951D62CBFD06FA932496" box="[1231,1347,742,765]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244646" pageId="2" pageNumber="81">KX244646</accessionNumber>
95-99-99
<taxonomicName id="FBF039F34425951D64CCFCE6FB192572" authority="Wadi Bani Khalid Oman 2578" authorityName="Wadi Bani Khalid Oman" authorityYear="2578" box="[712,1225,770,793]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="0E849E624425951D64CCFCE6FCAB2573" box="[712,891,773,792]" italics="true" pageId="2" pageNumber="81">Garra gallagheri</emphasis>
Wadi Bani Khalid
<collectingCountry id="44E702E04425951D6242FCE2FB5A2572" box="[1094,1162,770,793]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
2578
</taxonomicName>
<accessionNumber id="23A3DF934425951D62CBFCE2FA932572" box="[1231,1347,770,793]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244645" pageId="2" pageNumber="81">KX244645</accessionNumber>
<taxonomicName id="FBF039F34425951D64DCFCC1FC08255F" authority="Hotta" authorityName="Hotta" box="[728,984,797,820]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="0E849E624425951D64DCFCC1FC452558" box="[728,917,800,819]" italics="true" pageId="2" pageNumber="81">Garra longipinnis</emphasis>
Hotta
</taxonomicName>
<collectingCountry id="44E702E04425951D65DBFCFDFBF3255F" box="[991,1059,797,820]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
CN7572
<accessionNumber id="23A3DF934425951D6288FCFDFAD4255F" box="[1164,1284,797,820]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214755" pageId="2" pageNumber="81">KM214755</accessionNumber>
88-99-96
<taxonomicName id="FBF039F34425951D64E0FCDDFC34253B" authority="Jabal" authorityName="Jabal" box="[740,996,825,848]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="0E849E624425951D64E0FCDDFC712524" box="[740,929,828,847]" italics="true" pageId="2" pageNumber="81">Garra longipinnis</emphasis>
Jabal
</taxonomicName>
al Akhdar
<collectingCountry id="44E702E04425951D625DFCD9FB4D253B" box="[1113,1181,825,848]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
CN5897a
<accessionNumber id="23A3DF934425951D6311FCD9FA5D253B" box="[1301,1421,825,848]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214752" pageId="2" pageNumber="81">KM214752</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67E4FCB1FABB25B1" blockId="2.[360,1421,687,1704]" pageId="2" pageNumber="81">
74-71-90
<taxonomicName id="FBF039F34425951D64E0FCB9FC342507" authority="Jabal" authorityName="Jabal" box="[740,996,853,876]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="0E849E624425951D64E0FCB9FC712501" box="[740,929,855,875]" italics="true" pageId="2" pageNumber="81">Garra longipinnis</emphasis>
Jabal
</taxonomicName>
al Akhdar
<collectingCountry id="44E702E04425951D625DFCB5FB4D2507" box="[1113,1181,853,876]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
CN5897b
<accessionNumber id="23A3DF934425951D6311FCB5FA5D2507" box="[1301,1421,853,876]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214756" pageId="2" pageNumber="81">KM214756</accessionNumber>
<taxonomicName id="FBF039F34425951D64F9FC94FBF725EC" authority="Shawkah" authorityName="Shawkah" box="[765,1063,880,903]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E624425951D64F9FC94FC6925ED" box="[765,953,883,902]" italics="true" pageId="2" pageNumber="81">Garra barreimiae</emphasis>
Shawkah
</taxonomicName>
<collectingCountry id="44E702E04425951D622AFC90FBB125EC" box="[1070,1121,880,903]" name="United Arab Emirates" pageId="2" pageNumber="81">UAE</collectingCountry>
NHUAE-012
<accessionNumber id="23A3DF934425951D62FCFC90FAA125EC" box="[1272,1393,880,903]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214685" pageId="2" pageNumber="81">KM214685</accessionNumber>
<taxonomicName id="FBF039F34425951D64F3FC70FBF125C8" authority="Shawkah" authorityName="Shawkah" box="[759,1057,908,931]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E624425951D64F3FC70FC6325C9" box="[759,947,911,930]" italics="true" pageId="2" pageNumber="81">Garra barreimiae</emphasis>
Shawkah
</taxonomicName>
<collectingCountry id="44E702E04425951D622CFC6CFB8B25C8" box="[1064,1115,908,931]" name="United Arab Emirates" pageId="2" pageNumber="81">UAE</collectingCountry>
NHUAE-011
<accessionNumber id="23A3DF934425951D62F5FC6CFAB925C8" box="[1265,1385,908,931]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214758" pageId="2" pageNumber="81">KM214758</accessionNumber>
99-100-100
<taxonomicName id="FBF039F34425951D64F3FC4BFBF125D5" authority="Shawkah" authorityName="Shawkah" box="[759,1057,935,958]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E624425951D64F3FC4BFC6325D6" box="[759,947,938,957]" italics="true" pageId="2" pageNumber="81">Garra barreimiae</emphasis>
Shawkah
</taxonomicName>
<collectingCountry id="44E702E04425951D622CFC47FB8B25D5" box="[1064,1115,935,958]" name="United Arab Emirates" pageId="2" pageNumber="81">UAE</collectingCountry>
NHUAE-013
<accessionNumber id="23A3DF934425951D62F6FC47FABB25D5" box="[1266,1387,935,958]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214738" pageId="2" pageNumber="81">KM214738</accessionNumber>
<taxonomicName id="FBF039F34425951D64F3FC27FBF125B1" authority="Shawkah" authorityName="Shawkah" box="[759,1057,963,986]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E624425951D64F3FC27FC6325B2" box="[759,947,966,985]" italics="true" pageId="2" pageNumber="81">Garra barreimiae</emphasis>
Shawkah
</taxonomicName>
<collectingCountry id="44E702E04425951D622CFC23FB8B25B1" box="[1064,1115,963,986]" name="United Arab Emirates" pageId="2" pageNumber="81">UAE</collectingCountry>
NHUAE-014
<accessionNumber id="23A3DF934425951D62F6FC23FABB25B1" box="[1266,1387,963,986]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214779" pageId="2" pageNumber="81">KM214779</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67A0FC39FC0F22B8" blockId="2.[360,1421,687,1704]" pageId="2" pageNumber="81">
99-99-99
<taxonomicName id="FBF039F34425951D64F3FC03FBD2259D" authority="Wuray" authorityName="Wuray" box="[759,1026,991,1014]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E624425951D64F3FC03FC63259E" box="[759,947,994,1013]" italics="true" pageId="2" pageNumber="81">Garra barreimiae</emphasis>
Wuray
</taxonomicName>
<collectingCountry id="44E702E04425951D620DFC3FFBEC259D" box="[1033,1084,991,1014]" name="United Arab Emirates" pageId="2" pageNumber="81">UAE</collectingCountry>
NHUAE-001
<accessionNumber id="23A3DF934425951D62D7FC3FFA9C259D" box="[1235,1356,991,1014]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214734" pageId="2" pageNumber="81">KM214734</accessionNumber>
<taxonomicName id="FBF039F34425951D64F3FC1EFBD2227A" authority="Wuray" authorityName="Wuray" box="[759,1026,1018,1041]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E624425951D64F3FC1EFC63227B" box="[759,947,1021,1040]" italics="true" pageId="2" pageNumber="81">Garra barreimiae</emphasis>
Wuray
</taxonomicName>
<collectingCountry id="44E702E04425951D620DFC1AFBEC227A" box="[1033,1084,1018,1041]" name="United Arab Emirates" pageId="2" pageNumber="81">UAE</collectingCountry>
NHUAE-006
<accessionNumber id="23A3DF934425951D62D7FC1AFA9C227A" box="[1235,1356,1018,1041]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214767" pageId="2" pageNumber="81">KM214767</accessionNumber>
<taxonomicName id="FBF039F34425951D64DAFBFAFCBC2247" box="[734,876,1049,1068]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sahilia">
<emphasis id="0E849E624425951D64DAFBFAFCBC2247" box="[734,876,1049,1068]" italics="true" pageId="2" pageNumber="81">Garra sahilia</emphasis>
</taxonomicName>
SA NHKSA-015
<accessionNumber id="23A3DF934425951D622CFBF6FB702246" box="[1064,1184,1046,1069]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214774" pageId="2" pageNumber="81">KM214774</accessionNumber>
<taxonomicName id="FBF039F34425951D64DAFBD6FCBC222C" box="[734,876,1076,1096]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sahilia">
<emphasis id="0E849E624425951D64DAFBD6FCBC222C" box="[734,876,1076,1096]" italics="true" pageId="2" pageNumber="81">Garra sahilia</emphasis>
</taxonomicName>
<collectingRegion id="FE348C924425951D6576FBD1FC6A2223" box="[882,954,1073,1096]" country="Yemen" name="San'a'" pageId="2" pageNumber="81">Sanaa</collectingRegion>
<collectingCountry id="44E702E04425951D65C5FBD1FBDE2223" box="[961,1038,1073,1096]" name="Yemen" pageId="2" pageNumber="81">Yemen</collectingCountry>
CN872
<accessionNumber id="23A3DF934425951D626DFBD1FB322223" box="[1129,1250,1073,1096]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214689" pageId="2" pageNumber="81">KM214689</accessionNumber>
<taxonomicName id="FBF039F34425951D64DAFBB1FCBC2208" box="[734,876,1104,1123]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sahilia">
<emphasis id="0E849E624425951D64DAFBB1FCBC2208" box="[734,876,1104,1123]" italics="true" pageId="2" pageNumber="81">Garra sahilia</emphasis>
</taxonomicName>
SA NHKSA-014
<accessionNumber id="23A3DF934425951D622CFBADFB71220F" box="[1064,1185,1101,1124]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214786" pageId="2" pageNumber="81">KM214786</accessionNumber>
99-100-99
<taxonomicName id="FBF039F34425951D64BAFB8DFC9C2214" box="[702,844,1132,1151]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sahilia">
<emphasis id="0E849E624425951D64BAFB8DFC9C2214" box="[702,844,1132,1151]" italics="true" pageId="2" pageNumber="81">Garra sahilia</emphasis>
</taxonomicName>
SA NHKSA-016
