179 lines
21 KiB
XML
179 lines
21 KiB
XML
<document id="A9150FC9955002B1AA1B4F37D0D75924" ID-CLB-Dataset="20810" ID-DOI="10.11646/zootaxa.5094.1.4" ID-GBIF-Dataset="f9581dee-ff84-4f86-812e-fac15685f80d" ID-ISSN="1175-5326" ID-Zenodo-Dep="5964921" ID-ZooBank="2348120B-390B-4C6B-A217-315ED26EFBAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643789608299" checkinUser="plazi" docAuthor="Bartsch, Daniel & Sáfián, Szabolcs" docDate="2022" docId="03C60D60FFB3143CFF1D2E62A5F2300C" docLanguage="en" docName="zootaxa.5094.1.4.pdf" docOrigin="Zootaxa 5094 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tipulamima seyrigi Viette 1955" docType="treatment" docVersion="7" lastPageNumber="117" masterDocId="FFFF7518FFBD1432FF8A2D7CA4693666" masterDocTitle="Taxonomic changes and review of the genera Tipulamima Holland, 1893 and Macrotarsipodes Le Cerf, 1916 stat. rev. (Lepidoptera: Sesiidae: Sesiinae)" masterLastPageNumber="128" masterPageNumber="103" pageNumber="117" updateTime="1699306083117" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DF716C9478459A98AFCE8AC7098DB82E">Taxonomic changes and review of the genera Tipulamima Holland, 1893 and Macrotarsipodes Le Cerf, 1916 stat. rev. (Lepidoptera: Sesiidae: Sesiinae)</mods:title>
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<mods:namePart id="1B2B595917880CD20F427782462AD36C">Bartsch, Daniel</mods:namePart>
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<mods:affiliation id="C6FCB97C148AB2A8532D47566C707315">Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, 70191 Stuttgart.</mods:affiliation>
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<mods:namePart id="9E20BD7812E239E0364C6F7E9CB5F568">Sáfián, Szabolcs</mods:namePart>
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<mods:affiliation id="F3022439440D102A29E86C9DABA10D7C">African Natural History Research Trust Street Court, Kingsland, Leominster, Herefordshire, HR 6 9 QA, UK.</mods:affiliation>
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<mods:typeOfResource id="623313C77E43E50BC20215CBB47DA544">text</mods:typeOfResource>
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<mods:title id="EEE14115E07D06131D19EF480C9B91AB">Zootaxa</mods:title>
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<mods:date id="49272F07BA3748EE2B85A432D72505B2">2022</mods:date>
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<mods:number id="49FD58731585D728CA93BC47B2E08DB4">2022-02-02</mods:number>
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<treatment id="03C60D60FFB3143CFF1D2E62A5F2300C" ID-DOI="http://doi.org/10.5281/zenodo.6301198" ID-GBIF-Taxon="193266419" ID-Zenodo-Dep="6301198" LSID="urn:lsid:plazi:treatment:03C60D60FFB3143CFF1D2E62A5F2300C" httpUri="http://treatment.plazi.org/id/03C60D60FFB3143CFF1D2E62A5F2300C" lastPageNumber="117" pageId="14" pageNumber="117">
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<subSubSection id="C375EFFDFFB3143CFF1D2E62A646355E" box="[151,559,798,825]" pageId="14" pageNumber="117" type="nomenclature">
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<paragraph id="8BD0BC76FFB3143CFF1D2E62A646355E" blockId="14.[151,559,798,862]" box="[151,559,798,825]" pageId="14" pageNumber="117">
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<heading id="D0980B1AFFB3143CFF1D2E62A646355E" bold="true" box="[151,559,798,825]" fontSize="11" level="1" pageId="14" pageNumber="117" reason="1">
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<taxonomicName id="4C6FC7F5FFB3143CFF1D2E62A646355E" ID-CoL="576ZC" authority="Viette, [1955]" authorityName="Viette" authorityYear="1955" box="[151,559,798,825]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="species" species="seyrigi">
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<emphasis id="B91B6064FFB3143CFF1D2E62A646355E" bold="true" box="[151,559,798,825]" pageId="14" pageNumber="117">
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<emphasis id="B91B6064FFB3143CFF1D2E62A5EB355F" bold="true" box="[151,386,798,825]" italics="true" pageId="14" pageNumber="117">Tipulamima seyrigi</emphasis>
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Viette, [1955]
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C375EFFDFFB3143CFF1D2E3FA54E3538" box="[151,295,835,862]" pageId="14" pageNumber="117" type="description">
