321 lines
37 KiB
XML
321 lines
37 KiB
XML
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<mods:title id="D100EC110DDD891478F5E306F94472F2">The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution</mods:title>
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<mods:namePart id="7362F8304A217DB16705FE34B2F37656">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="5D500CC4868ECAC4E3AE999D33492A26">Gasca-Álvarez, Héctor J.</mods:namePart>
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<mods:namePart id="246C65BE1C696696D7F5B226BF7154BC">Amat-García, Germán</mods:namePart>
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<mods:date id="7E315A68F23279BDE5005E122AC47104">2015</mods:date>
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<mods:number id="32838A448691FDCE9A37D0130E4DA6CB">4048</mods:number>
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<treatment id="990DA529FF84FF9BFF4646B11580FE4A" ID-DOI="http://doi.org/10.5281/zenodo.5699384" ID-GBIF-Taxon="119645672" ID-Zenodo-Dep="5699384" LSID="urn:lsid:plazi:treatment:990DA529FF84FF9BFF4646B11580FE4A" httpUri="http://treatment.plazi.org/id/990DA529FF84FF9BFF4646B11580FE4A" lastPageId="28" lastPageNumber="479" pageId="26" pageNumber="477">
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<emphasis id="23D0C82DFF84FF9DFF4646B11553FAEE" bold="true" box="[151,778,1262,1288]" pageId="26" pageNumber="477">
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<taxonomicName id="D6A46FBCFF84FF9DFF4646B117EEFAEE" box="[151,439,1262,1288]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="bituberculatus">
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<emphasis id="23D0C82DFF84FF9DFF4646B117EEFAEE" bold="true" box="[151,439,1262,1288]" italics="true" pageId="26" pageNumber="477">Tomarus bituberculatus</emphasis>
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</taxonomicName>
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(Palisot de
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<bibRefCitation id="753569CEFF84FF9DFD9946B1155BFAEE" author="Beauvois" box="[584,770,1262,1288]" pageId="26" pageNumber="477" refString="Palisot de Beauvois, A. M. F. J. (1811) Insectes Recueillis en Afrique et em Amerique, dans les Royaumes d´Oware et de Benin, a Saint-Dominigue et dans les Etats-Unis, pendant les Anees 1786 - 1797. Levrault, Schoell, et Cie, Paris, France, Livraison 7, pp. 101 - 120." type="journal article" year="1811">Beauvois, 1811</bibRefCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="111B143FFF84FF9DFF46474E1765FACC" blockId="26.[151,778,1262,1322]" box="[151,316,1297,1322]" pageId="26" pageNumber="477">
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(
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<figureCitation id="899F08BAFF84FF9DFF4E474E176AFACC" box="[159,307,1297,1322]" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="26" pageNumber="477">Figs. 94–101</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF84FF9DFF4647061785FA0C" pageId="26" pageNumber="477" type="reference_group">
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<paragraph id="111B143FFF84FF9DFF46470617A2FA4A" blockId="26.[151,747,1369,1514]" pageId="26" pageNumber="477">
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<treatmentCitationGroup id="31B43311FF84FF9DFF46470617A2FA4A" pageId="26" pageNumber="477">
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<taxonomicName id="D6A46FBCFF84FF9DFF46470617A9FA89" ID-CoL="6Y84T" authority="Palisot" authorityName="Palisot" box="[151,496,1369,1391]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="bituberculatus">
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<emphasis id="23D0C82DFF84FF9DFF46470617FDFA89" box="[151,420,1369,1391]" italics="true" pageId="26" pageNumber="477">Scarabaeus bituberculatus</emphasis>
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Palisot
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</taxonomicName>
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de
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<treatmentCitation id="9005322EFF84FF9DFDC5470614BEFA89" author="Beauvois" box="[532,743,1369,1391]" page="103" pageId="26" pageNumber="477" year="1811">
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<bibRefCitation id="753569CEFF84FF9DFDC5470614ECFA89" author="Beauvois" box="[532,693,1369,1391]" pageId="26" pageNumber="477" refString="Palisot de Beauvois, A. M. F. J. (1811) Insectes Recueillis en Afrique et em Amerique, dans les Royaumes d´Oware et de Benin, a Saint-Dominigue et dans les Etats-Unis, pendant les Anees 1786 - 1797. Levrault, Schoell, et Cie, Paris, France, Livraison 7, pp. 101 - 120." type="journal article" year="1811">Beauvois, 1811</bibRefCitation>
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: 103
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</treatmentCitation>
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||
.
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<taxonomicName id="D6A46FBCFF84FF9DFF46472714C2FA68" authority="Burmeister, 1847: 100" authorityName="Burmeister" authorityPageNumber="100" authorityYear="1847" box="[151,667,1400,1422]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="scarabaeinus">
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<emphasis id="23D0C82DFF84FF9DFF46472717E9FA68" box="[151,432,1400,1422]" italics="true" pageId="26" pageNumber="477">Heteronychus scarabaeinus</emphasis>
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<treatmentCitation id="9005322EFF84FF9DFE67472714C2FA68" author="Burmeister" box="[438,667,1400,1422]" page="100" pageId="26" pageNumber="477" year="1847">
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<bibRefCitation id="753569CEFF84FF9DFE6747271431FA68" author="Burmeister" box="[438,616,1400,1422]" pageId="26" pageNumber="477" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia, Band 5. Enslin, Berlin, Germany." type="book" year="1847">Burmeister, 1847</bibRefCitation>
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: 100
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</treatmentCitation>
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</taxonomicName>
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.
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<taxonomicName id="D6A46FBCFF84FF9DFF4647C717AEFA4A" authority="Bates, 1888: 315" authorityName="Bates" authorityPageNumber="315" authorityYear="1888" box="[151,503,1430,1453]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="latifovea">
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<emphasis id="23D0C82DFF84FF9DFF4647C7171DFA4A" box="[151,324,1430,1453]" italics="true" pageId="26" pageNumber="477">Ligyrus latifovea</emphasis>
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<treatmentCitation id="9005322EFF84FF9DFE9847C817AEFA4A" author="Bates" box="[329,503,1430,1452]" page="315" pageId="26" pageNumber="477" year="1888">
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<bibRefCitation id="753569CEFF84FF9DFE9847C8179DFA4A" author="Bates" box="[329,452,1430,1452]" pageId="26" pageNumber="477" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, Family Dynastinae. In: Godman, F. D. & Salvin, O. (Eds), Biologia Centrali-Americana, Insecta, Coleoptera, Vol. II, Part 2, 296 - 342." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
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: 315
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="111B143FFF84FF9DFF4647E9145CFA2D" blockId="26.[151,747,1369,1514]" box="[151,517,1461,1483]" pageId="26" pageNumber="477">
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<treatmentCitationGroup id="31B43311FF84FF9DFF4647E9145CFA2D" box="[151,517,1461,1483]" pageId="26" pageNumber="477">
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<taxonomicName id="D6A46FBCFF84FF9DFF4647E91458FA2D" ID-CoL="6PZ9Q" authority="Arrow, 1913: 466" authorityName="Arrow" authorityPageNumber="466" authorityYear="1913" box="[151,513,1461,1483]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="maximus">
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<emphasis id="23D0C82DFF84FF9DFF4647E9171CFA2D" box="[151,325,1462,1483]" italics="true" pageId="26" pageNumber="477">Ligyrus maximus</emphasis>
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<treatmentCitation id="9005322EFF84FF9DFE9A47E91458FA2D" author="Arrow" box="[331,513,1461,1483]" page="466" pageId="26" pageNumber="477" year="1913">
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<bibRefCitation id="753569CEFF84FF9DFE9A47E91796FA2D" author="Arrow" box="[331,463,1461,1483]" pageId="26" pageNumber="477" refString="Arrow, G. J. (1913) Some new species of Lamellicorn Beetles from Brazil. The Annals and Magazine of natural History, including Zoology, Botany and Geology. London, series 8, volume 11, 456 - 466." type="journal article" year="1913">Arrow, 1913</bibRefCitation>
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||
: 466
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="111B143FFF84FF9DFF46478A1785FA0C" blockId="26.[151,747,1369,1514]" box="[151,476,1492,1514]" pageId="26" pageNumber="477">
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<treatmentCitationGroup id="31B43311FF84FF9DFF46478A1785FA0C" box="[151,476,1492,1514]" pageId="26" pageNumber="477">
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<taxonomicName id="D6A46FBCFF84FF9DFF46478A1781FA0C" ID-CoL="6PYXQ" authority="Arrow, 1914: 273" authorityName="Arrow" authorityPageNumber="273" authorityYear="1914" box="[151,472,1492,1514]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="latus">