<accessionNumber id="23A3DF934425951D620CFB89FB5122EB" box="[1032,1153,1129,1152]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214718" pageId="2" pageNumber="81">KM214718</accessionNumber>
<taxonomicName id="FBF039F34425951D64BCFB68FC9622F1" box="[696,838,1159,1178]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sahilia">
<emphasis id="0E849E624425951D64BCFB68FC9622F1" box="[696,838,1159,1178]" italics="true" pageId="2" pageNumber="81">Garra sahilia</emphasis>
</taxonomicName>
SA NHKSA-012
<accessionNumber id="23A3DF934425951D6206FB64FBAB22F0" box="[1026,1147,1156,1179]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214724" pageId="2" pageNumber="81">KM214724</accessionNumber>
<taxonomicName id="FBF039F34425951D64BCFB44FC9622DD" box="[696,838,1187,1206]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sahilia">
<emphasis id="0E849E624425951D64BCFB44FC9622DD" box="[696,838,1187,1206]" italics="true" pageId="2" pageNumber="81">Garra sahilia</emphasis>
</taxonomicName>
SA NHKSA-026
<accessionNumber id="23A3DF934425951D6206FB40FBAB22DC" box="[1026,1147,1184,1207]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214799" pageId="2" pageNumber="81">KM214799</accessionNumber>
<taxonomicName id="FBF039F34425951D67FDFB20FD7A22B9" box="[505,682,1215,1234]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="buettikerii">
<emphasis id="0E849E624425951D67FDFB20FD7A22B9" box="[505,682,1215,1234]" italics="true" pageId="2" pageNumber="81">Garra buettikerii</emphasis>
</taxonomicName>
SA NHKSA-021
<accessionNumber id="23A3DF934425951D6563FB5CFC0F22B8" box="[871,991,1212,1235]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214704" pageId="2" pageNumber="81">KM214704</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67FDFB3BFC0F2285" blockId="2.[360,1421,687,1704]" box="[505,991,1239,1262]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D67FDFB3BFD7A2286" box="[505,682,1242,1261]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="buettikerii">
<emphasis id="0E849E624425951D67FDFB3BFD7A2286" box="[505,682,1242,1261]" italics="true" pageId="2" pageNumber="81">Garra buettikerii</emphasis>
</taxonomicName>
SA NHKSA-022
<accessionNumber id="23A3DF934425951D6563FB37FC0F2285" box="[871,991,1239,1262]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214742" pageId="2" pageNumber="81">KM214742</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D679CFB07FC0F2361" blockId="2.[360,1421,687,1704]" box="[408,991,1255,1290]" pageId="2" pageNumber="81">
87-88-85
<taxonomicName id="FBF039F34425951D67FDFB17FD7A2362" box="[505,682,1270,1289]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="buettikerii">
<emphasis id="0E849E624425951D67FDFB17FD7A2362" box="[505,682,1270,1289]" italics="true" pageId="2" pageNumber="81">Garra buettikerii</emphasis>
</taxonomicName>
SA NHKSA-020
<accessionNumber id="23A3DF934425951D6563FB13FC0F2361" box="[871,991,1267,1290]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214723" pageId="2" pageNumber="81">KM214723</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D6795FAF3FC09234D" blockId="2.[360,1421,687,1704]" box="[401,985,1295,1318]" pageId="2" pageNumber="81">
91-93-91
<taxonomicName id="FBF039F34425951D67F7FAF3FD74234E" box="[499,676,1298,1317]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="buettikerii">
<emphasis id="0E849E624425951D67F7FAF3FD74234E" box="[499,676,1298,1317]" italics="true" pageId="2" pageNumber="81">Garra buettikerii</emphasis>
</taxonomicName>
SA NHKSA-024
<accessionNumber id="23A3DF934425951D6565FAEFFC09234D" box="[865,985,1295,1318]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214729" pageId="2" pageNumber="81">KM214729</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67F7FACEFB8B232A" blockId="2.[360,1421,687,1704]" box="[499,1115,1322,1345]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D67F7FACEFCF6232A" authority="Wadi Damad" authorityName="Wadi Damad" box="[499,806,1322,1345]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="tibanica">
<emphasis id="0E849E624425951D67F7FACEFD40232B" box="[499,656,1325,1344]" italics="true" pageId="2" pageNumber="81">Garra tibanica</emphasis>
Wadi Damad
</taxonomicName>
SA NHKSA-027
<accessionNumber id="23A3DF934425951D65E7FACAFB8B232A" box="[995,1115,1322,1345]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214686" pageId="2" pageNumber="81">KM214686</accessionNumber>
</paragraph>
</subSubSection>
<subSubSection id="74EA11FB4425951967F7FAAAFD6220AF" lastPageId="6" lastPageNumber="85" pageId="2" pageNumber="81" type="description">
<paragraph id="3C4F42704425951D67F7FAAAFB8B2336" blockId="2.[360,1421,687,1704]" box="[499,1115,1350,1373]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D67F7FAAAFCF62336" authority="Wadi Damad" authorityName="Wadi Damad" box="[499,806,1350,1373]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="tibanica">
<emphasis id="0E849E624425951D67F7FAAAFD402337" box="[499,656,1353,1372]" italics="true" pageId="2" pageNumber="81">Garra tibanica</emphasis>
Wadi Damad
</taxonomicName>
SA NHKSA-028
<accessionNumber id="23A3DF934425951D65E7FAA6FB8B2336" box="[995,1115,1350,1373]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214749" pageId="2" pageNumber="81">KM214749</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D6799FAB9FB8B2313" blockId="2.[360,1421,687,1704]" box="[413,1115,1369,1400]" pageId="2" pageNumber="81">
94-94-92
<taxonomicName id="FBF039F34425951D67F7FA86FCF62313" authority="Wadi Damad" authorityName="Wadi Damad" box="[499,806,1377,1400]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="tibanica">
<emphasis id="0E849E624425951D67F7FA86FD40231C" box="[499,656,1380,1400]" italics="true" pageId="2" pageNumber="81">Garra tibanica</emphasis>
Wadi Damad
</taxonomicName>
SA NHKSA-031
<accessionNumber id="23A3DF934425951D65E7FA81FB8B2313" box="[995,1115,1377,1400]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214789" pageId="2" pageNumber="81">KM214789</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D676CFA64FBFF23FF" blockId="2.[360,1421,687,1704]" box="[360,1071,1405,1433]" pageId="2" pageNumber="81">
97-100-100 100
<taxonomicName id="FBF039F34425951D640AFA61FD6523F8" box="[526,693,1408,1427]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="dunserei">
<emphasis id="0E849E624425951D640AFA61FD6523F8" box="[526,693,1408,1427]" italics="true" pageId="2" pageNumber="81">Garra dunserei</emphasis>
</taxonomicName>
Dhofar Oman CN7766
<accessionNumber id="23A3DF934425951D65BFFA9DFBFF23FF" box="[955,1071,1405,1428]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KP069477" pageId="2" pageNumber="81">KP069477</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67EAFA78FBFF23DB" blockId="2.[360,1421,687,1704]" box="[494,1071,1432,1456]" pageId="2" pageNumber="81">
<materialsCitation id="8C98482D4425951D67EAFA78FBFF23DB" ID-GBIF-Occurrence="1291640428" box="[494,1071,1432,1456]" country="Oman" location="Dhofar" pageId="2" pageNumber="81" specimenCount="1" stateProvince="Dhofar">
99
<taxonomicName id="FBF039F34425951D640AFA7DFD6523C4" box="[526,693,1436,1455]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="dunserei">
<emphasis id="0E849E624425951D640AFA7DFD6523C4" box="[526,693,1436,1455]" italics="true" pageId="2" pageNumber="81">Garra dunserei</emphasis>
</taxonomicName>
<collectingRegion id="FE348C924425951D64BFFA79FCD623DB" box="[699,774,1433,1456]" country="Oman" name="Zufar" pageId="2" pageNumber="81">Dhofar</collectingRegion>
<collectingCountry id="44E702E04425951D6509FA79FC8123DB" box="[781,849,1433,1456]" name="Oman" pageId="2" pageNumber="81">Oman</collectingCountry>
CN7771
<accessionNumber id="23A3DF934425951D65BFFA79FBFF23DB" box="[955,1071,1433,1456]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KP069476" pageId="2" pageNumber="81">KP069476</accessionNumber>
</materialsCitation>
</paragraph>
<paragraph id="3C4F42704425951D67EDFA49FDD823D1" blockId="2.[360,1421,687,1704]" box="[489,520,1449,1466]" pageId="2" pageNumber="81">100</paragraph>
<paragraph id="3C4F42704425951D6402FA58FBDD23A0" blockId="2.[360,1421,687,1704]" box="[518,1037,1460,1483]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D6402FA58FD3023A0" authority="Hasaq" authorityName="Hasaq" box="[518,736,1460,1483]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="smarti">
<emphasis id="0E849E624425951D6402FA58FD4123A1" box="[518,657,1463,1482]" italics="true" pageId="2" pageNumber="81">Garra smarti</emphasis>
Hasaq
</taxonomicName>
Oman CN5874
<accessionNumber id="23A3DF934425951D6590FA54FBDD23A0" box="[916,1037,1460,1483]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214744" pageId="2" pageNumber="81">KM214744</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67E8FA26FDD023BC" blockId="2.[360,1421,687,1704]" box="[492,512,1478,1495]" pageId="2" pageNumber="81">99</paragraph>
<paragraph id="3C4F42704425951D6797FA39FBDD238C" blockId="2.[360,1421,687,1704]" box="[403,1037,1488,1518]" pageId="2" pageNumber="81">
82-93-85 98
<taxonomicName id="FBF039F34425951D6402FA34FD30238C" authority="Hasaq" authorityName="Hasaq" box="[518,736,1488,1511]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="smarti">
<emphasis id="0E849E624425951D6402FA34FD41238D" box="[518,657,1491,1510]" italics="true" pageId="2" pageNumber="81">Garra smarti</emphasis>
Hasaq
</taxonomicName>
Oman CN5893
<accessionNumber id="23A3DF934425951D6590FA30FBDD238C" box="[916,1037,1488,1511]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214699" pageId="2" pageNumber="81">KM214699</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67E8FA13FC2F2068" blockId="2.[360,1421,687,1704]" box="[492,1023,1516,1540]" pageId="2" pageNumber="81">