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<paragraph id="8BD0BC76FFB3143CFF1D2E3FA54E3538" blockId="14.[151,559,798,862]" box="[151,295,835,862]" pageId="14" pageNumber="117">
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(
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<figureCitation id="1354A0F3FFB3143CFF152E3FA5763538" box="[159,287,835,862]" captionStart="FIGS 15–26" captionStartId="9.[151,210,1775,1799]" captionTargetBox="[152,1430,188,1728]" captionTargetId="figure-17@9.[151,1436,181,1750]" captionTargetPageId="9" captionText="FIGS 15–26. Specimens of Tipulamima. 15. T. flammipes (Hampson, 1910), lectotype ♂. 16–17. T. pyrosoma Hampson, 1919, holotype ♀. 18–20. Tipulamima hesperia sp. nov. 18–19. ♂ Guinea, Nimba Mts. (20) ♂ Ghana, Kumasi. 21–23. T. grandidieri (Le Cerf, 1917). (21) holotype ♂. 22–23. ♂ Madagascar Est, Moramanga, Andasibe. 24–26. T. seyrigi Viette, 1955. (24–25) holotype ♀. (26) ♀ Madagascar, Sambirano. Scale 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5964929" httpUri="https://zenodo.org/record/5964929/files/figure.png" pageId="14" pageNumber="117">Figs 24–26</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C375EFFDFFB3143CFF1D2EFBA5C635D8" pageId="14" pageNumber="117" type="reference_group">
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<paragraph id="8BD0BC76FFB3143CFF1D2EFBA5C635D8" blockId="14.[151,1437,903,958]" pageId="14" pageNumber="117">
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<treatmentCitationGroup id="AB7F9B58FFB3143CFF1D2EFBA5C635D8" pageId="14" pageNumber="117">
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<taxonomicName id="4C6FC7F5FFB3143CFF1D2EFBA59A35F9" ID-CoL="576ZC" authority="Viette, [1955]" authorityName="Viette" authorityYear="1955" box="[151,499,903,927]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="species" species="seyrigi">
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<emphasis id="B91B6064FFB3143CFF1D2EFBA53535F9" box="[151,348,903,927]" italics="true" pageId="14" pageNumber="117">Tipulamima seyrigi</emphasis>
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Viette, [1955]
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</taxonomicName>
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: 88; Viette 1957: 92, 1982: 21;
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<treatmentCitation id="0ACE9A67FFB3143CFCBC2EF4A0ED35F9" author="Heppner, J. B. & Duckworth, W. D." box="[822,1156,903,927]" page="41" pageId="14" pageNumber="117" year="1981">
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<bibRefCitation id="EFFEC187FFB3143CFCBC2EF4A0ED35F9" author="Heppner, J. B. & Duckworth, W. D." box="[822,1156,903,927]" pageId="14" pageNumber="117" pagination="1 - 144" refId="ref17844" refString="Heppner, J. B. & Duckworth, W. D. (1981) Classification of the Superfamily Sesioidea (Lepidoptera, Ditrysia). Smithsonian Contributions to Zoology, 314, 1 - 144. https: // doi. org / 10.5479 / si. 00810282.314" type="journal article" year="1981">Heppner & Duckworth 1981: 41</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0ACE9A67FFB3143CFB052EFBA48A35D8" author="Puhringer, F. & Kallies, A." page="33" pageId="14" pageNumber="117" year="2004">
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<bibRefCitation id="EFFEC187FFB3143CFB052EFBA48A35D8" author="Puhringer, F. & Kallies, A." pageId="14" pageNumber="117" pagination="1 - 85" refId="ref18417" refString="Puhringer, F. & Kallies, A. (2004) Provisional checklist of the Sesiidae of the world (Lepidoptera: Ditrysia). Mitteilungen der Entomologischen Arbeitsgemeinschaft Salzkammergut, 4, 1 - 85. Available from: https: // www. sesiidae. net / Checklst. htm (accessed 6 Oktober 2020)" type="journal article" year="2004">Pühringer & Kallies 2004: 33</bibRefCitation>
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</treatmentCitation>
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; Bartsch 2018: 174.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C375EFFDFFB3143CFF1D2E91A63A33BC" pageId="14" pageNumber="117" type="materials_examined">
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<paragraph id="8BD0BC76FFB3143CFF1D2E91A6A13228" blockId="14.[151,1437,1004,1642]" pageId="14" pageNumber="117">
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Type material examined.
