treatments-xml/data/69/F6/C0/69F6C070775DD62F8A4108041B2AAA77.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.193.2768" ID-GBIF-Dataset="ff21e5c7-e09d-4706-95db-77878b6c5031" ID-PMC="PMC3361139" ID-Pensoft-Pub="1313-2970-193-49" ID-PubMed="22679379" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-193-49" ModsDocOrigin="ZooKeys 193" ModsDocTitle="Review of the Southeast Asian species of the Aenictus javanus and Aenictus philippinensis species groups (Hymenoptera, Formicidae, Aenictinae)" checkinTime="1451249022177" checkinUser="pensoft" docAuthor="Jaitrong, Weeyawat &amp; Yamane, Seiki" docDate="2012" docId="69F6C070775DD62F8A4108041B2AAA77" docLanguage="en" docName="ZooKeys 193: 49-78" docOrigin="ZooKeys 193" docSource="http://dx.doi.org/10.3897/zookeys.193.2768" docTitle="Aenictus piercei Wheeler &amp; Chapman 1930" docType="treatment" docVersion="3" lastPageNumber="62" masterDocId="FFF3893EFF814C43FFE9B25BB50BFFA3" masterDocTitle="Review of the Southeast Asian species of the Aenictus javanus and Aenictus philippinensis species groups (Hymenoptera, Formicidae, Aenictinae)" masterLastPageNumber="78" masterPageNumber="49" pageNumber="61" updateTime="1668153898549" updateUser="ExternalLinkService">
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<mods:title>Review of the Southeast Asian species of the Aenictus javanus and Aenictus philippinensis species groups (Hymenoptera, Formicidae, Aenictinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jaitrong, Weeyawat</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yamane, Seiki</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2012</mods:date>
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<mods:number>193</mods:number>
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<mods:start>49</mods:start>
<mods:end>78</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.193.2768</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.193.2768</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-193-49</mods:identifier>
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<treatment ID-GBIF-Taxon="152035531" LSID="urn:lsid:plazi:treatment:69F6C070775DD62F8A4108041B2AAA77" httpUri="http://treatment.plazi.org/id/69F6C070775DD62F8A4108041B2AAA77" lastPageId="13" lastPageNumber="62" pageId="12" pageNumber="61">
<subSubSection pageId="12" pageNumber="61" type="nomenclature">
<paragraph pageId="12" pageNumber="61">
<taxonomicName LSID="http://species-id.net/wiki/Aenictus_piercei" authority="Wheeler &amp; Chapman, 1930" authorityName="Wheeler &amp; Chapman" authorityYear="1930" class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei Wheeler &amp; Chapman, 1930</taxonomicName>
Figs 67B
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="61" type="reference_group">
<paragraph pageId="12" pageNumber="61">
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
Wheeler &amp; Chapman, in 
<bibRefCitation author="Wheeler, WM" journalOrPublisher="Journal of the New York Entomological Society" pageId="25" pageNumber="74" pagination="193 - 212" title="Philippine ants of the genus Aenictus with descriptions of the females of two species." volume="38" year="1930">Wheeler 1930</bibRefCitation>
: 209, fig. 7 
<normalizedToken originalValue="e–g">e-g</normalizedToken>
; 
<bibRefCitation author="Wilson, EO" journalOrPublisher="Pacific Insects" pageId="25" pageNumber="74" pagination="427 - 483" title="The true army ants of the Indo-Australian area (Hymenoptera: Formicidae: Dorylinae)." volume="6" year="1964">Wilson 1964</bibRefCitation>
: 474, figs 61-62; 
<bibRefCitation author="Bolton, B" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" pageId="24" pageNumber="73" title="A New General Catalogue of the Ants of the World." year="1995">Bolton 1995</bibRefCitation>
: 60.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="61" type="types">
<paragraph pageId="12" pageNumber="61">Types.</paragraph>
<paragraph pageId="12" pageNumber="61">Two syntype workers on a pin, the Philippines, Negros, Cadiz, 2.VI.1924, leg. Dr. Pierce (MCZC, examined). The worker located below on the pin is selected as the lectotype (Fig. 6B).</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="61" type="measurements">
<paragraph pageId="12" pageNumber="61">Measurements.</paragraph>
<paragraph pageId="12" pageNumber="61">Lectotype: TL 2.15 mm; HL 0.53 mm; HW 0.48 mm; SL 0.28 mm; ML 0.70 mm; PL 0.20 mm; CI 90; SI 58.</paragraph>
<paragraph pageId="12" pageNumber="61">Paralectotypes (n = 2): TL 2.15 mm; HL 0.53-0.58 mm; HW 0.48-0.50 mm; SL 0.28-0.33 mm; ML 0.70-0.83 mm; PL 0.20-0.23 mm; CI 87-90; SI 58-65.</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="62" pageId="12" pageNumber="61" type="description">
<paragraph pageId="12" pageNumber="61">Description of worker</paragraph>
<paragraph lastPageId="13" lastPageNumber="62" pageId="12" pageNumber="61">
(lectotype, paralectotype and a non-type worker). Head in full-face view slightly longer than broad, subrectangular, with sides feebly convex and posterior margin almost straight; seen in profile occipital corner of head rounded. Antennal scape reaching midlength of head; antennal segment II longer and narrower than 
<pageBreakToken pageId="13" pageNumber="62" start="start">each</pageBreakToken>
of 
<normalizedToken originalValue="III–VI">III-VI</normalizedToken>
; terminal segment almost as long as VII+VIII+IX and 1.9 times as long as broad. Anterior margin of clypeus bearing 9-10 denticles (this observation is based on the single non-type worker, since in the lectotype mouth parts are buried in glue and the head of the paralectotype was missing). Masticatory margin of mandible with 3 acute teeth including large apical tooth; basal margin lacking denticles. Promesonotum in profile almost flat dorsally; in profile propodeum almost flat dorsally; propodeal junction angulate, right-angled; declivity of propodeum nearly flat, with blunt lateral carinae, but not demarcated basally by a transverse carina. Petiole almost as long as high, its dorsal outline convex; subpetiolar process well developed, subrectangular, its ventral margin slightly convex and longer than posterior margin; postpetiole almost as long as petiole.
</paragraph>
<paragraph pageId="13" pageNumber="62">Head including antennal scape entirely smooth and shiny. Mandible finely striate except along masticatory and outer margins. Pronotum entirely smooth and shiny except for anteriormost part microreticulate; mesonotum smooth and shiny; mesopleuron, metapleuron and propodeum microreticulate. Petiole entirely microreticulate. Postpetiole microreticulate except for a small smooth and shiny area on dorsal surface.</paragraph>
<paragraph pageId="13" pageNumber="62">Head and mesosoma dorsally with relatively sparse standing hairs mixed with sparse short hairs; longest pronotal hairs 0.09-0.10 mm long. Entire body yellowish brown or reddish brown; legs palest.</paragraph>
<caption pageId="13" pageNumber="62">
<paragraph pageId="13" pageNumber="62">
Figure 6. 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
(A, B, C, F, lectotype; D, E, non-type from the type locality). A Habitus in profile B lectotype and paralectotype designated in the present paper (arrow indicating the lectotype) C labels of lectotype D head in full-face view E mandible and anterior clypeal margin F propodeal declivity in dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="13" pageNumber="62" type="non-type material examined">
<paragraph pageId="13" pageNumber="62">Non-type material examined.</paragraph>
<paragraph pageId="13" pageNumber="62">
We examined a worker collected from the same place by Chapman but in a different year (2/6/29). It bears a small piece of white paper with handwriting 
<normalizedToken originalValue="“cotype”">&quot;cotype&quot;</normalizedToken>
, and a small piece of red paper. As this specimen was not mentioned in the original description, it is not part of the type series. However, all the three specimens belong to the same species without doubt.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="62" type="distribution">
<paragraph pageId="13" pageNumber="62">Distribution.</paragraph>
<paragraph pageId="13" pageNumber="62">Philippines (Negros and Mindanao) (Fig. 7B).</paragraph>
<caption pageId="13" pageNumber="62">
<paragraph pageId="13" pageNumber="62">
Figure 7. Distribution of the species of the 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus javanus" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="javanus">Aenictus javanus</taxonomicName>
group. A 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus doydeei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="doydeei">Aenictus doydeei</taxonomicName>
, 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus nishimurai" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="nishimurai">Aenictus nishimurai</taxonomicName>
, and 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus duengkaei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="duengkaei">Aenictus duengkaei</taxonomicName>
sp. n. B 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus javanus" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="javanus">Aenictus javanus</taxonomicName>
, 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus longinodus" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="longinodus">Aenictus longinodus</taxonomicName>
sp. n., and 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="13" pageNumber="62" type="bionomics">
<paragraph pageId="13" pageNumber="62">Bionomics.</paragraph>
<paragraph pageId="13" pageNumber="62">
Little is known about the bionomics of 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
. Nothing is mentioned by 
<bibRefCitation author="Wheeler, WM" journalOrPublisher="Journal of the New York Entomological Society" pageId="25" pageNumber="74" pagination="193 - 212" title="Philippine ants of the genus Aenictus with descriptions of the females of two species." volume="38" year="1930">Wheeler (1930)</bibRefCitation>
on it. However, judging from the hitherto known localities (Negros and Mindanao) (see 
<bibRefCitation author="Wilson, EO" journalOrPublisher="Pacific Insects" pageId="25" pageNumber="74" pagination="427 - 483" title="The true army ants of the Indo-Australian area (Hymenoptera: Formicidae: Dorylinae)." volume="6" year="1964">Wilson 1964</bibRefCitation>
) this species inhabits lowland (15-600 m) and is probably restricted to the Philippines. 
<bibRefCitation author="Wilson, EO" journalOrPublisher="Pacific Insects" pageId="25" pageNumber="74" pagination="427 - 483" title="The true army ants of the Indo-Australian area (Hymenoptera: Formicidae: Dorylinae)." volume="6" year="1964">Wilson (1964)</bibRefCitation>
cited India, Solon (ca. 1400 m), as a locality of 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
, but the identification should be reconfirmed.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="62" type="remarks">
<paragraph pageId="13" pageNumber="62">Remarks.</paragraph>
<paragraph pageId="13" pageNumber="62">
This species is most similar to 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus duengkaei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="duengkaei">Aenictus duengkaei</taxonomicName>
(see under 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus duengkaei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="duengkaei">Aenictus duengkaei</taxonomicName>
for differences). According to 
<bibRefCitation author="Wilson, EO" journalOrPublisher="Pacific Insects" pageId="25" pageNumber="74" pagination="427 - 483" title="The true army ants of the Indo-Australian area (Hymenoptera: Formicidae: Dorylinae)." volume="6" year="1964">Wilson (1964)</bibRefCitation>
the clypeus of 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
has an entire anterior margin without denticles. Following this information, 
<bibRefCitation author="Jaitrong, W" journalOrPublisher="Zootaxa" pageId="25" pageNumber="74" pagination="1 - 46" title="Synopsis of Aenictus species and revision of the A. currax and A. laeviceps groups in the eastern Oriental, Indo-Australian, and Australasian regions (Hymenoptera: Formicidae: Aenictinae)." volume="3128" year="2011">Jaitrong and Yamane (2011)</bibRefCitation>
treated 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
as a member of their 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
group (no denticles in this group). However, the non-type specimen mentioned above has nine denticles on the anterior clypeal margin. After carefully examining the type material of 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
we concluded that this species should be removed from the 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus piercei" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="piercei">Aenictus piercei</taxonomicName>
group and that it is a member of the 
<taxonomicName class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus javanus" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="javanus">Aenictus javanus</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>