97
<taxonomicName id="FBF039F34425951D6402FA10FD302068" authority="Hasaq" authorityName="Hasaq" box="[518,736,1516,1539]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="smarti">
<emphasis id="0E849E624425951D6402FA10FD41206A" box="[518,657,1518,1538]" italics="true" pageId="2" pageNumber="81">Garra smarti</emphasis>
Hasaq
</taxonomicName>
Oman CN773
<accessionNumber id="23A3DF934425951D6583FA0CFC2F2068" box="[903,1023,1516,1539]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM214732" pageId="2" pageNumber="81">KM214732</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D679BF9F3FBE52075" blockId="2.[360,1421,687,1704]" box="[415,1077,1543,1572]" pageId="2" pageNumber="81">
67-65-73
<taxonomicName id="FBF039F34425951D6410F9EBFC6B2075" authority="Wadi Andhur Oman 2559" authorityName="Wadi Andhur Oman" authorityYear="2559" box="[532,955,1543,1566]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624425951D6410F9EBFD4C2076" box="[532,668,1546,1565]" italics="true" pageId="2" pageNumber="81">Garra sindhi</emphasis>
Wadi Andhur Oman 2559
</taxonomicName>
<accessionNumber id="23A3DF934425951D65C5F9E7FBE52075" box="[961,1077,1543,1566]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244649" pageId="2" pageNumber="81">KX244649</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D6410F9C7FBE52051" blockId="2.[360,1421,687,1704]" box="[532,1077,1571,1594]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D6410F9C7FC6B2051" authority="Wadi Andhur Oman 2559" authorityName="Wadi Andhur Oman" authorityYear="2559" box="[532,955,1571,1594]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624425951D6410F9C7FD4C2052" box="[532,668,1574,1593]" italics="true" pageId="2" pageNumber="81">Garra sindhi</emphasis>
Wadi Andhur Oman 2559
</taxonomicName>
<accessionNumber id="23A3DF934425951D65C5F9C3FBE52051" box="[961,1077,1571,1594]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244650" pageId="2" pageNumber="81">KX244650</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D67BFF9A4FBE5203E" blockId="2.[360,1421,687,1704]" box="[443,1077,1598,1621]" pageId="2" pageNumber="81">
99-99-99
<taxonomicName id="FBF039F34425951D6410F9A2FC6B203E" authority="Wadi Andhur Oman 2559" authorityName="Wadi Andhur Oman" authorityYear="2559" box="[532,955,1598,1621]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624425951D6410F9A2FD4C203F" box="[532,668,1601,1620]" italics="true" pageId="2" pageNumber="81">Garra sindhi</emphasis>
Wadi Andhur Oman 2559
</taxonomicName>
<accessionNumber id="23A3DF934425951D65C5F9DEFBE5203E" box="[961,1077,1598,1621]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244651" pageId="2" pageNumber="81">KX244651</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D6410F9BEFBE5201A" blockId="2.[360,1421,687,1704]" box="[532,1077,1626,1649]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D6410F9BEFC6B201A" authority="Wadi Andhur Oman 2559" authorityName="Wadi Andhur Oman" authorityYear="2559" box="[532,955,1626,1649]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624425951D6410F9BEFD4C201B" box="[532,668,1629,1648]" italics="true" pageId="2" pageNumber="81">Garra sindhi</emphasis>
Wadi Andhur Oman 2559
</taxonomicName>
<accessionNumber id="23A3DF934425951D65C5F9BAFBE5201A" box="[961,1077,1626,1649]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX244652" pageId="2" pageNumber="81">KX244652</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D678EF99AFC5520E6" blockId="2.[360,1421,687,1704]" box="[394,901,1654,1677]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D678EF99AFCDE20E6" authority="Orontes Syria 1159" authorityName="Orontes Syria" authorityYear="1159" box="[394,782,1654,1677]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="0E849E624425951D678EF99AFDE120E0" box="[394,561,1656,1676]" italics="true" pageId="2" pageNumber="81">Garra variabilis</emphasis>
Orontes Syria 1159
</taxonomicName>
<accessionNumber id="23A3DF934425951D6511F996FC5520E6" box="[789,901,1654,1677]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ553391" pageId="2" pageNumber="81">KJ553391</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951D678EF975FC5520C3" blockId="2.[360,1421,687,1704]" box="[394,901,1681,1704]" pageId="2" pageNumber="81">
<taxonomicName id="FBF039F34425951D678EF975FCDE20C3" authority="Orontes Syria 1159" authorityName="Orontes Syria" authorityYear="1159" box="[394,782,1681,1704]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="81" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="0E849E624425951D678EF975FDE120CC" box="[394,561,1684,1703]" italics="true" pageId="2" pageNumber="81">Garra variabilis</emphasis>
Orontes Syria 1159
</taxonomicName>
<accessionNumber id="23A3DF934425951D6511F971FC5520C3" box="[789,901,1681,1704]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ553567" pageId="2" pageNumber="81">KJ553567</accessionNumber>
</paragraph>
<paragraph id="3C4F42704425951C6712F904FD442304" blockId="2.[278,319,1764,1784]" lastBlockId="3.[151,1437,682,1967]" lastPageId="3" lastPageNumber="82" pageId="2" pageNumber="81">
0.02
<emphasis id="0E849E624424951C66C3FD4AFE8924A8" bold="true" box="[199,345,682,707]" pageId="3" pageNumber="82">Description.</emphasis>
See
<figureCitation id="A4CB5EF54424951C6791FD4AFDCE24A8" box="[405,542,682,707]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="3.[151,249,621,643]" captionStartId-1="5.[151,249,1134,1156]" captionStartId-2="5.[151,249,1950,1972]" captionStartId-3="7.[151,249,1429,1451]" captionTargetBox-0="[151,1436,193,599]" captionTargetBox-1="[151,1436,193,1112]" captionTargetBox-2="[151,1436,1206,1928]" captionTargetBox-3="[151,1435,445,1408]" captionTargetId-0="figure@3.[151,1437,193,599]" captionTargetId-1="figure@5.[151,1436,193,1113]" captionTargetId-2="figure@5.[151,1436,1206,1929]" captionTargetId-3="figure@7.[151,1436,445,1408]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="7" captionText-0="FIGURE 2. Garra sindhi, ONHM 4196, holotype, 82.5 mm SL; Oman: Wadi Andhur." captionText-1="FIGURE 4. Garra sindhi, ONHM 4219, paratype. 48.7 mm SL; Oman: Wadi Andhur." captionText-2="FIGURE 5. Garra sindhi, about 45 mm SL, not preserved; Oman: Wadi Andhur." captionText-3="FIGURE 6. Pool in Wadi Andhur." httpUri-0="https://zenodo.org/record/256493/files/figure.png" httpUri-1="https://zenodo.org/record/256494/files/figure.png" httpUri-2="https://zenodo.org/record/256495/files/figure.png" httpUri-3="https://zenodo.org/record/256496/files/figure.png" pageId="3" pageNumber="82">Figures 2–6</figureCitation>
for general appearance and
<tableCitation id="717277CB4424951C6568FD4AFC1224A8" box="[876,962,682,707]" captionStart="TABLE 1" captionStartId="6.[151,239,152,175]" captionTargetBox="[151,1337,244,1175]" captionTargetPageId="6" captionText="TABLE 1. Morphometric data of Garra sindhi (holotype. ONHM 4196; paratypes, ONHM 4197, 4219, n = 34)." httpUri="http://table.plazi.org/id/688F12F8442195196693FF78FAE926C4" pageId="3" pageNumber="82" tableUuid="688F12F8442195196693FF78FAE926C4">Table 1</tableCitation>
for morphometric data. Body elongated, moderately compressed laterally, more compressed in region of caudal peduncle. Dorsal head profile rising gently, almost straight, more or less continuous with dorsal body profile to about middle between nape and dorsal-fin origin. Dorsal profile decreasing between middle between nape and dorsal-fin origin and about base of last dorsalfin ray. Slightly decreasing until caudal peduncle. Ventral profile more or less straight to anal-fin origin. Head small, with flat interorbital distance, height at nape less than head length, width at nape greater than depth at nape, without tubercles. Snout pointed, transverse groove absent. Proboscis not or only slightly elevated from depressed rostral surface. Depressed rostral fold poorly expressed. No groove between transverse lobe and lateral surface. Eye placed dorso-laterally in anterior half of head. Barbels in two pairs; rostral barbel antero-laterally located, shorter than eye diameter; maxillary barbel at corner of mouth, shorter than rostral barbel. Rostral cap welldeveloped, fimbriate, papillate on ventral surface. Upper lip present as a thin band of papillae. Upper jaw almost or completely covered by rostral cap. Lower jaw slightly curved or straight. Disc elliptical, shorter than wide or as long as wide and narrower than head width through base of maxillary barbel; papillae on anteromedian fold of same size, regularly arranged; groove between anteromedian fold and central callous-pad narrow and deep in some individuals or shallow and wide in others, papillae on latero-posterior flap of same size and distance as on papillae on anteromedian fold; anterior marginal surface of central callous pad without or with sparsely arranged small papillae; papillae on anteromedian fold and on latero-posterior flap almost or completely absent in some individuals; posterior margin of latero-posterior flap extending vertical to anterior or posterior margin of eye; in some individuals, only the posterior tip of latero-posterior flap is free while in others, the whole rim is free. Largest recorded specimen
<quantity id="FB08EF954424951C6774FAB6FE102305" box="[368,448,1366,1391]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="82" unit="mm" value="85.0">85 mm</quantity>
SL, not preserved.
</paragraph>
<caption id="688F12F84425951D6693F8F4FB8421ED" pageId="2" pageNumber="81">
<paragraph id="3C4F42704425951D6693F8F4FB8421ED" blockId="2.[151,1437,1812,1926]" pageId="2" pageNumber="81">
<emphasis id="0E849E624425951D6693F8F4FEC22142" bold="true" box="[151,274,1812,1834]" pageId="2" pageNumber="81">FIGURE 1.</emphasis>
Maximum likelihood estimation of the phylogenetic relationships based on the mitochondrial COI barcode region (Tamura 3-parameter model, discrete Gamma distribution for rate differences with 5 categories +
<emphasis id="0E849E624425951D62ACF8D3FB6A2123" box="[1192,1210,1843,1864]" italics="true" pageId="2" pageNumber="81">G</emphasis>
parameter = 0.3409. Nucleotide positions with less than 100 % site coverage were eliminated resulting in 652 analysed positions. Numbers of major nodes indicate bootstrap values from 1000 pseudoreplicates from the ML, NJ and MP method.
</paragraph>
</caption>
<caption id="688F12F84424951C6693FD8DFBD524E8" box="[151,1029,621,643]" httpUri="https://zenodo.org/record/256493/files/figure.png" pageId="3" pageNumber="82" targetBox="[151,1436,193,599]" targetPageId="3">
<paragraph id="3C4F42704424951C6693FD8DFBD524E8" blockId="3.[151,1029,621,643]" box="[151,1029,621,643]" pageId="3" pageNumber="82">
<emphasis id="0E849E624424951C6693FD8DFEC224E9" bold="true" box="[151,274,621,643]" pageId="3" pageNumber="82">FIGURE 2.</emphasis>
<taxonomicName id="FBF039F34424951C671CFD8EFE4B24E8" box="[280,411,621,643]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624424951C671CFD8EFE4B24E8" box="[280,411,621,643]" italics="true" pageId="3" pageNumber="82">Garra sindhi</emphasis>
</taxonomicName>
, ONHM 4196, holotype, 82.5 mm SL; Oman: Wadi Andhur.
</paragraph>
</caption>
<paragraph id="3C4F42704424951C66C3FA9AFD6321C5" blockId="3.[151,1437,682,1967]" pageId="3" pageNumber="82">
Dorsal fin with 3 simple and 7½ branched rays, last simple ray slightly shorter than or equally long as head length; distal margin concave; origin at middle between snout tip and last scale on caudal-fin base; inserted anterior to vertical from pelvic-fin origin; first branched ray longest, tip of last branched ray reaching vertical of anus. Pectoral fin with 1 simple and 12 branched rays, reaching midway or slightly beyond distance between pectoral- and pelvic-fin origin, length shorter than head length. Pelvic fin with 1 simple and 7–8 branched rays, reaching to, or slightly in front of anus, origin closer to anal-fin origin than to pectoral-fin origin, inserted below second or fourth branched dorsal-fin ray. Anal fin short, with 3 simple and 5½ branched rays; first branched ray longest, reaching to dark grey or black blotch on posterior caudal peduncle or slightly anterior to it; distal margin concave; origin almost in middle between caudal-fin base and pelvic-fin origin. 2 scales between anus and anal-fin origin. Caudal fin forked; with 9+8 branched rays; tip of lobes pointed; upper lobe longer than lower lobe. 8–12 small and very short gill rakers on lower limb of first gill arch. Lateral line complete, with 34 scales on body and 2 scales on anterior caudal fin. Transverse scale rows above lateral line 3½–4½; between lateral line and pelvic-fin origin 3½ and between lateral line and anal-fin origin 3½–4½. Circumpeduncular scale rows 14–16. Predorsal scales smaller than flank scales, irregularly arranged, anterior predorsal scales deeply embedded. Belly and breast behind pectoral-fin origin scaled, scales deeply embedded. One long axillary scale at base of pelvic fin, and 6
<emphasis id="0E849E624424951C6330F892FA9221E0" bold="true" box="[1332,1346,1906,1931]" pageId="3" pageNumber="82">–</emphasis>
8 scales between posteriormost pelvic-fin base and anus.
</paragraph>
<paragraph id="3C4F42704423951B6693F820FA4D21BD" blockId="4.[151,1437,1984,2006]" box="[151,1437,1984,2006]" pageId="4" pageNumber="83">
<emphasis id="0E849E624423951B6693F820FEC821BE" bold="true" box="[151,280,1984,2006]" pageId="4" pageNumber="83">FIGURE. 3.</emphasis>
<taxonomicName id="FBF039F34423951B671AF821FE7221BD" box="[286,418,1984,2006]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624423951B671AF821FE7221BD" box="[286,418,1984,2006]" italics="true" pageId="4" pageNumber="83">Garra sindhi</emphasis>
</taxonomicName>
, ONHM 4196,
<typeStatus id="E34BFCD24423951B6444F820FD4B21BD" box="[576,667,1984,2006]" pageId="4" pageNumber="83" type="holotype">holotype</typeStatus>
,
<quantity id="FB08EF954423951B64A1F820FD2E21BD" box="[677,766,1984,2006]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" pageId="4" pageNumber="83" unit="mm" value="82.5">82.5 mm</quantity>
SL, lateral, ventral and dorsal view of head;
<collectingCountry id="44E702E04423951B62C0F820FAD621BD" box="[1220,1286,1984,2006]" name="Oman" pageId="4" pageNumber="83">Oman</collectingCountry>
: Wadi Andhur.
</paragraph>
<caption id="688F12F84422951A6693FB8EFBD422EF" box="[151,1028,1134,1156]" httpUri="https://zenodo.org/record/256494/files/figure.png" pageId="5" pageNumber="84" targetBox="[151,1436,193,1112]" targetPageId="5">
<paragraph id="3C4F42704422951A6693FB8EFBD422EF" blockId="5.[151,1028,1134,1156]" box="[151,1028,1134,1156]" pageId="5" pageNumber="84">
<emphasis id="0E849E624422951A6693FB8EFEC222EF" bold="true" box="[151,274,1134,1156]" pageId="5" pageNumber="84">FIGURE 4.</emphasis>
<taxonomicName id="FBF039F34422951A671CFB8FFE4B22EF" box="[280,411,1134,1156]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624422951A671CFB8FFE4B22EF" box="[280,411,1134,1156]" italics="true" pageId="5" pageNumber="84">Garra sindhi</emphasis>
</taxonomicName>
, ONHM 4219, paratype. 48.7 mm SL; Oman: Wadi Andhur.
</paragraph>
</caption>
<caption id="688F12F84422951A6693F87EFC1F21DF" box="[151,975,1950,1972]" httpUri="https://zenodo.org/record/256495/files/figure.png" pageId="5" pageNumber="84" targetBox="[151,1436,1206,1928]" targetPageId="5">
<paragraph id="3C4F42704422951A6693F87EFC1F21DF" blockId="5.[151,975,1950,1972]" box="[151,975,1950,1972]" pageId="5" pageNumber="84">
<emphasis id="0E849E624422951A6693F87EFEC221DF" bold="true" box="[151,274,1950,1972]" pageId="5" pageNumber="84">FIGURE 5.</emphasis>
<taxonomicName id="FBF039F34422951A671CF87FFE4B21DF" box="[280,411,1950,1972]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624422951A671CF87FFE4B21DF" box="[280,411,1950,1972]" italics="true" pageId="5" pageNumber="84">Garra sindhi</emphasis>
</taxonomicName>
, about 45 mm SL, not preserved; Oman: Wadi Andhur.
</paragraph>
</caption>
<caption id="688F12F8442195196693FF78FAE926C4" ID-Table-UUID="688F12F8442195196693FF78FAE926C4" box="[151,1337,151,176]" httpUri="http://table.plazi.org/id/688F12F8442195196693FF78FAE926C4" pageId="6" pageNumber="85" targetBox="[151,1337,244,1175]" targetIsTable="true" targetPageId="6">
<paragraph id="3C4F4270442195196693FF78FAE926C4" blockId="6.[151,1337,151,224]" box="[151,1337,151,176]" pageId="6" pageNumber="85">
<emphasis id="0E849E62442195196693FF78FEDA26C4" bold="true" box="[151,266,152,175]" pageId="6" pageNumber="85">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="FBF039F3442195196405FF78FD5B26C4" box="[513,651,151,175]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="85" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E62442195196405FF78FD5B26C4" box="[513,651,151,175]" italics="true" pageId="6" pageNumber="85">Garra sindhi</emphasis>
</taxonomicName>
(holotype. ONHM 4196; paratypes, ONHM 4197, 4219, n=34).
</paragraph>
</caption>
<paragraph id="3C4F4270442195196463FF2AFC7D268B" blockId="6.[151,1337,151,224]" box="[615,941,202,224]" pageId="6" pageNumber="85">holotype paratypes (n=34)</paragraph>
<paragraph id="3C4F4270442195196505FF15FB5622FC" pageId="6" pageNumber="85">
<table id="4EF0B0D044216AE06693FF14FAE922FC" box="[151,1337,244,1175]" gridcols="4" gridrows="23" pageId="6" pageNumber="85">
<tr id="82C0403244216AE06693FF14FAE92761" box="[151,1337,244,266]" gridrow="0" pageId="6" pageNumber="85" rowspan-0="1">
<th id="C111294E44216AE06505FF14FCE82761" box="[769,824,244,266]" gridcol="1" gridrow="0" pageId="6" pageNumber="85">mean</th>
<th id="C111294E44216AE065A2FF14FC1D2761" box="[934,973,244,266]" gridcol="2" gridrow="0" pageId="6" pageNumber="85">SD</th>
<th id="C111294E44216AE06243FF14FAE92761" box="[1095,1337,244,266]" gridcol="3" gridrow="0" pageId="6" pageNumber="85">range</th>
</tr>
<tr id="82C0403244216AE06693FEFDFAE92758" box="[151,1337,285,307]" gridrow="1" pageId="6" pageNumber="85" rowspan-2="1">
<th id="C111294E44216AE06693FEFDFD432758" box="[151,659,285,307]" gridcol="0" gridrow="1" pageId="6" pageNumber="85">SL (mm) 82.5</th>
<td id="C111294E44216AE06505FEFDFCE82758" box="[769,824,285,307]" gridcol="1" gridrow="1" pageId="6" pageNumber="85">42.1</td>
<td id="C111294E44216AE06243FEFDFAE92758" box="[1095,1337,285,307]" gridcol="3" gridrow="1" pageId="6" pageNumber="85">29.2–84.7</td>
</tr>
<tr id="82C0403244216AE06693FEA6FAE92737" box="[151,1337,326,348]" gridrow="2" pageId="6" pageNumber="85" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="C111294E44216AE06693FEA6FD432737" box="[151,659,326,348]" gridcol="0" gridrow="2" pageId="6" pageNumber="85">In percent of standard length</th>
</tr>
<tr id="82C0403244216AE06693FE90FAE927ED" box="[151,1337,368,390]" gridrow="3" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FE90FD4327ED" box="[151,659,368,390]" gridcol="0" gridrow="3" pageId="6" pageNumber="85">Predorsal length 47.6</th>
<td id="C111294E44216AE06505FE90FCE827ED" box="[769,824,368,390]" gridcol="1" gridrow="3" pageId="6" pageNumber="85">51.4</td>
<td id="C111294E44216AE065A2FE90FC1D27ED" box="[934,973,368,390]" gridcol="2" gridrow="3" pageId="6" pageNumber="85">2.7</td>
<td id="C111294E44216AE06243FE90FAE927ED" box="[1095,1337,368,390]" gridcol="3" gridrow="3" pageId="6" pageNumber="85">45.6–56.8</td>
</tr>
<tr id="82C0403244216AE06693FE79FAE927C4" box="[151,1337,409,431]" gridrow="4" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FE79FD4327C4" box="[151,659,409,431]" gridcol="0" gridrow="4" pageId="6" pageNumber="85">Prepelvic length 52.2</th>
<td id="C111294E44216AE06505FE79FCE827C4" box="[769,824,409,431]" gridcol="1" gridrow="4" pageId="6" pageNumber="85">56.3</td>
<td id="C111294E44216AE065A2FE79FC1D27C4" box="[934,973,409,431]" gridcol="2" gridrow="4" pageId="6" pageNumber="85">2.2</td>
<td id="C111294E44216AE06243FE79FAE927C4" box="[1095,1337,409,431]" gridcol="3" gridrow="4" pageId="6" pageNumber="85">52.2–61.8</td>
</tr>
<tr id="82C0403244216AE06693FE22FAE927B3" box="[151,1337,450,472]" gridrow="5" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FE22FD4327B3" box="[151,659,450,472]" gridcol="0" gridrow="5" pageId="6" pageNumber="85">Preanal length 71.1</th>
<td id="C111294E44216AE06505FE22FCE827B3" box="[769,824,450,472]" gridcol="1" gridrow="5" pageId="6" pageNumber="85">73.0</td>
<td id="C111294E44216AE065A2FE22FC1D27B3" box="[934,973,450,472]" gridcol="2" gridrow="5" pageId="6" pageNumber="85">3.2</td>
<td id="C111294E44216AE06243FE22FAE927B3" box="[1095,1337,450,472]" gridcol="3" gridrow="5" pageId="6" pageNumber="85">61.3–80.9</td>
</tr>
<tr id="82C0403244216AE06693FE0CFAE92469" box="[151,1337,492,514]" gridrow="6" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FE0CFD432469" box="[151,659,492,514]" gridcol="0" gridrow="6" pageId="6" pageNumber="85">Distance between anus and anal-fin origin 2.9</th>
<td id="C111294E44216AE06505FE0CFCE82469" box="[769,824,492,514]" gridcol="1" gridrow="6" pageId="6" pageNumber="85">4.4</td>
<td id="C111294E44216AE065A2FE0CFC1D2469" box="[934,973,492,514]" gridcol="2" gridrow="6" pageId="6" pageNumber="85">0.9</td>
<td id="C111294E44216AE06243FE0CFAE92469" box="[1095,1337,492,514]" gridcol="3" gridrow="6" pageId="6" pageNumber="85">2.9–6.3</td>
</tr>
<tr id="82C0403244216AE06693FDF5FAE92440" box="[151,1337,533,555]" gridrow="7" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FDF5FD432440" box="[151,659,533,555]" gridcol="0" gridrow="7" pageId="6" pageNumber="85">Head length 22.3</th>
<td id="C111294E44216AE06505FDF5FCE82440" box="[769,824,533,555]" gridcol="1" gridrow="7" pageId="6" pageNumber="85">26.5</td>
<td id="C111294E44216AE065A2FDF5FC1D2440" box="[934,973,533,555]" gridcol="2" gridrow="7" pageId="6" pageNumber="85">1.6</td>
<td id="C111294E44216AE06243FDF5FAE92440" box="[1095,1337,533,555]" gridcol="3" gridrow="7" pageId="6" pageNumber="85">22.3–29.2</td>
</tr>
<tr id="82C0403244216AE06693FDDEFAE9243F" box="[151,1337,574,596]" gridrow="8" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FDDEFD43243F" box="[151,659,574,596]" gridcol="0" gridrow="8" pageId="6" pageNumber="85">Caudal peduncle length 14.7</th>
<td id="C111294E44216AE06505FDDEFCE8243F" box="[769,824,574,596]" gridcol="1" gridrow="8" pageId="6" pageNumber="85">14.5</td>
<td id="C111294E44216AE065A2FDDEFC1D243F" box="[934,973,574,596]" gridcol="2" gridrow="8" pageId="6" pageNumber="85">3.5</td>
<td id="C111294E44216AE06243FDDEFAE9243F" box="[1095,1337,574,596]" gridcol="3" gridrow="8" pageId="6" pageNumber="85">11.5–32.7</td>
</tr>
<tr id="82C0403244216AE06693FD88FAE92415" box="[151,1337,616,638]" gridrow="9" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FD88FD432415" box="[151,659,616,638]" gridcol="0" gridrow="9" pageId="6" pageNumber="85">Body depth at dorsal fin origin 22.9</th>
<td id="C111294E44216AE06505FD88FCE82415" box="[769,824,616,638]" gridcol="1" gridrow="9" pageId="6" pageNumber="85">23.1</td>
<td id="C111294E44216AE065A2FD88FC1D2415" box="[934,973,616,638]" gridcol="2" gridrow="9" pageId="6" pageNumber="85">1.5</td>
<td id="C111294E44216AE06243FD88FAE92415" box="[1095,1337,616,638]" gridcol="3" gridrow="9" pageId="6" pageNumber="85">19.9–26.1</td>
</tr>
<tr id="82C0403244216AE06693FD71FAE924CC" box="[151,1337,657,679]" gridrow="10" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FD71FD4324CC" box="[151,659,657,679]" gridcol="0" gridrow="10" pageId="6" pageNumber="85">Caudal peduncle depth 13.4</th>
<td id="C111294E44216AE06505FD71FCE824CC" box="[769,824,657,679]" gridcol="1" gridrow="10" pageId="6" pageNumber="85">11.7</td>
<td id="C111294E44216AE065A2FD71FC1D24CC" box="[934,973,657,679]" gridcol="2" gridrow="10" pageId="6" pageNumber="85">1.6</td>
<td id="C111294E44216AE06243FD71FAE924CC" box="[1095,1337,657,679]" gridcol="3" gridrow="10" pageId="6" pageNumber="85">9.7–20.0</td>
</tr>
<tr id="82C0403244216AE06693FD5AFAE924BB" box="[151,1337,698,720]" gridrow="11" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FD5AFD4324BB" box="[151,659,698,720]" gridcol="0" gridrow="11" pageId="6" pageNumber="85">Dorsal-fin length 27.7</th>
<td id="C111294E44216AE06505FD5AFCE824BB" box="[769,824,698,720]" gridcol="1" gridrow="11" pageId="6" pageNumber="85">23.7</td>
<td id="C111294E44216AE065A2FD5AFC1D24BB" box="[934,973,698,720]" gridcol="2" gridrow="11" pageId="6" pageNumber="85">1.9</td>
<td id="C111294E44216AE06243FD5AFAE924BB" box="[1095,1337,698,720]" gridcol="3" gridrow="11" pageId="6" pageNumber="85">18.4–27.7</td>
</tr>
<tr id="82C0403244216AE06693FD04FAE92491" box="[151,1337,740,762]" gridrow="12" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FD04FD432491" box="[151,659,740,762]" gridcol="0" gridrow="12" pageId="6" pageNumber="85">Anal-fin length 17.4</th>
<td id="C111294E44216AE06505FD04FCE82491" box="[769,824,740,762]" gridcol="1" gridrow="12" pageId="6" pageNumber="85">16.7</td>
<td id="C111294E44216AE065A2FD04FC1D2491" box="[934,973,740,762]" gridcol="2" gridrow="12" pageId="6" pageNumber="85">1.4</td>
<td id="C111294E44216AE06243FD04FAE92491" box="[1095,1337,740,762]" gridcol="3" gridrow="12" pageId="6" pageNumber="85">13.9–20.1</td>
</tr>
<tr id="82C0403244216AE06693FCEDFAE92548" box="[151,1337,781,803]" gridrow="13" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FCEDFD432548" box="[151,659,781,803]" gridcol="0" gridrow="13" pageId="6" pageNumber="85">Pelvic-fin length 18.2</th>
<td id="C111294E44216AE06505FCEDFCE82548" box="[769,824,781,803]" gridcol="1" gridrow="13" pageId="6" pageNumber="85">18.6</td>
<td id="C111294E44216AE065A2FCEDFC1D2548" box="[934,973,781,803]" gridcol="2" gridrow="13" pageId="6" pageNumber="85">1.5</td>
<td id="C111294E44216AE06243FCEDFAE92548" box="[1095,1337,781,803]" gridcol="3" gridrow="13" pageId="6" pageNumber="85">15.7–21.8</td>
</tr>
<tr id="82C0403244216AE06693FCD6FAE92527" box="[151,1337,822,844]" gridrow="14" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FCD6FD432527" box="[151,659,822,844]" gridcol="0" gridrow="14" pageId="6" pageNumber="85">Pectoral-fin length 18.0</th>
<td id="C111294E44216AE06505FCD6FCE82527" box="[769,824,822,844]" gridcol="1" gridrow="14" pageId="6" pageNumber="85">21.0</td>
<td id="C111294E44216AE065A2FCD6FC1D2527" box="[934,973,822,844]" gridcol="2" gridrow="14" pageId="6" pageNumber="85">1.6</td>
<td id="C111294E44216AE06243FCD6FAE92527" box="[1095,1337,822,844]" gridcol="3" gridrow="14" pageId="6" pageNumber="85">18.9–23.8</td>
</tr>
<tr id="82C0403244216AE06693FC80FAE9251D" box="[151,1337,864,886]" gridrow="15" pageId="6" pageNumber="85" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="C111294E44216AE06693FC80FD43251D" box="[151,659,864,886]" gridcol="0" gridrow="15" pageId="6" pageNumber="85">In percent of head length</th>
</tr>
<tr id="82C0403244216AE06693FC69FAE925F4" box="[151,1337,905,927]" gridrow="16" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FC69FD4325F4" box="[151,659,905,927]" gridcol="0" gridrow="16" pageId="6" pageNumber="85">Mouth width 43</th>
<td id="C111294E44216AE06505FC69FCE825F4" box="[769,824,905,927]" gridcol="1" gridrow="16" pageId="6" pageNumber="85">27</td>
<td id="C111294E44216AE065A2FC69FC1D25F4" box="[934,973,905,927]" gridcol="2" gridrow="16" pageId="6" pageNumber="85">6.9</td>
<td id="C111294E44216AE06243FC69FAE925F4" box="[1095,1337,905,927]" gridcol="3" gridrow="16" pageId="6" pageNumber="85">13–43</td>
</tr>
<tr id="82C0403244216AE06693FC52FAE925A3" box="[151,1337,946,968]" gridrow="17" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FC52FD4325A3" box="[151,659,946,968]" gridcol="0" gridrow="17" pageId="6" pageNumber="85">Oral disc length 22</th>
<td id="C111294E44216AE06505FC52FCE825A3" box="[769,824,946,968]" gridcol="1" gridrow="17" pageId="6" pageNumber="85">22</td>
<td id="C111294E44216AE065A2FC52FC1D25A3" box="[934,973,946,968]" gridcol="2" gridrow="17" pageId="6" pageNumber="85">4.1</td>
<td id="C111294E44216AE06243FC52FAE925A3" box="[1095,1337,946,968]" gridcol="3" gridrow="17" pageId="6" pageNumber="85">15–34</td>
</tr>
<tr id="82C0403244216AE06693FC3CFAE92599" box="[151,1337,988,1010]" gridrow="18" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FC3CFD432599" box="[151,659,988,1010]" gridcol="0" gridrow="18" pageId="6" pageNumber="85">Oral disc width 25</th>
<td id="C111294E44216AE06505FC3CFCE82599" box="[769,824,988,1010]" gridcol="1" gridrow="18" pageId="6" pageNumber="85">25</td>
<td id="C111294E44216AE065A2FC3CFC1D2599" box="[934,973,988,1010]" gridcol="2" gridrow="18" pageId="6" pageNumber="85">4.7</td>
<td id="C111294E44216AE06243FC3CFAE92599" box="[1095,1337,988,1010]" gridcol="3" gridrow="18" pageId="6" pageNumber="85">14–37</td>
</tr>
<tr id="82C0403244216AE06693FBE5FAE92270" box="[151,1337,1029,1051]" gridrow="19" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FBE5FD432270" box="[151,659,1029,1051]" gridcol="0" gridrow="19" pageId="6" pageNumber="85">Rostral barbel length 27</th>
<td id="C111294E44216AE06505FBE5FCE82270" box="[769,824,1029,1051]" gridcol="1" gridrow="19" pageId="6" pageNumber="85">20</td>
<td id="C111294E44216AE065A2FBE5FC1D2270" box="[934,973,1029,1051]" gridcol="2" gridrow="19" pageId="6" pageNumber="85">4.1</td>
<td id="C111294E44216AE06243FBE5FAE92270" box="[1095,1337,1029,1051]" gridcol="3" gridrow="19" pageId="6" pageNumber="85">9–29</td>
</tr>
<tr id="82C0403244216AE06693FBCEFAE9222F" box="[151,1337,1070,1092]" gridrow="20" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FBCEFD43222F" box="[151,659,1070,1092]" gridcol="0" gridrow="20" pageId="6" pageNumber="85">Maxillary barbel length 20</th>
<td id="C111294E44216AE06505FBCEFCE8222F" box="[769,824,1070,1092]" gridcol="1" gridrow="20" pageId="6" pageNumber="85">16</td>
<td id="C111294E44216AE065A2FBCEFC1D222F" box="[934,973,1070,1092]" gridcol="2" gridrow="20" pageId="6" pageNumber="85">3.7</td>
<td id="C111294E44216AE06243FBCEFAE9222F" box="[1095,1337,1070,1092]" gridcol="3" gridrow="20" pageId="6" pageNumber="85">12–22</td>
</tr>
<tr id="82C0403244216AE06693FBB8FAE92205" box="[151,1337,1112,1134]" gridrow="21" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FBB8FD432205" box="[151,659,1112,1134]" gridcol="0" gridrow="21" pageId="6" pageNumber="85">Eye diameter 21</th>
<td id="C111294E44216AE06505FBB8FCE82205" box="[769,824,1112,1134]" gridcol="1" gridrow="21" pageId="6" pageNumber="85">25</td>
<td id="C111294E44216AE065A2FBB8FC1D2205" box="[934,973,1112,1134]" gridcol="2" gridrow="21" pageId="6" pageNumber="85">3.4</td>
<td id="C111294E44216AE06243FBB8FAE92205" box="[1095,1337,1112,1134]" gridcol="3" gridrow="21" pageId="6" pageNumber="85">17–30</td>
</tr>
<tr id="82C0403244216AE06693FB61FAE922FC" box="[151,1337,1153,1175]" gridrow="22" pageId="6" pageNumber="85">
<th id="C111294E44216AE06693FB61FD4322FC" box="[151,659,1153,1175]" gridcol="0" gridrow="22" pageId="6" pageNumber="85">Interorbital distance 53</th>
<td id="C111294E44216AE06505FB61FCE822FC" box="[769,824,1153,1175]" gridcol="1" gridrow="22" pageId="6" pageNumber="85">52</td>
<td id="C111294E44216AE065A2FB61FC1D22FC" box="[934,973,1153,1175]" gridcol="2" gridrow="22" pageId="6" pageNumber="85">3.5</td>
<td id="C111294E44216AE06243FB61FAE922FC" box="[1095,1337,1153,1175]" gridcol="3" gridrow="22" pageId="6" pageNumber="85">46–58</td>
</tr>
</table>
</paragraph>
<paragraph id="3C4F42704421951966C3FB37FD6223A3" blockId="6.[151,1437,1239,2021]" pageId="6" pageNumber="85">
<emphasis id="0E849E624421951966C3FB37FE8F229B" bold="true" box="[199,351,1239,1264]" pageId="6" pageNumber="85">Colouration.</emphasis>
In living specimens: Head, back and flank brown with an olive green hue (
<figureCitation id="A4CB5EF54421951962A7FB37FB37229B" box="[1187,1255,1239,1264]" captionStart="FIGURE 6" captionStartId="7.[151,249,1429,1451]" captionTargetBox="[151,1435,445,1408]" captionTargetId="figure@7.[151,1436,445,1408]" captionTargetPageId="7" captionText="FIGURE 6. Pool in Wadi Andhur." httpUri="https://zenodo.org/record/256496/files/figure.png" pageId="6" pageNumber="85">Fig. 6</figureCitation>
). A faith yellow or golden mid-dorsal stripe on back. Ventral surface pale white. Flank darkest along midlateral line with an irregular shaped, wide, dark-brown stripe. Caudal, dorsal, pectoral and pelvic-fins with yellowish or greenish rays and hyaline membranes. Branched dorsal-fin ray 4–7½ with a black spot slightly above base, in some individuals black spot continuous with black dorsal-fin rays. Anal fin hyaline or with greenish pigments at its distal part. A dark-brown blotch behind upper edge of operculum. A dark-brown blotch at middle of posterior-most caudal peduncle in individuals smaller than
<quantity id="FB08EF95442195196430FA50FD5323AC" box="[564,643,1456,1480]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="85" unit="mm" value="40.0">40 mm</quantity>
SL.
</paragraph>
<paragraph id="3C4F42704421951966C3FA35FD6220AF" blockId="6.[151,1437,1239,2021]" pageId="6" pageNumber="85">
In ethanol-preserved specimens: Head, back and flank brown. Ventral surface pale grey. Flank darker brown along midlateral line than on upper and lower flank, scales with dark-brown margin and pale-brown centre giving the flank a slightly reticulate pattern. Individual flank scales whitish in centre. Caudal, dorsal, pectoral and pelvicfins with grey rays and hyaline membranes. Branched dorsal-fin ray 4–7½ with a black spot slightly above base, in some individuals, black spot continuous with black dorsal-fin rays. Anal fin hyaline or grey at its distal part. A dark-brown blotch behind upper edge of operculum. A dark-brown blotch at middle of posterior-most caudal peduncle in individuals smaller than
<quantity id="FB08EF95442195196430F94CFD5320AF" box="[564,643,1708,1732]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="85" unit="mm" value="40.0">40 mm</quantity>
SL.
</paragraph>
</subSubSection>
<subSubSection id="74EA11FB4421951866C3F92FFF0027E3" lastPageId="7" lastPageNumber="86" pageId="6" pageNumber="85" type="distribution">
<paragraph id="3C4F42704421951966C3F92FFCE7211C" blockId="6.[151,1437,1239,2021]" pageId="6" pageNumber="85">
<emphasis id="0E849E624421951966C3F92FFEB02083" bold="true" box="[199,352,1743,1768]" pageId="6" pageNumber="85">Distribution.</emphasis>
<taxonomicName id="FBF039F344219519676CF930FE2A2083" box="[360,506,1743,1768]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="85" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E6244219519676CF930FE2A2083" box="[360,506,1743,1768]" italics="true" pageId="6" pageNumber="85">Garra sindhi</emphasis>
</taxonomicName>
was found in a large pool (Fig. 7) in the lower Wadi Andhur in the Jebel Samhan Nature Reserve. Wadi Andhur is an endorheic drainage flowing from the Samhan plateau to the north of the Jebel Samhan escarpment (García et al. 2015). In areal pictures on Google satellite maps, pools with water are also visible higher up in Wadi Andhur as well as in adjacent wadis and
<taxonomicName id="FBF039F3442195196598F8DDFBD0213E" box="[924,1024,1852,1877]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="85" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E62442195196598F8DDFBD0213E" box="[924,1024,1852,1877]" italics="true" pageId="6" pageNumber="85">G. sindhi</emphasis>
</taxonomicName>
might be more widespread. Further research is recommended to clarify if this is in fact the case.
</paragraph>
<paragraph id="3C4F42704421951866C3F864FF0027E3" blockId="6.[151,1437,1239,2021]" lastBlockId="7.[151,1436,151,392]" lastPageId="7" lastPageNumber="86" pageId="6" pageNumber="85">
<emphasis id="0E849E624421951966C3F864FEFA21F6" bold="true" box="[199,298,1924,1949]" pageId="6" pageNumber="85">Habitat.</emphasis>
Wadi Andhur originates in the Samhan mountain and flows through narrow canyons north to the sand-gravel desert, which forms part of the Nejd. Wadi Andhur is situated in a hyper-arid region, with rainfall ranging from less than
<quantity id="FB08EF9544219519679AF82CFE2C218F" box="[414,508,1996,2020]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="85" unit="mm" value="100.0">100 mm</quantity>
to between
<quantity id="FB08EF95442195196483F82CFCCE218F" box="[647,798,1996,2021]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="6" pageNumber="85" unit="mm" value="175.0" valueMax="250.0" valueMin="100.0">100–250 mm</quantity>
per year (
<bibRefCitation id="58613F81442195196595F82CFB58218F" author="Kwarteng" box="[913,1160,1996,2021]" pageId="6" pageNumber="85" refString="Kwarteng, A. Y., Dorvlo, A. S. &amp; Vijaya Kumar, G. T. (2009) Analysis of a 27 - year rainfall data (1977 - 2003) in the Sultanate of Oman. Inernational Journal of Climatology, 29, 605 - 617. http: // dx. doi. org / 10.1002 / joc. 1727" type="journal article" year="2009">
Kwarteng
<emphasis id="0E849E6244219519620EF82DFB93218E" box="[1034,1091,1996,2021]" italics="true" pageId="6" pageNumber="85">et al.</emphasis>
2009
</bibRefCitation>
). Flow is infrequent and unpredictable. Flood events follow heavy storms and cyclones that occur erratically (
<bibRefCitation id="58613F81442095186254FF78FA9326DB" author="Kwarteng" box="[1104,1347,151,176]" pageId="7" pageNumber="86" refString="Kwarteng, A. Y., Dorvlo, A. S. &amp; Vijaya Kumar, G. T. (2009) Analysis of a 27 - year rainfall data (1977 - 2003) in the Sultanate of Oman. Inernational Journal of Climatology, 29, 605 - 617. http: // dx. doi. org / 10.1002 / joc. 1727" type="journal article" year="2009">
Kwarteng
<emphasis id="0E849E624420951862C3FF79FAD026DB" box="[1223,1280,151,176]" italics="true" pageId="7" pageNumber="86">et al.</emphasis>
2009
</bibRefCitation>
).
<taxonomicName id="FBF039F3442095186352FF78FF0B26BE" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E62442095186352FF78FF0B26BE" italics="true" pageId="7" pageNumber="86">Garra sindhi</emphasis>
</taxonomicName>
was found in a large pool of about 40 ×
<quantity id="FB08EF954420951864A4FF5CFD0926BF" box="[672,729,188,213]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="86" unit="m" value="35.0">35 m</quantity>
with stagnant water. The average depth of the pool was around
<quantity id="FB08EF95442095186693FF3FFF34269C" box="[151,228,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="86" unit="cm" value="60.0">60 cm</quantity>
with a maximum depth of about
<quantity id="FB08EF95442095186494FF00FD30269C" box="[656,736,224,248]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="86" unit="cm" value="80.0">80 cm</quantity>
. The water in the pool was clear and
<taxonomicName id="FBF039F34420951862C3FF00FAE6269C" box="[1223,1334,224,247]" class="Zygnematophyceae" family="Zygnemataceae" genus="Spirogyra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Zygnematales" pageId="7" pageNumber="86" phylum="Charophyta" rank="genus">
<emphasis id="0E849E624420951862C3FF00FAE6269C" box="[1223,1334,224,247]" italics="true" pageId="7" pageNumber="86">Spirogyra</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="FBF039F3442095186693FEE4FE112776" box="[151,449,260,285]" class="Magnoliopsida" family="Ceratophyllaceae" genus="Ceratophyllum" kingdom="Plantae" order="Ceratophyllales" pageId="7" pageNumber="86" phylum="Tracheophyta" rank="species" species="demersum">
<emphasis id="0E849E62442095186693FEE4FE112776" box="[151,449,260,285]" italics="true" pageId="7" pageNumber="86">Ceratophyllum demersum</emphasis>
</taxonomicName>
were growing along the edges of the bank, as well as several stands of
<taxonomicName id="FBF039F344209518635CFEE4FEF3272B" class="Liliopsida" family="Typhaceae" genus="Typha" kingdom="Plantae" order="Poales" pageId="7" pageNumber="86" phylum="Tracheophyta" rank="species" species="domingensis">
<emphasis id="0E849E6244209518635CFEE4FEF3272B" italics="true" pageId="7" pageNumber="86">Typha domingensis</emphasis>
</taxonomicName>
. The pool has a coarse gravel bed of round limestone pebbles. The pool is frequently visited by livestock and wildlife.
<taxonomicName id="FBF039F344209518679FFEACFDEA270F" box="[411,570,332,357]" class="Aves" family="Ardeidae" genus="Ardea" kingdom="Animalia" order="Pelecaniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="cinerea">
<emphasis id="0E849E6244209518679FFEACFDEA270F" box="[411,570,332,357]" italics="true" pageId="7" pageNumber="86">Ardea cinerea</emphasis>
</taxonomicName>
,
<taxonomicName id="FBF039F344209518644DFEACFCF7270F" box="[585,807,332,357]" class="Aves" family="Ardeidae" genus="Ixobrychus" kingdom="Animalia" order="Pelecaniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="0E849E6244209518644DFEACFCF7270F" box="[585,807,332,357]" italics="true" pageId="7" pageNumber="86">Ixobrychus sinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="FBF039F3442095186564FEACFB91270E" box="[864,1089,332,357]" class="Aves" family="Podicipedidae" genus="Podiceps" kingdom="Animalia" order="Podicipediformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="nigricollis">
<emphasis id="0E849E62442095186564FEACFB91270E" box="[864,1089,332,357]" italics="true" pageId="7" pageNumber="86">Podiceps nigricollis</emphasis>
</taxonomicName>
were observed foraging at the pool.
</paragraph>
</subSubSection>
<caption id="688F12F8442095186693FA75FE2623C0" box="[151,502,1429,1451]" httpUri="https://zenodo.org/record/256496/files/figure.png" pageId="7" pageNumber="86" targetBox="[151,1435,445,1408]" targetPageId="7">
<paragraph id="3C4F4270442095186693FA75FE2623C0" blockId="7.[151,502,1429,1451]" box="[151,502,1429,1451]" pageId="7" pageNumber="86">
<emphasis id="0E849E62442095186693FA75FEC223C0" bold="true" box="[151,274,1429,1451]" pageId="7" pageNumber="86">FIGURE 6.</emphasis>
Pool in Wadi Andhur.
</paragraph>
</caption>
<subSubSection id="74EA11FB4420951866C3FA38FDE820A2" pageId="7" pageNumber="86" type="description">
<paragraph id="3C4F42704420951866C3FA38FDE820A2" blockId="7.[151,1437,1496,2025]" pageId="7" pageNumber="86">
<emphasis id="0E849E624420951866C3FA38FE6B239A" bold="true" box="[199,443,1496,1521]" pageId="7" pageNumber="86">Conservation status.</emphasis>
Beside the pool, where
<taxonomicName id="FBF039F34420951864CEFA39FCFC239A" box="[714,812,1496,1521]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624420951864CEFA39FCFC239A" box="[714,812,1496,1521]" italics="true" pageId="7" pageNumber="86">G. sindhi</emphasis>
</taxonomicName>
was found, there are a few much smaller shallow pools upstream, several metres off the main pool. If the species is restricted to these pools, it would be one of the rarest freshwater fish species in the
<collectingRegion id="FE348C924420951867F3F9C1FD822053" box="[503,594,1569,1592]" country="American Samoa" name="Western" pageId="7" pageNumber="86">Western</collectingRegion>
Palaearctic. However, several other pools in the upper reaches of Wadi Andhur and in adjacent wadis are visible from areal pictures and the species might be more widespread than presently estimated.
<taxonomicName id="FBF039F3442095186787F989FDFD20EA" box="[387,557,1640,1665]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="dunsirei">
<emphasis id="0E849E62442095186787F989FDFD20EA" box="[387,557,1640,1665]" italics="true" pageId="7" pageNumber="86">Garra dunsirei</emphasis>
</taxonomicName>
and
<emphasis id="0E849E6244209518646DF989FD1E20EB" box="[617,718,1641,1664]" italics="true" pageId="7" pageNumber="86">G. s m a r t i</emphasis>
, two other species from the region, are also restricted to very small water-bodies. The population of
<taxonomicName id="FBF039F3442095186467F96DFD1B20CE" box="[611,715,1676,1701]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E62442095186467F96DFD1B20CE" box="[611,715,1676,1701]" italics="true" pageId="7" pageNumber="86">G. sindhi</emphasis>
</taxonomicName>
appears to be healthy and more than 1000 individuals were recorded at the time of the two visits.
</paragraph>
</subSubSection>
<subSubSection id="74EA11FB4420951866C3F934FEE8217A" pageId="7" pageNumber="86" type="etymology">
<paragraph id="3C4F42704420951866C3F934FEE8217A" blockId="7.[151,1437,1496,2025]" pageId="7" pageNumber="86">
<emphasis id="0E849E624420951866C3F934FE9E2086" bold="true" box="[199,334,1748,1773]" pageId="7" pageNumber="86">Etymology.</emphasis>
Named for Cynthia “
<taxonomicName id="FBF039F344209518643AF934FD5B2086" box="[574,651,1748,1773]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="sindhi">Sindhi</taxonomicName>
” Diane Powell in recognition of her support for the first authors work over a decade.
</paragraph>
</subSubSection>
<subSubSection id="74EA11FB4420951766C3F8FCFE7526BE" lastPageId="8" lastPageNumber="87" pageId="7" pageNumber="86" type="discussion">
<paragraph id="3C4F42704420951766C3F8FCFE7526BE" blockId="7.[151,1437,1496,2025]" lastBlockId="8.[151,1436,151,429]" lastPageId="8" lastPageNumber="87" pageId="7" pageNumber="86">
<emphasis id="0E849E624420951866C3F8FCFEEB215E" bold="true" box="[199,315,1820,1845]" pageId="7" pageNumber="86">Remarks.</emphasis>
<taxonomicName id="FBF039F3442095186741F8FDFE21215E" box="[325,497,1820,1845]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="dunsirei">
<emphasis id="0E849E62442095186741F8FDFE21215E" box="[325,497,1820,1845]" italics="true" pageId="7" pageNumber="86">Garra dunsirei</emphasis>
</taxonomicName>
,
<emphasis id="0E849E62442095186407F8FDFDB9215F" box="[515,617,1821,1844]" italics="true" pageId="7" pageNumber="86">G. s m a r t i</emphasis>
and
<taxonomicName id="FBF039F34420951864AFF8FDFCC1215E" box="[683,785,1820,1845]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="sindhi">
<emphasis id="0E849E624420951864AFF8FDFCC1215E" box="[683,785,1820,1845]" italics="true" pageId="7" pageNumber="86">G. sindhi</emphasis>
</taxonomicName>
are three closely related species endemic to the Omani
<collectingRegion id="FE348C92442095186693F8A0FF362132" box="[151,230,1856,1881]" country="Oman" name="Zufar" pageId="7" pageNumber="86">Dhofar</collectingRegion>
province. While
<taxonomicName id="FBF039F34420951867A3F8A1FDF02132" box="[423,544,1856,1881]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="dunsirei">
<emphasis id="0E849E624420951867A3F8A1FDF02132" box="[423,544,1856,1881]" italics="true" pageId="7" pageNumber="86">G. dunsirei</emphasis>
</taxonomicName>
and
<emphasis id="0E849E62442095186450F8A1FD662133" box="[596,694,1857,1880]" italics="true" pageId="7" pageNumber="86">G. s m a r t i</emphasis>
occur in drainage systems flowing to the sea, Wadi Andhur drains to the north, into the Nejd. The weak genetic differentiation between these species suggests a recent origin, which could be verified if geological data were available for this region. It is, however, not paralleled by limited morphological divergence among them, since all three species can be readily identified by their morphology. The existence of species with low genetic divergence corroborates the nature of evolution as a gradual process and highlights the key role taxonomists play in building and monitoring DNA barcode reference libraries used for specimen identification.
</paragraph>
</subSubSection>
<subSubSection id="74EA11FB442F951766C3FF3FFD4027C7" pageId="8" pageNumber="87" type="reference_group">
<paragraph id="3C4F4270442F951766C3FF3FFD4027C7" blockId="8.[151,1436,151,429]" pageId="8" pageNumber="87">
The
<taxonomicName id="FBF039F3442F951766FEFF00FE90269C" ID-CoL="4M9J" box="[250,320,224,247]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis id="0E849E62442F951766FEFF00FE90269C" box="[250,320,224,247]" italics="true" pageId="8" pageNumber="87">Garra</emphasis>
</taxonomicName>
population inhabiting Wadi Bani Khalid in the north of
<collectingCountry id="44E702E0442F951765BCFF00FC2D2693" box="[952,1021,224,248]" name="Oman" pageId="8" pageNumber="87">Oman</collectingCountry>
had been described as
<taxonomicName id="FBF039F3442F95176306FF00FEDC2776" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="subSpecies" species="barreimiae" subSpecies="gallagheri">
<emphasis id="0E849E62442F95176306FF00FEDC2776" italics="true" pageId="8" pageNumber="87">G. barreimiae gallagheri</emphasis>
</taxonomicName>
by
<bibRefCitation id="58613F81442F9517673DFEE5FE072777" author="Krupp" box="[313,471,260,284]" pageId="8" pageNumber="87" refString="Krupp, F. (1988) Freshwater fishes of the Wadi Batha drainage. Journal of Oman Studies Special Report, 3, 401 - 404." type="journal article" year="1988">Krupp (1988)</bibRefCitation>
. It has already been included in the molecular study by
<bibRefCitation id="58613F81442F95176253FEE4FA472777" box="[1111,1431,260,285]" pageId="8" pageNumber="87" refString="Kruckenhauser, L., Haring, E., Seemann, R. &amp; Sattmann, H. (2011) Genetic differentiation between cave and surface dwelling populations of Garra barreimiae (Cyprinidae) in Oman. BMC Evolutionary Biology, 11, 172 - 186. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 11 - 172" type="journal article">
Kruckenhauser
<emphasis id="0E849E62442F9517630DFEE5FA932776" box="[1289,1347,260,285]" italics="true" pageId="8" pageNumber="87">et al.</emphasis>
(2011)
</bibRefCitation>
, who did not draw any taxonomic conclusions from their study. We found
<taxonomicName id="FBF039F3442F95176213FEC8FB61272B" box="[1047,1201,295,320]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="0E849E62442F95176213FEC8FB61272B" box="[1047,1201,295,320]" italics="true" pageId="8" pageNumber="87">G. gallagheri</emphasis>
</taxonomicName>
to be related to
<taxonomicName id="FBF039F3442F95176380FEC8FEC5270E" ID-CoL="3FBKJ" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="0E849E62442F95176380FEC8FEC5270E" italics="true" pageId="8" pageNumber="87">G. longipinnis</emphasis>
</taxonomicName>
. The study by
<bibRefCitation id="58613F81442F951767C4FEACFD2D270F" box="[448,765,332,357]" pageId="8" pageNumber="87" refString="Kruckenhauser, L., Haring, E., Seemann, R. &amp; Sattmann, H. (2011) Genetic differentiation between cave and surface dwelling populations of Garra barreimiae (Cyprinidae) in Oman. BMC Evolutionary Biology, 11, 172 - 186. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 11 - 172" type="journal article">
Kruckenhauser
<emphasis id="0E849E62442F95176476FEADFD7C270E" box="[626,684,332,357]" italics="true" pageId="8" pageNumber="87">et al.</emphasis>
(2011)
</bibRefCitation>
suggests that the species diversity of
<taxonomicName id="FBF039F3442F951762AFFEADFB21270F" ID-CoL="4M9J" box="[1195,1265,333,356]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis id="0E849E62442F951762AFFEADFB21270F" box="[1195,1265,333,356]" italics="true" pageId="8" pageNumber="87">Garra</emphasis>
</taxonomicName>
in the north of
<collectingCountry id="44E702E0442F95176693FE90FF0C27E3" box="[151,220,368,392]" name="Oman" pageId="8" pageNumber="87">Oman</collectingCountry>
might be underestimated and more than the three actually recognised species (
<taxonomicName id="FBF039F3442F95176253FE90FB2227E3" ID-CoL="3FBGY" box="[1111,1266,367,392]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="0E849E62442F95176253FE90FB2227E3" box="[1111,1266,367,392]" italics="true" pageId="8" pageNumber="87">G. barreimiae</emphasis>
</taxonomicName>
,
<taxonomicName id="FBF039F3442F95176305FE90FA4527E3" box="[1281,1429,367,392]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="0E849E62442F95176305FE90FA4527E3" box="[1281,1429,367,392]" italics="true" pageId="8" pageNumber="87">G. gallagheri</emphasis>
</taxonomicName>
,
<taxonomicName id="FBF039F3442F95176693FE75FEE327C6" ID-CoL="3FBKJ" box="[151,307,404,429]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="0E849E62442F95176693FE75FEE327C6" box="[151,307,404,429]" italics="true" pageId="8" pageNumber="87">G. longipinnis</emphasis>
</taxonomicName>
) might exist in that small area.
</paragraph>
</subSubSection>
</treatment>
</document>
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