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<materialsCitation id="3B07B62BFFB3143CFE382E90A6A13228" ID-GBIF-Occurrence="3462234365" collectingDate="1932-12" collectionCode="MNHN" collectorName="A. Seyrig & P. Viette" country="Madagascar" location="Centre-Sud" municipality="Bekily" pageId="14" pageNumber="117" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
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||
<typeStatus id="54D402D4FFB3143CFE382E90A6733260" box="[434,538,1004,1030]" pageId="14" pageNumber="117" type="holotype">Holotype</typeStatus>
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||
<specimenCount id="9D6977FFFFB3143CFDAA2E91A65C3261" box="[544,565,1005,1031]" pageId="14" pageNumber="117" type="female">♀</specimenCount>
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||
(
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||
<figureCitation id="1354A0F3FFB3143CFDC92E90A6AE3261" box="[579,711,1004,1031]" captionStart="FIGS 15–26" captionStartId="9.[151,210,1775,1799]" captionTargetBox="[152,1430,188,1728]" captionTargetId="figure-17@9.[151,1436,181,1750]" captionTargetPageId="9" captionText="FIGS 15–26. Specimens of Tipulamima. 15. T. flammipes (Hampson, 1910), lectotype ♂. 16–17. T. pyrosoma Hampson, 1919, holotype ♀. 18–20. Tipulamima hesperia sp. nov. 18–19. ♂ Guinea, Nimba Mts. (20) ♂ Ghana, Kumasi. 21–23. T. grandidieri (Le Cerf, 1917). (21) holotype ♂. 22–23. ♂ Madagascar Est, Moramanga, Andasibe. 24–26. T. seyrigi Viette, 1955. (24–25) holotype ♀. (26) ♀ Madagascar, Sambirano. Scale 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5964929" httpUri="https://zenodo.org/record/5964929/files/figure.png" pageId="14" pageNumber="117">Figs 24–25</figureCitation>
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||
):
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<collectingCountry id="F378FCE6FFB3143CFD532E90A7093260" box="[729,864,1004,1030]" name="Madagascar" pageId="14" pageNumber="117">Madagascar</collectingCountry>
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||
<location id="8EB0EAADFFB3143CFCED2E90A7803260" LSID="urn:lsid:plazi:treatment:03C60D60FFB3143CFF1D2E62A5F2300C:8EB0EAADFFB3143CFCED2E90A7803260" box="[871,1001,1004,1030]" country="Madagascar" municipality="Bekily" name="Centre-Sud" pageId="14" pageNumber="117">Centre-Sud</location>
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,
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<collectingMunicipality id="6BB4260CFFB3143CFC7E2E90A0293260" box="[1012,1088,1004,1030]" pageId="14" pageNumber="117">Bekily</collectingMunicipality>
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,
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<date id="FFD19AB6FFB3143CFBC02E90A0D13260" box="[1098,1208,1004,1030]" pageId="14" pageNumber="117" value="1932-12">
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<collectingDate id="EF95635EFFB3143CFBC02E90A0D13260" box="[1098,1208,1004,1030]" pageId="14" pageNumber="117" value="1932-12">Dec.1932</collectingDate>
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</date>
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,
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<collectorName id="269AD9A0FFB3143CFB482E90A1443260" box="[1218,1325,1004,1030]" pageId="14" pageNumber="117">A. Seyrig</collectorName>
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leg.; with labels: “
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<collectingCountry id="F378FCE6FFB3143CFF7B296CA512324C" box="[241,379,1040,1066]" name="Madagascar" pageId="14" pageNumber="117">Madagascar</collectingCountry>
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/ Bekily / Reg Sud de i’Ile”; “
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<taxonomicName id="4C6FC7F5FFB3143CFD5B296CA735324C" box="[721,860,1040,1066]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="genus">Tipulamima</taxonomicName>
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/ seyrigi
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<taxonomicNameLabel id="A228DD1FFFB3143CFC4C296DA06A324D" box="[966,1027,1041,1067]" pageId="14" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
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/
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<typeStatus id="54D402D4FFB3143CFB93296CA0E8324C" box="[1049,1153,1040,1066]" pageId="14" pageNumber="117" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D6977FFFFB3143CFB03296DA0F7324D" box="[1161,1182,1041,1067]" pageId="14" pageNumber="117" type="female">♀</specimenCount>
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||
/
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<collectorName id="269AD9A0FFB3143CFB3F296DA171324C" box="[1205,1304,1040,1067]" pageId="14" pageNumber="117">P. Viette</collectorName>
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”; “Muséum Paris / XII.32. /
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<collectorName id="269AD9A0FFB3143CFEC02948A5D23228" box="[330,443,1076,1102]" pageId="14" pageNumber="117">A. Seyrig</collectorName>
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||
”; “
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||
<typeStatus id="54D402D4FFB3143CFE502948A6213228" box="[474,584,1076,1102]" pageId="14" pageNumber="117" type="holotype">Holotype</typeStatus>
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” (
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||
<collectionCode id="ED7E24B3FFB3143CFDEB2948A6D43228" box="[609,701,1076,1102]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="117">MNHN</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BD0BC76FFB3143CFF4D2924A63A33BC" blockId="14.[151,1437,1004,1642]" pageId="14" pageNumber="117">
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The only known specimen is the female
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<typeStatus id="54D402D4FFB3143CFD232924A7663214" box="[681,783,1112,1138]" pageId="14" pageNumber="117" type="holotype">holotype</typeStatus>
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||
. Analogous to the significant sexual dimorphism within other species of
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<taxonomicName id="4C6FC7F5FFB3143CFED92900A5B032F0" box="[339,473,1148,1174]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="genus">
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<emphasis id="B91B6064FFB3143CFED92900A5B032F0" box="[339,473,1148,1174]" italics="true" pageId="14" pageNumber="117">Tipulamima</emphasis>
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</taxonomicName>
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the conspecifity of
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<taxonomicName id="4C6FC7F5FFB3143CFD342901A74E32F0" authorityName="Viette" authorityYear="1955" box="[702,807,1149,1174]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="species" species="seyrigi">
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<emphasis id="B91B6064FFB3143CFD342901A74E32F0" box="[702,807,1149,1174]" italics="true" pageId="14" pageNumber="117">T. seyrigi</emphasis>
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||
</taxonomicName>
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||
and
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<taxonomicName id="4C6FC7F5FFB3143CFCEB2901A79632F0" box="[865,1023,1148,1174]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="species" species="grandidieri">
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<emphasis id="B91B6064FFB3143CFCEB2901A79632F0" box="[865,1023,1148,1174]" italics="true" pageId="14" pageNumber="117">T. grandidieri</emphasis>
|
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</taxonomicName>
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might be possible. A second female from Sambirano (
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<figureCitation id="1354A0F3FFB3143CFEEA29DCA5DA32DD" box="[352,435,1184,1211]" captionStart="FIGS 15–26" captionStartId="9.[151,210,1775,1799]" captionTargetBox="[152,1430,188,1728]" captionTargetId="figure-17@9.[151,1436,181,1750]" captionTargetPageId="9" captionText="FIGS 15–26. Specimens of Tipulamima. 15. T. flammipes (Hampson, 1910), lectotype ♂. 16–17. T. pyrosoma Hampson, 1919, holotype ♀. 18–20. Tipulamima hesperia sp. nov. 18–19. ♂ Guinea, Nimba Mts. (20) ♂ Ghana, Kumasi. 21–23. T. grandidieri (Le Cerf, 1917). (21) holotype ♂. 22–23. ♂ Madagascar Est, Moramanga, Andasibe. 24–26. T. seyrigi Viette, 1955. (24–25) holotype ♀. (26) ♀ Madagascar, Sambirano. Scale 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5964929" httpUri="https://zenodo.org/record/5964929/files/figure.png" pageId="14" pageNumber="117">Fig. 26</figureCitation>
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) (
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<collectionCode id="ED7E24B3FFB3143CFE4129DCA64D32DC" box="[459,548,1184,1210]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="117">MNHN</collectionCode>
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) listed by Viette ([1955]) differs significantly by the smaller transparent areas of both wings, the broader black apical area of the forewing, the larger hindwing discal spot and the much broader hindwing margin. It might further not even be conspecific with
|
||
<taxonomicName id="4C6FC7F5FFB3143CFCD02995A7A83364" authorityName="Viette" authorityYear="1955" box="[858,961,1257,1282]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="species" species="seyrigi">
|
||
<emphasis id="B91B6064FFB3143CFCD02995A7A83364" box="[858,961,1257,1282]" italics="true" pageId="14" pageNumber="117">T. seyrigi</emphasis>
|
||
</taxonomicName>
|
||
, given the extreme geographic disjunction of the two records and the completely different ecological setting. Another female from “Region d’Ivohibe, vallée de l’Iantara” listed by
|
||
<bibRefCitation id="EFFEC187FFB3143CFE1D284CA644332C" author="Viette, P." box="[407,557,1328,1354]" pageId="14" pageNumber="117" refId="ref18788" refString="Viette, P. (1982) Insectes. Lepidopteres Sesiidae. Fauna de Madagascar, 61, 64 pp., 40 figs." type="book" year="1982">Viette (1982)</bibRefCitation>
|
||
is here not taken into consideration. This specimen is superficially similar to
|
||
<taxonomicName id="4C6FC7F5FFB3143CFF1D2829A5693308" authorityName="Viette" authorityYear="1955" box="[151,256,1365,1390]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="species" species="seyrigi">
|
||
<emphasis id="B91B6064FFB3143CFF1D2829A5693308" box="[151,256,1365,1390]" italics="true" pageId="14" pageNumber="117">T. seyrigi</emphasis>
|
||
</taxonomicName>
|
||
, but does not belong to
|
||
<taxonomicName id="4C6FC7F5FFB3143CFD992828A6F03308" box="[531,665,1364,1390]" class="Insecta" family="Sesiidae" genus="Tipulamima" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91B6064FFB3143CFD992828A6F03308" box="[531,665,1364,1390]" italics="true" pageId="14" pageNumber="117">Tipulamima</emphasis>
|
||
</taxonomicName>
|
||
. It is smaller, generally more delicate; the orange parts of the wings paler, yellow; the transparent areas larger; the hindwing discal spot shorter, not reaching M
|
||
<subScript id="17EBBE33FFB3143CFB1528F6A0CE33FE" attach="left" box="[1183,1191,1418,1432]" fontSize="6" pageId="14" pageNumber="117">2</subScript>
|
||
; and the hindlegs not elongated, without black, or tufted scales. Because of these characters, we conclude that it belongs to another, so far undescribed species of
|
||
<taxonomicName id="4C6FC7F5FFB3143CFE1628BCA63933BC" authorityName="Niculescu" authorityYear="1964" box="[412,592,1472,1498]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="117" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375EFFDFFB3143CFF4D2898A5F2300C" pageId="14" pageNumber="117" type="description">
|
||
<paragraph id="8BD0BC76FFB3143CFF4D2898A5F2300C" blockId="14.[151,1437,1004,1642]" pageId="14" pageNumber="117">
|
||
<emphasis id="B91B6064FFB3143CFF4D2898A5563398" bold="true" box="[199,319,1508,1534]" pageId="14" pageNumber="117">Variation.</emphasis>
|
||
Unknown. The specimen from Sambirano has distal portion of the transparent areas of the forewings covered with orange scales; the margins of the hindwings three to four times wider; the discal spot of the hindwings somewhat larger, reaching common stalk of M
|
||
<subScript id="17EBBE33FFB3143CFD112B42A6CA302A" attach="left" box="[667,675,1598,1612]" fontSize="6" pageId="14" pageNumber="117">3</subScript>
|
||
and CuA
|
||
<subScript id="17EBBE33FFB3143CFC862B42A77D302A" attach="left" box="[780,788,1598,1612]" fontSize="6" pageId="14" pageNumber="117">1</subScript>
|
||
; the tarsus of the hindleg partially black; and the abdomen ventrally paler whitish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |