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<document id="70015B4B186661EF3C67E19813A622CA" ID-DOI="10.5281/zenodo.5340846" ID-ISSN="2345-7600" ID-Zenodo-Dep="5340846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630188033318" checkinUser="diego" docAuthor="Mendoza, Jose Christopher E. &amp; Ng, Peter K. L." docDate="2008" docId="0E16272EFF87FF8DFF17FCF2FBFBFD00" docLanguage="en" docName="56rbz385-404.pdf" docOrigin="Raffles Bulletin of Zoology 56 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Visayax osteodictyon Mendoza &amp; Ng 2008, new species" docType="treatment" docVersion="3" lastPageNumber="395" masterDocId="F22F5F56FF80FF87FFE2FFB3FFA0FFB8" masterDocTitle="New Genera And Species Of Euxanthine Crabs (Crustacea: Decapoda: Brachyura: Xanthidae) From The Bohol Sea, The Philippines" masterLastPageNumber="404" masterPageNumber="385" pageNumber="392" updateTime="1699180562169" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="57A525C0783E7B2EB3863C9BA2E353A6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C2E0E9945149CCC1E7B629AFEFD10990">
<mods:title id="701A101C77ACE7C78F31AF19A1429D0B">New Genera And Species Of Euxanthine Crabs (Crustacea: Decapoda: Brachyura: Xanthidae) From The Bohol Sea, The Philippines</mods:title>
</mods:titleInfo>
<mods:name id="9E31A3D0ADF2993A414E0BE2E921135B" type="personal">
<mods:role id="D32FEBC5E2C900AED09E9DB0279CBD37">
<mods:roleTerm id="523CC6942CA4A174F4F39129487C03E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B86EDAE4E29B7CAE3EA26429A491993D">Mendoza, Jose Christopher E.</mods:namePart>
</mods:name>
<mods:name id="B58488121116D1065DA11960C46671D9" type="personal">
<mods:role id="BAC09D970FFE3320C0589D231C9A29EB">
<mods:roleTerm id="C75444322F8E4FEF6F424392BF94B8C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F282FF44801D08BD3AD78540B75A055">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="55E426B477B1852BC947B0818E71188A">text</mods:typeOfResource>
<mods:relatedItem id="DD8D228B826C3368F3396F96B70410D2" type="host">
<mods:titleInfo id="B513DC55140921E6CF07B870689743FF">
<mods:title id="C8A356689F84C20A686E93B5389F054D">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="D2AD13F0B339A65AE2FB6D33EB3BDF52">
<mods:date id="F1D705A061F54AAB471AEA02C1DD9809">2008</mods:date>
<mods:detail id="3553D8B1CFF0114921CA81895190AF5B" type="pubDate">
<mods:number id="4662415C0A003E9C8F4093A9E1D0F7BD">2008-08-31</mods:number>
</mods:detail>
<mods:detail id="B2E80E400C83AB213A28EEAFC81D7AD0" type="volume">
<mods:number id="22614E6C46D25997B241B7CCCFB1761D">56</mods:number>
</mods:detail>
<mods:detail id="EB9C97F7F7451DC33E8D713D8003DE58" type="issue">
<mods:number id="1709895E73110638F85A695F000481BF">2</mods:number>
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<mods:extent id="D686F418CC8D49EC8DBF364C673D9488" unit="page">
<mods:start id="E0EDBB1351A2109C837A01D189352C6C">385</mods:start>
<mods:end id="ECD754090B758B422F2B51D2AAD3F2C4">404</mods:end>
</mods:extent>
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<mods:classification id="BB5706E2BF9A781CEC65A96A1C9B1ECE">journal article</mods:classification>
<mods:identifier id="C09A7A94F2495C084017D75367F99BFE" type="DOI">10.5281/zenodo.5340846</mods:identifier>
<mods:identifier id="7DB5DB3510707136B733067D98FC1A86" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="0C0DE92A37A989734EAAADC83130AA62" type="Zenodo-Dep">5340846</mods:identifier>
</mods:mods>
<treatment id="0E16272EFF87FF8DFF17FCF2FBFBFD00" LSID="urn:lsid:plazi:treatment:0E16272EFF87FF8DFF17FCF2FBFBFD00" httpUri="http://treatment.plazi.org/id/0E16272EFF87FF8DFF17FCF2FBFBFD00" lastPageId="10" lastPageNumber="395" pageId="7" pageNumber="392">
<subSubSection id="CEA5C5B3FF87FF80FF17FCF2FDCFFCE0" box="[245,623,832,857]" pageId="7" pageNumber="392" type="nomenclature">
<paragraph id="86009638FF87FF80FF17FCF2FDCFFCE0" blockId="7.[245,623,832,888]" box="[245,623,832,857]" pageId="7" pageNumber="392">
<heading id="DD482154FF87FF80FF17FCF2FDCFFCE0" bold="true" box="[245,623,832,857]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="392" reason="2">
<emphasis id="B4CB4A2AFF87FF80FF17FCF2FDCFFCE0" bold="true" box="[245,623,832,857]" pageId="7" pageNumber="392">
<taxonomicName id="41BFEDBBFF87FF80FF17FCF2FE7BFCE1" ID-CoL="5BK7P" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[245,475,833,857]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="392" phylum="Arthropoda" rank="species" species="osteodictyon" status="sp. nov.">
<emphasis id="B4CB4A2AFF87FF80FF17FCF2FE7BFCE1" bold="true" box="[245,475,833,857]" italics="true" pageId="7" pageNumber="392">Visayax osteodictyon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="AFF8F751FF87FF80FE08FCF3FDCFFCE0" box="[490,623,832,856]" pageId="7" pageNumber="392" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CEA5C5B3FF87FF80FEBDFCD2FDA5FCC0" box="[351,517,865,888]" pageId="7" pageNumber="392" type="description">
<paragraph id="86009638FF87FF80FEBDFCD2FDA5FCC0" blockId="7.[245,623,832,888]" box="[351,517,865,888]" pageId="7" pageNumber="392">
(
<figureCitation id="1E848ABDFF87FF80FE85FCD2FE14FCC0" box="[359,436,865,888]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="7" pageNumber="392">Figs. 3</figureCitation>
,
<figureCitation id="1E848ABDFF87FF80FE23FCD2FE70FCC0" box="[449,464,865,888]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="7" pageNumber="392">4</figureCitation>
,
<figureCitation id="1E848ABDFF87FF80FE3FFCD2FE5CFCC0" box="[477,508,865,888]" captionStart="Fig" captionStartId="16.[138,173,2006,2027]" captionTargetBox="[138,1476,921,1981]" captionTargetId="figure-398@16.[138,1476,921,1981]" captionTargetPageId="16" captionText="Fig. 9. Live or freshly killed colouration of: A, Rizalthus anconis, new genus, new species, female paratype (MNHN- B30700) (11.2 × 7.3 mm); B, Visayax osteodictyon, new genus and new species, paratype male (ZRC 2008.0219) (3.3 × 2.2 mm); C, Visayax estampadori, new genus and new species, male holotype (NMCR-27509) (6.7 × 4.6 mm); and D, Hepatoporus pumex, new species, male holotype (NMCR-27510) (8.0 × 5.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.5340864" httpUri="https://zenodo.org/record/5340864/files/figure.png" pageId="7" pageNumber="392">9B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CEA5C5B3FF87FF80FF91FC2CFBC9FD3E" pageId="7" pageNumber="392" type="materials_examined">
<paragraph id="86009638FF87FF80FF91FC2CFD51FC69" blockId="7.[115,753,927,977]" pageId="7" pageNumber="392">
<emphasis id="B4CB4A2AFF87FF80FF91FC2CFEE8FC0C" bold="true" box="[115,328,927,948]" italics="true" pageId="7" pageNumber="392">Material examined. –</emphasis>
<typeStatus id="5904289AFF87FF80FEADFC2CFE0CFC0C" box="[335,428,927,948]" pageId="7" pageNumber="392" type="holotype">Holotype</typeStatus>
:
<specimenCount id="90B95DB1FF87FF80FE55FC2CFE59FC0C" box="[439,505,927,948]" pageId="7" pageNumber="392" type="male">1 male</specimenCount>
(6.3 ×
<quantity id="41473BDDFF87FF80FDA3FC2CFD2DFC0C" box="[577,653,927,948]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="7" pageNumber="392" unit="mm" value="4.3">4.3 mm</quantity>
) (NMCR- 27508), Station S22, hard ground covered with sand,
<quantity id="41473BDDFF87FF80FD71FC0FFD4EFC69" box="[659,750,956,977]" metricMagnitude="1" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="7" pageNumber="392" unit="m" value="18.5" valueMax="22.0" valueMin="15.0">15–22 m</quantity>
,
</paragraph>
<paragraph id="86009638FF87FF80FCCCFF72FB67FF4B" blockId="7.[814,1453,193,243]" pageId="7" pageNumber="392">
<materialsCitation id="36D79C65FF87FF80FCCCFF72FB63FF4B" collectingDate="2004-06-21" collectorName="Pamilacan" county="Marine Biodiversity Project" latitude="9.49" location="Pamilacan Island" longLatPrecision="129" longitude="123.933334" municipality="Panglao" pageId="7" pageNumber="392" specimenCount="1">
<geoCoordinate id="E38BF0FFFF87FF80FCCCFF72FC25FF6E" box="[814,901,193,214]" degrees="9" direction="north" minutes="29.4" orientation="latitude" pageId="7" pageNumber="392" precision="92" value="9.49">9°29.4'N</geoCoordinate>
<geoCoordinate id="E38BF0FFFF87FF80FC6DFF72FBA0FF6E" box="[911,1024,193,214]" degrees="123" direction="east" minutes="56.0" orientation="longitude" pageId="7" pageNumber="392" precision="92" value="123.933334">123°56.0'E</geoCoordinate>
,
<location id="8360C0E3FF87FF80FBECFF72FB1DFF6E" LSID="urn:lsid:plazi:treatment:0E16272EFF87FF8DFF17FCF2FBFBFD00:8360C0E3FF87FF80FBECFF72FB1DFF6E" box="[1038,1213,193,214]" county="Marine Biodiversity Project" latitude="9.49" longLatPrecision="129" longitude="123.933334" municipality="Panglao" name="Pamilacan Island" pageId="7" pageNumber="392">
<collectorName id="2B4AF3EEFF87FF80FBECFF72FBD5FF6E" box="[1038,1141,193,214]" pageId="7" pageNumber="392">Pamilacan</collectorName>
Island
</location>
, coll. by
<collectingMunicipality id="66640C42FF87FF80FAC1FF72FAD3FF6E" box="[1315,1395,193,214]" pageId="7" pageNumber="392">Panglao</collectingMunicipality>
2004
<collectingCounty id="6F61EEB4FF87FF80FCCCFF6DFB9FFF4B" box="[814,1087,222,243]" pageId="7" pageNumber="392">Marine Biodiversity Project</collectingCounty>
,
<date id="F201B0F8FF87FF80FBABFF6DFB63FF4B" box="[1097,1219,222,243]" pageId="7" pageNumber="392" value="2004-06-21">
<collectingDate id="E2454910FF87FF80FBABFF6DFB63FF4B" box="[1097,1219,222,243]" pageId="7" pageNumber="392" value="2004-06-21">21 Jun.2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="86009638FF87FF80FCCCFEA4FBC9FD3E" blockId="7.[814,1454,279,646]" pageId="7" pageNumber="392">
<materialsCitation id="36D79C65FF87FF80FCCCFEA4FC42FE3B" collectingDate="2004-06-01" collectionCode="ZRC" collectorName="Station B" county="Marine Biodiversity Project" elevation="24" latitude="9.553333" location="Point" longLatPrecision="129" longitude="123.805" municipality="Panglao" pageId="7" pageNumber="392" specimenCode="ZRC 2008.0753" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="5904289AFF87FF80FCCCFEA4FC30FE94" box="[814,912,279,300]" pageId="7" pageNumber="392" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="90B95DB1FF87FF80FC7EFEA4FC5CFE94" box="[924,1020,279,300]" pageId="7" pageNumber="392" type="female">2 females</specimenCount>
(7.4 ×
<quantity id="41473BDDFF87FF80FBAEFEA4FB3AFE94" box="[1100,1178,279,300]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="7" pageNumber="392" unit="mm" value="5.3">5.3 mm</quantity>
) (
<specimenCode id="D6193E43FF87FF80FB52FEA4FAF5FE94" box="[1200,1365,279,300]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="392">ZRC 2008.0753</specimenCode>
),
<collectorName id="2B4AF3EEFF87FF80FA8AFEA4FC9FFEF1" pageId="7" pageNumber="392">Station B</collectorName>
14, reef slop with overhangs,
<quantity id="41473BDDFF87FF80FB88FE87FB37FEF1" box="[1130,1175,308,329]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="392" unit="m" value="24.0">
<elevation id="0D92710BFF87FF80FB88FE87FB37FEF1" box="[1130,1175,308,329]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="392" unit="m" value="24.0">24m</elevation>
</quantity>
,
<geoCoordinate id="E38BF0FFFF87FF80FB46FE87FB5AFEF1" box="[1188,1274,308,329]" degrees="9" direction="north" minutes="33.2" orientation="latitude" pageId="7" pageNumber="392" precision="92" value="9.553333">9°33.2'N</geoCoordinate>
<geoCoordinate id="E38BF0FFFF87FF80FAE1FE87FAD0FEF1" box="[1283,1392,308,329]" degrees="123" direction="east" minutes="48.3" orientation="longitude" pageId="7" pageNumber="392" precision="92" value="123.805">123°48.3'E</geoCoordinate>
, BBC
<location id="8360C0E3FF87FF80FCCCFEE2FCC3FEDE" LSID="urn:lsid:plazi:treatment:0E16272EFF87FF8DFF17FCF2FBFBFD00:8360C0E3FF87FF80FCCCFEE2FCC3FEDE" box="[814,867,337,358]" county="Marine Biodiversity Project" latitude="9.553333" longLatPrecision="129" longitude="123.805" municipality="Panglao" name="Point" pageId="7" pageNumber="392">Point</location>
,
<location id="8360C0E3FF87FF80FC8CFEE2FBA0FEDE" LSID="urn:lsid:plazi:treatment:0E16272EFF87FF8DFF17FCF2FBFBFD00:8360C0E3FF87FF80FC8CFEE2FBA0FEDE" box="[878,1024,337,358]" county="Marine Biodiversity Project" latitude="9.553333" longLatPrecision="129" longitude="123.805" municipality="Panglao" name="Panglao Island" pageId="7" pageNumber="392">Panglao Island</location>
, coll. by
<collectingMunicipality id="66640C42FF87FF80FBBFFEE2FB0AFEDE" box="[1117,1194,337,358]" pageId="7" pageNumber="392">Panglao</collectingMunicipality>
2004
<collectingCounty id="6F61EEB4FF87FF80FB0BFEE2FCD3FE3B" pageId="7" pageNumber="392">Marine Biodiversity Project</collectingCounty>
,
<date id="F201B0F8FF87FF80FC98FEDDFC42FE3B" box="[890,994,366,387]" pageId="7" pageNumber="392" value="2004-06-01">
<collectingDate id="E2454910FF87FF80FC98FEDDFC42FE3B" box="[890,994,366,387]" pageId="7" pageNumber="392" value="2004-06-01">1 Jun.2004</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="36D79C65FF87FF80FC08FEDDFC35FE61" collectingDate="2004-07-04" collectionCode="ZRC" collectorName="Station B" county="Marine Biodiversity Project" elevation="13" latitude="9.518333" location="Black Forest" longLatPrecision="129" longitude="123.68833" municipality="Panglao" pageId="7" pageNumber="392" specimenCode="ZRC 2008.0754" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="90B95DB1FF87FF80FC08FEDDFB89FE3B" box="[1002,1065,366,387]" pageId="7" pageNumber="392" type="male">1 male</specimenCount>
(6.6 ×
<quantity id="41473BDDFF87FF80FB8AFEDDFB10FE3B" box="[1128,1200,366,387]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="392" unit="mm" value="5.1">5.1 mm</quantity>
) (
<specimenCode id="D6193E43FF87FF80FB23FEDDFAFDFE3B" box="[1217,1373,366,387]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="392">ZRC 2008.0754</specimenCode>
),
<collectorName id="2B4AF3EEFF87FF80FA89FEDDFC9FFE27" pageId="7" pageNumber="392">Station B</collectorName>
6, coral patches,
<quantity id="41473BDDFF87FF80FC08FE39FBE4FE27" box="[1002,1092,394,415]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="7" pageNumber="392" unit="m" value="13.0" valueMax="14.0" valueMin="12.0">
<elevation id="0D92710BFF87FF80FC08FE39FBE4FE27" box="[1002,1092,394,415]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="7" pageNumber="392" unit="m" value="13.0" valueMax="14.0" valueMin="12.0">12–14 m</elevation>
</quantity>
,
<geoCoordinate id="E38BF0FFFF87FF80FBB3FE39FB06FE27" box="[1105,1190,394,415]" degrees="9" direction="north" minutes="31.1" orientation="latitude" pageId="7" pageNumber="392" precision="92" value="9.518333">9°31.1'N</geoCoordinate>
<geoCoordinate id="E38BF0FFFF87FF80FB4DFE39FABCFE27" box="[1199,1308,394,415]" degrees="123" direction="east" minutes="41.3" orientation="longitude" pageId="7" pageNumber="392" precision="92" value="123.68833">123°41.3'E</geoCoordinate>
,
<location id="8360C0E3FF87FF80FAC8FE39FA0AFE27" LSID="urn:lsid:plazi:treatment:0E16272EFF87FF8DFF17FCF2FBFBFD00:8360C0E3FF87FF80FAC8FE39FA0AFE27" box="[1322,1450,394,415]" county="Marine Biodiversity Project" latitude="9.518333" longLatPrecision="129" longitude="123.68833" municipality="Panglao" name="Black Forest" pageId="7" pageNumber="392">Black Forest</location>
,
<location id="8360C0E3FF87FF80FCCCFE14FC6AFE04" LSID="urn:lsid:plazi:treatment:0E16272EFF87FF8DFF17FCF2FBFBFD00:8360C0E3FF87FF80FCCCFE14FC6AFE04" box="[814,970,423,444]" county="Marine Biodiversity Project" latitude="9.518333" longLatPrecision="129" longitude="123.68833" municipality="Panglao" name="Balicasag Island" pageId="7" pageNumber="392">Balicasag Island</location>
, coll. by
<collectingMunicipality id="66640C42FF87FF80FBFFFE14FBC9FE04" box="[1053,1129,423,444]" pageId="7" pageNumber="392">Panglao</collectingMunicipality>
2004
<collectingCounty id="6F61EEB4FF87FF80FB40FE14FA0AFE04" box="[1186,1450,423,444]" pageId="7" pageNumber="392">Marine Biodiversity Project</collectingCounty>
,
<date id="F201B0F8FF87FF80FCCCFE77FC35FE61" box="[814,917,452,473]" pageId="7" pageNumber="392" value="2004-07-04">
<collectingDate id="E2454910FF87FF80FCCCFE77FC35FE61" box="[814,917,452,473]" pageId="7" pageNumber="392" value="2004-07-04">4 Jul.2004</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="36D79C65FF87FF80FC7DFE77FC0AFD97" collectingDate="2004-06-19" collectionCode="ZRC" collectorName="Station B" county="Marine Biodiversity Project" elevation="12" latitude="9.625" location="Bingag" longLatPrecision="129" longitude="123.78167" municipality="Panglao" pageId="7" pageNumber="392" specimenCode="ZRC 2008.0218" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="90B95DB1FF87FF80FC7DFE77FC40FE61" box="[927,992,452,473]" pageId="7" pageNumber="392" type="male">1 male</specimenCount>
(2.3 x
<quantity id="41473BDDFF87FF80FBC7FE77FBD1FE61" box="[1061,1137,452,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="392" unit="mm" value="1.7">1.7 mm</quantity>
) (
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17, reef wall with small caves,
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, coll. by
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<collectingDate id="E2454910FF87FF80FCCCFDA9FC0AFD97" box="[814,938,538,559]" pageId="7" pageNumber="392" value="2004-06-19">19 Jun.2004</collectingDate>
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;
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(3.3 x
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<collectorName id="2B4AF3EEFF87FF80FA8AFDA9FC9EFDF4" pageId="7" pageNumber="392">Station B</collectorName>
39, reef wall with small caves,
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1,
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, coll. by
<collectingMunicipality id="66640C42FF87FF80FB3BFDE7FA86FDD1" box="[1241,1318,596,617]" pageId="7" pageNumber="392">Panglao</collectingMunicipality>
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,
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<collectingDate id="E2454910FF87FF80FC1FFDC2FBC5FD3E" box="[1021,1125,625,646]" pageId="7" pageNumber="392" value="2004-07-02">2 Jul.2004</collectingDate>
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.
</paragraph>
</subSubSection>
<subSubSection id="CEA5C5B3FF87FF8EFCCCFD1CFA3FF840" lastPageId="9" lastPageNumber="394" pageId="7" pageNumber="392" type="description">
<paragraph id="86009638FF87FF8EFCCCFD1CFF6AFC60" blockId="7.[814,1454,687,966]" lastBlockId="9.[115,754,193,984]" lastPageId="9" lastPageNumber="394" pageId="7" pageNumber="392">
<emphasis id="B4CB4A2AFF87FF80FCCCFD1CFC65FD7F" bold="true" box="[814,965,687,711]" italics="true" pageId="7" pageNumber="392">Description. –</emphasis>
Carapace (
<figureCitation id="1E848ABDFF87FF80FBD9FD1CFB2CFD7E" box="[1083,1164,687,710]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="7" pageNumber="392">Fig. 3A</figureCitation>
) broader than long; regions well-defined; 1M, 2M and 2F fused and raised, culminating in a dorsally directed tubercle on 2M; sections of branchial region—3L and 5L—raised into dorsally-projecting tubercles; 3M and 1P distinct, 4M indistinct; entire dorsal surface covered by network of fused granules forming a reticulate pattern, with short setae sparsely distributed at their bases; suborbital region eroded with several small depressions; subhepatic and pterygostomial regions with low, discrete granules. Front deflexed ventrally, appearing truncate, shallowly bilobed from dorsal view; lobes broadly triangular, projecting medially, separated by a wide U-shaped median cleft that continues as a shallow median groove in the frontal region; laterally curving toward inner orbital angle, separated from it by a short groove. Supraorbital region granulose; margin mildly serrated, relatively short, not clearly meeting anterolateral margin, without distinct external orbital tooth. Suborbital margin mildly serrated, outer edge connected to anterolateral margin by a ridge consisting of fused granules. Orbits small, width about 0.1 times carapace width; eyestalks short, distal edge with cornea lined with small, tooth-like granules; corneas well developed. Anterolateral margin strongly arcuate, granulose, appearing irregularly serrated; anteriormost part not meeting supraorbital margin but curving into pterygostomial region where it eventually disappears close to anterolateral corner of buccal cavity; posterior end forming distinct, angular junction with deeply concave posterolateral margin. Anterior portion of posterolateral margin subparallel to frontal margin, and continues as a curve to meet posterior margin. Last ambulatory leg coapted against concavity in posterolateral margin. Junction with posterior carapace margin defined by a granular, upwardly and outwardly projecting tooth; posterior carapace margin straight.
</paragraph>
<caption id="D2C0C6B0FF87FF80FF91F7A1FDA1F7FC" ID-DOI="http://doi.org/10.5281/zenodo.5340852" ID-Zenodo-Dep="5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="7" pageNumber="392" startId="7.[115,150,2066,2087]" targetBox="[115,1453,1024,2040]" targetPageId="7">
<paragraph id="86009638FF87FF80FF91F7A1FDA1F7FC" blockId="7.[115,1453,2066,2116]" pageId="7" pageNumber="392">
Fig. 3.
<taxonomicName id="41BFEDBBFF87FF80FF5AF7A1FE23F79F" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[184,387,2066,2087]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="392" phylum="Arthropoda" rank="species" species="osteodictyon" status="new genus, new species">
<emphasis id="B4CB4A2AFF87FF80FF5AF7A1FE23F79F" box="[184,387,2066,2087]" italics="true" pageId="7" pageNumber="392">Visayax osteodictyon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="AFF8F751FF87FF80FE72F7A1FDDAF79F" box="[400,634,2066,2087]" pageId="7" pageNumber="392" rank="species">new genus, new species</taxonomicNameLabel>
, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface.
</paragraph>
</caption>
<caption id="D2C0C6B0FF88FF8FFF68F846FC31F7FC" ID-DOI="http://doi.org/10.5281/zenodo.5340854" ID-Zenodo-Dep="5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="8" pageNumber="393" startId="8.[138,173,2037,2058]" targetBox="[157,1455,197,1987]" targetPageId="8">
<paragraph id="86009638FF88FF8FFF68F846FC31F7FC" blockId="8.[138,1476,2037,2116]" pageId="8" pageNumber="393">
Fig. 4.
<taxonomicName id="41BFEDBBFF88FF8FFF32F846FE3BF7B2" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[208,411,2037,2058]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="393" phylum="Arthropoda" rank="species" species="osteodictyon" status="new genus, new species">
<emphasis id="B4CB4A2AFF88FF8FFF32F846FE3BF7B2" box="[208,411,2037,2058]" italics="true" pageId="8" pageNumber="393">Visayax osteodictyon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="AFF8F751FF88FF8FFE4BF846FD34F7B2" box="[425,660,2037,2058]" pageId="8" pageNumber="393" rank="species">new genus, new species</taxonomicNameLabel>
, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm.
</paragraph>
</caption>
<paragraph id="86009638FF89FF8EFF91FBB2FEC7FAA0" blockId="9.[115,754,1025,1304]" pageId="9" pageNumber="394">
Antennules (
<figureCitation id="1E848ABDFF89FF8EFEE7FBB2FECBFBA0" box="[261,363,1025,1048]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="9" pageNumber="394">Figs. 3C</figureCitation>
,
<figureCitation id="1E848ABDFF89FF8EFE99FBB2FE3EFBA0" box="[379,414,1025,1048]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="9" pageNumber="394">4A</figureCitation>
) folding almost transversely. Basal antennal segment large, subrectangular, granulose, almost occupying entire space between antennular fossa and internal orbital angle, orbital hiatus completely filled; long flagellum arising from distal margin, almost reaching outer edge of orbit. Posterior edge of epistome (
<figureCitation id="1E848ABDFF89FF8EFDDFFB12FD32FB00" box="[573,658,1185,1208]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="9" pageNumber="394">Fig. 4D</figureCitation>
) slightly undulate, with very shallow, rounded median projection, lateral regions with slight depressions. Endostome without any discernible ridges.
</paragraph>
<paragraph id="86009638FF89FF8EFF91FAF2FDD1F900" blockId="9.[115,754,1345,1720]" pageId="9" pageNumber="394">
Outer surface of third maxilliped (
<figureCitation id="1E848ABDFF89FF8EFE05FAF2FDE6FAE0" box="[487,582,1345,1368]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="9" pageNumber="394">Figs. 3B</figureCitation>
,
<figureCitation id="1E848ABDFF89FF8EFDB1FAF2FDD3FAE0" box="[595,627,1345,1368]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="9" pageNumber="394">4B</figureCitation>
) granulose, eroded. Merus subquadrate, median length about half that of ischium, with 2 shallow depressions on either side of a submedian, granular ridge; margins with small granules, anterior and external margins slightly concave, internal margin straight; anteroexternal angle rounded. Ischium subrectangular, inner margin with granules and short, stiff setae; with deep, longitudinal median sulcus, with smaller longitudinal depression anterolateral to it, adjacent to anteroexternal margin; separated from basis by distinct suture. Exopod granulose, tapering toward distal end which almost reaches anterior edge of merus, flagellum long.
</paragraph>
<paragraph id="86009638FF89FF8EFF91F952FB35FF40" blockId="9.[115,754,1761,2072]" lastBlockId="9.[814,1453,193,248]" pageId="9" pageNumber="394">
Surface of male thoracic sternum (
<figureCitation id="1E848ABDFF89FF8EFE1EF952FDF3F940" box="[508,595,1761,1784]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="9" pageNumber="394">Fig. 3B</figureCitation>
) eroded, with reticulate pattern of ridges and depressions similar to dorsal carapace surface; large distinct depression just anterior to telson of abdomen, no discernible suture within, smaller depressions seen on either side of abdomen; anterior region relatively elongate. Sternites 1 and 2 completely fused into triangular plate, separated from sternite 3 by deep transverse suture. Sternites 3 and 4 partially fused, with partial suture seen near anterior edge of cheliped coxae. Abdominal cavity deep, sternal condyle on sternite 5 slightly nearer to suture with sternite 4, abdomen reaching to imaginary line joining posterior edge of cheliped coxae.
</paragraph>
<paragraph id="86009638FF89FF8EFCCCFE92FA22FCC0" blockId="9.[814,1453,289,888]" pageId="9" pageNumber="394">
Chelipeds (
<figureCitation id="1E848ABDFF89FF8EFC4AFE92FB95FE80" box="[936,1077,289,312]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="9" pageNumber="394">Figs. 3A, 3D</figureCitation>
) similar, subequal. Fingers shorter than palm, cutting edges with 4 or 5 teeth, tips pointed. Dactylus slightly curved, with three rows of granules, continuing distally as ridges and separated by shallow grooves along its length, base granulose. Fixed finger, slightly deflexed with 2 rows of granules continuing as ridges and broad submarginal groove on outer surface continuing from palm; base granulose and pigmented. Palm outer surface eroded in reticulate pattern similar to dorsal carapace surface, upper margin irregular, wedge-like and granulose, lower margin slightly convex and granulose; inner surface relatively smoother, with fewer and smaller granules; pigmentation on base of fixed finger continues into lower portions of palm outer and inner surfaces forming a dark patch. Carpus short, dorsal and ventral surface eroded, reticulate, inner distal angle with low triangular tooth. Inner surface of fingers, palm and carpus coapted against anterolateral margin of carapace. Merus granulose, slightly longer than carpus, with rectangular, ventro-distal tooth apposed against carpus.
</paragraph>
<paragraph id="86009638FF89FF8EFCCCFC12FA2EFAA0" blockId="9.[814,1453,929,1304]" pageId="9" pageNumber="394">
Ambulatory legs (
<figureCitation id="1E848ABDFF89FF8EFC1BFC12FBFDFC00" box="[1017,1117,929,952]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="9" pageNumber="394">Figs. 3A</figureCitation>
,
<figureCitation id="1E848ABDFF89FF8EFB8EFC12FB2DFC00" box="[1132,1165,929,952]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="9" pageNumber="394">4C</figureCitation>
) relatively short, first leg longest, coxa-to-dactylus length about 0.7 times carapace width, sparsely setose. Merus subrectangular, subtriangular in cross-section; anterior and posterior edges, especially fourth ambulatory leg, lined with large, pointed granules; dorsal surface in fourth ambulatory leg with reticulate pattern of fused granules, others finely granular. Dorsal surface of carpus reticulate, with row of pointed granules on serrated anterior edge. Propodus subrectangular, rugose, anterior and posterior edges serrated in most legs. Dactylus slightly curved inwards, dorsal surface and edges covered with small spines, short setae; terminates distally in curved chitinous claw.
</paragraph>
<paragraph id="86009638FF89FF8EFCCCFAF2FB18F9A0" blockId="9.[814,1453,1345,1560]" pageId="9" pageNumber="394">
Male abdomen and telson (
<figureCitation id="1E848ABDFF89FF8EFBBBFAF2FB0CFAE0" box="[1113,1196,1345,1368]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="9" pageNumber="394">Fig. 4E</figureCitation>
) with reticulate pattern of fused granules and depressions. Telson rounded; lateral margins convex. Segment 6 subquadrate, about 1.5 times length of telson, lateral margins convex. Segments 3–5 completely fused, without any trace of sutures; lateral margins markedly concave. Segment 2 subtrapezoidal, segment 1 trapezoidal without transverse ridge.
</paragraph>
<paragraph id="86009638FF89FF8EFCCCF9F2FAFCF960" blockId="9.[814,1453,1601,1752]" pageId="9" pageNumber="394">
G1 (
<figureCitation id="1E848ABDFF89FF8EFC82F9F2FC72F9E0" box="[864,978,1601,1624]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="9" pageNumber="394">Fig. 4F–G</figureCitation>
) long and slender, mostly straight, curving outward and tapering distally; terminally located opening bordered by 2 lamellar folds; several long, thick and stiff simple setae located subdistally; basal external region with row of stiff simple setae, surrounded by finer setae.
</paragraph>
<paragraph id="86009638FF89FF8EFCCCF8B2FA3FF840" blockId="9.[814,1453,1793,2040]" pageId="9" pageNumber="394">
<emphasis id="B4CB4A2AFF89FF8EFCCCF8B2FBA0F8A1" bold="true" box="[814,1024,1793,1817]" italics="true" pageId="9" pageNumber="394">Live colouration. –</emphasis>
The carapace and pereiopods (
<figureCitation id="1E848ABDFF89FF8EFAB0F8B2FA04F8A0" box="[1362,1444,1793,1816]" captionStart="Fig" captionStartId="16.[138,173,2006,2027]" captionTargetBox="[138,1476,921,1981]" captionTargetId="figure-398@16.[138,1476,921,1981]" captionTargetPageId="16" captionText="Fig. 9. Live or freshly killed colouration of: A, Rizalthus anconis, new genus, new species, female paratype (MNHN- B30700) (11.2 × 7.3 mm); B, Visayax osteodictyon, new genus and new species, paratype male (ZRC 2008.0219) (3.3 × 2.2 mm); C, Visayax estampadori, new genus and new species, male holotype (NMCR-27509) (6.7 × 4.6 mm); and D, Hepatoporus pumex, new species, male holotype (NMCR-27510) (8.0 × 5.7 mm)." figureDoi="http://doi.org/10.5281/zenodo.5340864" httpUri="https://zenodo.org/record/5340864/files/figure.png" pageId="9" pageNumber="394">Fig. 9B</figureCitation>
) are basically white. The fused granules forming the reticulate pattern on the carapace, chelipeds and ambulatory legs are variably tinged light pink. There are also small, discrete spots or smudges of dark orange-brown on carapace, palm of chelae, and anterior edge of merus and carpus of ambulatory legs. The lower part of outer and inner surfaces of palm of the chelae is black while both fingers of the chelae are white.
</paragraph>
</subSubSection>
<subSubSection id="CEA5C5B3FF8AFF8DFF68FF72FE2DFEC0" pageId="10" pageNumber="395" type="etymology">
<paragraph id="86009638FF8AFF8DFF68FF72FE2DFEC0" blockId="10.[138,777,193,376]" pageId="10" pageNumber="395">
<emphasis id="B4CB4A2AFF8AFF8DFF68FF72FE83FF61" bold="true" box="[138,291,193,217]" italics="true" pageId="10" pageNumber="395">Etymology. –</emphasis>
The specific epithet
<taxonomicName id="41BFEDBBFF8AFF8DFDFDFF72FD0DFF60" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[543,685,193,216]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="osteodictyon">
<emphasis id="B4CB4A2AFF8AFF8DFDFDFF72FD0DFF60" box="[543,685,193,216]" italics="true" pageId="10" pageNumber="395">osteodictyon</emphasis>
</taxonomicName>
derives from two Greek words, “osteon” which means bone, and “dictyon” which means net, characterizing the reticulate sculpturing—reminiscent of spongy bone tissue—on the carapace, pereiopods and other surfaces of this species. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="CEA5C5B3FF8AFF8DFF68FE12FBFBFD00" pageId="10" pageNumber="395" type="discussion">
<paragraph id="86009638FF8AFF8DFF68FE12FC5AFE20" blockId="10.[138,777,417,920]" lastBlockId="10.[837,1476,193,408]" pageId="10" pageNumber="395">
<emphasis id="B4CB4A2AFF8AFF8DFF68FE12FE40FE00" box="[138,480,417,441]" italics="true" pageId="10" pageNumber="395">
<emphasis id="B4CB4A2AFF8AFF8DFF68FE12FEA2FE01" bold="true" box="[138,258,417,441]" italics="true" pageId="10" pageNumber="395">Remarks. –</emphasis>
<taxonomicName id="41BFEDBBFF8AFF8DFEE5FE12FE40FE00" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[263,480,417,440]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="osteodictyon" status="sp. nov.">Visayax osteodictyon</taxonomicName>
</emphasis>
,
<taxonomicNameLabel id="AFF8F751FF8AFF8DFE09FE12FDCAFE00" box="[491,618,417,440]" pageId="10" pageNumber="395" rank="species">new species</taxonomicNameLabel>
, is superficially similar to the various species of
<taxonomicName id="41BFEDBBFF8AFF8DFE15FE72FD33FE60" authorityName="A. Milne-Edwards, 1879 [in A. Milne-Edwards, 1873-" authorityYear="1880" box="[503,659,449,472]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="B4CB4A2AFF8AFF8DFE15FE72FD33FE60" box="[503,659,449,472]" italics="true" pageId="10" pageNumber="395">Glyptoxanthus</emphasis>
</taxonomicName>
A. Milne- Edwards, 1879, particularly in the sculpturing of the carapace and pereiopods (cf.
<bibRefCitation id="E22EEBC9FF8AFF8DFE04FDB2FD71FDA0" author="Rathbun, M" box="[486,721,513,536]" pageId="10" pageNumber="395" pagination="1 - 609" refId="ref16111" refString="Rathbun, M. J., 1930. The cancroid crabs of America of the families Euryalidae, Portunidae, Atelecyclidae, Cancridae and Xanthidae. United States National Museum Bulletin, 152: 1 - 609, Pls. 1 - 230." type="journal article" year="1930">Rathbun, 1930: 263</bibRefCitation>
, pls. 107–109). It is easily distinguished from the
<typeStatus id="5904289AFF8AFF8DFD9CFD92FD0CFD80" box="[638,684,545,568]" pageId="10" pageNumber="395">type</typeStatus>
species,
<taxonomicName id="41BFEDBBFF8AFF8DFF68FDF2FDC9FDE0" authority="(Stimpson, 1859)" baseAuthorityName="Stimpson" baseAuthorityYear="1859" box="[138,617,577,600]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="erosus">
<emphasis id="B4CB4A2AFF8AFF8DFF68FDF2FE2BFDE0" box="[138,395,577,600]" italics="true" pageId="10" pageNumber="395">Glyptoxanthus erosus</emphasis>
(
<bibRefCitation id="E22EEBC9FF8AFF8DFE7DFDF2FDC1FDE0" author="Stimpson, W" box="[415,609,577,600]" pageId="10" pageNumber="395" pagination="49 - 93" refId="ref16589" refString="Stimpson, W., 1859. Notes on North American Crustacea. Annals of the Lyceum of Natural History of New York, 7: 49 - 93, Pl. 1." type="journal article" year="1859">Stimpson, 1859</bibRefCitation>
)
</taxonomicName>
, by having a networked, anastomosing pattern to the fused granules (
<figureCitation id="1E848ABDFF8AFF8DFD31FDD2FF72FD20" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="10" pageNumber="395">Figs. 3A, D</figureCitation>
,
<figureCitation id="1E848ABDFF8AFF8DFF03FD32FE85FD20" box="[225,293,641,664]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="10" pageNumber="395">4C, E</figureCitation>
), whereas in
<taxonomicName id="41BFEDBBFF8AFF8DFE22FD32FD8AFD20" baseAuthorityName="Stimpson" baseAuthorityYear="1859" box="[448,554,641,664]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="erosus">
<emphasis id="B4CB4A2AFF8AFF8DFE22FD32FD8AFD20" box="[448,554,641,664]" italics="true" pageId="10" pageNumber="395">G. erosus</emphasis>
</taxonomicName>
, the fused granules tend to take a more vermicular pattern and are more closely spaced. The front of
<taxonomicName id="41BFEDBBFF8AFF8DFE8AFD72FE6EFD60" baseAuthorityName="Stimpson" baseAuthorityYear="1859" box="[360,462,705,728]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="erosus">
<emphasis id="B4CB4A2AFF8AFF8DFE8AFD72FE6EFD60" box="[360,462,705,728]" italics="true" pageId="10" pageNumber="395">G. erosus</emphasis>
</taxonomicName>
has two deep lateral notches, giving it a quadridentate appearance (vs. two simple, broadly triangular lobes in
<taxonomicName id="41BFEDBBFF8AFF8DFEBCFCB2FDA3FCA0" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[350,515,769,792]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="osteodictyon">
<emphasis id="B4CB4A2AFF8AFF8DFEBCFCB2FDA3FCA0" box="[350,515,769,792]" italics="true" pageId="10" pageNumber="395">V. osteodictyon</emphasis>
</taxonomicName>
,
<figureCitation id="1E848ABDFF8AFF8DFDF6FCB2FDCCFCA0" box="[532,620,769,792]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="10" pageNumber="395">Fig. 3A</figureCitation>
); there are no dorsally directed tubercles on the carapace (vs. present,
<figureCitation id="1E848ABDFF8AFF8DFD3DFC92FF0EFCE0" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="10" pageNumber="395">Fig. 3A</figureCitation>
); there are oblique ridges in the endostome (vs. absent); there is a visible median longitudinal suture on male thoracic sternite 4, just anterior to the telson (vs. absent,
<figureCitation id="1E848ABDFF8AFF8DFD47FC32FD5BFC20" box="[677,763,897,920]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="10" pageNumber="395">Fig. 3B</figureCitation>
); the palm of the chela is stouter and more robust (vs. more slender,
<figureCitation id="1E848ABDFF8AFF8DFC40FF52FBBDFF40" box="[930,1053,225,248]" captionStart="Fig" captionStartId="7.[115,150,2066,2087]" captionTargetBox="[115,1453,1024,2040]" captionTargetId="figure-221@7.[115,1453,1024,2040]" captionTargetPageId="7" captionText="Fig. 3. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, dorsal view; B, ventral view; C, frontal view; D, left chela, outer surface." figureDoi="http://doi.org/10.5281/zenodo.5340852" httpUri="https://zenodo.org/record/5340852/files/figure.png" pageId="10" pageNumber="395">Fig. 3A, D</figureCitation>
); and while the G1 is similar to those seen in species of
<taxonomicName id="41BFEDBBFF8AFF8DFBF0FEB1FB23FEA1" authorityName="Dana" authorityYear="1851" box="[1042,1155,258,281]" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="B4CB4A2AFF8AFF8DFBF0FEB1FB23FEA1" box="[1042,1155,258,281]" italics="true" pageId="10" pageNumber="395">Euxanthus</emphasis>
</taxonomicName>
: it is much longer, the distal end is blunt, with only one subterminal seta, and the terminus folds over the opening forming a triangular flap (vs. distal end tapering, with several subterminal setae and no triangular flap,
<figureCitation id="1E848ABDFF8AFF8DFC99FE32FC4EFE20" box="[891,1006,385,408]" captionStart="Fig" captionStartId="8.[138,173,2037,2058]" captionTargetBox="[157,1455,197,1987]" captionTargetId="figure-14@8.[137,1474,178,2021]" captionTargetPageId="8" captionText="Fig. 4. Visayax osteodictyon, new genus, new species, male holotype (NMCR-27508) (6.3 × 4.3 mm): A, frontal view, left side, showing antenna, antennule and orbit; B, left third maxilliped, external view; C, left fourth ambulatory leg; D, epistome; E, abdomen and telson; F, left G1, external view; G, left G1, internal view. Scale bars: A–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5340854" httpUri="https://zenodo.org/record/5340854/files/figure.png" pageId="10" pageNumber="395">Fig. 4F–G</figureCitation>
).
</paragraph>
<paragraph id="86009638FF8AFF8DFCA7FE71FBFBFD00" blockId="10.[837,1476,449,696]" pageId="10" pageNumber="395">
<taxonomicName id="41BFEDBBFF8AFF8DFCA7FE71FB87FE61" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[837,1063,450,473]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="osteodictyon">
<emphasis id="B4CB4A2AFF8AFF8DFCA7FE71FB87FE61" box="[837,1063,450,473]" italics="true" pageId="10" pageNumber="395">Visayax osteodictyon</emphasis>
</taxonomicName>
is thus far known only from the
<typeStatus id="5904289AFF8AFF8DFA77FE72FA63FE60" box="[1429,1475,449,472]" pageId="10" pageNumber="395">type</typeStatus>
locality, Pamilacan Island, and nearby islands of Panglao and Balicasag, all of which are in the
<collectingRegion id="447B58DAFF8AFF8DFB14FDB2FA98FDA0" box="[1270,1336,513,536]" country="Philippines" name="Bohol" pageId="10" pageNumber="395">Bohol</collectingRegion>
Sea, central
<collectingCountry id="FEA8D6A8FF8AFF8DFCA7FD92FC61FD80" box="[837,961,545,568]" name="Philippines" pageId="10" pageNumber="395">Philippines</collectingCountry>
. It has been collected at depths ranging between
<quantity id="41473BDDFF8AFF8DFCA7FDF2FC10FDE0" box="[837,944,577,600]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="2.5" metricValueMin="0.3" pageId="10" pageNumber="395" unit="m" value="14.0" valueMax="25.0" valueMin="3.0">3 to 25 m</quantity>
, in sandy substrate and in coral reef walls. It has been observed to occur together with
<taxonomicName id="41BFEDBBFF8AFF8DFB2DFDD1FA2BFDC1" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[1231,1419,610,633]" class="Malacostraca" family="Xanthidae" genus="Rizalthus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="anconis" status="sp. nov.">
<emphasis id="B4CB4A2AFF8AFF8DFB2DFDD1FA2BFDC1" box="[1231,1419,610,633]" italics="true" pageId="10" pageNumber="395">Rizalthus anconis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="AFF8F751FF8AFF8DFA75FDD2FC35FD20" pageId="10" pageNumber="395" rank="species">new species</taxonomicNameLabel>
, and
<taxonomicName id="41BFEDBBFF8AFF8DFC2EFD31FBCBFD21" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" box="[972,1131,642,665]" class="Malacostraca" family="Xanthidae" genus="Visayax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="395" phylum="Arthropoda" rank="species" species="estampadori" status="sp. nov.">
<emphasis id="B4CB4A2AFF8AFF8DFC2EFD31FBCBFD21" box="[972,1131,642,665]" italics="true" pageId="10" pageNumber="395">V. estampadori</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="AFF8F751FF8AFF8DFB9AFD32FB5BFD20" box="[1144,1275,641,664]" pageId="10" pageNumber="395" rank="species">new species</taxonomicNameLabel>
, in reef walls from at least one station (B17).
</paragraph>
</subSubSection>
</treatment>
</document>
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