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<emphasis id="23D0C82DFF84FF9DFF46478A1745FA0C" box="[151,284,1492,1514]" italics="true" pageId="26" pageNumber="477">Ligyrus latus</emphasis>
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<treatmentCitation id="9005322EFF84FF9DFEF3478B1781FA0C" author="Arrow" box="[290,472,1492,1514]" page="273" pageId="26" pageNumber="477" year="1914">
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<bibRefCitation id="753569CEFF84FF9DFEF3478B17FCFA0C" author="Arrow" box="[290,421,1492,1514]" pageId="26" pageNumber="477" refString="Arrow, G. J. (1914) Some further notes on lamellicorn beetles of the subfamily Dynastinae. Annals and Magazine of Natural History, series 8, volume 14, 257 - 276, 360." type="journal article" year="1914">Arrow, 1914</bibRefCitation>
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: 273
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="59BE47B4FF84FF9CFF464449170CFEA6" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477" type="description">
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<paragraph id="111B143FFF84FF9CFF464449170CFEA6" blockId="26.[151,1437,1558,2015]" lastBlockId="27.[151,1436,151,320]" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
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<emphasis id="23D0C82DFF84FF9DFF4644491770F9C9" bold="true" box="[151,297,1558,1583]" pageId="26" pageNumber="477">Description.</emphasis>
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Habitus as in
|
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<figureCitation id="899F08BAFF84FF9DFE1744491441F9C9" box="[454,536,1558,1583]" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="26" pageNumber="477">Fig. 94</figureCitation>
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. Length 26.0–
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<quantity id="D65CB9DAFF84FF9DFD6844491542F9C8" box="[697,795,1558,1583]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" pageId="26" pageNumber="477" unit="mm" value="28.5">28.5 mm</quantity>
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||
(♂), 25.0–30.0 mm (♀). Width 13.0–15.0 mm (♂),
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<quantity id="D65CB9DAFF84FF9DFA8C444916A4F9B4" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.45" metricValueMin="1.35" pageId="26" pageNumber="477" unit="mm" value="14.0" valueMax="14.5" valueMin="13.5">13.5– 14.5 mm</quantity>
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(♀). Color black to dark brown, venter lighter.
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<emphasis id="23D0C82DFF84FF9DFCF544651535F9B5" bold="true" box="[804,876,1594,1619]" italics="true" pageId="26" pageNumber="477">Head:</emphasis>
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Frons flat; surface strongly rugose and grooved; base nearly smooth, with sparse, deep punctures. Frontoclypeal suture only visible on the sides, interrupted by 2 frontal tubercles. Tubercles transverse and triangular, separated by about 4.5–5.0 times a tubercle diameter (
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<figureCitation id="899F08BAFF84FF9DFABE44DD16EEF959" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="26" pageNumber="477">Fig. 97</figureCitation>
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). Clypeus strongly rugose and grooved; shape triangular, apex narrowed with 2 reflexed teeth. Interocular distance equals 3.5–3.7 times the transverse ocular diameter. Antennal subequal in length to antennomeres 2–7. Mandibles with 2 apical teeth and one basal rounded lobe.
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<emphasis id="23D0C82DFF84FF9DFCE744B015EAF8E0" bold="true" box="[822,947,1775,1798]" italics="true" pageId="26" pageNumber="477">Pronotum:</emphasis>
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Surface with shallow and small punctures, denser and deeper on anterior and posterior angles. Apical tubercle prominent, apex strongly acute and slightly recurved (
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<figureCitation id="899F08BAFF84FF9DFEC345691728F8A9" box="[274,369,1846,1871]" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="26" pageNumber="477">Fig. 96</figureCitation>
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||
). Subapical fovea transversally oval, deep, rugose and finely punctate (
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||
<figureCitation id="899F08BAFF84FF9DFB23456913C9F8A9" box="[1266,1424,1846,1871]" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="26" pageNumber="477">Figs. 95, 97</figureCitation>
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||
).
|
||
<taxonomicName id="D6A46FBCFF84FF9DFF4645051728F895" authority="Surface" authorityName="Surface" box="[151,369,1882,1907]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="26" pageNumber="477" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="23D0C82DFF84FF9DFF464505174CF895" bold="true" box="[151,277,1882,1907]" italics="true" pageId="26" pageNumber="477">Scutellum:</emphasis>
|
||
Surface
|
||
</taxonomicName>
|
||
with small to minute, sparse punctures.
|
||
<emphasis id="23D0C82DFF84FF9DFC8D450515F4F895" bold="true" box="[860,941,1882,1907]" italics="true" pageId="26" pageNumber="477">Elytra:</emphasis>
|
||
Surface with deep, large punctures, some smaller between intervals; 3 pairs of double rows of punctures; elytral suture present.
|
||
<emphasis id="23D0C82DFF84FF9DFB4C4521134CF871" bold="true" box="[1181,1301,1918,1943]" italics="true" pageId="26" pageNumber="477">
|
||
<taxonomicName id="D6A46FBCFF84FF9DFB4C45211354F871" box="[1181,1293,1918,1943]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="26" pageNumber="477" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
Basal third strongly punctate and rugose. Disc with sparser punctures. Surface weakly convex in males, nearly flat in females.
|
||
<emphasis id="23D0C82DFF84FF9DFF2D45981762F838" bold="true" box="[252,315,1991,2014]" italics="true" pageId="26" pageNumber="477">Legs:</emphasis>
|
||
Protibia tridentate, teeth spaced by a subequal distance. Male protarsus simple, not enlarged. Apex of metatibia strongly crenulate; apical margin with 15–17 spinules. Apex of first metatarsomere subtruncate.
|
||
<emphasis id="23D0C82DFF85FF9CFEF742E31723FF35" bold="true" box="[294,378,188,211]" italics="true" pageId="27" pageNumber="478">Venter:</emphasis>
|
||
Prosternal process long, extending beyond procoxae; apex flat, longitudinally oval, posterior half bordered by long yellow setae.
|
||
<emphasis id="23D0C82DFF85FF9CFDF942BF14F7FF1E" bold="true" box="[552,686,223,248]" pageId="27" pageNumber="478">
|
||
<emphasis id="23D0C82DFF85FF9CFDF942BF14FDFF11" bold="true" box="[552,676,224,247]" italics="true" pageId="27" pageNumber="478">Parameres</emphasis>
|
||
:
|
||
</emphasis>
|
||
Base broad; lateral teeth small and acute; apex broad, with 2 small lobed projections on each apex (
|
||
<figureCitation id="899F08BAFF85FF9CFDD4435B14D6FEFA" box="[517,655,260,285]" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="27" pageNumber="478">Figs. 98–99</figureCitation>
|
||
).
|
||
<emphasis id="23D0C82DFF85FF9CFD73435C1524FEFA" bold="true" box="[674,893,259,284]" italics="true" pageId="27" pageNumber="478">Spiculum gastrale:</emphasis>
|
||
Base larger than lateral branches, apex strongly broad (
|
||
<figureCitation id="899F08BAFF85FF9CFF3743781711FEA6" box="[230,328,295,320]" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="27" pageNumber="478">Fig. 100</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="45DB44B7FF85FF9CFF4646D8175BFB3D" httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="27" pageNumber="478" targetBox="[162,1433,376,1132]" targetPageId="27">
|
||
<paragraph id="111B143FFF85FF9CFF4646D8175BFB3D" blockId="27.[151,1436,1159,1243]" pageId="27" pageNumber="478">
|
||
<emphasis id="23D0C82DFF85FF9CFF4646D81738FB7A" bold="true" box="[151,353,1159,1182]" pageId="27" pageNumber="478">FIGURES 94–101.</emphasis>
|
||
<taxonomicName id="D6A46FBCFF85FF9CFEB646D6140FFB7B" box="[359,598,1159,1182]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="bituberculatus">
|
||
<emphasis id="23D0C82DFF85FF9CFEB646D6140FFB7B" box="[359,598,1159,1182]" italics="true" pageId="27" pageNumber="478">Tomarus bituberculatus</emphasis>
|
||
</taxonomicName>
|
||
. 94) habitus in dorsal view, 95
|
||
<emphasis id="23D0C82DFF85FF9CFC4146D715C4FB7B" bold="true" box="[912,925,1160,1181]" pageId="27" pageNumber="478">–</emphasis>
|
||
96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="59BE47B4FF85FF9BFF16475D124FFF32" lastPageId="28" lastPageNumber="479" pageId="27" pageNumber="478" type="diagnosis">
|
||
<paragraph id="111B143FFF85FF9CFF16475D1240FAD9" blockId="27.[151,1437,1282,1991]" pageId="27" pageNumber="478">
|
||
<emphasis id="23D0C82DFF85FF9CFF16475D1718FAFD" bold="true" box="[199,321,1282,1307]" pageId="27" pageNumber="478">Diagnosis.</emphasis>
|
||
This species is similar to
|
||
<taxonomicName id="D6A46FBCFF85FF9CFDB5475C14E9FAFC" box="[612,688,1283,1306]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="gyas">
|
||
<emphasis id="23D0C82DFF85FF9CFDB5475C14E9FAFC" box="[612,688,1283,1306]" italics="true" pageId="27" pageNumber="478">T. gyas</emphasis>
|
||
</taxonomicName>
|
||
by the size of the pronotal tubercle and fovea. However, the fovea of
|
||
<taxonomicName id="D6A46FBCFF85FF9CFF644778172BFAD9" box="[181,370,1318,1343]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="bituberculatus">
|
||
<emphasis id="23D0C82DFF85FF9CFF644778172BFAD9" box="[181,370,1318,1343]" italics="true" pageId="27" pageNumber="478">T. bituberculatus</emphasis>
|
||
</taxonomicName>
|
||
is oval and wider, and the protibia has no fourth basal tooth.
|
||
</paragraph>
|
||
<paragraph id="111B143FFF85FF9CFF16471515F0F821" blockId="27.[151,1437,1282,1991]" pageId="27" pageNumber="478">
|
||
<emphasis id="23D0C82DFF85FF9CFF16471517CEFA85" bold="true" box="[199,407,1354,1379]" pageId="27" pageNumber="478">Locality records.</emphasis>
|
||
(
|
||
<figureCitation id="899F08BAFF85FF9CFE7D4715144BFA85" box="[428,530,1354,1379]" captionStart="FIGURES 94 – 101" captionStartId="27.[151,264,1159,1182]" captionTargetBox="[162,1433,376,1132]" captionTargetId="figure@27.[151,1436,373,1138]" captionTargetPageId="27" captionText="FIGURES 94 – 101. Tomarus bituberculatus. 94) habitus in dorsal view, 95 – 96) variations in shape pronotal fovea, 97) habitus in lateral view, 98) parameres in frontal view, 99) parameres in lateral view, 100) spiculum gastrale, 101) locality records in Colombia." httpUri="https://zenodo.org/record/234036/files/figure.png" pageId="27" pageNumber="478">Fig. 101</figureCitation>
|
||
)
|
||
<specimenCount id="07A2DFB6FF85FF9CFDF64715148FFA85" box="[551,726,1354,1379]" pageId="27" pageNumber="478" type="generic">216 specimens</specimenCount>
|
||
,
|
||
<specimenCount id="07A2DFB6FF85FF9CFD3647141570FA85" box="[743,809,1355,1379]" pageId="27" pageNumber="478" type="generic">104♂</specimenCount>
|
||
, 112♀. Specimens were seen from ANDES, BCRC, CEUA, CEUN, CIUQ, CTI, HJG, IAvH, ICN, LEUC, MEFLG, MLS, MPUJ, UNAB, UNSM, UPN.
|
||
<emphasis id="23D0C82DFF85FF9CFACE47311696FA4D" bold="true" pageId="27" pageNumber="478">Amazonas (17):</emphasis>
|
||
Leticia (15). Leticia,
|
||
<collectingCountry id="69B354AFFF85FF9CFE1447CD141DFA4D" box="[453,580,1426,1451]" name="Macedonia" pageId="27" pageNumber="478">Macedonia</collectingCountry>
|
||
, Resguardo indígena Ticuna (1). Tarapacá, Río Tacana (1).
|
||
<emphasis id="23D0C82DFF85FF9CFB3547CD13C5FA4D" bold="true" box="[1252,1436,1426,1451]" pageId="27" pageNumber="478">Antioquia (12):</emphasis>
|
||
Apartadó (2). El Carmen de Viboral (1). Medellín (4). Puerto Berrío (4).
|
||
<collectingCountry id="69B354AFFF85FF9CFC1D47E8123DFA29" box="[972,1124,1462,1487]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="27" pageNumber="478">Santa Helena</collectingCountry>
|
||
, Vda. El Mazo (1).
|
||
<emphasis id="23D0C82DFF85FF9CFA9247E91699FA15" bold="true" pageId="27" pageNumber="478">Arauca (3):</emphasis>
|
||
Alred. Río Arauca (3).
|
||
<emphasis id="23D0C82DFF85FF9CFE0347851402FA15" bold="true" box="[466,603,1498,1523]" pageId="27" pageNumber="478">Boyacá (2):</emphasis>
|
||
Villa de Leyva, SFF Iguaque (2).
|
||
<emphasis id="23D0C82DFF85FF9CFC3547851230FA15" bold="true" box="[996,1129,1498,1523]" pageId="27" pageNumber="478">Caldas (6):</emphasis>
|
||
La Dorada (2). Manizales,
|
||
<collectingCountry id="69B354AFFF85FF9CFF4647A116BAF9F1" box="[151,227,1534,1559]" name="Cyprus" pageId="27" pageNumber="478">Chipre</collectingCountry>
|
||
(1). Manizales, Vda. Hoyo Frío, Reserva Planalto CENICAFÉ (1). Manizales, Vda. Maltería, Granja Tesorito (1).
|
||
<collectingCountry id="69B354AFFF85FF9CFEF9447D17C9F9DD" box="[296,400,1570,1595]" name="Palestine" pageId="27" pageNumber="478">Palestina</collectingCountry>
|
||
, Vda. Santágueda, Granja Montelindo, Valle de Santagueda (1).
|
||
<emphasis id="23D0C82DFF85FF9CFBB7447D12A5F9DD" bold="true" box="[1126,1276,1570,1595]" pageId="27" pageNumber="478">Caquetá (2):</emphasis>
|
||
Florencia (2).
|
||
<emphasis id="23D0C82DFF85FF9CFF464419171EF9B9" bold="true" box="[151,327,1606,1631]" pageId="27" pageNumber="478">Casanare (20):</emphasis>
|
||
Tauramena, Vda. Villa Rosa, Fca. Barcelona (2). Yopal, Agua Azul, Hacienda Los Rosales (2). W. Agua Azul 1 KM (2). Yopal (14).
|
||
<emphasis id="23D0C82DFF85FF9CFDF14435154DF965" bold="true" box="[544,788,1642,1667]" pageId="27" pageNumber="478">Cundinamarca (12):</emphasis>
|
||
Bogotá (3). Fusagasugá, Cuenca Río
|
||
<collectingCountry id="69B354AFFF85FF9CFB1744351348F965" box="[1222,1297,1642,1667]" name="Jordan" pageId="27" pageNumber="478">Jordán</collectingCountry>
|
||
(1). Guasca (4). La Calera, Vda. La Jagada (1). La Mesa, Vda. Acacías (2). Tocaima (1).
|
||
<emphasis id="23D0C82DFF85FF9CFC2144D112C9F941" bold="true" box="[1008,1168,1678,1703]" pageId="27" pageNumber="478">Guaviare (2):</emphasis>
|
||
Caño Mocuare (1). San José del Guaviare, Fca. La Floresta (1).
|
||
<emphasis id="23D0C82DFF85FF9CFD8644ED1492F92D" bold="true" box="[599,715,1714,1739]" pageId="27" pageNumber="478">Huila (2):</emphasis>
|
||
San Agustín (2).
|
||
<emphasis id="23D0C82DFF85FF9CFC4344ED1210F92D" bold="true" box="[914,1097,1714,1739]" pageId="27" pageNumber="478">Magdalena (1):</emphasis>
|
||
Santa Ana (1).
|
||
<emphasis id="23D0C82DFF85FF9CFB2B44ED132EF92D" bold="true" box="[1274,1399,1714,1739]" pageId="27" pageNumber="478">Meta (15):</emphasis>
|
||
La Macarena (1). Puerto López (4). Puerto López, Vda. Menegua (1). Puerto López, Colegio INSTIVAL (2). Puerto Gaitán (1). Puerto Gaitán, Altamira, Los Llaneros (2). Villavicencio (1); Correg. Remolinos, Cafam Llanos (3).
|
||
<emphasis id="23D0C82DFF85FF9CFF464541171CF8D1" bold="true" box="[151,325,1822,1847]" pageId="27" pageNumber="478">Putumayo (1):</emphasis>
|
||
Puerto Asís (1).
|
||
<emphasis id="23D0C82DFF85FF9CFDDD454114EDF8D1" bold="true" box="[524,692,1822,1847]" pageId="27" pageNumber="478">Risaralda (1):</emphasis>
|
||
Pereira (1).
|
||
<emphasis id="23D0C82DFF85FF9CFC97454115AFF8D1" bold="true" box="[838,1014,1822,1847]" pageId="27" pageNumber="478">Santander (5):</emphasis>
|
||
Carare-Opón (1). Cimitarra, Correg. Puerto Olaya, Central Termocentro Isagen (3). Vélez, Campo Capote (1).
|
||
<emphasis id="23D0C82DFF85FF9CFC0E451D122BF8BD" bold="true" box="[991,1138,1858,1883]" pageId="27" pageNumber="478">Tolima (46):</emphasis>
|
||
Honda (45). Valencia (1).
|
||
<emphasis id="23D0C82DFF85FF9CFF46453917CDF899" bold="true" box="[151,404,1894,1919]" pageId="27" pageNumber="478">Valle del Cauca (26):</emphasis>
|
||
Buga (1). Bugalagrande (1). Cali (1). Calipuerto (5). “Mares”, Cordillera Occidental (12). Palmira (5). Roldanillo (1).
|
||
<emphasis id="23D0C82DFF85FF9CFE0445D51430F845" bold="true" box="[469,617,1930,1955]" pageId="27" pageNumber="478">Vichada (6):</emphasis>
|
||
Puerto Carreño, Fca. El Toro (1). Puerto Carreño, Barrio El Centro (4). Santa Rosalia (1).
|
||
<emphasis id="23D0C82DFF85FF9CFEB445F11430F821" bold="true" box="[357,617,1966,1991]" pageId="27" pageNumber="478">Ambiguous data (36):</emphasis>
|
||
<collectingCountry id="69B354AFFF85FF9CFDA145F11487F821" box="[624,734,1966,1991]" name="Colombia" pageId="27" pageNumber="478">Colombia</collectingCountry>
|
||
(4). No data (32).
|
||
</paragraph>
|
||
<paragraph id="111B143FFF82FF9BFF1642C8124FFF32" blockId="28.[151,1436,151,429]" pageId="28" pageNumber="479">
|
||
<emphasis id="23D0C82DFF82FF9BFF1642C8178FFF56" bold="true" box="[199,470,151,176]" pageId="28" pageNumber="479">Temporal distribution.</emphasis>
|
||
January (6), February (6), March (21), April (18), May (8), June (6), July (13), August (5), September (10), October (14), November (18), December (4). No data (87).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59BE47B4FF82FF9BFF1642801580FE4A" pageId="28" pageNumber="479" type="distribution">
|
||
<paragraph id="111B143FFF82FF9BFF16428017DAFE82" blockId="28.[151,1436,151,429]" pageId="28" pageNumber="479">
|
||
<emphasis id="23D0C82DFF82FF9BFF1642801739FF1E" bold="true" box="[199,352,223,248]" pageId="28" pageNumber="479">Distribution.</emphasis>
|
||
<collectingCountry id="69B354AFFF82FF9BFEA142801791FF1E" box="[368,456,223,248]" name="Mexico" pageId="28" pageNumber="479">Mexico</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFE0C42801400FF1E" box="[477,601,223,248]" name="Guatemala" pageId="28" pageNumber="479">Guatemala</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFDBC428014A2FF1E" box="[621,763,223,248]" name="El Salvador" pageId="28" pageNumber="479">El Salvador</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFCDE42801526FF1E" box="[783,895,223,248]" name="Honduras" pageId="28" pageNumber="479">Honduras</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFC4242801250FF1E" box="[915,1033,223,248]" name="Nicaragua" pageId="28" pageNumber="479">Nicaragua</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFBCC42BF12FAFF1E" box="[1053,1187,223,248]" name="Costa Rica" pageId="28" pageNumber="479">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFB6642BF134AFF11" box="[1207,1299,224,247]" name="Panama" pageId="28" pageNumber="479">Panama</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFAF6428013C1FF1E" box="[1319,1432,223,248]" name="Colombia" pageId="28" pageNumber="479">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFF46435B1754FEFB" box="[151,269,260,285]" name="Venezuela" pageId="28" pageNumber="479">Venezuela</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFEC8435B1722FEFB" box="[281,379,260,285]" name="Trinidad and Tobago" pageId="28" pageNumber="479">Trinidad</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFE59435B17BEFEFB" box="[392,487,260,285]" name="Ecuador" pageId="28" pageNumber="479">Ecuador</collectingCountry>
|
||
,
|
||
<collectingCountry id="69B354AFFF82FF9BFE25435A1473FEFA" box="[500,554,261,284]" name="Peru" pageId="28" pageNumber="479">Peru</collectingCountry>
|
||
, and
|
||
<collectingCountry id="69B354AFFF82FF9BFDB6435B14F2FEFB" box="[615,683,260,285]" name="Brazil" pageId="28" pageNumber="479">Brazil</collectingCountry>
|
||
(
|
||
<bibRefCitation id="753569CEFF82FF9BFD6D435B1502FEFB" author="Endrodi" box="[700,859,260,285]" pageId="28" pageNumber="479" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen Museum fur Tierkunde, Dresden, 37, 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="753569CEFF82FF9BFCB9435B1236FEFB" author="Ratcliffe" box="[872,1135,260,285]" pageId="28" pageNumber="479" refString="Ratcliffe, B. C. & Cave, R. D. (2006) The Dynastinae Scarab Beetles of Honduras, Nicaragua and El Salvador. Bulletin of the University of Nebraska State Museum, 21, 1 - 424." type="journal article" year="2006">Ratcliffe & Cave 2006</bibRefCitation>
|
||
). In
|
||
<collectingCountry id="69B354AFFF82FF9BFB72435B134DFEFB" box="[1187,1300,260,285]" name="Colombia" pageId="28" pageNumber="479">Colombia</collectingCountry>
|
||
, the species has been found also in Chocó (
|
||
<bibRefCitation id="753569CEFF82FF9BFDD0437814FAFEA6" author="Endrodi" box="[513,675,295,320]" pageId="28" pageNumber="479" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen Museum fur Tierkunde, Dresden, 37, 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
|
||
), Cauca (
|
||
<bibRefCitation id="753569CEFF82FF9BFCC04378121CFEA6" author="Pardo-Locarno" box="[785,1093,295,320]" pageId="28" pageNumber="479" refString="Pardo-Locarno, L. C., Montoya, J. & Schoonhoven, A. (2005) Abundancia de chisas rizofagas (Coleoptera: Melolonthidae) en agroecosistemas de Caldono y Buenos Aires, Cauca, Colombia. Revista Colombiana de Entomologia, 29, 177 - 184." type="journal article" year="2005">
|
||
Pardo-Locarno
|
||
<emphasis id="23D0C82DFF82FF9BFC15437615A0FEA6" box="[964,1017,295,320]" italics="true" pageId="28" pageNumber="479">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
), Cesar, and Córdoba (Pardo- Locarno
|
||
<emphasis id="23D0C82DFF82FF9BFF2A43121772FE83" box="[251,299,332,357]" italics="true" pageId="28" pageNumber="479">et al</emphasis>
|
||
. 2012).
|
||
</paragraph>
|
||
<paragraph id="111B143FFF82FF9BFF1643301580FE4A" blockId="28.[151,1436,151,429]" pageId="28" pageNumber="479">
|
||
<emphasis id="23D0C82DFF82FF9BFF16433017D5FE6E" bold="true" box="[199,396,367,392]" pageId="28" pageNumber="479">Natural history.</emphasis>
|
||
Specimens were collected with light traps, including black light, in different habitats including premontane and gallery forests. They were taken at
|
||
<quantity id="D65CB9DAFF82FF9BFC9D43CB158CFE4A" box="[844,981,404,429]" metricMagnitude="3" metricUnit="m" metricValue="1.415" metricValueMax="2.8" metricValueMin="0.03" pageId="28" pageNumber="479" unit="m" value="1415.0" valueMax="2800.0" valueMin="30.0">
|
||
30
|
||
<emphasis id="23D0C82DFF82FF9BFCB943CB152FFE4B" bold="true" box="[872,886,404,429]" pageId="28" pageNumber="479">–</emphasis>
|
||
2,800 m
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |