169 lines
19 KiB
XML
169 lines
19 KiB
XML
<document id="22E5822DA24F7D6B786EB9ADF4F03F72" ID-DOI="10.5281/zenodo.5328899" ID-ISSN="0253-116X" ID-Zenodo-Dep="5328899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630160147779" checkinUser="carolina" docAuthor="Casale, A., D. W & P, Antalya" docDate="2012" docId="03C63E676804FFA9FF000222FDC901E3" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1111-1127.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Laemostenus (Pristonychus) eggeri Casale & D.W & P 2012, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1120" masterDocId="FFFF461F680CFFA0FFB2033BFFE2035B" masterDocTitle="New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)" masterLastPageNumber="1127" masterPageNumber="1111" pageNumber="1119" updateTime="1699169291185" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="F35051F0059FD546364CDC9D4C6CD236">
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<mods:title id="E4A5879D31D8A948731187413D67D088">New or little known Laemostenus species from the Near and Middle East (Coleoptera, Carabidae: Sphodrini)</mods:title>
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</mods:titleInfo>
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<mods:name id="99947DB22FE0A5591F9238BD27F02DCA" type="personal">
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<mods:role id="4F14394552EB9A5E6A3245159553079D">
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<mods:roleTerm id="EE71B2BFDF9864D81A2C3005BFC56742">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6D6D12933852A4453ACCF05F31B90D95">Casale, A.</mods:namePart>
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</mods:name>
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<mods:name id="9BAA2D9FAF7233E89E1C499AACC7ABDB" type="personal">
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<mods:role id="8019266981735A698B6A01F974F41510">
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<mods:roleTerm id="465ECCA7B8C514255F4173C610A00590">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="016F50CCA40AD21A3BAE7888A86F393D">D. W</mods:namePart>
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</mods:name>
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<mods:name id="35E7258616C085C628E7F53575EAAA8F" type="personal">
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<mods:role id="33BC755BE3A3D8652FB55EBB1AA9FF1E">
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<mods:roleTerm id="C45D66E5DF5BFA67083266744D515898">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9770864A41328E0DF0C59E4E04ADFB3B">P, Antalya</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="25A40EFD66EBCE78225C4CBB2CA67D3A">text</mods:typeOfResource>
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<mods:titleInfo id="520EC037E37ABA50D3C12EEF34DC2BF2">
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<mods:title id="F0E6AC655C636804FA793F27F5E75E97">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="2194BCD4C4C5EBE0F4C93DF0440D233A">
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<mods:date id="3A9818D6B29A96486466A0F1217360FE">2012</mods:date>
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<mods:detail id="E4A3D28F55F83A0E61A1E92ED85C616F" type="pubDate">
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<mods:number id="F26DB9BDD0A51AC944A42C831C8DAA05">2012-12-28</mods:number>
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</mods:detail>
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<mods:detail id="0779BF4A7E1922B621C2F878C8DE0A9E" type="volume">
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<mods:number id="C46ED2D96234F994214998C749D25091">44</mods:number>
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</mods:detail>
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<mods:detail id="8D1B57C1B42A8449CC590419396EA03F" type="issue">
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<mods:number id="C149CB2AE27ACB4C41AFB1DA4BB61149">2</mods:number>
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</mods:detail>
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<mods:extent id="6EDEF9397BF94414067464306E72CBC2" unit="page">
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<mods:start id="190A2B852133001DD06B8164EA2E8F06">1111</mods:start>
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<mods:end id="2D7E0F3A00AE522A3814F055659D3D14">1127</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="25D9ADAF67EA10E0BAA88BC0E7C4BF98">journal article</mods:classification>
