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<document id="A4CBC8ABA84A40E8D3C9D1EAA39B55E1" ID-DOI="10.5281/zenodo.187429" ID-GBIF-Dataset="54ec63fe-608a-49b0-b247-93dc91321877" ID-ISSN="1175-5326" ID-Zenodo-Dep="187429" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461070156554" checkinUser="plazi" docAuthor="Das, Indraneil &amp; Lim, Kelvin K. P." docDate="2009" docId="03F2922DFFF4565FFF693AF6FC23E7D9" docLanguage="en" docName="zt02088p023.pdf" docOrigin="Zootaxa 2088" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dibamus tebal Das &amp; Lim, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="FFCBEA55FFF55659FFFE3F15FF9BE726" masterDocTitle="A new species of Dibamus (Squamata: Dibamidae) from Pulau Simeuleu, Mentawai Archipelago, Indonesia" masterLastPageNumber="23" masterPageNumber="15" pageNumber="16" updateTime="1698584208895" updateUser="plazi">
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<mods:title id="7EF090ADD3F594C75541E027AB0C09E3">A new species of Dibamus (Squamata: Dibamidae) from Pulau Simeuleu, Mentawai Archipelago, Indonesia</mods:title>
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<mods:namePart id="A9120096C081DFBB5730D37A39C94897">Das, Indraneil</mods:namePart>
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<mods:namePart id="6D701486B010913B7219C18797317CCF">Lim, Kelvin K. P.</mods:namePart>
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<mods:title id="8820F8DF51DA2F2574F1FF8EF10CC937">Zootaxa</mods:title>
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<mods:date id="993B58573C505CB5D259B3C2B5F5BA99">2009</mods:date>
<mods:detail id="C5FD99F1E31A915647B4F1237F85AC72" type="volume">
<mods:number id="A7AAB8EEB18E5D313A8222C7F22BCBA9">2088</mods:number>
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<mods:identifier id="30CC7B75ECC90FD2B8F0C8FE4A8C0171" type="DOI">10.5281/zenodo.187429</mods:identifier>
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<treatment id="03F2922DFFF4565FFF693AF6FC23E7D9" ID-DOI="http://doi.org/10.5281/zenodo.5690668" ID-GBIF-Taxon="119615606" ID-Zenodo-Dep="5690668" LSID="urn:lsid:plazi:treatment:03F2922DFFF4565FFF693AF6FC23E7D9" httpUri="http://treatment.plazi.org/id/03F2922DFFF4565FFF693AF6FC23E7D9" lastPageId="6" lastPageNumber="20" pageId="1" pageNumber="16">
<subSubSection id="C34170B0FFF45658FF693AF6FE8CE106" pageId="1" pageNumber="16" type="nomenclature">
<paragraph id="8BE4233BFFF45658FF693AF6FE30E2DA" blockId="1.[151,427,1506,1568]" box="[151,427,1506,1533]" pageId="1" pageNumber="16">
<heading id="D0AC9457FFF45658FF693AF6FE30E2DA" bold="true" box="[151,427,1506,1533]" fontSize="11" level="1" pageId="1" pageNumber="16" reason="1">
<emphasis id="B92FFF29FFF45658FF693AF6FE30E2DA" bold="true" box="[151,427,1506,1533]" pageId="1" pageNumber="16">
<taxonomicName id="4C5B58B8FFF45658FF693AF6FEDDE2DB" box="[151,326,1507,1533]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="16" phylum="Chordata" rank="species" species="tebal" status="sp. nov.">
<emphasis id="B92FFF29FFF45658FF693AF6FEDDE2DB" bold="true" box="[151,326,1507,1533]" italics="true" pageId="1" pageNumber="16">Dibamus tebal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C4252FFF45658FEB33AF7FE30E2DA" box="[333,427,1506,1532]" pageId="1" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE4233BFFF45658FF693913FE8CE106" blockId="1.[151,427,1506,1568]" box="[151,279,1542,1568]" pageId="1" pageNumber="16">
(
<figureCitation id="13603FBEFFF45658FF613913FE94E106" box="[159,271,1542,1568]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,255,1826,1850]" captionStartId-1="3.[151,257,1911,1935]" captionStartId-2="4.[151,255,650,674]" captionStartId-3="4.[151,255,1703,1727]" captionTargetBox-0="[151,1434,879,1800]" captionTargetBox-1="[315,1271,194,1885]" captionTargetBox-2="[449,1136,186,625]" captionTargetBox-3="[268,1318,746,1678]" captionTargetId-0="figure@2.[151,1436,879,1801]" captionTargetId-1="figure@3.[315,1272,194,1887]" captionTargetId-2="figure@4.[449,1138,186,626]" captionTargetId-3="figure@4.[268,1319,722,1679]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="FIGURE 1. Holotype of Dibamus tebal sp. nov. (ZMA 15500). Marker = 10 mm." captionText-1="FIGURE 2. Head of holotype of Dibamus tebal sp. nov. (ZMA 15500) in dorsal (top), lateral (middle) and ventral (bottom) views. Marker = 2 mm." captionText-2="FIGURE 3. Anal region of holotype of Dibamus tebal sp. nov. (ZMA 15500), showing flap-like hind limbs. Marker = 1 mm." captionText-3="FIGURE 4. Radiograph of holotype of Dibamus tebal sp. nov. (ZMA 15500)." httpUri-0="https://zenodo.org/record/187430/files/figure.png" httpUri-1="https://zenodo.org/record/187431/files/figure.png" httpUri-2="https://zenodo.org/record/187432/files/figure.png" httpUri-3="https://zenodo.org/record/187433/files/figure.png" pageId="1" pageNumber="16">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34170B0FFF45658FF693946FC72E14C" box="[151,1001,1619,1642]" pageId="1" pageNumber="16" type="description">
<paragraph id="8BE4233BFFF45658FF693946FC72E14C" blockId="1.[151,1001,1619,1642]" box="[151,1001,1619,1642]" pageId="1" pageNumber="16">
<emphasis id="B92FFF29FFF45658FF693946FE00E14C" box="[151,411,1619,1642]" italics="true" pageId="1" pageNumber="16">
<taxonomicName id="4C5B58B8FFF45658FF693946FF6CE14C" box="[151,247,1619,1642]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="16" phylum="Chordata" rank="genus">Dibamus</taxonomicName>
novae-guineae
</emphasis>
: N. de
<bibRefCitation id="EFCA5ECAFFF45658FE173946FDF9E14C" author="Rooj" box="[489,610,1619,1642]" pageId="1" pageNumber="22" refString="de Rooij, N. (1922) Fauna Simalurensis. Reptilia. Zoologische Mededeelingen, 6, 217 - 238." type="journal article" year="1922">Rooj. 1922</bibRefCitation>
.
<emphasis id="B92FFF29FFF45658FD933946FC02E14C" box="[621,921,1619,1642]" italics="true" pageId="1" pageNumber="16">Zoölogische Mededeelingen</emphasis>
6, 218.
</paragraph>
</subSubSection>
<subSubSection id="C34170B0FFF45658FF69398CFA12E00E" pageId="1" pageNumber="16" type="materials_examined">
<paragraph id="8BE4233BFFF45658FF69398CFA12E00E" blockId="1.[151,1437,1689,2025]" pageId="1" pageNumber="16">
<emphasis id="B92FFF29FFF45658FF69398CFE8EE195" bold="true" box="[151,277,1689,1715]" pageId="1" pageNumber="16">
<typeStatus id="54E09D99FFF45658FF69398CFE8BE195" box="[151,272,1689,1715]" pageId="1" pageNumber="16" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED4ABBFEFFF45658FEE3398FFEC4E192" box="[285,351,1690,1716]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="1" pageNumber="16">ZMA</collectionCode>
15500, Sinabang (
<geoCoordinate id="EE6F45FCFFF45658FDBA398FFD4DE192" box="[580,726,1690,1716]" direction="north" orientation="latitude" pageId="1" pageNumber="16" precision="15" value="2.4833333">02º28’60&quot;N</geoCoordinate>
,
<geoCoordinate id="EE6F45FCFFF45658FD1A398FFCE9E192" box="[740,882,1690,1716]" direction="east" orientation="longitude" pageId="1" pageNumber="16" precision="15" value="96.38334">96º22’60&quot;E</geoCoordinate>
; 0 m asl),
<collectingCountry id="F34C63ABFFF45658FC0C398FFBADE192" box="[1010,1078,1690,1716]" name="Iceland" pageId="1" pageNumber="16">Pulau</collectingCountry>
Simeuleu, Daerah Istimewah Aceh Propinsi,
<collectingCountry id="F34C63ABFFF45658FEB239D5FDD5E1FC" box="[332,590,1728,1754]" name="Indonesia" pageId="1" pageNumber="16">Republic of Indonesia</collectingCountry>
(
<figureCitation id="13603FBEFFF45658FDA339D5FD3FE1FC" box="[605,676,1728,1754]" captionStart="FIGURE 5. A" captionStartId="5.[151,257,1547,1571]" captionTargetBox="[325,1261,777,1518]" captionTargetId="figure@5.[325,1261,777,1193]" captionTargetPageId="5" captionText="FIGURE 5. A. Map of South-east Asia (top), showing location of northern Sumatra, the Mentawai Archipelago and other islands off south-west Sumatra; area enlarged indicated. B. Map of northern Sumatra (bottom), showing the location of Pulau Simeuleu, the type locality of Dibamus tebal sp. nov." httpUri="https://zenodo.org/record/187434/files/figure.png" pageId="1" pageNumber="16">Fig. 5</figureCitation>
); sex: adult male, collector: Edward Richard Jacobson; sampling details unavailable, except that a paper label associated with the specimen bears the pencilled date &quot;
<date id="FFE505FBFFF45658FF5E381BFE87E00E" box="[160,284,1806,1832]" bridgedPair="-" pageId="1" pageNumber="16" value="1913-07">VII–1913</date>
&quot;. Jacobson is known to have visited Simeulue in 1913 (see
<bibRefCitation id="EFCA5ECAFFF45658FC2C381BFB25E00E" author="Fransen" box="[978,1214,1806,1832]" pageId="1" pageNumber="22" refString="Fransen, C. H. J. M., Holthuis, L. B. &amp; Adema, J. P. H. M. (1997) Type-catalogue of the Decapoda Crustacea in the collections of the Nationaal Natuurhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Mededeelingen, 311, i - xvi + 1 - 344." type="journal article" year="1997">
Fransen
<emphasis id="B92FFF29FFF45658FBC9381BFBEAE001" box="[1079,1137,1806,1831]" italics="true" pageId="1" pageNumber="16">et al.</emphasis>
, 1997
</bibRefCitation>
; de
<bibRefCitation id="EFCA5ECAFFF45658FB13381BFAE7E00E" author="Rooij" box="[1261,1404,1806,1832]" pageId="1" pageNumber="22" refString="de Rooij, N. (1922) Fauna Simalurensis. Reptilia. Zoologische Mededeelingen, 6, 217 - 238." type="journal article" year="1922">Rooij, 1922</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34170B0FFF4565BFF3B3821FD1BE465" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16" type="diagnosis">
<paragraph id="8BE4233BFFF4565BFF3B3821FB06E794" blockId="1.[151,1437,1689,2025]" lastBlockId="2.[151,1437,152,835]" lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
<emphasis id="B92FFF29FFF45658FF3B3821FED2E068" bold="true" box="[197,329,1844,1870]" pageId="1" pageNumber="16">Diagnosis:</emphasis>
A mid-sized (SVL
<quantity id="4CA38EDEFFF45658FDC93821FD28E068" box="[567,691,1844,1870]" metricMagnitude="-1" metricUnit="m" metricValue="1.335" pageId="1" pageNumber="16" unit="mm" value="133.5">133.5 mm</quantity>
) species of
<taxonomicName id="4C5B58B8FFF45658FCBC3821FC35E06B" box="[834,942,1844,1869]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B92FFF29FFF45658FCBC3821FC35E06B" box="[834,942,1844,1869]" italics="true" pageId="1" pageNumber="16">Dibamus</emphasis>
</taxonomicName>
, with relatively robust body (body width 4.9% SVL), separable from congeners using the following combination of characters: postoculars two; scales around neck 24; scales around midbody 24; scales around vent 15; ventrals 186; subcaudals 42; frontonasal single; rostral suture incomplete; labial and nasals sutures complete; two pairs of preanal pores; single tibial pore located between two scales on inner distal end of each hind limb; nuchal collar or body band absent; presacral vertebrae 119; postsacral vertebrae 26; and a relatively long tail (18.7% SVL).
</paragraph>
<paragraph id="8BE4233BFFF7565BFF3B3FABFD1BE465" blockId="2.[151,1437,152,835]" pageId="2" pageNumber="17">
In showing two postoculars, the new species differs from
<taxonomicName id="4C5B58B8FFF7565BFC6C3FAAFB6AE7FE" authority="Darevsky, 1992" authorityName="Darevsky" authorityYear="1992" box="[914,1265,190,216]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="bogadeki">
<emphasis id="B92FFF29FFF7565BFC6C3FAAFBB3E7FE" box="[914,1064,191,216]" italics="true" pageId="2" pageNumber="17">D. bogadeki</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFBCF3FABFB6AE7FE" author="Darevsky" box="[1073,1265,190,216]" pageId="2" pageNumber="22" refString="Darevsky, I. S. (1992) Two new species of worm-like lizard Dibamus (Sauria: Dibamidae), with remarks on the distribution and ecology of Dibamus in Vietnam. Asiatic Herpetological Research, 4, 1 - 12." type="journal article" year="1992">Darevsky, 1992</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F34C63ABFFF7565BFAFD3FABFA03E7FE" box="[1283,1432,190,216]" name="Hong Kong" pageId="2" pageNumber="17">Hong Kong</collectingCountry>
, eastern
<collectingCountry id="F34C63ABFFF7565BFF0F3FF0FEA7E7D9" box="[241,316,229,255]" name="China" pageId="2" pageNumber="17">China</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFEAE3FF0FD7AE7D9" authority="Das and Yaakob, 2003" authorityName="Das and Yaakob" authorityYear="2003" box="[336,737,229,255]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="booliati">
<emphasis id="B92FFF29FFF7565BFEAE3FF0FE4BE7D8" box="[336,464,229,254]" italics="true" pageId="2" pageNumber="17">D. booliati</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFE293FF0FD7AE7D9" author="Das" box="[471,737,229,255]" pageId="2" pageNumber="22" refString="Das, I. &amp; Yaakob, N. S. (2003) A new species of Dibamus (Squamata: Dibamidae) from Peninsular Malaysia. Raffles Bulletin of Zoology, 51, 143 - 147." type="journal article" year="2003">Das and Yaakob, 2003</bibRefCitation>
</taxonomicName>
(northern Malay Peninsula),
<taxonomicName id="4C5B58B8FFF7565BFBC53FF0FACCE7D9" authority="Angel, 1935" authorityName="Angel" authorityYear="1935" box="[1083,1367,229,255]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="B92FFF29FFF7565BFBC53FF0FB25E7D8" box="[1083,1214,229,254]" italics="true" pageId="2" pageNumber="17">D. bourreti</emphasis>
Angel, 1935
</taxonomicName>
(Tam Dao,
<collectingCountry id="F34C63ABFFF7565BFF263E19FEA0E600" box="[216,315,268,294]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFEA83E19FD05E600" authority="Ineich, 1999" authorityName="Ineich" authorityYear="1999" box="[342,670,268,294]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="deharvengi">
<emphasis id="B92FFF29FFF7565BFEA83E19FD9BE603" box="[342,512,268,293]" italics="true" pageId="2" pageNumber="17">D. deharvengi</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFDF73E19FD05E600" author="Ineich" box="[521,670,268,294]" pageId="2" pageNumber="22" refString="Ineich, I. (1999) Une nouvelle espece de Dibamus (Reptilia, Squamata, Dibamidae) du Vietnam. Bulletin de la Societe Zoologique de France, 124, 279 - 286." type="journal article" year="1999">Ineich, 1999</bibRefCitation>
</taxonomicName>
(Binh Châu,
<collectingCountry id="F34C63ABFFF7565BFCBF3E19FC31E600" box="[833,938,268,294]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFC413E19FAE8E600" authority="Darevsky, 1992" authorityName="Darevsky" authorityYear="1992" box="[959,1395,268,294]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="greeri">
<emphasis id="B92FFF29FFF7565BFC413E19FBB7E603" box="[959,1068,268,293]" italics="true" pageId="2" pageNumber="17">D. greeri</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFBCA3E19FB6BE600" author="Darevsky" box="[1076,1264,268,294]" pageId="2" pageNumber="22" refString="Darevsky, I. S. (1992) Two new species of worm-like lizard Dibamus (Sauria: Dibamidae), with remarks on the distribution and ecology of Dibamus in Vietnam. Asiatic Herpetological Research, 4, 1 - 12." type="journal article" year="1992">Darevsky, 1992</bibRefCitation>
(
<collectingCountry id="F34C63ABFFF7565BFB013E19FAF1E600" box="[1279,1386,268,294]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="4C5B58B8FFF7565BFA813E19FF67E66D" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="leucurus">
<emphasis id="B92FFF29FFF7565BFA813E19FF67E66D" italics="true" pageId="2" pageNumber="17">D. leucurus</emphasis>
</taxonomicName>
(Bleeker, 1860; Sumatra, Borneo, and possibly islands of southern
<collectingCountry id="F34C63ABFFF7565BFBEB3E27FB05E66A" box="[1045,1182,306,332]" name="Philippines" pageId="2" pageNumber="17">Philippines</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFB4F3E27FF49E655" authority="Smith, 1921" authorityName="Smith" authorityYear="1921" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B92FFF29FFF7565BFB4F3E27FADCE66D" box="[1201,1351,306,331]" italics="true" pageId="2" pageNumber="17">D. montanus</emphasis>
Smith, 1921
</taxonomicName>
(Le Bosquet, Langbian Plateau,
<collectingCountry id="F34C63ABFFF7565BFDAD3E4CFD20E655" box="[595,699,345,371]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFD313E4CFBAFE655" authority="Honda et al., 1997" authorityName="Honda et al." authorityYear="1997" box="[719,1076,345,371]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="somsaki">
<emphasis id="B92FFF29FFF7565BFD313E4CFCC9E654" box="[719,850,345,370]" italics="true" pageId="2" pageNumber="17">D. somsaki</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFCA73E4CFBAFE655" author="Honda" box="[857,1076,345,371]" pageId="2" pageNumber="22" refString="Honda, M., Nabhitabhata, J., Ota, H. &amp; Hikida, T. (1997) A new species of Dibamus (Squamata: Dibamidae) from Thailand. Raffles Bulletin of Zoology, 45, 275 - 279." type="journal article" year="1997">
Honda
<emphasis id="B92FFF29FFF7565BFC513E4CFC71E654" box="[943,1002,345,370]" italics="true" pageId="2" pageNumber="17">et al.</emphasis>
, 1997
</bibRefCitation>
</taxonomicName>
(Khao Soi Dao National Park, Chanthaburi Province,
<collectingCountry id="F34C63ABFFF7565BFE593E95FD8AE6BC" box="[423,529,384,410]" name="Thailand" pageId="2" pageNumber="17">Thailand</collectingCountry>
) and
<taxonomicName id="4C5B58B8FFF7565BFDAD3E95FC4BE6BC" authority="Diaz et al., 2004" authorityName="Diaz et al." authorityYear="2004" box="[595,976,384,410]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="tiomanensis">
<emphasis id="B92FFF29FFF7565BFDAD3E95FC98E6BF" box="[595,771,384,409]" italics="true" pageId="2" pageNumber="17">D. tiomanensis</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFCF53E95FC4BE6BC" author="Diaz" box="[779,976,384,410]" pageId="2" pageNumber="22" refString="Diaz, R. E., Leong, T. - M., Grismer, L. L. &amp; Yaakob, N. S. (2004) A new species of Dibamus (Squamata: Dibamidae) from West Malaysia. Asiatic Herpetological Research, 10, 1 - 7." type="journal article" year="2004">
Diaz
<emphasis id="B92FFF29FFF7565BFCB43E95FCE5E6BF" box="[842,894,384,409]" italics="true" pageId="2" pageNumber="17">et al</emphasis>
., 2004
</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F34C63ABFFF7565BFC213E95FBB8E6BC" box="[991,1059,384,410]" name="Iceland" pageId="2" pageNumber="17">Pulau</collectingCountry>
Tioman, Pahang,
<collectingCountry id="F34C63ABFFF7565BFB043E95FAF1E6BC" box="[1274,1386,384,410]" name="Malaysia" pageId="2" pageNumber="17">Malaysia</collectingCountry>
), all of which have a single postocular. The complete and separate rostral suture distinguishes it from
<taxonomicName id="4C5B58B8FFF7565BFAF73EB2FEF8E6C1" authority="Das &amp; Lim, 2005" authorityName="Das &amp; Lim" authorityYear="2005" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="dezwaani">
<emphasis id="B92FFF29FFF7565BFAF73EB2FA07E6E6" box="[1289,1436,423,448]" italics="true" pageId="2" pageNumber="17">D. dezwaani</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFF693ED8FEF8E6C1" author="Das" box="[151,355,461,487]" pageId="2" pageNumber="22" refString="Das, I. &amp; Lim, K. K. P. (2005) New species of Dibamus (Squamata: Dibamidae) from Pulau Nias, Indonesia. Journal of Herpetology, 39, 113 - 117." type="journal article" year="2005">Das &amp; Lim, 2005</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F34C63ABFFF7565BFE8F3ED8FE2DE6C1" box="[369,438,461,487]" name="Iceland" pageId="2" pageNumber="17">Pulau</collectingCountry>
Nias,
<collectingCountry id="F34C63ABFFF7565BFDFE3ED8FDECE6C1" box="[512,631,461,487]" name="Indonesia" pageId="2" pageNumber="17">Indonesia</collectingCountry>
). The complete nasal suture separates the new species from
<taxonomicName id="4C5B58B8FFF7565BFACF3ED8FEBFE528" authority="Greer, 1985" authorityName="Greer" authorityYear="1985" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="smithi">
<emphasis id="B92FFF29FFF7565BFACF3ED8FA07E6C0" box="[1329,1436,461,486]" italics="true" pageId="2" pageNumber="17">D. smithi</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFF693EE1FEBFE528" author="Greer" box="[151,292,500,526]" pageId="2" pageNumber="22" refString="Greer, A. E. (1985) The relationships of the lizard genera Anelytropsis and Dibamus. Journal of Herpetology, 19, 116 - 156." type="journal article" year="1985">Greer, 1985</bibRefCitation>
</taxonomicName>
(Daban, Langbian Plateau,
<collectingCountry id="F34C63ABFFF7565BFD8D3EE1FD41E528" box="[627,730,500,526]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
) and
<taxonomicName id="4C5B58B8FFF7565BFCE23EE1FB72E528" authority="Das and Lim, 2003 (Borneo)" authorityName="Das and Lim, 2003 (Borneo" box="[796,1257,500,526]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="vorisi">
<emphasis id="B92FFF29FFF7565BFCE23EE1FC1EE52B" box="[796,901,500,525]" italics="true" pageId="2" pageNumber="17">D. vorisi</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFC733EE1FBE9E528" author="Das" box="[909,1138,500,526]" pageId="2" pageNumber="22" refString="Das, I. &amp; Lim, K. K. P. (2003) Two new species of Dibamus (Squamata: Dibamidae) from Borneo. Raffles Bulletin of Zoology, 51, 137 - 141." type="journal article" year="2003">Das and Lim, 2003</bibRefCitation>
(Borneo)
</taxonomicName>
, both of which show reduced nasal sutures. Finally, the contact of the first infralabial with three scales differentiates it from
<taxonomicName id="4C5B58B8FFF7565BFF693D54FC9CE57D" authority="Dumeril and Bibron, 1839" authorityName="Dumeril and Bibron" authorityYear="1839" box="[151,775,577,603]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="B92FFF29FFF7565BFF693D54FE23E57C" box="[151,440,577,602]" italics="true" pageId="2" pageNumber="17">Dibamus novaeguineae</emphasis>
Duméril and Bibron, 1839
</taxonomicName>
(western New
<collectingCountry id="F34C63ABFFF7565BFC3A3D54FBBAE57D" box="[964,1057,577,603]" name="Guinea-Bissau" pageId="2" pageNumber="17">Guinea</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFBC63D54FB62E57C" box="[1080,1273,577,602]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="nicobaricum">
<emphasis id="B92FFF29FFF7565BFBC63D54FB62E57C" box="[1080,1273,577,602]" italics="true" pageId="2" pageNumber="17">D. nicobaricum</emphasis>
</taxonomicName>
(Fitzinger in Steindachner, 1867) (
<collectingCountry id="F34C63ABFFF7565BFE593D7DFD94E5A4" box="[423,527,616,642]" name="India" pageId="2" pageNumber="17">Nicobar</collectingCountry>
Archipelago,
<collectingCountry id="F34C63ABFFF7565BFD3B3D7DFC97E5A4" box="[709,780,616,642]" name="India" pageId="2" pageNumber="17">India</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFCDD3D7DFBDBE5A4" authority="Greer, 1985" authorityName="Greer" authorityYear="1985" box="[803,1088,616,642]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B92FFF29FFF7565BFCDD3D7DFC04E5A7" box="[803,927,616,641]" italics="true" pageId="2" pageNumber="17">D. taylori</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFC573D7DFBDBE5A4" author="Greer" box="[937,1088,616,642]" pageId="2" pageNumber="22" refString="Greer, A. E. (1985) The relationships of the lizard genera Anelytropsis and Dibamus. Journal of Herpetology, 19, 116 - 156." type="journal article" year="1985">Greer, 1985</bibRefCitation>
</taxonomicName>
(Sumba,
<collectingCountry id="F34C63ABFFF7565BFB453D7DFA95E5A4" box="[1211,1294,616,642]" name="Indonesia" pageId="2" pageNumber="17">Flores</collectingCountry>
, Wetar and Lombok),
<taxonomicName id="4C5B58B8FFF7565BFEE53D9AFC9DE58E" authority="Das and Lim, 2003 (Borneo)" authorityName="Das and Lim, 2003 (Borneo" box="[283,774,654,680]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="B92FFF29FFF7565BFEE53D9AFE16E58E" box="[283,397,655,680]" italics="true" pageId="2" pageNumber="17">D. ingeri</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFE683D9BFD13E58E" author="Das" box="[406,648,654,680]" pageId="2" pageNumber="22" refString="Das, I. &amp; Lim, K. K. P. (2003) Two new species of Dibamus (Squamata: Dibamidae) from Borneo. Raffles Bulletin of Zoology, 51, 137 - 141." type="journal article" year="2003">Das and Lim, 2003</bibRefCitation>
(Borneo)
</taxonomicName>
,
<taxonomicName id="4C5B58B8FFF7565BFCED3D9AFB52E58E" authority="Honda et al., 2001" authorityName="Honda et al." authorityYear="2001" box="[787,1225,654,680]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="kondaoensis">
<emphasis id="B92FFF29FFF7565BFCED3D9AFC4EE58E" box="[787,981,655,680]" italics="true" pageId="2" pageNumber="17">D. kondaoensis</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFC203D9BFB52E58E" author="Honda" box="[990,1225,654,680]" pageId="2" pageNumber="22" refString="Honda, M., Ota, H., Hikida, T. &amp; Darevsky, I. S. (2001) A new species of the worm-like lizard, Dibamus Dumeril &amp; Bibron 1839 (Squamata Dibamidae), from Vietnam. Tropical Zoology, 14, 119 - 125." type="journal article" year="2001">
Honda
<emphasis id="B92FFF29FFF7565BFBC73D9AFBE1E58E" box="[1081,1146,655,680]" italics="true" pageId="2" pageNumber="17">et al.</emphasis>
, 2001
</bibRefCitation>
</taxonomicName>
(Kondao Island,
<collectingCountry id="F34C63ABFFF7565BFF693DA0FF64E5E9" box="[151,255,693,719]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
),
<taxonomicName id="4C5B58B8FFF7565BFEED3DA0FDFDE5E9" authority="Schlegel, 1858" authorityName="Schlegel" authorityYear="1858" box="[275,614,693,719]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="celebensis">
<emphasis id="B92FFF29FFF7565BFEED3DA0FE2BE5E8" box="[275,432,693,718]" italics="true" pageId="2" pageNumber="17">D. celebensis</emphasis>
Schlegel, 1858
</taxonomicName>
(Sulawesi, eastern
<collectingCountry id="F34C63ABFFF7565BFCB43DA0FC25E5E9" box="[842,958,693,719]" name="Indonesia" pageId="2" pageNumber="17">Indonesia</collectingCountry>
) and
<taxonomicName id="4C5B58B8FFF7565BFC013DA0FAA2E5E9" authority="Greer, 1985" authorityName="Greer" authorityYear="1985" box="[1023,1337,693,719]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="seramensis">
<emphasis id="B92FFF29FFF7565BFC013DA0FB3DE5E8" box="[1023,1190,693,718]" italics="true" pageId="2" pageNumber="17">D. seramensis</emphasis>
<bibRefCitation id="EFCA5ECAFFF7565BFB533DA0FAA2E5E9" author="Greer" box="[1197,1337,693,719]" pageId="2" pageNumber="22" refString="Greer, A. E. (1985) The relationships of the lizard genera Anelytropsis and Dibamus. Journal of Herpetology, 19, 116 - 156." type="journal article" year="1985">Greer, 1985</bibRefCitation>
</taxonomicName>
(Seram, Makulu, eastern
<collectingCountry id="F34C63ABFFF7565BFEA43DC9FE4AE5D0" box="[346,465,732,758]" name="Indonesia" pageId="2" pageNumber="17">Indonesia</collectingCountry>
), all of which have a single scale bounding Infralabial I. In showing a thick body, at ca. 5% SVL, the new species can be differentiated from congeners, which show a range of 2.4–4.4%, except
<taxonomicName id="4C5B58B8FFF7565BFF693C3CFE9AE464" box="[151,257,809,834]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="greeri">
<emphasis id="B92FFF29FFF7565BFF693C3CFE9AE464" box="[151,257,809,834]" italics="true" pageId="2" pageNumber="17">D. greeri</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCA5ECAFFF7565BFEEE3C3CFDE8E465" author="Darevsky" box="[272,627,809,835]" pageId="2" pageNumber="22" refString="Darevsky, I. S. (1992) Two new species of worm-like lizard Dibamus (Sauria: Dibamidae), with remarks on the distribution and ecology of Dibamus in Vietnam. Asiatic Herpetological Research, 4, 1 - 12." type="journal article" year="1992">Darevsky, 1992; Das, unpubl.</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2473B3FFF7565BFF693837FB9DE01F" box="[151,1030,1826,1850]" httpUri="https://zenodo.org/record/187430/files/figure.png" pageId="2" pageNumber="17" targetBox="[151,1434,879,1800]" targetPageId="2">
<paragraph id="8BE4233BFFF7565BFF693837FB9DE01F" blockId="2.[151,1030,1826,1850]" box="[151,1030,1826,1850]" pageId="2" pageNumber="17">
<emphasis id="B92FFF29FFF7565BFF693837FE82E01C" bold="true" box="[151,281,1826,1850]" pageId="2" pageNumber="17">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="4C5B58B8FFF7565BFE583837FDD9E01F" box="[422,578,1826,1849]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="tebal" status="sp. nov.">
<emphasis id="B92FFF29FFF7565BFE583837FDD9E01F" box="[422,578,1826,1849]" italics="true" pageId="2" pageNumber="17">Dibamus tebal</emphasis>
</taxonomicName>
<emphasis id="B92FFF29FFF7565BFDB73837FD04E01C" bold="true" box="[585,671,1826,1850]" pageId="2" pageNumber="17">
<taxonomicNameLabel id="A21C4252FFF7565BFDB73837FD04E01C" box="[585,671,1826,1850]" pageId="2" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ZMA 15500). Marker = 10 mm.
</paragraph>
</caption>
<subSubSection id="C34170B0FFF7565CFF3B3872FCD7E068" lastPageId="5" lastPageNumber="20" pageId="2" pageNumber="17" type="description">
<paragraph id="8BE4233BFFF7565CFF3B3872FBA9E7D9" blockId="2.[151,1436,1895,2037]" lastBlockId="5.[151,1436,152,719]" lastPageId="5" lastPageNumber="20" pageId="2" pageNumber="17">
<emphasis id="B92FFF29FFF7565BFF3B3872FEC4E0A7" bold="true" box="[197,351,1895,1921]" pageId="2" pageNumber="17">Description:</emphasis>
SVL
<quantity id="4CA38EDEFFF7565BFE5A3872FD85E0A7" box="[420,542,1895,1921]" metricMagnitude="-1" metricUnit="m" metricValue="1.335" pageId="2" pageNumber="17" unit="mm" value="133.5">133.5 mm</quantity>
, TL
<quantity id="4CA38EDEFFF7565BFDAD3872FD25E0A7" box="[595,702,1895,1921]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="2" pageNumber="17" unit="mm" value="24.9">24.9 mm</quantity>
; snout distinctly acute, slightly rounded, distinctly conical (IN/ IO ratio 0.52), projecting well beyond lower jaws; nostril laterally oriented, oval, situated closer to tip of snout than to orbit (E-N/E-S ratio 0.66); head wider than long (HL/HW ratio 0.81), not flattened (HL/HD ratio 0.89); rostral pad single, with a large number of evenly distributed sensory papillae; rostral sutures incomplete, traversing about half the distance between the edge of the maxilla and nostrils; nasal and labial sutures complete, extending from ocular to nostril; posterior border of rostral curved; frontonasal single, width
<quantity id="4CA38EDEFFF1565DFF1F382FFEA5E072" box="[225,318,1850,1876]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="19" unit="mm" value="2.1">2.1 mm</quantity>
, depth
<quantity id="4CA38EDEFFF1565DFE6F382FFE75E072" box="[401,494,1850,1876]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="19" unit="mm" value="0.7">0.7 mm</quantity>
; anterior frontonasal suture with a sinous border; frontal single, width 2.8, depth
<quantity id="4CA38EDEFFF1565DFF693874FF6FE05D" box="[151,244,1889,1915]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="19" unit="mm" value="1.6">1.6 mm</quantity>
, x 2.3 times wider than frontonasal; interparietal single, not enlarged relative to scales on the sides or posteriorly, narrower (width
<quantity id="4CA38EDEFFF1565DFE18389DFDD9E084" box="[486,578,1928,1954]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="19" unit="mm" value="0.1">0.1 mm</quantity>
) than frontonasal (width
<quantity id="4CA38EDEFFF1565DFC96389DFC5FE084" box="[872,964,1928,1954]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="19" unit="mm" value="0.7">0.7 mm</quantity>
) and frontal (width
<quantity id="4CA38EDEFFF1565DFB52389DFA92E084" box="[1196,1289,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="19" unit="mm" value="1.4">1.4 mm</quantity>
), posteriorly bordered by three subequal nuchal scales; postoculars two; supralabial single, elongate, bordering ocular ventrally; first infralabial lanceolate, length
<quantity id="4CA38EDEFFF1565DFD6738C0FD69E0C9" box="[665,754,2005,2031]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="19" unit="mm" value="3.3">3.3 mm</quantity>
(infralabial length/HW ratio 0.69), the pair separated by a smaller, elongate trapezoid mental; three scales border posterior edge of first infralabial; eyes dimly visible through ocular scale; ocular scale deeper than anterior body scales; tongue broad, fleshy and covered with transverse lamellae; tongue tip lacking lamellae, unnotched; teeth small, acute.
</paragraph>
<caption id="DF2473B3FFF6565AFF693862FE6DE097" httpUri="https://zenodo.org/record/187431/files/figure.png" pageId="3" pageNumber="18" targetBox="[315,1271,194,1885]" targetPageId="3">
<paragraph id="8BE4233BFFF6565AFF693862FE6DE097" blockId="3.[151,1436,1911,1969]" pageId="3" pageNumber="18">
<emphasis id="B92FFF29FFF6565AFF693862FE85E0A9" bold="true" box="[151,286,1911,1935]" pageId="3" pageNumber="18">FIGURE 2.</emphasis>
Head of holotype of
<taxonomicName id="4C5B58B8FFF6565AFDF03862FD2BE0A8" box="[526,688,1911,1934]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="18" phylum="Chordata" rank="species" species="tebal" status="sp. nov.">
<emphasis id="B92FFF29FFF6565AFDF03862FD2BE0A8" box="[526,688,1911,1934]" italics="true" pageId="3" pageNumber="18">Dibamus tebal</emphasis>
</taxonomicName>
<emphasis id="B92FFF29FFF6565AFD463862FC8AE0A9" bold="true" box="[696,785,1911,1935]" pageId="3" pageNumber="18">
<taxonomicNameLabel id="A21C4252FFF6565AFD463862FC8AE0A9" box="[696,785,1911,1935]" pageId="3" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ZMA 15500) in dorsal (top), lateral (middle) and ventral (bottom) views. Marker = 2 mm.
</paragraph>
</caption>
<caption id="DF2473B3FFF1565DFF693D9FFF53E5E2" httpUri="https://zenodo.org/record/187432/files/figure.png" pageId="4" pageNumber="19" targetBox="[449,1136,186,625]" targetPageId="4">
<paragraph id="8BE4233BFFF1565DFF693D9FFF53E5E2" blockId="4.[151,1436,650,708]" pageId="4" pageNumber="19">
<emphasis id="B92FFF29FFF1565DFF693D9FFE82E584" bold="true" box="[151,281,650,674]" pageId="4" pageNumber="19">FIGURE 3.</emphasis>
Anal region of holotype of
<taxonomicName id="4C5B58B8FFF1565DFDBC3D9EFD45E584" box="[578,734,651,674]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="19" phylum="Chordata" rank="species" species="tebal" status="sp. nov.">
<emphasis id="B92FFF29FFF1565DFDBC3D9EFD45E584" box="[578,734,651,674]" italics="true" pageId="4" pageNumber="19">Dibamus tebal</emphasis>
</taxonomicName>
<emphasis id="B92FFF29FFF1565DFD1A3D9FFCA1E584" bold="true" box="[740,826,650,674]" pageId="4" pageNumber="19">
<taxonomicNameLabel id="A21C4252FFF1565DFD1A3D9FFCA1E584" box="[740,826,650,674]" pageId="4" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ZMA 15500), showing flap-like hind limbs. Marker = 1 mm.
</paragraph>
</caption>
<caption id="DF2473B3FFF1565DFF6939B2FC46E198" box="[151,989,1703,1727]" httpUri="https://zenodo.org/record/187433/files/figure.png" pageId="4" pageNumber="19" targetBox="[268,1318,746,1678]" targetPageId="4">
<paragraph id="8BE4233BFFF1565DFF6939B2FC46E198" blockId="4.[151,989,1703,1727]" box="[151,989,1703,1727]" pageId="4" pageNumber="19">
<emphasis id="B92FFF29FFF1565DFF6939B2FE82E199" bold="true" box="[151,281,1703,1727]" pageId="4" pageNumber="19">FIGURE 4.</emphasis>
Radiograph of holotype of
<taxonomicName id="4C5B58B8FFF1565DFDBE39BDFD47E199" box="[576,732,1704,1727]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="19" phylum="Chordata" rank="species" species="tebal" status="sp. nov.">
<emphasis id="B92FFF29FFF1565DFDBE39BDFD47E199" box="[576,732,1704,1727]" italics="true" pageId="4" pageNumber="19">Dibamus tebal</emphasis>
</taxonomicName>
<emphasis id="B92FFF29FFF1565DFD1D39B2FCA2E199" bold="true" box="[739,825,1703,1727]" pageId="4" pageNumber="19">
<taxonomicNameLabel id="A21C4252FFF1565DFD1D39B2FCA2E199" box="[739,825,1703,1727]" pageId="4" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(ZMA 15500).
</paragraph>
</caption>
<paragraph id="8BE4233BFFF0565CFF3B3E19FD12E5E9" blockId="5.[151,1436,152,719]" pageId="5" pageNumber="20">
Body relatively robust, BW
<quantity id="4CA38EDEFFF0565CFDE53E19FDECE600" box="[539,631,268,294]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="20" unit="mm" value="6.6">6.6 mm</quantity>
(BW/SVL ratio 0.05); head distinct from neck; tail relatively long (TL/SVL ratio 0.19), tip rounded, not wider than rest of tail; tail base slightly thickened (TW/TL ratio 0.21); body scales smooth, subcycloid, including near preanal region; body and tail covered with cycloid scales that are unnotched posteriorly; scales on dorsal surface of body larger than those on tail; transverse scale rows posterior to head 25, at midbody 24, and anterior to vent 15; ventrals 186; subcaudals 42; presacral vertebrae 119; postsacral vertebrae 26 (tail complete); hind limbs present, measuring
<quantity id="4CA38EDEFFF0565CFBE43ED8FBEEE6C1" box="[1050,1141,461,487]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="20" unit="mm" value="4.8">4.8 mm</quantity>
in length and
<quantity id="4CA38EDEFFF0565CFADC3ED8FAE6E6C1" box="[1314,1405,461,487]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="20" unit="mm" value="1.6">1.6 mm</quantity>
in width (left) and
<quantity id="4CA38EDEFFF0565CFEAB3EE1FE34E528" box="[341,431,500,526]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="20" unit="mm" value="5.1">5.1 mm</quantity>
in length and
<quantity id="4CA38EDEFFF0565CFDA93EE1FD2AE528" box="[599,689,500,526]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="20" unit="mm" value="2.2">2.2 mm</quantity>
in width (right), comprising reduced (3.59 and 3.82% SVL, for left and right limbs, respectively) flap-like structures on either sides of vent, covered with four paired scales, terminating in a single scale with a rounded tip; a pair of enlarged scales on preanal region, separated by a median scale, overlapping those on sides, followed by three rows of regular-sized scales up to the anal region; preanal pores present; tibial pores located between two scales on inner distal end of hind limb; and postanal scales not reduced, relative to body scales.
</paragraph>
<caption id="DF2473B3FFF0565CFF69391EFC15E14E" httpUri="https://zenodo.org/record/187434/files/figure.png" pageId="5" pageNumber="20" targetBox="[325,1261,777,1518]" targetPageId="5">
<paragraph id="8BE4233BFFF0565CFF69391EFC15E14E" blockId="5.[151,1436,1547,1640]" pageId="5" pageNumber="20">
<emphasis id="B92FFF29FFF0565CFF69391EFE86E105" bold="true" box="[151,285,1547,1571]" pageId="5" pageNumber="20">FIGURE 5.</emphasis>
A. Map of South-east Asia (top), showing location of northern Sumatra, the Mentawai Archipelago and other islands off south-west Sumatra; area enlarged indicated. B. Map of northern Sumatra (bottom), showing the location of Pulau Simeuleu, the type locality of
<taxonomicName id="4C5B58B8FFF0565CFD6B3944FCAAE14E" box="[661,817,1617,1640]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="20" phylum="Chordata" rank="species" species="tebal" status="sp. nov.">
<emphasis id="B92FFF29FFF0565CFD6B3944FCAAE14E" box="[661,817,1617,1640]" italics="true" pageId="5" pageNumber="20">Dibamus tebal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C4252FFF0565CFCC63945FC15E14E" box="[824,910,1616,1640]" pageId="5" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BE4233BFFF0565CFF3B398CFE25E1FC" blockId="5.[151,1437,1689,2025]" pageId="5" pageNumber="20">
<emphasis id="B92FFF29FFF0565CFF3B398CFE63E195" bold="true" box="[197,504,1689,1715]" pageId="5" pageNumber="20">Measurements (in mm):</emphasis>
SVL 133.5; BW 6.6; TL 24.9; TW 5.3; E-N 2.1; E-S 3.2; IN 1.6; IO 3.1; HL 3.9; HW 4.8 and HD 4.4.
</paragraph>
<paragraph id="8BE4233BFFF0565CFF3B39F2FCD7E068" blockId="5.[151,1437,1689,2025]" pageId="5" pageNumber="20">
<emphasis id="B92FFF29FFF0565CFF3B39F2FEFDE027" bold="true" box="[197,358,1767,1793]" pageId="5" pageNumber="20">Colouration:</emphasis>
Colour in life unknown. After nearly a century in preservative, dorsum is a discoloured pale brown, each scale centre dark; lacking scattered light patches; venter slightly paler than dorsum; and tip of snout, including supralabials, throat and anal region cream.
</paragraph>
</subSubSection>
<subSubSection id="C34170B0FFF0565FFF3B384EFC23E7D9" lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20" type="etymology">
<paragraph id="8BE4233BFFF0565CFF3B384EFAAFE053" blockId="5.[151,1437,1689,2025]" box="[197,1332,1883,1909]" pageId="5" pageNumber="20">
<emphasis id="B92FFF29FFF0565CFF3B384EFECDE053" bold="true" box="[197,342,1883,1909]" pageId="5" pageNumber="20">Etymology:</emphasis>
From the Bahasa
<collectingCountry id="F34C63ABFFF0565CFDD5384EFD06E053" box="[555,669,1883,1909]" name="Indonesia" pageId="5" pageNumber="20">Indonesia</collectingCountry>
/
<collectingCountry id="F34C63ABFFF0565CFD58384EFC8BE053" box="[678,784,1883,1909]" name="Malaysia" pageId="5" pageNumber="20">Malaysia</collectingCountry>
('tebal'), meaning thick, for the robust habitus.
</paragraph>
<paragraph id="8BE4233BFFF0565FFF3B3894FC23E7D9" blockId="5.[151,1437,1689,2025]" lastBlockId="6.[151,1436,152,255]" lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20">
<emphasis id="B92FFF29FFF0565CFF3B3894FE01E0BD" bold="true" box="[197,410,1921,1947]" pageId="5" pageNumber="20">Natural history:</emphasis>
Nothing is on record of the natural history of this species. Species of
<taxonomicName id="4C5B58B8FFF0565CFAFE3897FAF5E0BD" box="[1280,1390,1922,1947]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="B92FFF29FFF0565CFAFE3897FAF5E0BD" box="[1280,1390,1922,1947]" italics="true" pageId="5" pageNumber="20">Dibamus</emphasis>
</taxonomicName>
are fossorial and associated with lowland rainforest leaf litter. They are presumed to be predators of litter arthropods and earthworms (
<bibRefCitation id="EFCA5ECAFFF0565CFDF738DAFD66E0CF" author="Pough" box="[521,765,1999,2025]" pageId="5" pageNumber="22" refString="Pough, F. H., Andrews, R. M., Cadle, J. E., Crump, M. L., Savitzky, A. H. &amp; Wells, K. D. (2001). Herpetology. Second edition. Prentice Hall, Upper Saddle River, New Jersey." type="book" year="2001">
Pough
<emphasis id="B92FFF29FFF0565CFD9B38DAFD3CE0CE" box="[613,679,1999,2024]" italics="true" pageId="5" pageNumber="20">et al.</emphasis>
, 2001
</bibRefCitation>
;
<bibRefCitation id="EFCA5ECAFFF0565CFCF538DAFC7AE0CF" author="Zug" box="[779,993,1999,2025]" pageId="5" pageNumber="22" refString="Zug, G. R., Vitt, L. J. &amp; Caldwell, J. P. (2001) Herpetology. An Introductory Biology of Amphibians and Reptiles. Second Edition. Academic Press, San Diego." type="book" year="2001">
Zug
<emphasis id="B92FFF29FFF0565CFCB938DAFC1AE0CE" box="[839,897,1999,2024]" italics="true" pageId="5" pageNumber="20">et al</emphasis>
., 2001
</bibRefCitation>
). The endemicity of the Mentawai Archipelago, within which
<collectingCountry id="F34C63ABFFF3565FFE293F8DFD81E794" box="[471,538,152,178]" name="Iceland" pageId="6" pageNumber="21">Pulau</collectingCountry>
Simeuleu lies, is here reemphasized with the diagnosis of the new
<taxonomicName id="4C5B58B8FFF3565FFAD53F8DFA0EE797" box="[1323,1429,152,177]" class="Reptilia" family="Dibamidae" genus="Dibamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="B92FFF29FFF3565FFAD53F8DFA0EE797" box="[1323,1429,152,177]" italics="true" pageId="6" pageNumber="21">Dibamus</emphasis>
</taxonomicName>
. A number of vertebrates are now known to be confined to these islands, which lie in close proximity to the island of Sumatra, but separated from it by deep waters (Das, 2005).
</paragraph>
</subSubSection>
</treatment>
</document>
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