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<document id="ED292BE838D5B324B76AB8A284C9B2F2" ID-CLB-Dataset="22156" ID-DOI="10.11646/zootaxa.4778.2.4" ID-GBIF-Dataset="64190d5b-0175-4254-8d8f-c3657b2aceaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3856586" ID-ZooBank="5ACCAA9D-0C9B-4435-B665-15A0C79E2D6E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589444738901" checkinUser="plazi" docAuthor="Lan, Tianqi, Yao, Zhiyuan, Zheng, Guo, Wongprom, Prasit &amp; Li, Shuqiang" docDate="2020" docId="971C0A2AFFD2F470FF2AE4CBFCAFB5CA" docLanguage="en" docName="zootaxa.4778.2.4.pdf" docOrigin="Zootaxa 4778 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Savarna chiangmai Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="318" masterDocId="6B257252FFD6F477FFBDE71EFFF5B705" masterDocTitle="Five new species of the spider genus Savarna Huber, 2005 (Araneae: Pholcidae) from Thailand" masterLastPageNumber="328" masterPageNumber="311" pageNumber="315" updateTime="1698839606252" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C95AA931ADD09698FF2AF5C1D80DCC8B">Five new species of the spider genus Savarna Huber, 2005 (Araneae: Pholcidae) from Thailand</mods:title>
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<mods:name id="B7795C81F5D8DCDB39B05C6BECCB1F93" type="personal">
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<mods:namePart id="8FA212CE10E79BA98AEE35A82F16F269">Lan, Tianqi</mods:namePart>
<mods:affiliation id="77C6CB2CE59E64412539DD04593E1FDC">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; ltq 0217 @ outlook. com; https: // orcid. org / 0000 - 0002 - 9491 - 9813</mods:affiliation>
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<mods:namePart id="6629134F93D7A5CD79B5F54865E33822">Yao, Zhiyuan</mods:namePart>
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<mods:namePart id="4A2577F5DF10B2CD3D8F6E8247CDC5DA">Zheng, Guo</mods:namePart>
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<mods:namePart id="357C4E9E20175589CC682661C16BE3D5">Wongprom, Prasit</mods:namePart>
<mods:affiliation id="5AD9202F24E25DB5BB9E4A076CFC65A7">Faculty of Forestry, Kasetsart University, Ladyao, Jatujak, Bangkok 10903, Thailand</mods:affiliation>
</mods:name>
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<mods:namePart id="50A63D9EEFDBFB778ECDF85430D8B186">Li, Shuqiang</mods:namePart>
<mods:affiliation id="0BDD6503E9A7739A6225DFBD85D98F08">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title id="9EF626E6D5796A4D8B714C4028B9CC1E">Zootaxa</mods:title>
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<mods:date id="4EEA89EF2FDD1040C956B9580E027983">2020</mods:date>
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<mods:number id="FA2143B6091B6CC09351B474BF4FDD2E">4778</mods:number>
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<treatment id="971C0A2AFFD2F470FF2AE4CBFCAFB5CA" ID-DOI="http://doi.org/10.5281/zenodo.3856758" ID-GBIF-Taxon="164133560" ID-Zenodo-Dep="3856758" LSID="urn:lsid:plazi:treatment:971C0A2AFFD2F470FF2AE4CBFCAFB5CA" httpUri="http://treatment.plazi.org/id/971C0A2AFFD2F470FF2AE4CBFCAFB5CA" lastPageId="7" lastPageNumber="318" pageId="4" pageNumber="315">
<subSubSection id="57AFE8B7FFD2F473FF2AE4CBFD96B4EA" box="[151,611,981,1008]" pageId="4" pageNumber="315" type="nomenclature">
<paragraph id="1F0ABB3CFFD2F473FF2AE4CBFD96B4EA" blockId="4.[151,611,981,1043]" box="[151,611,981,1008]" pageId="4" pageNumber="315">
<heading id="44420C50FFD2F473FF2AE4CBFD96B4EA" bold="true" box="[151,611,981,1008]" fontSize="11" level="1" pageId="4" pageNumber="315" reason="1">
<emphasis id="2DC1672EFFD2F473FF2AE4CBFD96B4EA" bold="true" box="[151,611,981,1008]" pageId="4" pageNumber="315">
<taxonomicName id="D8B5C0BFFFD2F473FF2AE4CBFE0BB4EA" ID-CoL="948Q9" authority="Yao &amp; Li" authorityName="Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li" authorityYear="2020" box="[151,510,981,1008]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="315" phylum="Arthropoda" rank="species" species="chiangmai" status="sp. nov.">
<emphasis id="2DC1672EFFD2F473FF2AE4CBFE70B4F5" bold="true" box="[151,389,981,1008]" italics="true" pageId="4" pageNumber="315">Savarna chiangmai</emphasis>
Yao &amp; Li
</taxonomicName>
<taxonomicNameLabel id="36F2DA55FFD2F473FDB8E4CBFD96B4EA" box="[517,611,981,1007]" pageId="4" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="57AFE8B7FFD2F473FF2AE4E7FF0DB316" box="[151,248,1017,1043]" pageId="4" pageNumber="315" type="description">
<paragraph id="1F0ABB3CFFD2F473FF2AE4E7FF0DB316" blockId="4.[151,611,981,1043]" box="[151,248,1017,1043]" pageId="4" pageNumber="315">
<figureCitation id="878EA7B9FFD2F473FF2AE4E7FF0DB316" box="[151,248,1017,1043]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1899,1924]" captionStartId-1="6.[151,250,1902,1927]" captionTargetBox-0="[226,1369,181,1864]" captionTargetBox-1="[217,1370,181,1874]" captionTargetId-0="figure@5.[218,1369,181,1873]" captionTargetId-1="figure@6.[217,1370,181,1876]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.3856592" figureDoi-1="http://doi.org/10.5281/zenodo.3856594" httpUri-0="https://zenodo.org/record/3856592/files/figure.png" httpUri-1="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Figs 34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="57AFE8B7FFD2F473FF2AE35FFC6AB3A6" pageId="4" pageNumber="315" type="materials_examined">
<paragraph id="1F0ABB3CFFD2F473FF2AE35FFC6AB3A6" blockId="4.[151,1437,1088,2015]" pageId="4" pageNumber="315">
<emphasis id="2DC1672EFFD2F473FF2AE35FFEB7B35E" bold="true" box="[151,322,1088,1115]" pageId="4" pageNumber="315">Type material.</emphasis>
<materialsCitation id="AFDDB161FFD2F473FEFBE35FFC06B37A" ID-GBIF-Occurrence="2618695308" collectingDate="2014-06-27" collectorName="P. Wongprom" country="Thailand" elevation="662" latitude="19.567465" location="Tham Mongkhon Wissuti Taeb Cave" longLatPrecision="1" longitude="99.16848" pageId="4" pageNumber="315" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="2DC1672EFFD2F473FEFBE35FFE49B35E" bold="true" box="[326,444,1089,1115]" pageId="4" pageNumber="315">
<typeStatus id="C00E059EFFD2F473FEFBE35FFE42B35E" box="[326,439,1089,1115]" pageId="4" pageNumber="315" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="09B370B5FFD2F473FE7DE35CFE20B359" box="[448,469,1090,1116]" pageId="4" pageNumber="315" type="male"></specimenCount>
(IZCAS-Ar39730),
<location id="1A6AEDE7FFD2F473FD09E35FFBB1B35E" LSID="urn:lsid:plazi:treatment:971C0A2AFFD2F470FF2AE4CBFCAFB5CA:1A6AEDE7FFD2F473FD09E35FFBB1B35E" box="[692,1092,1089,1115]" country="Thailand" latitude="19.567465" longLatPrecision="1" longitude="99.16848" name="Tham Mongkhon Wissuti Taeb Cave" pageId="4" pageNumber="315">Tham Mongkhon Wissuti Taeb Cave</location>
(
<geoCoordinate id="7A81DDFBFFD2F473FBEDE35FFB07B35E" box="[1104,1266,1089,1115]" degrees="19" direction="north" minutes="34" orientation="latitude" pageId="4" pageNumber="315" precision="1" seconds="2.87" value="19.567465">19°342.87”N</geoCoordinate>
,
<geoCoordinate id="7A81DDFBFFD2F473FB47E35EFA62B35E" box="[1274,1431,1088,1115]" degrees="99" direction="east" minutes="10" orientation="longitude" pageId="4" pageNumber="315" precision="1" seconds="6.53" value="99.16848">99°106.53”E</geoCoordinate>
,
<elevation id="94985C0FFFD2F473FF2AE37BFEA6B37A" box="[151,339,1125,1151]" metricMagnitude="2" metricUnit="m" metricValue="6.62" pageId="4" pageNumber="315" unit="m" value="662.0">
elevation
<quantity id="D84D16D9FFD2F473FEBAE37BFEA6B37A" box="[263,339,1125,1151]" metricMagnitude="2" metricUnit="m" metricValue="6.62" pageId="4" pageNumber="315" unit="m" value="662.0">662 m</quantity>
</elevation>
),
<emphasis id="2DC1672EFFD2F473FED7E37AFE1BB37A" bold="true" box="[362,494,1124,1151]" pageId="4" pageNumber="315">Chiangmai</emphasis>
,
<emphasis id="2DC1672EFFD2F473FE43E37AFD9EB37A" bold="true" box="[510,619,1124,1151]" pageId="4" pageNumber="315">
<collectingCountry id="67A2FBACFFD2F473FE43E37AFD9EB37A" box="[510,619,1124,1151]" name="Thailand" pageId="4" pageNumber="315">Thailand</collectingCountry>
</emphasis>
,
<date id="6B0B9DFCFFD2F473FDC1E37BFCECB37A" box="[636,793,1125,1151]" pageId="4" pageNumber="315" value="2014-06-27">
<collectingDate id="7B4F6414FFD2F473FDC1E37BFCECB37A" box="[636,793,1125,1151]" pageId="4" pageNumber="315" value="2014-06-27">27 June 2014</collectingDate>
</date>
,
<collectorName id="B240DEEAFFD2F473FC9BE37BFC4AB37A" box="[806,959,1125,1151]" pageId="4" pageNumber="315">P. Wongprom</collectorName>
leg.
</materialsCitation>
<emphasis id="2DC1672EFFD2F473FC46E37BFB88B37A" bold="true" box="[1019,1149,1125,1151]" pageId="4" pageNumber="315">
<typeStatus id="C00E059EFFD2F473FC46E37BFB82B37A" box="[1019,1143,1125,1151]" pageId="4" pageNumber="315" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="09B370B5FFD2F473FB3BE37BFB4BB37A" box="[1158,1214,1125,1151]" pageId="4" pageNumber="315" type="male">2♂♂</specimenCount>
(IZCAS-Ar39731 Ar39732),
<materialsCitation id="AFDDB161FFD2F473FEAFE397FC69B3A6" ID-GBIF-Occurrence="2618695305" box="[274,924,1161,1187]" collectingDate="2014-06-27" collectorName="P. Wongprom" country="Thailand" elevation="662" latitude="19.567465" location="Tham Mongkhon Wissuti Taeb Cave" longLatPrecision="1" longitude="99.16848" pageId="4" pageNumber="315" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="09B370B5FFD2F473FEAFE397FEBFB3A6" box="[274,330,1161,1187]" pageId="4" pageNumber="315" type="female">2♀♀</specimenCount>
(IZCAS-Ar39733Ar39734), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="57AFE8B7FFD2F473FF7AE3B3FB8EB3C2" box="[199,1147,1197,1223]" pageId="4" pageNumber="315" type="etymology">
<paragraph id="1F0ABB3CFFD2F473FF7AE3B3FB8EB3C2" blockId="4.[151,1437,1088,2015]" box="[199,1147,1197,1223]" pageId="4" pageNumber="315">
<emphasis id="2DC1672EFFD2F473FF7AE3B3FEBBB3C2" bold="true" box="[199,334,1197,1223]" pageId="4" pageNumber="315">Etymology.</emphasis>
The specific name is a noun in apposition and refers to the
<typeStatus id="C00E059EFFD2F473FC57E3B3FBEFB3C2" box="[1002,1050,1197,1223]" pageId="4" pageNumber="315">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="57AFE8B7FFD2F473FF7AE3CEFAC3B2C6" pageId="4" pageNumber="315" type="diagnosis">
<paragraph id="1F0ABB3CFFD2F473FF7AE3CEFAC3B2C6" blockId="4.[151,1437,1088,2015]" pageId="4" pageNumber="315">
<emphasis id="2DC1672EFFD2F473FF7AE3CEFEB7B3EE" bold="true" box="[199,322,1232,1259]" pageId="4" pageNumber="315">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="D8B5C0BFFFD2F473FDE8E3CFFD20B3EE" authority="Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li, 2020" authorityName="Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li" authorityYear="2020" box="[597,725,1233,1259]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="315" phylum="Arthropoda" rank="species" species="bannang" status="sp. nov.">
<emphasis id="2DC1672EFFD2F473FDE8E3CFFD20B3EE" box="[597,725,1233,1259]" italics="true" pageId="4" pageNumber="315">S. bannang</emphasis>
</taxonomicName>
<emphasis id="2DC1672EFFD2F473FD60E3CFFCCDB3EE" bold="true" box="[733,824,1233,1259]" pageId="4" pageNumber="315">
<taxonomicNameLabel id="36F2DA55FFD2F473FD60E3CFFCCDB3EE" box="[733,824,1233,1259]" pageId="4" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="878EA7B9FFD2F473FCF5E3CFFC58B3EE" box="[840,941,1233,1259]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1897,1922]" captionStartId-1="3.[151,250,1900,1925]" captionTargetBox-0="[217,1370,181,1867]" captionTargetBox-1="[218,1370,181,1874]" captionTargetId-0="figure@2.[217,1371,176,1872]" captionTargetId-1="figure@3.[218,1370,181,1875]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Savarna bannang sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at long, spine-shaped subdistal sclerite dorsally, arrow 2 points at curved distal apophysis, arrows 3 and 4 point at two large, distal membranous processes respectively; D. Retrolateral view). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 2. Savarna bannang sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.3856588" figureDoi-1="http://doi.org/10.5281/zenodo.3856590" httpUri-0="https://zenodo.org/record/3856588/files/figure.png" httpUri-1="https://zenodo.org/record/3856590/files/figure.png" pageId="4" pageNumber="315">Figs 12</figureCitation>
) with similar embolus (
<figureCitation id="878EA7B9FFD2F473FB03E3CFFAE2B3EE" box="[1214,1303,1233,1259]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4C</figureCitation>
), male chelicerae (
<figureCitation id="878EA7B9FFD2F473FF4EE3EBFEB8B20A" box="[243,333,1269,1295]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4D</figureCitation>
) and epigynum (
<figureCitation id="878EA7B9FFD2F473FDACE3EBFD9FB20A" box="[529,618,1269,1295]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4A</figureCitation>
) and
<taxonomicName id="D8B5C0BFFFD2F473FD10E3EBFB15B20A" authority="Wongprom &amp; Wiwatwitaya, 2015" authorityName="Wongprom &amp; Wiwatwitaya" authorityYear="2015" box="[685,1248,1268,1295]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="315" phylum="Arthropoda" rank="species" species="kraburiensis">
<emphasis id="2DC1672EFFD2F473FD10E3EBFCA2B20A" box="[685,855,1269,1295]" italics="true" pageId="4" pageNumber="315">S. kraburiensis</emphasis>
<bibRefCitation id="7B24C6CDFFD2F473FCDDE3EBFB15B20A" author="Wongprom, P. &amp; Wiwatwitaya, D." box="[864,1248,1268,1295]" pageId="4" pageNumber="315" pagination="1 - 5" refId="ref8024" refString="Wongprom, P. &amp; Wiwatwitaya, D. (2015) One new species of the genus Savarna Huber, 2005 (Araneae, Pholcidae) from southern Thailand. ZooKeys, 498, 1 - 5. https: // doi. org / 10.3897 / zookeys. 498.9269" type="journal article" year="2015">Wongprom &amp; Wiwatwitaya, 2015</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="7B24C6CDFFD2F473FAA2E3EBFE5CB236" author="Wongprom, P. &amp; Wiwatwitaya, D." pageId="4" pageNumber="315" pagination="1 - 5" refId="ref8024" refString="Wongprom, P. &amp; Wiwatwitaya, D. (2015) One new species of the genus Savarna Huber, 2005 (Araneae, Pholcidae) from southern Thailand. ZooKeys, 498, 1 - 5. https: // doi. org / 10.3897 / zookeys. 498.9269" type="journal article" year="2015">Wongprom &amp; Wiwatwitaya 2015: 2</bibRefCitation>
, figs 12) with similar male chelicerae (
<figureCitation id="878EA7B9FFD2F473FCCEE207FC39B236" box="[883,972,1305,1331]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4D</figureCitation>
) but can be distinguished by presence of angular subdistal sclerite dorsally (arrow
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<figureCitation id="878EA7B9FFD2F473FD20E223FD06B252" box="[669,755,1341,1367]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Fig. 3C</figureCitation>
), short distal spine prolaterally (arrow
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<figureCitation id="878EA7B9FFD2F473FB60E223FAC7B252" box="[1245,1330,1341,1367]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Fig. 3C</figureCitation>
) and two small, distal membranous processes (arrows 1 and
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<figureCitation id="878EA7B9FFD2F473FCB4E27FFC95B27E" box="[777,864,1377,1403]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Fig. 3D</figureCitation>
) on procursus and by presence of wavy vulval anterior arch and long elliptic vulval pore plates (
<figureCitation id="878EA7B9FFD2F473FD15E29BFCF5B29A" box="[680,768,1413,1439]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4B</figureCitation>
); also distinguished from
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<emphasis id="2DC1672EFFD2F473FB99E29BFB39B29A" box="[1060,1228,1413,1439]" italics="true" pageId="4" pageNumber="315">S. kraburiensis</emphasis>
</taxonomicName>
by short and wide embolus (
<figureCitation id="878EA7B9FFD2F473FEB8E2B7FEA8B2C6" box="[261,349,1449,1475]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4C</figureCitation>
), by slightly protruding epigynum (
<figureCitation id="878EA7B9FFD2F473FD51E2B7FCB1B2C6" box="[748,836,1449,1475]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4A</figureCitation>
) and by elliptic vulval pore plates (
<figureCitation id="878EA7B9FFD2F473FB6CE2B7FADCB2C6" box="[1233,1321,1449,1475]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="57AFE8B7FFD2F470FF7AE2D2FF22B5AE" lastPageId="7" lastPageNumber="318" pageId="4" pageNumber="315" type="description">
<paragraph id="1F0ABB3CFFD2F470FF7AE2D2FD08B61A" blockId="4.[151,1437,1088,2015]" lastBlockId="7.[151,1437,153,719]" lastPageId="7" lastPageNumber="318" pageId="4" pageNumber="315">
<emphasis id="2DC1672EFFD2F473FF7AE2D2FE6AB2E2" bold="true" box="[199,415,1484,1511]" pageId="4" pageNumber="315">Description. Male</emphasis>
(
<emphasis id="2DC1672EFFD2F473FE0FE2D3FDECB2E2" bold="true" box="[434,537,1485,1511]" pageId="4" pageNumber="315">
<typeStatus id="C00E059EFFD2F473FE0FE2D3FDECB2E2" box="[434,537,1485,1511]" pageId="4" pageNumber="315" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS-Ar39730): Total length 2.49 (2.63 with clypeus), prosoma 0.90 long, 0.96 wide, opisthosoma 1.59 long, 1.08 wide. Leg I: 27.11 (7.02 + 0.41 + 6.58 + 10.54 + 2.56), leg II: 17.44 (4.99 + 0.38 + 4.17 + 6.67 + 1.23), leg III: 12.95 (3.83 + 0.35 + 3.14 + 4.72 + 0.91), leg IV: 14.21 (4.16 + 0.36 + 3.32 + 5.43 + 0.94); tibia I L/d: 59; leg formula: I-II-IV-III. Distance PME-PME 0.15, diameter PME 0.12, distance PME- ALE 0.05, AME absent. Sternum wider than long (0.74/0.56). Habitus as in
<figureCitation id="878EA7B9FFD2F473FC41E143FBB7B172" box="[1020,1090,1629,1655]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Figs 4</figureCitation>
EF. Dorsal shield of prosoma yellowish, with brown margins and wide, brown median stripe behind ocular area; ocular area brown, but yellowish medially; clypeus brown; sternum black. Legs brownish, but slightly whitish on distal parts of femora and tibiae, with darker rings on subdistal parts of femora and proximal, median and subdistal parts of tibiae. Opisthosoma yellowish, with black subcuticular spots lying above deeper white spots dorsally and laterally and distinct ventral pattern consisting of three interconnected black marks. Ocular area elevated, each eye triad on top of short, laterally directed hump. Thoracic furrow deep. Chelicerae as in
<figureCitation id="878EA7B9FFD2F473FCB3E02BFC9DB04A" box="[782,872,1845,1871]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4D</figureCitation>
, with pair of proximo-lateral apophyses. Palps as in
<figureCitation id="878EA7B9FFD2F473FF09E047FF02B076" box="[180,247,1881,1907]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Figs 3</figureCitation>
AB, trochanter with small dorsal apophysis (arrow in
<figureCitation id="878EA7B9FFD2F473FCDFE047FC4FB076" box="[866,954,1881,1907]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Fig. 3A</figureCitation>
) and long ventral apophysis proximally attached to femur (
<figureCitation id="878EA7B9FFD2F473FEEEE063FE5FB092" box="[339,426,1917,1943]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Fig. 3B</figureCitation>
); procursus simple proximally but complex distally, with angular subdistal sclerite dorsally (arrow
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), short distal spine prolaterally (arrow
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<figureCitation id="878EA7B9FFD2F473FCF1E0BFFC55B0BE" box="[844,928,1953,1979]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Fig. 3C</figureCitation>
) and two small, distal membranous processes (arrows 1 and
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<figureCitation id="878EA7B9FFD2F473FECEE0DBFE3BB0DA" box="[371,462,1989,2015]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[226,1369,181,1864]" captionTargetId="figure@5.[218,1369,181,1873]" captionTargetPageId="5" captionText="FIGURE 3. Savarna chiangmai sp. nov., holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="4" pageNumber="315">Fig. 3D</figureCitation>
); bulb (
<figureCitation id="878EA7B9FFD2F473FD9AE0DBFD8AB0DA" box="[551,639,1989,2015]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="4" pageNumber="315">Fig. 4C</figureCitation>
) with proximal sclerite and sclerotized embolus (provided with large, sclerotized distal apophysis, arrow
<quantity id="D84D16D9FFD1F470FD9FE787FDB8B7B6" box="[546,589,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="318" unit="in" value="1.0">1 in</quantity>
<figureCitation id="878EA7B9FFD1F470FDE9E787FD5FB7B6" box="[596,682,153,179]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="7" pageNumber="318">Fig. 4C</figureCitation>
) apparently containing sperm duct (sperm duct opening visible, arrow
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<figureCitation id="878EA7B9FFD1F470FF46E7A3FEA6B7D2" box="[251,339,189,215]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="7" pageNumber="318">Fig. 4C</figureCitation>
). Retrolateral trichobothrium of tibia I at 8% proximally; prolateral trichobothrium absent on tibia I, present on other tibiae; legs with vertical hairs in higher than usual density on all tibiae, without short spines and curved hairs; tarsus I with 19 distinct pseudosegments.
</paragraph>
<caption id="4BCAEBB4FFD3F472FF2AE075FB49B0EA" ID-DOI="http://doi.org/10.5281/zenodo.3856592" ID-Zenodo-Dep="3856592" httpUri="https://zenodo.org/record/3856592/files/figure.png" pageId="5" pageNumber="316" startId="5.[151,250,1899,1924]" targetBox="[226,1369,181,1864]" targetPageId="5">
<paragraph id="1F0ABB3CFFD3F472FF2AE075FB49B0EA" blockId="5.[151,1436,1899,2032]" pageId="5" pageNumber="316">
<emphasis id="2DC1672EFFD3F472FF2AE075FEE1B081" bold="true" box="[151,276,1899,1924]" pageId="5" pageNumber="316">FIGURE 3.</emphasis>
<taxonomicName id="D8B5C0BFFFD3F472FEA1E072FE17B086" authority="Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li, 2020" authorityName="Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li" authorityYear="2020" box="[284,482,1899,1923]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="chiangmai" status="sp. nov.">
<emphasis id="2DC1672EFFD3F472FEA1E072FE17B086" box="[284,482,1899,1923]" italics="true" pageId="5" pageNumber="316">Savarna chiangmai</emphasis>
</taxonomicName>
<emphasis id="2DC1672EFFD3F472FE57E075FDC8B086" bold="true" box="[490,573,1899,1923]" pageId="5" pageNumber="316">
<taxonomicNameLabel id="36F2DA55FFD3F472FE57E075FDC8B086" box="[490,573,1899,1923]" pageId="5" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Palp (A. Prolateral view, arrow points at dorsal apophysis; B. Retrolateral view); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at angular subdistal sclerite dorsally, arrow 2 points at short distal spine prolaterally; D. Retrolateral view, arrows 1 and 2 point at two small, distal membranous processes respectively). b = bulb, e = embolus, pr = procursus, tr = trochanter. Scale bars: 0.10 (AB), 0.05 (CD).
</paragraph>
</caption>
<caption id="4BCAEBB4FFD0F471FF2AE070FB8BB0F6" ID-DOI="http://doi.org/10.5281/zenodo.3856594" ID-Zenodo-Dep="3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="6" pageNumber="317" startId="6.[151,250,1902,1927]" targetBox="[217,1370,181,1874]" targetPageId="6">
<paragraph id="1F0ABB3CFFD0F471FF2AE070FB8BB0F6" blockId="6.[151,1436,1902,2035]" pageId="6" pageNumber="317">
<emphasis id="2DC1672EFFD0F471FF2AE070FEE6B082" bold="true" box="[151,275,1902,1927]" pageId="6" pageNumber="317">FIGURE 4.</emphasis>
<taxonomicName id="D8B5C0BFFFD0F471FEA7E071FE15B082" authority="Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li, 2020" authorityName="Lan &amp; Yao &amp; Zheng &amp; Wongprom &amp; Li" authorityYear="2020" box="[282,480,1903,1927]" class="Arachnida" family="Pholcidae" genus="Savarna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="317" phylum="Arthropoda" rank="species" species="chiangmai" status="sp. nov.">
<emphasis id="2DC1672EFFD0F471FEA7E071FE15B082" box="[282,480,1903,1927]" italics="true" pageId="6" pageNumber="317">Savarna chiangmai</emphasis>
</taxonomicName>
<emphasis id="2DC1672EFFD0F471FE5AE071FDCCB083" bold="true" box="[487,569,1903,1927]" pageId="6" pageNumber="317">
<taxonomicNameLabel id="36F2DA55FFD0F471FE5AE071FDCCB083" box="[487,569,1903,1927]" pageId="6" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH).
</paragraph>
</caption>
<paragraph id="1F0ABB3CFFD1F470FF7AE637FDABB6F2" blockId="7.[151,1437,153,719]" pageId="7" pageNumber="318">
<emphasis id="2DC1672EFFD1F470FF7AE637FEEBB646" bold="true" box="[199,286,297,323]" pageId="7" pageNumber="318">Female</emphasis>
(
<emphasis id="2DC1672EFFD1F470FE8DE637FE6EB646" bold="true" box="[304,411,297,323]" pageId="7" pageNumber="318">
<typeStatus id="C00E059EFFD1F470FE8DE637FE6EB646" box="[304,411,297,323]" pageId="7" pageNumber="318" type="paratype">paratype</typeStatus>
</emphasis>
, IZCAS-Ar39733): Similar to male, habitus as in
<figureCitation id="878EA7B9FFD1F470FC68E637FBEEB646" box="[981,1051,297,323]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="7" pageNumber="318">Figs 4</figureCitation>
GH. Total length 2.72 (2.84 with clypeus), prosoma 0.88 long, 0.88 wide, opisthosoma 1.84 long, 1.48 wide. Legs I and IV missing (see variation), leg II: 14.43 (4.16 + 0.35 + 3.44 + 5.23 + 1.25), leg III: 10.09 (3.11 + 0.31 + 2.34 + 3.52 + 0.81). Distance PME- PME 0.14, diameter PME 0.11, distance PME-ALE 0.04, AME absent. Sternum slightly wider than long (0.62/0.52). Ocular area black, but brown medially. Epigynum (
<figureCitation id="878EA7B9FFD1F470FD6FE6A7FCDDB6D6" box="[722,808,441,467]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="7" pageNumber="318">Fig. 4A</figureCitation>
) slightly protruding. Vulva (
<figureCitation id="878EA7B9FFD1F470FBDBE6A7FB49B6D6" box="[1126,1212,441,467]" captionStart="FIGURE 4" captionStartId="6.[151,250,1902,1927]" captionTargetBox="[217,1370,181,1874]" captionTargetId="figure@6.[217,1370,181,1876]" captionTargetPageId="6" captionText="FIGURE 4. Savarna chiangmai sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulbal process, prolateral view, arrow 1 points at large, sclerotized distal apophysis, arrow 2 points at sperm duct opening; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, e = embolus, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD), 0.50 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3856594" httpUri="https://zenodo.org/record/3856594/files/figure.png" pageId="7" pageNumber="318">Fig. 4B</figureCitation>
) with wavy anterior arch and pair of long elliptic pore plates.
</paragraph>
<paragraph id="1F0ABB3CFFD1F470FF7AE51EFDD5B566" blockId="7.[151,1437,153,719]" pageId="7" pageNumber="318">
<emphasis id="2DC1672EFFD1F470FF7AE51EFEB4B51E" bold="true" box="[199,321,512,539]" pageId="7" pageNumber="318">Variation:</emphasis>
Tibia I in one male
<typeStatus id="C00E059EFFD1F470FDA1E51FFD89B51E" box="[540,636,513,539]" pageId="7" pageNumber="318" type="paratype">paratype</typeStatus>
(leg I missing in another specimen): 6.01 (IZCAS-Ar39731). In another female
<typeStatus id="C00E059EFFD1F470FF56E53BFEBEB53A" box="[235,331,549,575]" pageId="7" pageNumber="318" type="paratype">paratype</typeStatus>
(IZCAS-Ar39734): Leg I: 22.23 (5.19 + 0.37 + 5.08 + 9.12 + 2.47), leg IV: 13.72 (4.14 + 0.33 + 3.31 + 5.10 + 0.84); tibia I L/d: 46.
</paragraph>
<paragraph id="1F0ABB3CFFD1F470FF7AE573FF22B5AE" blockId="7.[151,1437,153,719]" pageId="7" pageNumber="318">
<emphasis id="2DC1672EFFD1F470FF7AE573FE70B582" bold="true" box="[199,389,620,647]" pageId="7" pageNumber="318">Natural history.</emphasis>
The species was found in the twilight zone of the Tham Mongkhon Wissuti Taeb Cave (karst area).
</paragraph>
</subSubSection>
<subSubSection id="57AFE8B7FFD1F470FF7AE5AAFCAFB5CA" box="[199,858,692,719]" pageId="7" pageNumber="318" type="distribution">
<paragraph id="1F0ABB3CFFD1F470FF7AE5AAFCAFB5CA" blockId="7.[151,1437,153,719]" box="[199,858,692,719]" pageId="7" pageNumber="318">
<emphasis id="2DC1672EFFD1F470FF7AE5AAFE94B5CA" bold="true" box="[199,353,692,719]" pageId="7" pageNumber="318">Distribution.</emphasis>
Known only from the
<typeStatus id="C00E059EFFD1F470FDDDE5ABFD65B5CA" box="[608,656,693,719]" pageId="7" pageNumber="318">type</typeStatus>
locality (
<figureCitation id="878EA7B9FFD1F470FD46E5ABFCB9B5CA" box="[763,844,693,719]" captionStart="FIGURE 11" captionStartId="16.[151,250,1318,1343]" captionTargetBox="[151,1436,181,1293]" captionTargetId="figure@16.[151,1436,181,1293]" captionTargetPageId="16" captionText="FIGURE 11. Known distribution records of new Savarna species from Thailand. 1. Savarna bannang sp. nov.; 2. S. chiangmai sp. nov.; 3. S. huahin sp. nov.; 4. S. satun sp. nov.; 5. S. thungsong sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3856608" httpUri="https://zenodo.org/record/3856608/files/figure.png" pageId="7" pageNumber="318">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>