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<mods:identifier id="20D4B64BA204BBF1D501287A589F5DF5" type="DOI">10.5281/zenodo.5328899</mods:identifier>
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<mods:identifier id="F3DA84FCB4716F5549C8DAE08B2465FF" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="5314C4AF8938BE58D682A0188ECBDB42" type="Zenodo-Dep">5328899</mods:identifier>
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</mods:mods>
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<treatment id="03C63E676804FFA9FF000222FDC901E3" LSID="urn:lsid:plazi:treatment:03C63E676804FFA9FF000222FDC901E3" httpUri="http://treatment.plazi.org/id/03C63E676804FFA9FF000222FDC901E3" lastPageId="9" lastPageNumber="1120" pageId="8" pageNumber="1119">
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<subSubSection id="C375DCFA6804FFA8FF000222FDDD02D7" pageId="8" pageNumber="1119" type="nomenclature">
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<paragraph id="8BD08F716804FFA8FF000222FD970275" blockId="8.[178,629,280,302]" box="[178,629,280,302]" pageId="8" pageNumber="1119">
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<heading id="D098381D6804FFA8FF000222FD970275" box="[178,629,280,302]" fontSize="8" level="2" pageId="8" pageNumber="1119" reason="2">
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<emphasis id="B91B53636804FFA8FF000222FD970275" bold="true" box="[178,629,280,302]" pageId="8" pageNumber="1119">
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<taxonomicName id="4C6FF4F26804FFA8FF000222FDC00275" ID-CoL="8MJ8C" authority="Casale & D.W & P, 2012" authorityName="Casale & D.W & P" authorityYear="2012" box="[178,546,280,302]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1119" phylum="Arthropoda" rank="species" species="eggeri" status="sp. nov." subGenus="Pristonychus">
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<emphasis id="B91B53636804FFA8FF000222FEDE0275" bold="true" box="[178,316,281,302]" italics="true" pageId="8" pageNumber="1119">Laemostenus</emphasis>
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(
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<emphasis id="B91B53636804FFA8FEF80223FE300275" bold="true" box="[330,466,280,302]" italics="true" pageId="8" pageNumber="1119">Pristonychus</emphasis>
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)
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<emphasis id="B91B53636804FFA8FE500222FDC00275" bold="true" box="[482,546,281,302]" italics="true" pageId="8" pageNumber="1119">eggeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A228EE186804FFA8FD9A0223FD970275" box="[552,629,280,302]" pageId="8" pageNumber="1119" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD08F716804FFA8FF000273FDDD02D7" blockId="8.[178,1087,328,396]" pageId="8" pageNumber="1119">
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T y p e m a t e r i a l:
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<typeStatus id="54D431D36804FFA8FE1B0273FE180207" box="[425,506,328,348]" pageId="8" pageNumber="1119" type="holotype">Holotype</typeStatus>
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, labelled: "
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<collectingCountry id="F378CFE16804FFA8FDCB0273FD2E0207" box="[633,716,328,348]" name="Syria" pageId="8" pageNumber="1119">SYRIEN</collectingCountry>
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, nord-west
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<collectingRegion id="49AB41936804FFA8FC870273FC950207" box="[821,887,328,348]" country="Syria" name="Halab" pageId="8" pageNumber="1119">Aleppo</collectingRegion>
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, ca.
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<quantity id="4C9722946804FFA8FC100273FBE70207" box="[930,1029,328,348]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="8" pageNumber="1119" unit="m" value="525.0" valueMax="550.0" valueMin="500.0">500-550 m</quantity>
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, umg. HARIM u. BASHMISHLI, I. Puchner 1.V.-20.V.99" (cWR).
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<typeStatus id="54D431D36804FFA8FCB5025BFCBD022F" box="[775,863,352,372]" pageId="8" pageNumber="1119" type="paratype">Paratypes</typeStatus>
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: 12, 15, same as
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<typeStatus id="54D431D36804FFA8FF6D0243FEC902D7" box="[223,299,376,396]" pageId="8" pageNumber="1119" type="holotype">holotype</typeStatus>
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(BOL, cCA, cEG, cHE, cWR).
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</paragraph>
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</subSubSection>
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<subSubSection id="C375DCFA6804FFA9FF0002A0FDC901E3" lastPageId="9" lastPageNumber="1120" pageId="8" pageNumber="1119" type="description">
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<paragraph id="8BD08F716804FFA8FF0002A0FBEF0295" blockId="8.[178,1085,411,462]" pageId="8" pageNumber="1119">
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D i a g n o s i s: Amedium-sized,piceousblack
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<taxonomicName id="4C6FF4F26804FFA8FD0102A0FC3302EB" authority="" box="[691,977,411,433]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1119" phylum="Arthropoda" rank="subGenus" subGenus="Pristonychus">
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<emphasis id="B91B53636804FFA8FD0102A0FCD502EB" box="[691,823,411,432]" italics="true" pageId="8" pageNumber="1119">Laemostenus</emphasis>
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(
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<emphasis id="B91B53636804FFA8FCF402A0FC2802EA" box="[838,970,411,433]" italics="true" pageId="8" pageNumber="1119">Pristonychus</emphasis>
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)
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</taxonomicName>
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species of the
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<taxonomicName id="4C6FF4F26804FFA8FF650282FEA80295" baseAuthorityName="Herbst" baseAuthorityYear="1784" box="[215,330,440,462]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1119" phylum="Arthropoda" rank="species" species="terricola">
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<emphasis id="B91B53636804FFA8FF650282FEA80295" box="[215,330,440,462]" italics="true" pageId="8" pageNumber="1119">L. terricola</emphasis>
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</taxonomicName>
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species group (in the widest sense of
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<bibRefCitation id="EFFEF2806804FFA8FD780283FCBA0295" author="CASALE A" box="[714,856,440,462]" pageId="8" pageNumber="1119" refId="ref8179" refString="CASALE A. (1988): Revisione degli Sphodrina (Coleoptera, Carabidae, Sphodrini). - Museo regionale di Scienze naturali, Torino, Monografie 5: 1024 pp." type="book" year="1988">CASALE 1988</bibRefCitation>
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). Habitus
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<figureCitation id="135493F46804FFA8FC0D0283FBEF0295" box="[959,1037,440,462]" captionStart="Figs 22-25" captionStartId="16.[178,218,1506,1526]" captionTargetBox="[266,976,232,1475]" captionTargetId="figure-64@16.[178,1085,212,1495]" captionTargetPageId="16" captionText="Figs 22-25: Laemostenus, habitus. (22) L. croyi nov.sp., paratype. (23) L. phoenicius nov.sp., paratype (Maїfouq). (24) L. eggeri nov.sp., paratype. (25) L. bellicosus nov.sp., holotype (various magnifications)." figureDoi="http://doi.org/10.5281/zenodo.5328905" httpUri="https://zenodo.org/record/5328905/files/figure.png" pageId="8" pageNumber="1119">Fig. 24.</figureCitation>
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</paragraph>
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<paragraph id="8BD08F716804FFA8FF0002E5FDA00175" blockId="8.[178,1085,477,558]" pageId="8" pageNumber="1119">
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E t y m o l o g y: Dedicated to our friend and active coleopterist Manfred Egger (Wattens,
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<collectingCountry id="F378CFE16804FFA8FF5A02C1FED4014B" box="[232,310,506,528]" name="Austria" pageId="8" pageNumber="1119">Austria</collectingCountry>
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), with great experiences in collecting cave beetles who kindly offered to us for study the specimens described here.
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</paragraph>
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<paragraph id="8BD08F716804FFA8FF000105FDF7012B" blockId="8.[178,1085,573,624]" pageId="8" pageNumber="1119">
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D e s c r i p t i o n: Body medium-sized (TL: 14.0-
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<quantity id="4C9722946804FFA8FD6D0106FCA20108" box="[735,832,573,595]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="8" pageNumber="1119" unit="mm" value="16.5">16.5 mm</quantity>
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,
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<typeStatus id="54D431D36804FFA8FCE20106FC4B0108" box="[848,937,573,595]" pageId="8" pageNumber="1119" type="holotype">holotype</typeStatus>
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<quantity id="4C9722946804FFA8FC070106FBF50108" box="[949,1047,573,595]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="1119" unit="mm" value="15.5">15.5 mm</quantity>
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; L: 15.0-18.0 mm,
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<typeStatus id="54D431D36804FFA8FEFF0161FE44012B" box="[333,422,602,624]" pageId="8" pageNumber="1119" type="holotype">holotype</typeStatus>
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<quantity id="4C9722946804FFA8FE1E0161FDE8012B" box="[428,522,602,624]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="8" pageNumber="1119" unit="mm" value="16.8">16.8 mm</quantity>
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).
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</paragraph>
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<paragraph id="8BD08F716804FFA8FF000144FE3401E8" blockId="8.[178,1085,639,691]" pageId="8" pageNumber="1119">Colour: Piceous black; labrum, mandibles and palpomeres brown-blackish. Surface shining, elytra not sericeous.</paragraph>
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<paragraph id="8BD08F716804FFA8FF0001F9FDF301AE" blockId="8.[178,1085,706,757]" pageId="8" pageNumber="1119">Microsculpure: Head and pronotum with transverse, very indistinct microlines, elytra with distinct, isodiametric meshes.</paragraph>
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<paragraph id="8BD08F716804FFA8FF00003EFEC2000E" blockId="8.[178,1086,773,853]" pageId="8" pageNumber="1119">Head small, narrow, smooth or with very shallow transverse wrinkles; eyes small, shorter than tempora, but prominent outside; frontal impressions very short, narrow, moderately impressed.</paragraph>
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<paragraph id="8BD08F716804FFA8FF00005EFE360771" blockId="8.[178,1086,868,1066]" pageId="8" pageNumber="1119">Pronotum (Fig. 10) transverse (LP/WP: 0.85), slightly cordate, its lateral sides reflexed, markedly widened in front, posteriorly briefly sinuate and markedly constricted to the basolateral angles, which are small but prominent laterally; anterolateral angles small, but prominent; base fully beaded. Disc flattened, with shallow transverse wrinkles; basal impressions small, elongate, deep, each with some large, deep punctures extended, in some individuals, to the basal area and the lateral furrow; anterolateral and basolateral setiferous punctures present.</paragraph>
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<paragraph id="8BD08F716804FFA8FF000701FE3D0714" blockId="8.[178,479,1081,1103]" box="[178,479,1081,1103]" pageId="8" pageNumber="1119">Mesosternum not denticulate.</paragraph>
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<paragraph id="8BD08F716804FFA8FF000764FCD90797" blockId="8.[178,1086,1119,1228]" pageId="8" pageNumber="1119">Elytra ovate, short (LE/WE: 1.50), markedly depressed at base, subconvex in the apical half. Base narrow; basal ridge weakly concave; humeral tooth and shoulders vanished. Striae deep, punctuate; intervals subconvex. Chaetotaxy: Basal pore present; umbilicate series with 18-19 setiferous punctures; 2 setae at apex of stria 7.</paragraph>
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<paragraph id="8BD08F716804FFA8FF0007E7FC0206FA" blockId="8.[178,1085,1244,1441]" pageId="8" pageNumber="1119">Legs long and slender; profemora on ventral side longitudinally shallowly concave for entire length, their outer margin smooth or with some small teeth and a few (2-5) short setae; mesotibiae straight in females, markedly curved in males; metatibiae each with apical brush of rather long, yellow-reddish setae; tarsomeres with dorsal pubescence sparse and short; metatarsomeres very thin and elongate; males with protarsomeres 1-3 dilated and with ventral, biseriate adhesive vestiture. Tarsal claws with a series of very small teeth, reduced in number (5-6), along the basal half of the internal margin.</paragraph>
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<paragraph id="8BD08F716804FFA8FF00068AFE0B0544" blockId="8.[178,1085,1457,1567]" pageId="8" pageNumber="1119">Male genitalia (Figs 11-14): Aedeagus with median lobe rather short, arcuate, medially inflated; apex very short, regularly rounded, dorsally depressed, not prominent on the dorsal side. Apical orifice wide. Right paramere rounded apically; left paramere with short apical membranous lobe.</paragraph>
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<paragraph id="8BD08F716804FFA9FF000515FE6B02E6" blockId="8.[178,509,1582,1604]" lastBlockId="9.[178,1085,219,445]" lastPageId="9" lastPageNumber="1120" pageId="8" pageNumber="1119">
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Female genitalia: Not examined. C o m p a r i s o n s: The new species is related to both
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<taxonomicName id="4C6FF4F26805FFA9FCA303E0FF0F0255" authority="(G. MULLER 1931)" baseAuthorityName="G. MULLER" baseAuthorityYear="1931" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1120" phylum="Arthropoda" rank="species" species="pisidicus" subGenus="Pristonychus">
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<emphasis id="B91B53636805FFA9FCA303E0FCC403AB" box="[785,806,219,240]" italics="true" pageId="9" pageNumber="1120">L.</emphasis>
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(
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<emphasis id="B91B53636805FFA9FC8703E0FCA903AB" box="[821,843,219,240]" italics="true" pageId="9" pageNumber="1120">P.</emphasis>
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)
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<emphasis id="B91B53636805FFA9FCE903E0FC5703AA" box="[859,949,219,241]" italics="true" pageId="9" pageNumber="1120">pisidicus</emphasis>
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(G. MÜLLER 1931)
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</taxonomicName>
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and
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<emphasis id="B91B53636805FFA9FE9403C2FED60255" box="[294,308,249,270]" italics="true" pageId="9" pageNumber="1120">L</emphasis>
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. (
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<emphasis id="B91B53636805FFA9FEFC03C2FEBF0255" box="[334,349,249,270]" italics="true" pageId="9" pageNumber="1120">P</emphasis>
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.)
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<emphasis id="B91B53636805FFA9FEC503C3FE5F0255" box="[375,445,248,270]" italics="true" pageId="9" pageNumber="1120">sciakyi</emphasis>
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<bibRefCitation id="EFFEF2806805FFA9FE7A03C3FCDE0255" author="CASALE A & VIGNA TAGLIANTI" box="[456,828,248,270]" pageId="9" pageNumber="1120" pagination="277 - 406" refId="ref8290" refString="CASALE A. & A. VIGNA TAGLIANTI (1999): Caraboid beetles (excl. Cicindelidae) of Anatolia, and their biogeographical significance (Coleoptera, Caraboidea). - Biogeographia 20: 277 - 406." type="journal article" year="1999">CASALE & VIGNA TAGLIANTI 1999</bibRefCitation>
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, Anatolian species from which it is recognized by the more reduced although prominent eyes, the flattened, more transverse pronotum, widened in front, the shorter, more ovate elytra, with shoulders and humeral tooth fully vanished, the markedly curved mesotibiae in males, the more slender and more elongate metatarsomeres, with shorter and sparse dorsal pubescence, the tarsal claws with more reduced teeth on the internal side, and the different shape of the median lobe of the aedeagus.
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</paragraph>
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<paragraph id="8BD08F716805FFA9FF0002F6FD220146" blockId="9.[178,1085,461,541]" pageId="9" pageNumber="1120">
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D i s t r i b u t i o n a n d h a b i t a t: An epigean species, known so far only from the
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<collectingRegion id="49AB41936805FFA9FF6502D1FE8B015B" box="[215,361,490,512]" country="Syria" name="Halab" pageId="9" pageNumber="1120">Aleppo region</collectingRegion>
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(North-Western
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<collectingCountry id="F378CFE16805FFA9FDA402D1FDAD015B" box="[534,591,490,512]" name="Syria" pageId="9" pageNumber="1120">Syria</collectingCountry>
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), at
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<quantity id="4C9722946805FFA9FDC802D1FD0A015B" box="[634,744,490,512]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="9" pageNumber="1120" unit="m" value="525.0" valueMax="550.0" valueMin="500.0">500-550 m</quantity>
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a.s.l. The specimens were caught by traps but we have no information on the biotope.
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</paragraph>
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<paragraph id="8BD08F716805FFA9FF000116FDC901E3" blockId="9.[178,1085,557,696]" pageId="9" pageNumber="1120">
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R e m a r k s: As in the case of
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<taxonomicName id="4C6FF4F26805FFA9FDA40116FCEA0119" authority="Casale & D.W & P, 2012" authorityName="Casale & D.W & P" authorityYear="2012" box="[534,776,557,579]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1120" phylum="Arthropoda" rank="species" species="croyi" status="sp. nov." subGenus="Laemostenus">
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<emphasis id="B91B53636805FFA9FDA40116FDC60119" box="[534,548,557,578]" italics="true" pageId="9" pageNumber="1120">L</emphasis>
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. (
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<emphasis id="B91B53636805FFA9FD8F0116FD230119" box="[573,705,557,578]" italics="true" pageId="9" pageNumber="1120">Laemostenus</emphasis>
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)
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<emphasis id="B91B53636805FFA9FD660116FCEA0119" box="[724,776,557,578]" italics="true" pageId="9" pageNumber="1120">croyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A228EE186805FFA9FCA00116FCBE0119" box="[786,860,557,578]" pageId="9" pageNumber="1120" rank="species">nov.sp.</taxonomicNameLabel>
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described above, the discovery of a new, apparently not rare taxon in an area well investigated since many years was unexpected, and confirms the high diversity and the importance in biogeographic aspects of the carabid fauna in the Eastern Mediterranean area as a whole (
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<bibRefCitation id="EFFEF2806805FFA9FF0B0199FDC201E3" author="CASALE A & VIGNA TAGLIANTI" box="[185,544,674,696]" pageId="9" pageNumber="1120" pagination="277 - 406" refId="ref8290" refString="CASALE A. & A. VIGNA TAGLIANTI (1999): Caraboid beetles (excl. Cicindelidae) of Anatolia, and their biogeographical significance (Coleoptera, Caraboidea). - Biogeographia 20: 277 - 406." type="journal article" year="1999">CASALE & VIGNA TAGLIANTI 1999</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |